Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.
58-463: Cervus is a genus of deer that primarily are native to Eurasia , although one species occurs in northern Africa and another in North America . In addition to the species presently placed in this genus, it has included a whole range of other species now commonly placed in other genera. Additionally, the species-level taxonomy is in a state of flux. Until the 1970s, Cervus also included
116-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as
174-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,
232-714: A quarter of all paddy fields in Japan. Chiba Prefecture is famous for its peanuts, also being the largest producer in Japan. Rare species of the lichen genus Menegazzia are found only in Honshu. Most of Japan's tea and silk is from Honshu. Japan's three largest industrial regions are all located on Honshu: the Keihin region , the Hanshin Industrial Region , and the Chūkyō Industrial Area . Honshu
290-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for
348-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;
406-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and
464-436: A yearly breeding season where they display sensory exploitation, intrasexual competition, and weaponry. Females will fight for optimal mating opportunities and sexually selection for males with larger antler size and/or greater roar quality. The degree of polygyny and female aggregation is dependent on the level of food distribution. Females aggregating in areas with more food leading to larger harems Female distribution influences
522-573: Is an indicator of mate potential. Red deer can distinguish the calls of the males in their harem, others and their offspring. Females select for males with larger antlers which indirectly benefits them. Large antler size in males is a sign of health and strength. The visual display is a reliable indicator of mate quality, providing indirect benefits. The females are not directly affected by these characteristics, but they will produce more viable and fit offspring. Males with large antlers mate and sire more offspring than smaller, younger males. Large antler size
580-536: Is correlated with overall health, fitness and an increase in sperm production and quality. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of
638-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms
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#1732764994703696-472: Is home to a large portion of Japan's minimal mineral reserves, including small oil and coal deposits. Several coal deposits are located in the northern part of the island, concentrated in Fukushima Prefecture and Niigata Prefecture , though Honshu's coal production is negligible in comparison to Hokkaido and Kyushu . Most of Japan's oil reserves are also located in northern Honshu, along
754-517: Is often overlooked because it is not as extravagant as male antler combat. Female conflicts occur so the winner has first access to the harem male at the start of the mating season before he is exhausted or low on sperm storage. While an emphasis in observations of sexual selection is placed on combat using antlers, males with higher roaring rates are also being selected for. During the breeding season males will make calls to attract mates and compete with other males. Like antler size, mating call quality
812-513: Is roughly 1,300 km (810 mi) long and ranges from 50 to 230 km (31 to 143 mi) wide, and its total area is 227,960 km (88,020 sq mi), making it slightly larger than the island of Great Britain . Its land area has been increasing with land reclamation and coastal uplift in the north due to plate tectonics with a convergent boundary . Honshu has 10,084 kilometres (6,266 mi) of coastline. Mountainous and volcanic, Honshu experiences frequent earthquakes (such as
870-412: Is seismically active, and is home to 40 active volcanoes. In 2011, an earthquake of magnitude 9.0–9.1 occurred off the coast of Honshu, generating tsunami waves up to 40.5 meters (133 ft) high and killing 19,747. It was the most powerful earthquake ever recorded in Japan , and the fourth most powerful earthquake in the world since modern record-keeping began in 1900. The tsunami subsequently led to
928-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,
986-701: Is the largest and most populous island of Japan . It is located south of Hokkaidō across the Tsugaru Strait , north of Shikoku across the Inland Sea , and northeast of Kyūshū across the Kanmon Straits . The island separates the Sea of Japan , which lies to its north and west, from the North Pacific Ocean to the south and east. It is the seventh-largest island in the world, and
1044-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with
1102-657: The Caucasus and northwestern Iran , and the elk/wapiti ( C. canadensis ) to North America and the Asian regions of the Tian Shan , Altai , and Great Khingan . Alternatively, the barbarus group species are subspecies of the "true" red deer, while the C. wallichii and C. xanthopygus groups are subspecies of the elk/wapiti. It has been proposed that the sika deer should be split into four species based on genetics, morphology and voice, although this may be premature based on
