131-525: The chestnuts are the deciduous trees and shrubs in the genus Castanea , in the beech family Fagaceae . The name also refers to the edible nuts they produce. They are native to temperate regions of the Northern Hemisphere. Chestnut trees are of moderate growth rate (for the Chinese chestnut tree) to fast-growing for American and European species. Their mature heights vary from
262-489: A coleoptile that forms the first leaf while the radicle is covered with a coleorhiza that connects to the primary root and adventitious roots form the sides. Here the hypocotyl is a rudimentary axis between radicle and plumule. The seeds of corn are constructed with these structures; pericarp, scutellum (single large cotyledon) that absorbs nutrients from the endosperm, plumule, radicle, coleoptile, and coleorhiza – these last two structures are sheath-like and enclose
393-508: A zygote . The embryo within a seed develops from the zygote and grows within the mother plant to a certain size before growth is halted. The formation of the seed is the defining part of the process of reproduction in seed plants ( spermatophytes ). Other plants such as ferns , mosses and liverworts , do not have seeds and use water-dependent means to propagate themselves. Seed plants now dominate biological niches on land, from forests to grasslands both in hot and cold climates . In
524-480: A 10 kg (22 lb) yield for a 10-year-old tree is a reliable conservative estimate, though some exceptional specimens of that age have yielded 100 kg (220 lb). So, most producers earn less than $ 5,000 per year, with a third of them not having sold anything so far. Moreover, the plantings have so far been mostly of Chinese species, but the products are not readily available. The American Chestnut Foundation in collaboration with many partners ( SUNY ESF ,
655-399: A behavior known as " marcescence " whereby dead leaves are not shed in the fall and remain on the tree until being blown off by the weather. This is caused by incomplete development of the abscission layer. It is mainly seen in the seedling and sapling stage, although mature trees may have marcescence of leaves on the lower branches. A number of deciduous plants remove nitrogen and carbon from
786-422: A curved megagametophyte often giving the seed a tight "C" shape. The last ovule shape is called amphitropous , where the ovule is partly inverted and turned back 90 degrees on its stalk (the funicle or funiculus ). In the majority of flowering plants, the zygote's first division is transversely oriented in regards to the long axis, and this establishes the polarity of the embryo. The upper or chalazal pole becomes
917-411: A few will end in a favorable place for growth. Herbaceous perennials and woody plants often have larger seeds; they can produce seeds over many years, and larger seeds have more energy reserves for germination and seedling growth and produce larger, more established seedlings after germination. Seeds serve several functions for the plants that produce them. Key among these functions are nourishment of
1048-550: A germ with two cotyledons connected to creamy-white flesh throughout. Some varieties have consistently only one embryo per fruit (nut) or have only one large fruit per burr, well rounded (no flat face). The name of varieties with these characteristics may start with "marron" for example marron de Lyon in France, or Marrone di Mugello in Italy. Chestnut fruit may not exhibit epigeal dormancy. It may germinate right upon falling to
1179-602: A greater rate of transpiration (and hence CO 2 uptake as this occurs when stomata are open) during the summer growth period. The deciduous characteristic has developed repeatedly among woody plants. Trees include maple , many oaks and nothofagus , elm , beech , aspen , and birch , among others, as well as a number of coniferous genera , such as larch and Metasequoia . Deciduous shrubs include honeysuckle , viburnum , and many others. Most temperate woody vines are also deciduous, including grapes , poison ivy , Virginia creeper , wisteria , etc. The characteristic
1310-550: A heavy, sweet odor that some people find too sweet or unpleasant. Other catkins have these pollen-bearing flowers, but also carry near the twig from which these spring, small clusters of female or fruit-producing flowers. Two or three flowers together form a four-lobed prickly calybium , which ultimately grows completely together to make the brown hull, or husk , covering the fruits. Chestnut flowers are not self-compatible, so two trees are required for pollination . All Castanea species readily hybridize with each other. The fruit
1441-415: A location and be there at a time favorable for germination and growth. When the fruits open and release their seeds in a regular way, it is called dehiscent , which is often distinctive for related groups of plants; these fruits include capsules , follicles , legumes , silicles and siliques . When fruits do not open and release their seeds in a regular fashion, they are called indehiscent, which include
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#17327728581711572-421: A patchwork of leafy and leafless trees. Seed In botany , a seed is a plant embryo and nutrient reserve enclosed in a seed coat, a protective outer covering called a testa. More generally, the term "seed" means anything that can be sown , which may include seed and husk or tuber . Seeds are the product of the ripened ovule , after the embryo sac is fertilized by sperm from pollen , forming
1703-400: A relatively low initial investment; this is a factor in the small size of the present production operations, with half of them being between 3 and 10 acres (12,000 and 40,000 m). Another determining factor in the small productivity of the sector is that most orchards have been created less than 10 years ago, so have young trees which are as now barely entering commercial production. Assuming
1834-670: A song that celebrates chestnuts, as well as " Jeongseokga ", a song from the Goryeo period. Gongju , one of Baekje 's former capitals, is renowned for its chestnuts, with an annual chestnut festival that takes place in the winter. In the Samgukji(Records Of The Three Kingdoms) , a book that was compiled during the Jin dynasty about the Three Kingdoms , chestnuts are used in the description of Mahan,
1965-714: A staple food in southern Europe, Turkey, and southwestern and eastern Asia for millennia, largely replacing cereals where these would not grow well, if at all, in mountainous Mediterranean areas. Evidence of its cultivation by humans is found since around 2000 BC. Alexander the Great and the Romans planted chestnut trees across Europe while on their various campaigns. A Greek army is said to have survived their retreat from Asia Minor in 401–399 BC thanks to their stores of chestnuts. Ancient Greeks, such as Dioscorides and Galen, wrote of chestnuts to comment on their medicinal properties—and of
