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101-510: Tusker or Tuskers may refer to: Animals [ edit ] an elephant with large tusks (discussed at Tusk#Size ) an adult male wild boar , particularly the American razorback population Military unit nicknames [ edit ] 4th Battalion, 64th Armor Regiment , a United States Army unit No. 5 Squadron IAF , an Indian Air Force squadron 413 Transport and Rescue Squadron ,

202-644: A 60% loss. In 2012, a large upsurge in ivory poaching was reported, with about 70% of the product flowing to China. China was the biggest market for poached ivory but announced that it would phase out the legal domestic manufacture and sale of ivory products in May 2015. Conflicts between elephants and a growing human population are a major issue in elephant conservation. Human encroachment into natural areas where bush elephants occur or their increasing presence in adjacent areas has spurred research into methods of safely driving groups of elephants away from humans. Playback of

303-546: A Canadian Armed Forces air force squadron Sports teams [ edit ] Sri Lanka rugby union team nickname "The Tuskers” East Africa rugby union team , who tour under the nickname "Tuskers" Tusker F.C. , a Kenyan football club owned by the East African Breweries Ballari Tuskers , a Karnataka Premier League cricket franchise Kochi Tuskers Kerala , a former Indian Premier League cricket franchise Florida Tuskers ,

404-409: A Y. The female's clitoris may be 40 cm (16 in). The vulva is found lower than in other herbivores, between the hind legs instead of under the tail. Determining pregnancy status can be difficult due to the animal's large belly. The female's mammary glands occupy the space between the front legs, which puts the suckling calf within reach of the female's trunk. Elephants have a unique organ,

505-449: A day. Elephants tend to stay near water sources. They have morning, afternoon, and nighttime feeding sessions. At midday, elephants rest under trees and may doze off while standing. Sleeping occurs at night while the animal is lying down. Elephants average 3–4 hours of sleep per day. Both males and family groups typically move no more than 20 km (12 mi) a day, but distances as far as 180 km (112 mi) have been recorded in

606-499: A distinct species and called it the 'forest elephant'; later authors considered it to be a subspecies . Morphological and genetic analyses have since provided evidence for species-level differences between the African bush elephant and the African forest elephant. In 1907, Richard Lydekker proposed six African elephant subspecies based on the different sizes and shapes of their ears. They are all considered synonymous with

707-504: A faster gait similar to running. In walking, the legs act as pendulums, with the hips and shoulders moving up and down while the foot is planted on the ground. The fast gait does not meet all the criteria of running, since there is no point where all the feet are off the ground, although the elephant uses its legs much like other running animals, and can move faster by quickening its stride. Fast-moving elephants appear to 'run' with their front legs, but 'walk' with their hind legs and can reach

808-433: A female and her herd for about a month before moving on in search of another mate. Less than a third of the population of female elephants will be in estrus at any given time and the gestation period of an elephant is long, so it makes more evolutionary sense for a male to search for as many females as possible rather than stay with one group. Both species are threatened by habitat loss and fragmentation, and poaching for

909-631: A former United Football League franchise Matabeleland Tuskers , a Zimbabwean cricket franchise Tuskers women's cricket team , a Zimbabwean women's cricket team Other uses [ edit ] Tusker (beer) , a Kenyan beer bottled by East African Breweries Tusker Rock , a rock in the Bristol Channel, Wales a character in the Japanese anime series Kiba a member of the DC Comics Justice League of Earth

1010-418: A highly intelligent species that is thought to be equal to cetaceans and primates . The African elephant’s cognitive complexity includes behaviors indicative of empathy, problem-solving, and cooperative group behaviors. These traits underscore the evolutionary convergence of intelligence across species, similar to that seen in primates and cetaceans. African elephants are at their most fertile between

1111-494: A human brain. It is the largest of all terrestrial mammals. While the elephant brain is larger overall, it is proportionally smaller than the human brain . At birth, an elephant's brain already weighs 30–40% of its adult weight. The cerebrum and cerebellum are well developed, and the temporal lobes are so large that they bulge out laterally. Their temporal lobes are proportionally larger than those of other animals, including humans. The throat of an elephant appears to contain

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1212-500: A loud, low way to attract males from far away. Male elephants can also smell the hormones of a female ready for breeding. This leads males to compete with each other to mate, which results in the females mating with older, healthier males. Females choose to a point who they mate with, since they are the ones who try to get males to compete to guard them. However, females are not guarded in the early and late stages of estrus, which may permit mating by younger males not in musth. Males over

1313-453: A physical and behavioral condition that is characterized by elevated testosterone, aggression and more sexual activity. Musth also serves a purpose of calling attention to the females that they are of good quality, and it cannot be mimicked as certain calls or noises may be. Males sire few offspring in periods when they are not in musth. During the middle of estrus, female elephants look for males in musth to guard them. The females will yell, in