1160-543: The Great Kantō earthquake , which heavily damaged Tokyo in September 1923; and the earthquake of March 2011 , which moved the northeastern part of the island by varying amounts of as much as 5.3 m (17 ft) while causing devastating tsunamis). The highest peak is the active volcano Mount Fuji at 3,776 m (12,388 ft), which makes Honshu the world's 7th highest island . There are many rivers, including
1218-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to
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#17327649947031276-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;
1334-691: The Kashmir stag , which has not been sampled in recent studies. If it is included in the Central Asian red deer, the scientific name of that species is C. hanglu . If it is not included, the scientific name of that species is C. yarkandensis , and the Kashmir stag ( C. hanglu ) may represent a separate monotypic species. The Central Asian red deer was considered its own species (including the Yarkand deer, Kashmir stag and Bactrian deer as subspecies) by
1392-728: The Kurushima Kaikyō Bridge ; Shimotsui-Seto Bridge , Hitsuishijima Bridge , Iwakurojima Bridge , Yoshima Bridge , Kita Bisan-Seto Bridge , and the Minami Bisan-Seto Bridge ), the Seikan Tunnel connects Honshu with Hokkaidō, and the Kanmonkyo Bridge and Kanmon Tunnel connect Honshu with Kyūshū . These are notable flora and fauna of Honshu. Being on the Ring of Fire , the island of Honshu
1450-533: The Shinano River , Japan's longest. The Japanese Alps span the width of Honshu, from the 'Sea of Japan' coast to the Pacific shore. The climate is generally humid subtropical in western Japan and humid continental in the north. Honshu has a total population of 104 million people, according to a 2017 estimate, 81.3% of the entire population of Japan. The largest city is Tokyo (population: 13,988,129),
1508-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,
1566-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,
1624-582: The second-most populous after the Indonesian island of Java . Honshu had a population of 104 million as of 2017 , constituting 81.3% of the entire population of Japan, and mostly concentrated in the coastal areas and plains. Approximately 30% of the total population resides in the Greater Tokyo Area on the Kantō Plain . As the historical center of Japanese cultural and political power,
1682-607: The IUCN in 2017, and by the American Society of Mammalogists in 2021. Others members of the red deer group, which may represent separate species, are C. corsicanus , C. wallichi and C. xanthopygus . If so, C. corsicanus includes the subspecies C. c. barbarus (perhaps a synonym of corsicanus ), and is restricted to Maghreb in North Africa, Corsica and Sardinia . C. wallichi would probably include
1740-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of
1798-540: The capital of Japan and part of the Greater Tokyo Area , the most populous metropolitan area in the world. Honshu is connected to the islands of Hokkaidō, Kyūshū and Shikoku by tunnels and bridges. Three bridge systems have been built across the islands of the Inland Sea between Honshu and Shikoku ( Akashi Kaikyo Bridge and the Ōnaruto Bridge ; Shin-Onomichi Bridge , Innoshima Bridge , Ikuchi Bridge , Tatara Bridge , Ōmishima Bridge , Hakata–Ōshima Bridge , and
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1856-719: The economy along the northwestern Sea of Japan coast is largely based on fishing and agriculture. The island is linked to the other three major Japanese islands by a number of bridges and tunnels. The island primarily shares two climates, with Northern Honshu having a mainly humid continental climate while the south has a humid subtropical climate . The name of the island, Honshū ( 本州 ) , directly translates to "main province" or "original land" in English. Humans first arrived in Honshu approximately 37,000 years ago, and likely earlier. The first humans to arrive in Honshu were Stone Age hunter-gatherers from Northeast Asia, likely following
1914-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this
1972-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,
2030-516: The internal phylogeny of Cervus to be as follows: C. elaphus ( Red deer ) C. hanglu ( Central Asian red deer ) C. albirostris ( Thorold's deer ) C. nippon ( Sika deer ) C. canadensis ( Elk ) Rusa The remains of Cervus are known from the early-mid Pliocene of China. Members of the genus Cervus have polygynous mating systems within harems. These harems consist of several males, numerous females and their young offspring 1–3 years in age Members of this genus have
2088-597: The island includes several past Japanese capitals, including Kyōto , Nara , and Kamakura . Much of the island's southern shore forms part of the Taiheiyō Belt , a megalopolis that spans several of the Japanese islands. Honshu contains Japan's highest mountain, Mount Fuji, and its largest lake, Lake Biwa . Most of Japan's industry is located in a belt running along Honshu's southern coast, from Tokyo to Nagoya , Kyōto , Osaka , Kobe , and Hiroshima ; by contrast,
2146-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,