2096-435: A supply of nutrients for the embryo in most monocotyledons and the endospermic dicotyledons. Seeds have been considered to occur in many structurally different types (Martin 1946). These are based on a number of criteria, of which the dominant one is the embryo-to-seed size ratio. This reflects the degree to which the developing cotyledons absorb the nutrients of the endosperm, and thus obliterate it. Six types occur amongst
2227-550: A tegmen from the inner integument while unitegmic seeds have only one integument. Usually, parts of the testa or tegmen form a hard protective mechanical layer. The mechanical layer may prevent water penetration and germination. Amongst the barriers may be the presence of lignified sclereids . The outer integument has a number of layers, generally between four and eight organised into three layers: (a) outer epidermis, (b) outer pigmented zone of two to five layers containing tannin and starch, and (c) inner epidermis. The endotegmen
2358-405: A way for the species to survive dry or cold seasons. Ephemeral plants are usually annuals that can go from seed to seed in as few as six weeks. Seed germination is a process by which a seed embryo develops into a seedling. It involves the reactivation of the metabolic pathways that lead to growth and the emergence of the radicle or seed root and plumule or shoot. The emergence of the seedling above
2489-435: Is frost -tender; bud -burst is later than most other fruit trees, so late frosts can be damaging to young buds. Trees can be found at altitudes between 200 and 1000 m above sea level; some mention between 300 and 750 m altitude, while the famous Hundred Horse Chestnut on Mount Etna stands at 1200 metres. They can tolerate maritime exposure, although growth is reduced. Seeds germinate in late winter or early spring, but
2620-606: Is a deciduous woody shrub found in Japan . Forests where a majority of the trees lose their foliage at the end of the typical growing season are called deciduous forests. These forests are found in many areas worldwide and have distinctive ecosystems, understory growth, and soil dynamics. Two distinctive types of deciduous forests are found growing around the world. Temperate deciduous forest biomes are plant communities distributed in North and South America, Asia, Southern slopes of
2751-423: Is a hard, shiny, brown outer hull or husk , called the pericarpus; the industry calls this the "peel". Underneath the pericarpus is another, thinner skin, called the pellicle or episperm. The pellicle closely adheres to the seed itself, following the grooves usually present at the surface of the fruit. These grooves are of variable sizes and depths according to the species and variety. The fruit inside these shows
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#17327728581712882-513: Is a traditional beverage, and it is gaining popularity with the tourists and in continental Portugal. Always served as part of the New Year's menu in Japan, chestnuts represent both success and hard times—mastery and strength. The Japanese chestnut ( kuri ) was in cultivation before rice and the Chinese chestnut ( C. mollissima ) possibly for 2,000 to 6,000 years. During British colonial rule in
3013-509: Is at a suitable temperature with proper soil moisture. This true dormancy or innate dormancy is therefore caused by conditions within the seed that prevent germination. Thus dormancy is a state of the seed, not of the environment. Induced dormancy, enforced dormancy or seed quiescence occurs when a seed fails to germinate because the external environmental conditions are inappropriate for germination, mostly in response to conditions being too dark or light, too cold or hot, or too dry. Seed dormancy
3144-519: Is at the modern village of Keramidi . The town probably took its name, though, from the trees growing around it. In the Mediterranean climate zone, chestnut trees are rarer in Greece because the chalky soil is not conducive to the tree's growth. Kastania is located on one of the relatively few sedimentary or siliceous outcrops. They grow so abundantly there that their presence would have determined
3275-481: Is caused by conditions outside the embryo, including: Endogenous dormancy is caused by conditions within the embryo itself, including: The following types of seed dormancy do not involve seed dormancy, strictly speaking, as lack of germination is prevented by the environment, not by characteristics of the seed itself (see Germination ): Not all seeds undergo a period of dormancy. Seeds of some mangroves are viviparous; they begin to germinate while still attached to
3406-700: Is completed (marked by the transition from bright green spring leaves to dark green summer ones) the chlorophyll level in the leaves remains stable until cool temperatures arrive in autumn. When autumn arrives and the days are shorter or when plants are drought-stressed, the chlorophyll steadily breaks down, allowing other pigments present in the leaf to become apparent and resulting in non-green colored foliage. The brightest leaf colors are produced when days grow short and nights are cool, but remain above freezing. These other pigments include carotenoids that are yellow, brown, and orange. Anthocyanin pigments produce red and purple colors, though they are not always present in
3537-412: Is contained in a spiny (very sharp) cupule 5–11 cm in diameter, also called "bur" or " burr ". The burrs are often paired or clustered on the branch and contain one to seven nuts according to the different species , varieties , and cultivars . Around the time the fruits reach maturity, the burrs turn yellow-brown and split open in two or four sections. They can remain on the tree longer than they hold
3668-681: Is derived from an earlier English term "chesten nut", which descends from the Old French word chastain (Modern French, châtaigne ). The French word in turn derives from Latin Castanea (also the scientific name of the tree), which traces to the Ancient Greek word κάστανον (sweet chestnut). A possible source of the Greek word is the ancient town of Casthanaea in Magnesia . Its location
3799-402: Is derived from the inner epidermis of the inner integument, the exotegmen from the outer surface of the inner integument. The endotesta is derived from the inner epidermis of the outer integument, and the outer layer of the testa from the outer surface of the outer integument is referred to as the exotesta . If the exotesta is also the mechanical layer, this is called an exotestal seed, but if