1414-548: A pouch where it can store water for later use. The larynx of the elephant is the largest known among mammals. The vocal folds are anchored close to the epiglottis base. When comparing an elephant's vocal folds to those of a human, an elephant's are proportionally longer, thicker, with a greater cross-sectional area. In addition, they are located further up the vocal tract with an acute slope. The heart of an elephant weighs 12–21 kg (26–46 lb). Its apex has two pointed ends, an unusual trait among mammals. In addition,

1515-418: A protective crust. Elephants have difficulty releasing heat through the skin because of their low surface-area-to-volume ratio , which is many times smaller than that of a human. They have even been observed lifting up their legs to expose their soles to the air. Elephants only have sweat glands between the toes, but the skin allows water to disperse and evaporate, cooling the animal. In addition, cracks in

1616-703: A result of the trade ban, African elephant populations recovered in Southern African range countries. The African Elephant Specialist Group has set up a Human-Elephant Conflict Task Force with the aim to develop conflict mitigation strategies. In 2005, the West African Elephant Memorandum of Understanding was signed by 12 West African countries. The Convention on the Conservation of Migratory Species of Wild Animals provided financial support for four years to implement

1717-541: A single species, Loxodonta africana , but molecular studies have affirmed their status as separate species. Mammoths ( Mammuthus ) are nested within living elephants as they are more closely related to Asian elephants than to African elephants. Another extinct genus of elephant, Palaeoloxodon , is also recognised, which appears to have close affinities with African elephants and to have hybridised with African forest elephants. Over 180 extinct members of order Proboscidea have been described. The earliest proboscideans,

1818-468: A specific time; however, they are less likely to reproduce in times of drought than when water is plentiful. The gestation period of an elephant is 22 months and fertile females usually give birth every 3–6 years, so if they live to around 50 years of age, they may produce 7 offspring. Females are a scarce and mobile resource for the males so there is intense competition to gain access to estrous females. Post sexual maturity, males begin to experience musth ,

1919-440: A strong grasp. The flexibility of the trunk is aided by the numerous wrinkles in the skin. The African elephants have two finger-like extensions at the tip of the trunk that allow them to pluck small food. The Asian elephant has only one and relies more on wrapping around a food item. Asian elephant trunks have better motor coordination . The trunk's extreme flexibility allows it to forage and wrestle other elephants with it. It

2020-661: A top speed of 25 km/h (16 mph). At this speed, most other quadrupeds are well into a gallop, even accounting for leg length. Spring-like kinetics could explain the difference between the motion of elephants and other animals. The cushion pads expand and contract, and reduce both the pain and noise that would come from a very heavy animal moving. Elephants are capable swimmers: they can swim for up to six hours while completely waterborne, moving at 2.1 km/h (1 mph) and traversing up to 48 km (30 mi) continuously. The brain of an elephant weighs 4.5–5.5 kg (10–12 lb) compared to 1.6 kg (4 lb) for

2121-432: A year. As the tusk develops, it is topped with smooth, cone-shaped enamel that eventually wanes. The dentine is known as ivory and has a cross-section of intersecting lines, known as "engine turning", which create diamond-shaped patterns. Being living tissue, tusks are fairly soft and about as dense as the mineral calcite . The tusk protrudes from a socket in the skull, and most of it is external. At least one-third of

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2222-512: Is Loxodonta cookei , with remains of the species known from around 7–5 million years ago, from remains found in Chad, Kenya, Uganda, and South Africa. Analysis of nuclear DNA sequences indicates that the genetic divergence between African bush and forest elephants dates 2.6 – 5.6 million years ago. The African forest elephant was found to have a high degree of genetic diversity , likely reflecting periodic fragmentation of their habitat during

2323-655: Is prehensile , bringing food and water to the mouth and grasping objects. Tusks, which are derived from the incisor teeth, serve both as weapons and as tools for moving objects and digging. The large ear flaps assist in maintaining a constant body temperature as well as in communication. African elephants have larger ears and concave backs, whereas Asian elephants have smaller ears and convex or level backs. Elephants are scattered throughout sub-Saharan Africa , South Asia, and Southeast Asia and are found in different habitats, including savannahs , forests, deserts, and marshes . They are herbivorous , and they stay near water when it

2424-409: Is accessible. They are considered to be keystone species , due to their impact on their environments. Elephants have a fission–fusion society , in which multiple family groups come together to socialise. Females (cows) tend to live in family groups, which can consist of one female with her calves or several related females with offspring. The leader of a female group, usually the oldest cow, is known as

2525-456: Is covered with sparse, bristled dark-brown to black hair. Short tactile hair grows on the trunk, which has two finger-like processes at the tip, whereas Asian elephants only have one. Their large ears help to reduce body heat. Flapping them creates air currents and exposes the ears' inner sides where large blood vessels increase heat loss during hot weather. The trunk is a prehensile elongation of its upper lip and nose. This highly sensitive organ

2626-718: Is derived from the Latin word elephas ( genitive elephantis ) ' elephant ' , which is the Latinised form of the ancient Greek ἐλέφας ( elephas ) (genitive ἐλέφαντος ( elephantos )), probably from a non- Indo-European language , likely Phoenician . It is attested in Mycenaean Greek as e-re-pa (genitive e-re-pa-to ) in Linear B syllabic script. As in Mycenaean Greek, Homer used