2204-446: The level of polygyny. Female-female competition has been observed within harems in the red deer species ( Cervus elaphus ) prior to and during the mating season. Aggression is displayed through nose threats, kicking, and displacements. Elevated aggression has only been observed during the breeding season . Competition can be for access to mates or reproductive resources such as food, or nesting areas. Female-female aggression in ungulates
2262-396: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Honshu Honshu ( 本州 , Honshū , pronounced [hoꜜɰ̃ɕɯː] ; lit. ' main island ' ) , historically called Akitsushima ( 秋津島 , lit. ' Dragonfly island ' ) ,
2320-617: The meltdown of 3 nuclear reactors at the Fukushima Daiichi Nuclear Power Plant , leading to the Fukushima nuclear disaster . Honshu island generates around US$ 3.5 trillion or more than 80% of Japan's GDP . Fruit, vegetables, grains, rice and cotton make up the main produce grown in Honshu. The Tohoku region, spanning the north-eastern part of the island, is notable for its rice production, with 65% of cultivated land being rice paddy fields – almost
2378-471: The members of the genera Axis , Dama , and Elaphurus , and until the late 1980s, it included members of Rucervus and Rusa . In the third edition of Mammal Species of the World from 2005, only the red deer ( C. elaphus ) and sika deer ( C. nippon ) were recognized as species in the genus Cervus . Genetic and morphological evidence suggest more species should be recognized. For example,
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2436-786: The migration of ice age megafauna . Surviving artifacts from this period include finely-crafted stone blades, similar to those found in Siberia . Honshu was the target of devastating air raids during the Pacific War of World War II . The first air raid to strike Honshu and the other home islands was the Doolittle Raid . With the introduction of the Boeing B-29 Superfortress , the firebombing of Tokyo culminated in Operation Meetinghouse ,
2494-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus
2552-640: The most destructive air raid in human history, which destroyed 16 square miles (41 km ; 10,000 acres) of central Tokyo , leaving an estimated 100,000 civilians dead, and over one million homeless. The war ended with the atomic bombings of Hiroshima and Nagasaki shortly before Japan's surrender and signing of the Japanese Instrument of Surrender on September 2, 1945, on board the USS ; Missouri (BB-63) in Tokyo Bay . The island
2610-741: The most heavily used. The San'yō Shinkansen , connects the two largest cities in western Japan, Shin-Osaka in Osaka with Hakata Station in Fukuoka . Both the Tokaido Shinkansen and the Sanyo Shinkansen help form a continuous high-speed railway through the Taiheiyō Belt megalopolis. The island is divided into five nominal regions and contains 34 prefectures, including metropolitan Tokyo. Administratively, some smaller islands are included within these prefectures, notably including
2668-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this
2726-543: The presently available evidence. If split, the potential species are C. yesoensis from northern and central Japan ( Hokkaido and northern and central Honshu ), C. nippon of southern Japan (southern Honshu, Shikoku , Kyushu , Okinawa , Tsushima and other small islands), C. hortulorum of mainland Asia (the Russian Far East , Korea , central and eastern China and northern Vietnam ), and C. taiouanus of Taiwan . A 2014 mitochondrial DNA study showed
2784-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for
2842-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or
2900-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in
2958-622: The species Cervus canadensis (elk/wapiti) is considered a separate species. Within the red deer species group , some sources have recommended the Central Asian red deer ( Cervus hanglu ) should be treated as a species. If the Central Asian red deer (from the Caspian Sea to western China ) is recognized as a species, it includes the Yarkand deer and Bactrian deer (the two may be synonymous ), but it could possibly also include
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#17327649947033016-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,
3074-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being
3132-427: The subspecies C. w. kansuensis and C. w. macneilli (both are perhaps synonyms of C. w. wallichi ), and would be found from Tibet to central China. C. xanthopygus would probably include the subspecies C. x. alashanicus (perhaps a synonym of C. x. xanthopygus ), and would be found from the Russian Far East to northeastern China. This would restrict the "true" red deer ( C. elaphus ) to Europe, Anatolia ,
3190-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for
3248-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up
3306-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in
3364-404: The west coast, spanning Niigata, Yamagata , and Akita Prefectures. Most of Japan's copper, lead, zinc and chromite is located on Honshu, along with smaller, scattered deposits of gold, silver, arsenic, sulfur and pyrite . The Tokaido Shinkansen , opened in 1964 between Tokyo and Shin-Ōsaka , is Japan's first high-speed rail line. It is the world's oldest high-speed rail line and one of
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