3930-400: Is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf; it consists of layers of cells that can separate from each other. The cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. When auxin coming from the leaf is produced at a rate consistent with that from the body of
4061-741: Is low, varying between 5 and 20 mg/(kg·h) depending on the temperature. Most chestnut wood production is done by coppice systems, cut on a 12-year rotation to provide small timber which does not split as badly as large logs. In southern England (particularly in Kent ), sweet chestnut has traditionally been grown as coppices, being recut every 10 years or so on rotation for poles used for firewood, and fencing (fence posts and chestnut paling). An excellent soil-enriching understory in pine forests, sustainable forest management incorporates more mixed plantings of proven efficiency, as opposed to monosylviculture. A study presented in 1997 has evaluated positively
Chestnut - Misplaced Pages Continue
4192-444: Is not the same as seed persistence in the soil or on the plant, though even in scientific publications dormancy and persistence are often confused or used as synonyms. Often, seed dormancy is divided into four major categories: exogenous; endogenous; combinational; and secondary. A more recent system distinguishes five classes: morphological, physiological, morphophysiological, physical, and combinational dormancy. Exogenous dormancy
4323-535: Is protection against disease. Seeds protect and nourish the embryo or young plant. They usually give a seedling a faster start than a sporeling from a spore, because of the larger food reserves in the seed and the multicellularity of the enclosed embryo. Unlike animals, plants are limited in their ability to seek out favorable conditions for life and growth. As a result, plants have evolved many ways to disperse their offspring by dispersing their seeds (see also vegetative reproduction ). A seed must somehow "arrive" at
4454-517: Is recommended at least during summer and early autumn. Once established, they resist droughts well. In addition to being consumed fresh, chestnuts can also be canned, pureed, or preserved in sugar or syrup ( marrons glacés ). Shelled and cooked nuts should be covered, refrigerated, and used within 3–4 days. Cooked chestnuts, either whole, chopped, or pureed, may be frozen in an airtight container and held up to 9 months. Because of their high water content, transpiration rates, and consequent loss weight,
4585-474: Is required later. The optimum rainfall for chestnut trees is 800 mm (31 in) or more, ideally in even distribution throughout the year. Mulching during summer is recommended. Rainfall below 700 mm (28 in) per year needs be complemented with, for example, a drip irrigation system. This should water the soil at the outer half of the circle formed by the drip line to encourage root growth. Independently from annual rainfall, watering young trees
4716-723: Is the Mugello region ; in 1996, the European Community granted the fruit Protected Geographic Indication (equivalent to the French Appellation d'Origine Contrôlée ) status to the Mugello sweet chestnut . It is markedly sweet, peels easily, is not excessively floury or astringent , and has notes of vanilla , hazelnut , and, more subtly, fresh bread. It has no unpleasant aroma, such as yeast , fungus , mold , or paper, which sometimes occur with other chestnuts. The main regions in France for chestnut production are
4847-446: Is the ability of the embryo to germinate and is affected by a number of different conditions. Some plants do not produce seeds that have functional complete embryos, or the seed may have no embryo at all, often called empty seeds. Predators and pathogens can damage or kill the seed while it is still in the fruit or after it is dispersed. Environmental conditions like flooding or heat can kill the seed before or during germination. The age of
4978-411: Is then aborted or absorbed during early development. The seed is composed of the embryo (the result of fertilization) and tissue from the mother plant, which also form a cone around the seed in coniferous plants such as pine and spruce . Seeds are very diverse, and as such there are many terms are used to describe them. A typical seed includes two basic parts: In addition, the endosperm forms
5109-530: Is useful in plant identification; for instance in parts of Southern California and the American Southeast , deciduous and evergreen oak species may grow side by side. Periods of leaf fall often coincide with seasons: winter in the case of cool-climate plants or the dry-season in the case of tropical plants, however there are no deciduous species among tree-like monocotyledonous plants, e.g. palms , yuccas , and dracaenas . The hydrangea hirta
5240-419: Is usually triploid , and is rich in oil or starch , and protein . In gymnosperms, such as conifers , the food storage tissue (also called endosperm) is part of the female gametophyte , a haploid tissue. The endosperm is surrounded by the aleurone layer (peripheral endosperm), filled with proteinaceous aleurone grains. Originally, by analogy with the animal ovum , the outer nucellus layer ( perisperm )
5371-580: The Mississippi River were made from it. In 1911, the food book The Grocer's Encyclopedia noted that a cannery in Holland included in its "vegetables-and-meat" ready-cooked combinations, a "chestnuts and sausages" casserole beside the more classic "beef and onions" and "green peas and veal". This celebrated the chestnut culture that would bring whole villages out in the woods for three weeks each autumn (and keep them busy all winter), and deplored
Chestnut - Misplaced Pages Continue
5502-471: The Pacific Northwest . Due to disease, American chestnut wood almost disappeared from the market for decades, although quantities can still be obtained as reclaimed lumber . Today, they only survive as single trees separated from any others (very rare), and as living stumps , or "stools", with only a few growing enough shoots to produce seeds shortly before dying. This is just enough to preserve
5633-617: The Romagna region, roasted chestnuts are often served with a traditional wine, the Cagnina di Romagna . It is traditional to eat roasted chestnuts in Portugal on St. Martin's Day . Their popularity declined during the last few centuries, partly due to their reputation of "food for poor people". Many people did not want to take chestnut bread as " bread " because chestnut flour does not rise. Some slandered chestnut products in such words as