2727-677: Is innervated primarily by the trigeminal nerve , and is thought to be manipulated by about 40,000–60,000 muscles. Because of this muscular structure, the trunk is so strong that elephants can use it to lift about 3% of their own body weight. They use it for smelling, touching, feeding, drinking, dusting, producing sounds, loading, defending and attacking. Elephants sometimes swim underwater and use their trunks as snorkels . Both male and female African elephants have tusks that grow from deciduous teeth called tushes, which are replaced by tusks when calves are about one year old. Tusks are composed of dentin , which forms small diamond-shaped structures in

2828-542: Is larger than that of any other terrestrial animal. The elephant's brain is similar to a human brain in terms of structure and complexity; the elephant's cortex has as many neurons as that of a human brain, suggesting convergent evolution . Elephants exhibit a wide variety of behaviours, including those associated with grief , learning , mimicry , art , play , a sense of humor , altruism , use of tools , compassion , cooperation , self-awareness , memory and possibly language . All of these behaviors point to

2929-620: Is led by an older cow known as the matriarch . African forest elephant groups are less cohesive than African bush elephant groups, probably because of the lack of predators. When separate family units bond, they form kinship or bond groups. After puberty , male elephants tend to form close alliances with other males. While females are the most active members of African elephant groups, both male and female elephants are capable of distinguishing between hundreds of different low-frequency infrasonic calls to communicate with and identify each other. Elephants use some vocalisations that are beyond

3030-417: Is powerful enough to lift up to 350 kg (770 lb), but it also has the precision to crack a peanut shell without breaking the seed. With its trunk, an elephant can reach items up to 7 m (23 ft) high and dig for water in the mud or sand below. It also uses it to clean itself. Individuals may show lateral preference when grasping with their trunks: some prefer to twist them to the left, others to

3131-421: Is the largest terrestrial animal . Cows are 2.2–2.6 m (7.2–8.5 ft) tall at the shoulder and weigh 2,160–3,232 kg (4,762–7,125 lb), while bulls are 3.2–4 m (10–13 ft) tall and weigh 4,700–6,048 kg (10,362–13,334 lb). Its back is concave-shaped, while the back of the African forest elephant is nearly straight. The largest recorded individual stood 3.96 metres (13.0 ft) at

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3232-728: Is the third largest terrestrial animal. African elephants are distributed in Sub-Saharan Africa , where they inhabit Sahelian scrubland and arid regions, tropical rainforests , mopane and miombo woodlands . African forest elephant populations occur only in Central and West Africa. Elephants are the animals with the lowest sleep times, especially African elephants. Research has found their average sleep to be only 2 hours in 24-hour cycles. Both African elephant species live in family units comprising several adult cows, their daughters and their subadult sons. Each family unit

3333-600: Is typically more worn down, as it is shorter and blunter. For African elephants, tusks are present in both males and females and are around the same length in both sexes, reaching up to 300 cm (9 ft 10 in), but those of males tend to be more massive. In the Asian species, only the males have large tusks. Female Asians have very small tusks, or none at all. Tuskless males exist and are particularly common among Sri Lankan elephants . Asian males can have tusks as long as Africans', but they are usually slimmer and lighter;

3434-571: The African bush elephant ( Loxodonta africana ), the African forest elephant ( L. cyclotis ), and the Asian elephant ( Elephas maximus ). They are the only surviving members of the family Elephantidae and the order Proboscidea ; extinct relatives include mammoths and mastodons . Distinctive features of elephants include a long proboscis called a trunk, tusks , large ear flaps, pillar-like legs, and tough but sensitive grey skin. The trunk

3535-564: The Bering Strait into the Holocene, with their latest survival being on Wrangel Island , where they persisted until around 4,000 years ago. Over the course of their evolution, probobscideans grew in size. With that came longer limbs and wider feet with a more digitigrade stance, along with a larger head and shorter neck. The trunk evolved and grew longer to provide reach. The number of premolars, incisors, and canines decreased, and

3636-848: The Convention on International Trade in Endangered Species of Wild Fauna and Flora listed the African elephant on CITES Appendix I . This listing banned international trade of African elephants and their body parts by countries that signed the CITES agreement. Hunting elephants is banned in the Central African Republic, Democratic Republic of Congo, Gabon, Côte d'Ivoire, and Senegal. After the ban came into force in 1990, retail sales of ivory carvings in South Africa have plummeted by more than 95% within 10 years. As

3737-842: The Early Pleistocene , all non-elephantid probobscidean genera outside of the Americas became extinct with the exception of Stegodon , with gomphotheres dispersing into South America as part of the Great American interchange , and mammoths migrating into North America around 1.5 million years ago. At the end of the Early Pleistocene, around 800,000 years ago the elephantid genus Palaeoloxodon dispersed outside of Africa, becoming widely distributed in Eurasia. Proboscideans were represented by around 23 species at