5764-553: The autumn ; to the shedding of petals , after flowering; and to the shedding of ripe fruit . The antonym of deciduous in the botanical sense is evergreen . Generally, the term "deciduous" means "the dropping of a part that is no longer needed or useful" and the "falling away after its purpose is finished". In plants, it is the result of natural processes. "Deciduous" has a similar meaning when referring to animal parts, such as deciduous antlers in deer , deciduous teeth (baby teeth) in some mammals (including humans); or decidua ,
5895-417: The cone scales as they develop in some species of conifer . Angiosperm (flowering plants) seeds consist of three genetically distinct constituents: (1) the embryo formed from the zygote, (2) the endosperm, which is normally triploid, (3) the seed coat from tissue derived from the maternal tissue of the ovule. In angiosperms, the process of seed development begins with double fertilization , which involves
6026-548: The dormant period. Frosts and snowfalls are beneficial rather than harmful to the trees. The dormant plant is very cold-hardy in Britain, to the Royal Horticultural Society 's H6 hardiness rating, to -20 °C. Chestnut is hardy to USDA zone 5 , which is −29 °C (−20 °F) lower in average minimal temperature than London in zone 9 . The young growth in spring, even on mature plants, however,
6157-456: The dry season or other seasons, depending on variations in rainfall . The converse of deciduous is evergreen , where foliage is shed on a different schedule from deciduous plants, therefore appearing to remain green year round because not all the leaves are shed at the same time. Plants that are intermediate may be called semi-deciduous ; they lose old foliage as new growth begins. Other plants are semi-evergreen and lose their leaves before
6288-412: The embryo , dispersal to a new location, and dormancy during unfavorable conditions. Seeds fundamentally are means of reproduction, and most seeds are the product of sexual reproduction which produces a remixing of genetic material and phenotype variability on which natural selection acts. Plant seeds hold endophytic microorganisms that can perform various functions, the most important of which
6419-421: The flowering plants , the ovary ripens into a fruit which contains the seed and serves to disseminate it. Many structures commonly referred to as "seeds" are actually dry fruits. Sunflower seeds are sometimes sold commercially while still enclosed within the hard wall of the fruit, which must be split open to reach the seed. Different groups of plants have other modifications, the so-called stone fruits (such as
6550-503: The foliage before they are shed and store them in the form of proteins in the vacuoles of parenchyma cells in the roots and the inner bark. In the spring, these proteins are used as a nitrogen source during the growth of new leaves or flowers. Plants with deciduous foliage have advantages and disadvantages compared to plants with evergreen foliage. Since deciduous plants lose their leaves to conserve water or to better survive winter weather conditions, they must regrow new foliage during
6681-424: The legumes (such as beans and peas ), trees such as the oak and walnut , vegetables such as squash and radish , and sunflowers . According to Bewley and Black (1978), Brazil nut storage is in hypocotyl and this place of storage is uncommon among seeds. All gymnosperm seeds are albuminous. The seed coat develops from the maternal tissue, the integuments , originally surrounding the ovule. The seed coat in
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#17327728581716812-416: The peach ) have a hardened fruit layer (the endocarp ) fused to and surrounding the actual seed. Nuts are the one-seeded, hard-shelled fruit of some plants with an indehiscent seed, such as an acorn or hazelnut . The first land plants evolved around 468 million years ago, and reproduced using spores. The earliest seed bearing plants to appear were the gymnosperms , which have no ovaries to contain
6943-613: The American Chestnut Cooperators' Foundation and many others from education, research, and industry sectors contributing to the program) are in the last stages of developing a variety that is as close as possible to the American chestnut, while having incorporated the blight-resistant gene of the Asiatic species. Considering the additional advantage that chestnut trees can be easily grown organically, and assuming
7074-534: The American chestnut species, mainly C. dentata and some others, long before European immigrants introduced their stock to America, and before the arrival of chestnut blight . In some places, such as the Appalachian Mountains , one-quarter of hardwoods were chestnuts. Mature trees often grew straight and branch-free for 50 ft (15 m), up to 100 ft, averaging up to 5 ft in diameter. For three centuries, most barns and homes east of
7205-536: The American chestnuts is not completely resolved, particularly between the chinkapins ( Castanea ozarkensis and Castanea pumila ), which are sometimes considered to be the same species. Genetics have indicated the California native "golden chinkapin" ( Chrysolepis chrysophylla ) is worthy of inclusion in a different genus along with a species from Coastal China. There is also another chestnut, Castanea alabamensis , which may be its own species. The name "chestnut"
7336-844: The Himalayas, Europe and for cultivation purposes in Oceania. They have formed under climatic conditions which have great seasonable temperature variability. Growth occurs during warm summers, leaf drop in autumn, and dormancy during cold winters. These seasonally distinctive communities have diverse life forms that are impacted greatly by the seasonality of their climate, mainly temperature and precipitation rates. These varying and regionally different ecological conditions produce distinctive forest plant communities in different regions. Tropical and subtropical deciduous forest biomes have developed in response not to seasonal temperature variations but to seasonal rainfall patterns. During prolonged dry periods
7467-606: The Indian subcontinent. China has about 300 chestnut cultivars . Moreover, the 'Dandong' chestnut (belonging to the Japanese chestnut C. crenata ) is a major cultivar in Liaoning Province . In South Korea , roasted chestnuts ( gunbam ) are a popular winter snack, and serve as a symbol of abundance in ancestral rituals. Roasted chestnuts are also included in folk songs of Korea , which include " Gunbam Taryeong " ,
7598-471: The United States imports 7.5 million pounds of non-organic chestnuts per year. A study of the sector in 2005 found that U.S. producers are mainly part-timers diversifying an existing agricultural business, or hobbyists. Another recent study indicates that investment in a new plantation takes 13 years to break even, at least within the current Australian market. Starting a small-scale operation requires
7729-434: The bread which "gives a sallow complexion" written in 1770, or in 1841 "this kind of mortar which is called a soup". The last decades' worldwide renewal may have profited from the huge reforestation efforts started in the 1930s in the United States to establish varieties of C. sativa which may be resistant to chestnut blight, as well as to relieve the strain on cereal supplies. The main region in Italy for chestnut production
7860-636: The cells are filled with starch , as for instance cereal grains , or not (non-farinaceous). The endosperm may also be referred to as "fleshy" or "cartilaginous" with thicker soft cells such as coconut , but may also be oily as in Ricinus (castor oil), Croton and Poppy . The endosperm is called "horny" when the cell walls are thicker such as date and coffee , or "ruminated" if mottled, as in nutmeg , palms and Annonaceae . In most monocotyledons (such as grasses and palms ) and some ( endospermic or albuminous ) dicotyledons (such as castor beans )
7991-405: The cells of the outer epidermis enlarge radially and their walls thicken, with nucleus and cytoplasm compressed into the outer layer. these cells which are broader on their inner surface are called palisade cells. In the inner epidermis, the cells also enlarge radially with plate like thickening of the walls. The mature inner integument has a palisade layer, a pigmented zone with 15–20 layers, while
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#17327728581718122-668: The chestnut oak ( Quercus prinus ) and the American beech ( Fagus grandifolia ), both of which are also in the Fagaceae family. Brazil nuts , called "Brasil chestnuts" ( castañas de Brasil in Spanish) or "chestnuts from Pará" ( castanha-do-Pará in Portuguese) are also unrelated. Chestnuts belong to the family Fagaceae , which also includes oaks and beeches . The four main species groups are commonly known as American, European, Chinese, and Japanese chestnuts. The taxonomy of
8253-478: The chestnut tree is practically the sole subsistence of our highlanders", while in 1879 it was said that it almost exclusively fed whole populations for half the year, as "a temporary but complete substitution for cereals". In Britain, boundary records compiled in the reign of King John already showed the famous Tortworth Chestnut in South Gloucestershire , as a landmark; it was also known by
8384-574: The cold in their first winter, and also from mice and squirrels . Chestnuts are considered self-sterile, so at least two trees are needed for pollination. Castanea grows best in a soil with good drainage and adequate moisture. The tree prefers sloping, deep soils; it does not like shallow or heavy soils with impermeable, clay subsoils . The Chinese chestnut prefers a fertile, well-drained soil, but it grows well in fairly dry, rocky, or poor soils. Although Castanea can grow in very acidic soil, and while these soils are reasonably well tolerated,
8515-468: The country has nearly 350 growers, annually producing around 1,200 metric tons of chestnuts, of which 80% come from northeast Victoria . The produce is mostly sold to the domestic fresh fruit market. Chestnuts are slowly gaining popularity in Australia. A considerable increase in production is expected in the next 10 years, due to the increase in commercial plantings during the last 15 to 25 years. By far,
8646-425: The development of brands in the market and everything else being equal, home-grown products would reach higher prices than imports, the high volume of which indicates a market with expanding prospects. As of 2008, the price for chestnuts sold fresh in the shell ranges from $ 1.50/lb ($ 3.30/kg) wholesale to about $ 5/lb ($ 11/kg) retail, depending mainly on the size. The Australian gold rush of the 1850s and 1860s led to
8777-618: The départements of Ardèche , with the famous "Châtaigne d'Ardèche" (A.O.C) Archived 2012-02-27 at the Wayback Machine , of the Var (Eastern Provence), of the Cévennes (Gard and Lozère départements) and of the Lyon region. France annually produces over 1,000 metric tons, but still imports about 8,000 metric tons, mainly from Italy. In Portugal's archipelago of Madeira, chestnut liquor
8908-428: The earliest trees to lose their leaves in the fall. In sub-Arctic climates such as Alaska , leaves begin turning colors as early as August. However, for most temperate regions it takes place in late September through early November and in subtropical climates such as the southern US, it may be November into December. Leaf drop or abscission involves complex physiological signals and changes within plants. When leafout
9039-407: The early 19th century, influential works being: Angiosperm seeds are "enclosed seeds", produced in a hard or fleshy structure called a fruit that encloses them for protection. Some fruits have layers of both hard and fleshy material. In gymnosperms, no special structure develops to enclose the seeds, which begin their development "naked" on the bracts of cones. However, the seeds do become covered by
9170-440: The elaiosomes are eaten. The remainder of the seed, which is hard and inedible to the ants, then germinates either within the nest or at a removal site where the seed has been discarded by the ants. This dispersal relationship is an example of mutualism , since the plants depend upon the ants to disperse seeds, while the ants depend upon the plants seeds for food. As a result, a drop in numbers of one partner can reduce success of
9301-429: The embryo is embedded in the endosperm (and nucellus), which the seedling will use upon germination . In the non-endospermic dicotyledons the endosperm is absorbed by the embryo as the latter grows within the developing seed, and the cotyledons of the embryo become filled with stored food. At maturity, seeds of these species have no endosperm and are also referred to as exalbuminous seeds. The exalbuminous seeds include
9432-701: The fall months and the Western United States as it has more evergreen and fewer deciduous plants, combined with the West Coast and its maritime climate. ( See also : Autumn leaf color ) Most of the Southern Hemisphere lacks deciduous plants due to its milder winters and smaller landmass, most of which is nearer the equator with only far southern South America and the southern island of New Zealand producing distinct fall colors. The beginnings of leaf drop starts when an abscission layer