3838-538: The Etosha region of Namibia. Elephants go on seasonal migrations in response to changes in environmental conditions. In northern Botswana, they travel 325 km (202 mi) to the Chobe River after the local waterholes dry up in late August. Loxodonta African elephants are members of the genus Loxodonta comprising two living elephant species , the African bush elephant ( L. africana ) and

3939-756: The International Union for Conservation of Nature (IUCN). One of the biggest threats to elephant populations is the ivory trade , as the animals are poached for their ivory tusks. Other threats to wild elephants include habitat destruction and conflicts with local people. Elephants are used as working animals in Asia. In the past, they were used in war; today, they are often controversially put on display in zoos, or employed for entertainment in circuses . Elephants have an iconic status in human culture and have been widely featured in art, folklore, religion, literature, and popular culture. The word elephant

4040-705: The Sahel in the north to the Highveld in the south. Decrease of suitable habitat was the major cause for the decline of elephant populations until the 1950s. Hunting African elephants for the ivory trade accelerated the decline from the 1970s onwards. The carrying capacity of remaining suitable habitats was estimated at 8,985,000 elephants at most by 1987. In the 1970s and 1980s, the price of ivory rose, and poaching for ivory increased, particularly in Central African range countries where access to elephant habitats

4141-522: The changes in the Pleistocene . Gene flow between the two African elephant species was examined at 21 locations. The analysis revealed that several African bush elephants carried mitochondrial DNA of African forest elephants, indicating they hybridised in the savanna-forest transition zone in ancient times. However, despite the hybridisation at the contact zone between the two species, there appears to have been little effective gene flow between

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4242-405: The incisors , known as the tusks ; 12 deciduous premolars ; and 12 molars . Unlike most mammals, teeth are not replaced by new ones emerging from the jaws vertically. Instead, new teeth start at the back of the mouth and push out the old ones. The first chewing tooth on each side of the jaw falls out when the elephant is two to three years old. This is followed by four more tooth replacements at

4343-443: The lozenge -shaped enamel of the molar teeth , which differs significantly from the rounded shape of the Asian elephant 's molar enamel. The first scientific description of the African elephant was written in 1797 by Johann Friedrich Blumenbach , who proposed the scientific name Elephas africanus . Loxodonte was proposed as a generic name for the African elephant by Frédéric Cuvier in 1825. An anonymous author used

4444-507: The matriarch . Males (bulls) leave their family groups when they reach puberty and may live alone or with other males. Adult bulls mostly interact with family groups when looking for a mate. They enter a state of increased testosterone and aggression known as musth , which helps them gain dominance over other males as well as reproductive success. Calves are the centre of attention in their family groups and rely on their mothers for as long as three years. Elephants can live up to 70 years in

4545-423: The pleural cavity . This may allow the animal to deal with the pressure differences when its body is underwater and its trunk is breaking the surface for air. Elephants breathe mostly with the trunk but also with the mouth. They have a hindgut fermentation system, and their large and small intestines together reach 35 m (115 ft) in length. Less than half of an elephant's food intake gets digested, despite

4646-487: The temporal gland , located on both sides of the head. This organ is associated with sexual behaviour, and males secrete a fluid from it when in musth . Females have also been observed with these secretions. Elephants are herbivorous and will eat leaves, twigs, fruit, bark, grass, and roots. African elephants mostly browse , while Asian elephants mainly graze . They can eat as much as 300 kg (660 lb) of food and drink 40 L (11 US gal) of water in

4747-432: The ventricles of the heart split towards the top, a trait also found in sirenians. When upright, the elephant's heart beats around 28 beats per minute and actually speeds up to 35 beats when it lies down. The blood vessels are thick and wide and can hold up under high blood pressure. The lungs are attached to the diaphragm , and breathing relies less on the expanding of the ribcage. Connective tissue exists in place of

4848-596: The "shovel tuskers" like Platybelodon ), choerolophodontids and stegodontids . Around 10 million years ago, the earliest members of the family Elephantidae emerged in Africa, having originated from gomphotheres. Elephantids are distinguished from earlier proboscideans by a major shift in the molar morphology to parallel lophs rather than the cusps of earlier proboscideans, allowing them to become higher-crowned (hypsodont) and more efficient in consuming grass. The Late Miocene saw major climactic changes, which resulted in

4949-480: The 1980s, it was difficult to carry out systematic surveys in several East African range countries due to civil wars . In 1987, it was estimated that the African elephant population had declined to 760,000 individuals. In 1989, only 608,000 African elephants were estimated to have survived. In 1989, the Kenyan Wildlife Service burned a stockpile of tusks in protest against the ivory trade. When

5050-528: The African Eritherium and Phosphatherium are known from the late Paleocene . The Eocene included Numidotherium , Moeritherium , and Barytherium from Africa. These animals were relatively small and, some, like Moeritherium and Barytherium were probably amphibious. Later on, genera such as Phiomia and Palaeomastodon arose; the latter likely inhabited more forested areas. Proboscidean diversification changed little during