9563-588: The farmers and landowners to plant four trees yearly, among which was a chestnut tree – plus olive , fig and mulberry trees. Many communities owe their origin and former richness to the ensuing chestnut woods. In France, the marron glacé , a candied chestnut involving 16 different processes in a typically French cooking style, is always served at Christmas and New Year's time. In Modena , Italy, they are soaked in wine before roasting and serving, and are also traditionally eaten on Saint Simon's Day in Tuscany . In
9694-452: The fertilised ovule, an immature plant from which a new plant will grow under proper conditions. The embryo has one cotyledon or seed leaf in monocotyledons , two cotyledons in almost all dicotyledons and two or more in gymnosperms. In the fruit of grains (caryopses) the single monocotyledon is shield shaped and hence called a scutellum . The scutellum is pressed closely against the endosperm from which it absorbs food and passes it to
9825-645: The first few years of their lives deriving energy from the fungi and do not produce green leaves. At up to 55 pounds (25 kilograms) the largest seed is the coco de mer (Lodoicea maldivica). This indicates a 25 Billion fold difference in seed weight. Plants that produce smaller seeds can generate many more seeds per flower, while plants with larger seeds invest more resources into those seeds and normally produce fewer seeds. Small seeds are quicker to ripen and can be dispersed sooner, so autumn all blooming plants often have small seeds. Many annual plants produce great quantities of smaller seeds; this helps to ensure at least
9956-422: The first recorded plantings of European chestnut trees, brought from Europe by settlers. Along the years, most chestnut tree plantations were C. sativa stock, which is still the dominant species. Some of these remain today. Some trees in northern Victoria are around 120 years old and up to 60 m tall. Chestnuts grow well in southwest Western Australia , which has cold winters and warm to hot summers. As of 2008,
10087-553: The flatulence induced by eating too much of it. To the early Christians, chestnuts symbolized chastity . Until the introduction of the potato , whole forest-dwelling communities which had scarce access to wheat flour relied on chestnuts as their main source of carbohydrates . In some parts of Italy, a cake made of chestnuts is used as a substitute for potatoes. In 1583, Charles Estienne and Jean Liébault wrote, "an infinity of people live on nothing else but (the chestnut)". In 1802, an Italian agronomist said of Tuscany that "the fruit of
10218-591: The flowers can be damaged by frost or, in dry season regions, result in water stress on the plant. Spring leafout and fall leaf drop are triggered by a combination of daylight and air temperatures. The exact conditions required will vary with the species, but generally more cold-tolerant genera such as Salix will leaf-out earlier and lose their leaves later, while genera such as Fraxinus and Juglans can only grow in warm, frost-free conditions so they need at least 13 hours of daylight and air temperatures of around 70 °F (21 °C) to leaf out. They will be among
10349-426: The foliage is dropped to conserve water and prevent death from drought. Leaf drop is not seasonally dependent as it is in temperate climates. It can occur any time of year and varies by region of the world. Even within a small local area there can be variations in the timing and duration of leaf drop; different sides of the same mountain and areas that have high water tables or areas along streams and rivers can produce
10480-662: The former land of Baekje. In the Philippines , the endemic talakatak or Philippine chestnut ( Castanopsis philippinensis ) is not cultivated commercially, though its nuts are harvested from the wild and consumed locally. Imported chestnuts (known as kastanyas in Tagalog , from Spanish castañas ) are traditionally sold as street food in the Philippines during the Christmas season . Native Americans were eating
10611-541: The fruit was a local staple food in the early 17th century. These synonyms are or have been in use: Fagus Castanea (used by Linnaeus in first edition of Species Plantarum , 1753), Sardian nut, Jupiter's nut, husked nut, and Spanish chestnut (U.S.). The tree is noted for attracting wildlife. The nuts are an important food for jays , pigeons , wild boar , deer, and squirrels . American and Chinese chinquapins ( C. pumila and C. henryi ) have very small nuts that are an important source of food for wildlife. It has been
10742-467: The fruit, but more often achieve complete opening and release the fruits only after having fallen on the ground; opening is partly due to soil humidity . The chestnut fruit has a pointed end with a small tuft at its tip (called "flame" in Italian ), and at the other end, a hilum – a pale brown attachment scar. In many varieties, the fruit is flattened on one or two sides. It has two skins. The first one
10873-507: The fruits achenes , caryopses , nuts , samaras , and utricles . Other seeds are enclosed in fruit structures that aid wind dispersal in similar ways: Myrmecochory is the dispersal of seeds by ants . Foraging ants disperse seeds which have appendages called elaiosomes (e.g. bloodroot , trilliums , acacias , and many species of Proteaceae ). Elaiosomes are soft, fleshy structures that contain nutrients for animals that eat them. The ants carry such seeds back to their nest, where
11004-408: The fusion of two male gametes with the egg cell and the central cell to form the primary endosperm and the zygote. Right after fertilization, the zygote is mostly inactive, but the primary endosperm divides rapidly to form the endosperm tissue. This tissue becomes the food the young plant will consume until the roots have developed after germination . After fertilization, the ovules develop into
11135-518: The genetic material used to engineer an American chestnut tree with the minimal necessary genetic input from any of the disease-immune Asiatic species. Efforts started in the 1930s are still ongoing to repopulate the country with these trees, in Massachusetts and many places elsewhere in the United States. In the 1970s, geneticist Charles Burnham began back-breeding Asian chestnut into American chestnut populations to confer blight resistance with
11266-446: The grasses, are not distinct structures, but are fused with the fruit wall to form a pericarp .) The testae of both monocots and dicots are often marked with patterns and textured markings, or have wings or tufts of hair. When the seed coat forms from only one layer, it is also called the testa, though not all such testae are homologous from one species to the next. The funiculus abscisses (detaches at fixed point – abscission zone),
11397-629: The ground in the autumn, with the roots emerging from the seed right away and the leaves and stem the following spring. The germ can lose viability soon after ripening and under drying conditions. The superior fruiting varieties among European chestnuts have good size, sweet taste, and easy-to-remove inner skins. American chestnuts are usually very small (around 5 g ), but sweet-tasting with easy-to-remove pellicles. Some Japanese varieties have very large nuts (around 40 g ), with typically difficult-to-remove pellicles. Chinese chestnut pellicles are usually easy to remove, and their sizes vary greatly according to
11528-431: The growing parts. Embryo descriptors include small, straight, bent, curved, and curled. Within the seed, there usually is a store of nutrients for the seedling that will grow from the embryo. The form of the stored nutrition varies depending on the kind of plant. In angiosperms, the stored food begins as a tissue called the endosperm , which is derived from the mother plant and the pollen via double fertilization . It
11659-417: The humid maritime climate and lower overall species diversity . It is also a factor that the continental United States and southern Canada are at a lower latitude than northern Europe, so the sun during the fall months is higher and stronger. This combination of strong sun and cool temperatures leads to more intense fall colors. The Southern United States also has poor fall colors due to warm temperatures during
11790-434: The inner epidermis may remain a single layer, it may also divide to produce two to three layers and accumulates starch, and is referred to as the colourless layer. By contrast, the outer epidermis becomes tanniferous . The inner integument may consist of eight to fifteen layers. As the cells enlarge, and starch is deposited in the outer layers of the pigmented zone below the outer epidermis, this zone begins to lignify, while
11921-444: The innermost layer is known as the fringe layer. In gymnosperms, which do not form ovaries, the ovules and hence the seeds are exposed. This is the basis for their nomenclature – naked seeded plants. Two sperm cells transferred from the pollen do not develop the seed by double fertilization, but one sperm nucleus unites with the egg nucleus and the other sperm is not used. Sometimes each sperm fertilizes an egg cell and one zygote
12052-590: The lack of food diversity in the United States's shop shelves. Soon after that, however, the American chestnuts were nearly wiped out by chestnut blight. The discovery of the blight fungus on some Asian chestnut trees planted on Long Island , New York , was made public in 1904. Within 40 years, the nearly four billion-strong American chestnut population in North America was devastated; only a few clumps of trees remained in Michigan, Wisconsin, California , and
12183-424: The latter example these hairs are the source of the textile crop cotton . Other seed appendages include the raphe (a ridge), wings, caruncles (a soft spongy outgrowth from the outer integument in the vicinity of the micropyle), spines, or tubercles. A scar also may remain on the seed coat, called the hilum , where the seed was attached to the ovary wall by the funicle. Just below it is a small pore, representing
12314-470: The leaves. Rather, they are produced in the foliage in late summer, when sugars are trapped in the leaves after the process of abscission begins. Parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. The New England region of the United States and southeastern Canada tend to produce particularly good autumn colors for this reason, with Europe producing generally poorer colors due to
12445-500: The life length is short. If kept moist, they can be stored in a cool place for a few months, but must be checked regularly for signs of germination. Low temperature prolongs dormancy. Sowing them as soon as ripe is better, either in cold frames or seedbeds outdoors, where they can be left in situ for one to two years before being planted in their permanent positions, or in pots, where the plants can be put out into their permanent positions in summer or autumn. They must be protected from
12576-402: The main area of growth of the embryo, while the lower or micropylar pole produces the stalk-like suspensor that attaches to the micropyle. The suspensor absorbs and manufactures nutrients from the endosperm that are used during the embryo's growth. The main components of the embryo are: Monocotyledonous plants have two additional structures in the form of sheaths. The plumule is covered with
12707-422: The mature seed can be a paper-thin layer (e.g. peanut ) or something more substantial (e.g. thick and hard in honey locust and coconut ), or fleshy as in the sarcotesta of pomegranate . The seed coat helps protect the embryo from mechanical injury, predators, and drying out. Depending on its development, the seed coat is either bitegmic or unitegmic . Bitegmic seeds form a testa from the outer integument and
12838-514: The mechanical layer is the endotegmen, then the seed is endotestal. The exotesta may consist of one or more rows of cells that are elongated and pallisade like (e.g. Fabaceae ), hence 'palisade exotesta'. In addition to the three basic seed parts, some seeds have an appendage, an aril , a fleshy outgrowth of the funicle ( funiculus ), (as in yew and nutmeg ) or an oily appendage, an elaiosome (as in Corydalis ), or hairs (trichomes). In
12969-456: The micropyle of the ovule. Seeds are very diverse in size. The dust-like orchid seeds are the smallest, with about one million seeds per gram; they are often embryonic seeds with immature embryos and no significant energy reserves. Orchids and a few other groups of plants are mycoheterotrophs which depend on mycorrhizal fungi for nutrition during germination and the early growth of the seedling. Some terrestrial orchid seedlings, in fact, spend
13100-668: The mid-1700s to 1947, the sweet chestnut, C. sativa , was widely introduced in the temperate parts of the Indian subcontinent , mainly in the lower to middle Himalayas . They are widely found in British-founded hill stations in northern India , and to a lesser extent in Bhutan and Nepal . They are mainly used as an ornamental tree and are found in almost all British-founded botanical gardens and official governmental compounds (such as larger official residences) in temperate parts of