5151-763: The African bush elephant. A third species, the West African elephant, has also been proposed but needs confirmation. It is thought that this lineage has been isolated from the others for 2.4 million years. Between the late 18th and 21st centuries, the following extinct African elephants were described on the basis of fossil remains: Relationships of living and extinct elephants based on DNA, after Palkopoulou et al. 2018. Loxodonta (African elephants) [REDACTED] † Palaeoloxodon (straight-tusked elephants) [REDACTED] Elephas (Asian elephants) [REDACTED] † Mammuthus (mammoths) [REDACTED] The oldest species of Loxodonta known

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5252-410: The African elephant also allows for the digestion of various plant parts, including fibrous stems, bark and roots. African elephants are highly intelligent. They have a very large and highly convoluted neocortex , a trait they share with humans , apes and some dolphin species. They are among the world's most intelligent species. With a mass of just over 5 kg (11 lb), the elephant brain

5353-665: The Greek word to mean ivory , but after the time of Herodotus , it also referred to the animal. The word elephant appears in Middle English as olyfaunt ( c.  1300 ) and was borrowed from Old French oliphant (12th century). Orycteropodidae [REDACTED] Macroscelididae [REDACTED] Chrysochloridae [REDACTED] Tenrecidae [REDACTED] Procaviidae [REDACTED] Elephantidae [REDACTED] Dugongidae [REDACTED] Trichechidae [REDACTED] Elephants belong to

5454-585: The Latinized spelling Loxodonta in 1827. This author was recognized as authority by the International Code of Zoological Nomenclature in 1999. Elephas ( Loxodonta ) cyclotis was proposed by Paul Matschie in 1900, who described three African elephant zoological specimens from Cameroon whose skulls differed in shape from those of elephant skulls collected elsewhere in Africa. In 1936, Glover Morrill Allen considered this elephant to be

5555-660: The Oligocene. One notable species of this epoch was Eritreum melakeghebrekristosi of the Horn of Africa , which may have been an ancestor to several later species. early proboscideans, e.g. Moeritherium [REDACTED] Deinotheriidae [REDACTED] Mammutidae [REDACTED] Gomphotheriidae [REDACTED] Stegodontidae [REDACTED] Loxodonta [REDACTED] Palaeoloxodon [REDACTED] Mammuthus [REDACTED] Elephas [REDACTED] A major event in proboscidean evolution

5656-501: The University of Nebraska's former Holland Computing Center high memory cluster the name of a DHR B Class steam locomotive the sports teams of Somers High School (New York) , a public school See also [ edit ] Tusk Topics referred to by the same term [REDACTED] This disambiguation page lists articles associated with the title Tusker . If an internal link led you here, you may wish to change

5757-412: The age of 25 compete strongly for females in estrus, and are more successful the larger and more aggressive they are. Bigger males tend to sire bigger offspring. Wild males begin breeding in their thirties when they are at a size and weight that is competitive with other adult males. Male reproductive success is maximal in mid-adulthood and then begins to decline. However, this can depend on the ranking of

5858-589: The ages of 25 and 45. Calves are born after a gestation period of up to nearly two years. The calves are cared for by their mother and other young females in the group, known as allomothering . African elephants show sexual dimorphism in weight and shoulder height by age 20, due to the rapid early growth of males. By age 25, males are double the weight of females; however, both sexes continue to grow throughout their lives. Female African elephants are able to start reproducing at around 10 to 12 years of age, and are in estrus for about 2 to 7 days. They do not mate at

5959-425: The ages of four to six, 9–15, 18–28, and finally in their early 40s. The final (usually sixth) set must last the elephant the rest of its life. Elephant teeth have loop-shaped dental ridges, which are more diamond-shaped in African elephants. The tusks of an elephant are modified second incisors in the upper jaw. They replace deciduous milk teeth at 6–12 months of age and keep growing at about 17 cm (7 in)

6060-586: The beginning of the Late Pleistocene . Proboscideans underwent a dramatic decline during the Late Pleistocene as part of the Late Pleistocene extinctions of most large mammals globally, with all remaining non-elephantid proboscideans (including Stegodon , mastodons , and the American gomphotheres Cuvieronius and Notiomastodon ) and Palaeoloxodon becoming extinct, with mammoths only surviving in relict populations on islands around

6161-453: The cheek teeth (molars and premolars) became longer and more specialised. The incisors developed into tusks of different shapes and sizes. Several species of proboscideans became isolated on islands and experienced insular dwarfism , some dramatically reducing in body size, such as the 1 m (3 ft 3 in) tall dwarf elephant species Palaeoloxodon falconeri . Elephants are the largest living terrestrial animals. Some species of

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6262-402: The decline and extinction of many proboscidean groups. The earliest members of the modern genera of Elephantidae appeared during the latest Miocene–early Pliocene around 5 million years ago. The elephantid genera Elephas (which includes the living Asian elephant) and Mammuthus (mammoths) migrated out of Africa during the late Pliocene, around 3.6 to 3.2 million years ago. Over the course of

6363-420: The ear is considerably thinner. Elephants are typically grey, but African elephants look brown or reddish after rolling in coloured mud. Asian elephants have some patches of depigmentation, particularly on the head. Calves have brownish or reddish hair, with the head and back being particularly hairy. As elephants mature, their hair darkens and becomes sparser, but dense concentrations of hair and bristles remain on

6464-567: The elephant loses the last of its molars and will likely die of starvation which is a common cause of death. African elephants have 24 teeth in total, six on each quadrant of the jaw. The enamel plates of the molars are fewer in number than in Asian elephants. The enamel of the molar teeth wears into a distinctive lozenge/loxodont ( <> ) shape characteristic to all members of the genus Loxodonta . While some extinct species of Loxodonta retained permanent premolar teeth, these have been lost in both living species. The African bush elephant

6565-525: The elephant uses its trunk as a snorkel . The trunk also acts as a sense organ. Its sense of smell may be four times greater than a bloodhound 's nose. The infraorbital nerve , which makes the trunk sensitive to touch, is thicker than both the optic and auditory nerves. Whiskers grow all along the trunk, and are particularly packed at the tip, where they contribute to its tactile sensitivity. Unlike those of many mammals, such as cats and rats, elephant whiskers do not move independently ("whisk") to sense

6666-452: The elephant's lifetime. The dental formula of elephants is 1.0.3.3 0.0.3.3  × 2 = 26 . Elephants have four molars ; each weighs about 5 kg (11 lb) and measures about 30 cm (12 in) long. As the front pair wears down and drops out in pieces, the back pair moves forward, and two new molars emerge in the back of the mouth. Elephants replace their teeth four to six times in their lifetimes. At around 40 to 60 years of age,

6767-402: The environment; the trunk itself must move to bring the whiskers into contact with nearby objects. Whiskers grow in rows along each side on the ventral surface of the trunk, which is thought to be essential in helping elephants balance objects there, whereas they are more evenly arranged on the dorsal surface. The number and patterns of whiskers are distinctly different between species. Damaging

6868-410: The extinct elephant genus Palaeoloxodon considerably exceeded modern elephants in size making them among the largest land mammals ever. The skeleton is made up of 326–351 bones. The vertebrae are connected by tight joints, which limit the backbone's flexibility. African elephants have 21 pairs of ribs, while Asian elephants have 19 or 20 pairs. The skull contains air cavities ( sinuses ) that reduce

6969-585: The family Elephantidae , the sole remaining family within the order Proboscidea . Their closest extant relatives are the sirenians ( dugongs and manatees ) and the hyraxes , with which they share the clade Paenungulata within the superorder Afrotheria . Elephants and sirenians are further grouped in the clade Tethytheria . Three species of living elephants are recognised; the African bush elephant ( Loxodonta africana ), forest elephant ( Loxodonta cyclotis ), and Asian elephant ( Elephas maximus ). African elephants were traditionally considered

7070-519: The front and hind limbs can support an elephant's weight, although 60% is borne by the front. The position of the limbs and leg bones allows an elephant to stand still for extended periods of time without tiring. Elephants are incapable of turning their manus as the ulna and radius of the front legs are secured in pronation . Elephants may also lack the pronator quadratus and pronator teres muscles or have very small ones. The circular feet of an elephant have soft tissues, or "cushion pads" beneath

7171-440: The hearing range of humans, to communicate across large distances. Elephant mating rituals include the gentle entwining of trunks. Bulls were believed to be solitary animals, becoming independent once reaching maturity. New research suggests that bulls maintain ecological knowledge for the herd, facilitating survival when searching for food and water, which also benefits the young bulls who associate with them. Bulls only return to

7272-423: The herd to breed or to socialize; they do not provide prenatal care to their offspring, but rather play a fatherly role to younger bulls to show dominance. While feeding, the African elephant uses its trunk to pluck leaves and its tusks to tear at branches, which can cause enormous damage to foliage. Fermentation of the food takes place in the hindgut , enabling large food intakes. The large size and hindgut of

7373-657: The illegal ivory trade in several range countries. Loxodonta is one of two extant genera in the family Elephantidae . The name refers to the lozenge-shaped enamel of their molar teeth. Fossil remains of Loxodonta species have been found in Africa , spanning from the Late Miocene (from around 7–6 million years ago) onwards. The name Loxodonta comes from the Ancient Greek words λοξός ( loxós, "slanting", "crosswise") and ὀδούς ( odoús , "tooth"), referring to

7474-422: The illegal ivory trade is a threat in several range countries as well. The African bush elephant is listed as Endangered and the African forest elephant as Critically Endangered on the respective IUCN Red Lists . Based on vegetation types that provide suitable habitat for African elephants, it was estimated that in the early 19th century a maximum of 26,913,000 African elephants might have been present from

7575-552: The international ivory trade reopened in 2006, the demand and price for ivory increased in Asia. In Chad's Zakouma National Park , more than 3,200 elephants were killed between 2005 and 2010. The park did not have sufficient guards to combat poaching and their weapons were outdated. Well organized networks facilitated smuggling the ivory through Sudan. The government of Tanzania estimated that more than 85,000 elephants were lost to poaching in Tanzania between 2009 and 2014, representing

7676-542: The largest ear flaps. The ossicles are adapted for hearing low frequencies, being most sensitive at 1 kHz . Lacking a lacrimal apparatus (tear duct), the eye relies on the harderian gland in the orbit to keep it moist. A durable nictitating membrane shields the globe. The animal's field of vision is compromised by the location and limited mobility of the eyes. Elephants are dichromats and they can see well in dim light but not in bright light. The elongated and prehensile trunk, or proboscis , consists of both

7777-402: The largest recorded was 302 cm (9 ft 11 in) long and weighed 39 kg (86 lb). Hunting for elephant ivory in Africa and Asia has resulted in an effective selection pressure for shorter tusks and tusklessness. An elephant's skin is generally very tough, at 2.5 cm (1 in) thick on the back and parts of the head. The skin around the mouth, anus , and inside of

7878-456: The link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Tusker&oldid=1193104145 " Category : Disambiguation pages Hidden categories: Short description is different from Wikidata All article disambiguation pages All disambiguation pages Elephant Elephants are the largest living land animals. Three living species are currently recognised:

7979-438: The male within their group, as higher-ranking males maintain a higher rate of reproduction. Most observed matings are by males in musth over 35 years of age. Twenty-two long observations showed that age and musth are extremely important factors; "… older males had markedly elevated paternity success compared with younger males, suggesting the possibility of sexual selection for longevity in this species." Males usually stay with

8080-449: The manus or pes , which allow them to bear the animal's great mass. They appear to have a sesamoid , an extra "toe" similar in placement to a giant panda 's extra "thumb", that also helps in weight distribution. As many as five toenails can be found on both the front and hind feet. Elephants can move both forward and backward, but are incapable of trotting , jumping , or galloping . They can move on land only by walking or ambling :

8181-425: The middle with a thinner tip and supported by a thicker base. They contain numerous blood vessels called capillaries . Warm blood flows into the capillaries, releasing excess heat into the environment. This effect is increased by flapping the ears back and forth. Larger ear surfaces contain more capillaries, and more heat can be released. Of all the elephants, African bush elephants live in the hottest climates and have

8282-413: The nose and upper lip, which fuse in early fetal development. This versatile appendage contains up to 150,000 separate muscle fascicles , with no bone and little fat. These paired muscles consist of two major types: superficial (surface) and internal. The former are divided into dorsal, ventral , and lateral muscles, while the latter are divided into transverse and radiating muscles. The muscles of

8383-408: The nuclear genome of P. antiquus deriving from L. cyclotis. This ancestry is closer to modern west African populations than to central African populations of forest elephants. Analysis of Chinese Palaeoloxodon mitogenomes suggests that this forest elephant ancestry was widely shared among Palaeoloxodon species. African elephants have grey folded skin up to 30 mm (1.2 in) thick that

8484-423: The process lasting a day. An elephant's bladder can store up to 18 litres of urine and its kidneys can produce more than 50 litres of urine per day. A male elephant's testes, like other Afrotheria , are internally located near the kidneys. The penis can be as long as 100 cm (39 in) with a 16 cm (6 in) wide base. It curves to an 'S' when fully erect and has an orifice shaped like

8585-608: The recorded sounds of angry Western honey bees has been found to be remarkably effective at prompting elephants to flee an area. Farmers have tried scaring elephants away by more aggressive means such as fire or the use of chili peppers along fences to protect their crops. In 1986, the African Elephant Database was initiated with the aim to monitor the status of African elephant populations. This database includes results from aerial surveys, dung counts, interviews with local people and data on poaching. In 1989,

8686-493: The right. Elephant trunks are capable of powerful siphoning. They can expand their nostrils by 30%, leading to a 64% greater nasal volume, and can breathe in almost 30 times faster than a human sneeze, at over 150 m/s (490 ft/s). They suck up water, which is squirted into the mouth or over the body. The trunk of an adult Asian elephant is capable of retaining 8.5 L (2.2 US gal) of water. They will also sprinkle dust or grass on themselves. When underwater,

8787-484: The shoulder, and is estimated to have weighed 10,400 kg (22,900 lb). The tallest recorded individual stood 4.21 m (13.8 ft) at the shoulder and weighed 8,000 kg (18,000 lb). The African forest elephant is smaller with a weight of up to 4,000 kg (8,800 lb) and a shoulder height of 1.8–2.4 m (5 ft 11 in – 7 ft 10 in) in females and 2.4–3 m (7 ft 10 in – 9 ft 10 in) in males. It

8888-401: The skin may reduce dehydration and allow for increased thermal regulation in the long term. To support the animal's weight, an elephant's limbs are positioned more vertically under the body than in most other mammals. The long bones of the limbs have cancellous bones in place of medullary cavities . This strengthens the bones while still allowing haematopoiesis (blood cell creation). Both

8989-456: The skull, which serve as protection. Because of the size of the head, the neck is relatively short to provide better support. Elephants are homeotherms and maintain their average body temperature at ~ 36 °C (97 °F), with a minimum of 35.2 °C (95.4 °F) during the cool season, and a maximum of 38.0 °C (100.4 °F) during the hot dry season. Elephant ear flaps, or pinnae , are 1–2 mm (0.039–0.079 in) thick in

9090-576: The smaller African forest elephant ( L. cyclotis ). Both are social herbivores with grey skin . However, they differ in the size and colour of their tusks as well as the shape and size of their ears and skulls . Both species are at a pertinent risk of extinction according to the IUCN Red List ; as of 2021, the bush elephant is considered endangered while the forest elephant is considered critically endangered . They are threatened by habitat loss and fragmentation, along with poaching for

9191-472: The tip of the tail and parts of the head and genitals. Normally, the skin of an Asian elephant is covered with more hair than its African counterpart. Their hair is thought to help them lose heat in their hot environments. Although tough, an elephant's skin is very sensitive and requires mud baths to maintain moisture and protection from burning and insect bites. After bathing, the elephant will usually use its trunk to blow dust onto its body, which dries into

9292-514: The trunk connect to a bony opening in the skull. The nasal septum consists of small elastic muscles between the nostrils, which are divided by cartilage at the base. A unique proboscis nerve – a combination of the maxillary and facial nerves – lines each side of the appendage. As a muscular hydrostat , the trunk moves through finely controlled muscle contractions, working both with and against each other. Using three basic movements: bending, twisting, and longitudinal stretching or retracting,

9393-422: The trunk has near unlimited flexibility. Objects grasped by the end of the trunk can be moved to the mouth by curving the appendage inward. The trunk can also bend at different points by creating stiffened "pseudo-joints". The tip can be moved in a way similar to the human hand. The skin is more elastic on the dorsal side of the elephant trunk than underneath; allowing the animal to stretch and coil while maintaining

9494-508: The trunk would be detrimental to an elephant's survival, although in rare cases, individuals have survived with shortened ones. One trunkless elephant has been observed to graze using its lips with its hind legs in the air and balancing on its front knees. Floppy trunk syndrome is a condition of trunk paralysis recorded in African bush elephants and involves the degeneration of the peripheral nerves and muscles. The disorder has been linked to lead poisoning. Elephants usually have 26 teeth:

9595-433: The tusk contains the pulp , and some have nerves that stretch even further. Thus, it would be difficult to remove it without harming the animal. When removed, ivory will dry up and crack if not kept cool and wet. Tusks function in digging, debarking, marking, moving objects, and fighting. Elephants are usually right- or left-tusked, similar to humans, who are typically right- or left-handed . The dominant, or "master" tusk,

9696-410: The tusk's center that become larger at its periphery. Tusks are primarily used to dig for roots and strip the bark from trees for food, for fighting each other during the mating season, and for defending themselves against predators. The tusks weigh from 23 to 45 kg (51–99 lb) and can be from 1.5 to 2.4 m (5–8 ft) long. They are curved forward and continue to grow throughout

9797-490: The two species since their initial split. DNA from the European straight-tusked elephant ( Palaeoloxodon antiquus ) indicates that the extinct elephant genus Palaeoloxodon is more closely related to African elephants than to Asian elephants or mammoths. Analysis of the genome of P. antiquus also shows that Palaeloxodon extensively hybridised with African forest elephants, with the mitochondrial genome and over 30% of

9898-419: The weight of the skull while maintaining overall strength. These cavities give the inside of the skull a honeycomb -like appearance. By contrast, the lower jaw is dense. The cranium is particularly large and provides enough room for the attachment of muscles to support the entire head. The skull is built to withstand great stress, particularly when fighting or using the tusks. The brain is surrounded by arches in

9999-463: The wild. They communicate by touch, sight, smell, and sound; elephants use infrasound and seismic communication over long distances. Elephant intelligence has been compared with that of primates and cetaceans . They appear to have self-awareness , and possibly show concern for dying and dead individuals of their kind. African bush elephants and Asian elephants are listed as endangered and African forest elephants as critically endangered by

10100-438: Was facilitated by logging and petroleum mining industries. Between 1976 and 1980, about 830 t (820 long tons; 910 short tons) raw ivory was exported from Africa to Hong Kong and Japan , equivalent to tusks of about 222,000 African elephants. The first continental elephant census was carried out in 1976. At the time, 1.34 million elephants were estimated to range over 7,300,000 km (2,800,000 sq mi). In

10201-670: Was the collision of Afro-Arabia with Eurasia, during the Early Miocene, around 18–19 million years ago, allowing proboscideans to disperse from their African homeland across Eurasia and later, around 16–15 million years ago into North America across the Bering Land Bridge . Proboscidean groups prominent during the Miocene include the deinotheres , along with the more advanced elephantimorphs , including mammutids (mastodons), gomphotheres , amebelodontids (which includes

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