13231-483: The mid-20th century, most of the US imports are from Southern Italy , with the large, meaty, and richly flavored Sicilian chestnuts being considered among the best quality for bulk sale and supermarket retail. Some imports come from Portugal and France. The next two largest sources of imports are China and South Korea. The French varieties of marrons are highly favored and sold at high prices in gourmet shops. As of 2024,
13362-567: The minimum difference in genes. In the 1950s, the Dunstan chestnut was developed in Greensboro, N.C., and constitutes the majority of blight-free chestnuts produced in the United States annually. Today, the demand for the nut outstrips supply . The United States imported 4,056 metric tons of European in-shell chestnuts worth $ 10 million in 2007. The U.S. chestnut industry is in its infancy, producing less than 1% of total world production. Since
13493-399: The monocotyledons, ten in the dicotyledons, and two in the gymnosperms (linear and spatulate). This classification is based on three characteristics: embryo morphology, amount of endosperm and the position of the embryo relative to the endosperm. In endospermic seeds, there are two distinct regions inside the seed coat, an upper and larger endosperm and a lower smaller embryo. The embryo is
13624-489: The most common species in Australia is the European chestnut, but small numbers of the other species, as well as some hybrids, have been planted. The Japanese chestnut ( C. crenata ) does well in wet and humid weather and in hot summers (about 30 °C); and was introduced to New Zealand in the early 1900s, more so in the upper North Island region. Chestnuts produce a better crop when subjected to chill temperatures during
13755-410: The need for light. In 2020, world production of cultivated chestnuts was 2,322 tonnes , led by China with 75% of the total (table). Deciduous In the fields of horticulture and botany , the term deciduous ( / d ɪ ˈ s ɪ dʒ u . ə s / ) means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves , usually in
13886-566: The need for sun for optimal growth. The butt of the tree is sometimes painted with white paint to protect the tree from sunburn until it has developed enough canopy . Wide spacing between the trees encourages low, broad crowns with maximum exposure to sunshine to increase fruit production. Where chestnut trees touch, virtually no fruit is produced. Current industrial planting spacings can range from 7 x 7 to 20 x 20 m. The closer plantings, which are more popular, mean quicker increases in short-term production, but heavy pruning or even tree removal
14017-412: The next growing season, retaining some during winter or dry periods. Many deciduous plants flower during the period when they are leafless, as this increases the effectiveness of pollination . The absence of leaves improves wind transmission of pollen for wind-pollinated plants and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as
14148-783: The next suitable growing season; this uses resources which evergreens do not need to expend. Evergreens suffer greater water loss during the winter and they also can experience greater predation pressure , especially when small. Deciduous trees experience much less branch and trunk breakage from glaze ice storms when leafless, and plants can reduce water loss due to the reduction in availability of liquid water during cold winter days. Losing leaves in winter may reduce damage from insects; repairing leaves and keeping them functional may be more costly than just losing and regrowing them. Removing leaves also reduces cavitation which can damage xylem vessels in plants. This then allows deciduous plants to have xylem vessels with larger diameters and therefore
14279-478: The nuts react as fresh fruits (not as nuts). They should be kept cool at all times, including in shops when on display for sale. To preserve their freshness for a few months with no artificial refrigeration, the chestnuts can be soaked in cold water for about 20 hours immediately after harvest , after which they are dried in the shade, then layered in dry sand. Chestnuts behave similarly to seeds in that they produce very little ethylene , and their respiration rate
14410-403: The optimal conditions for survival of the resulting seedling; the second is spreading germination of a batch of seeds over time so a catastrophe (e.g. late frosts, drought, herbivory ) does not result in the death of all offspring of a plant ( bet-hedging ). Seed dormancy is defined as a seed failing to germinate under environmental conditions optimal for germination, normally when the environment
14541-477: The other. In South Africa , the Argentine ant ( Linepithema humile ) has invaded and displaced native species of ants. Unlike the native ant species, Argentine ants do not collect the seeds of Mimetes cucullatus or eat the elaiosomes. In areas where these ants have invaded, the numbers of Mimetes seedlings have dropped. Seed dormancy has two main functions: the first is synchronizing germination with
14672-410: The parent. The large, heavy root allows the seed to penetrate into the ground when it falls. Many garden plant seeds will germinate readily as soon as they have water and are warm enough; though their wild ancestors may have had dormancy, these cultivated plants lack it. After many generations of selective pressure by plant breeders and gardeners, dormancy has been selected out. For annuals , seeds are
14803-542: The place's name. Still others take the name as coming from the Greek name of Sardis glans (Sardis acorn) – Sardis being the capital of Lydia , Asia Minor, from where the fruit had spread. The name is cited twice in the King James Version of the Bible. In one instance, Jacob puts peeled twigs in the water troughs to promote healthy offspring of his livestock. Although it may indicate another tree, it indicates
14934-469: The plant, the cells of the abscission layer remain connected; in autumn, or when under stress, the auxin flow from the leaf decreases or stops, triggering cellular elongation within the abscission layer. The elongation of these cells breaks the connection between the different cell layers, allowing the leaf to break away from the plant. It also forms a layer that seals the break, so the plant does not lose sap. Some trees, particularly oaks and beeches, exhibit
15065-455: The plumule and radicle, acting as a protective covering. The maturing ovule undergoes marked changes in the integuments, generally a reduction and disorganization but occasionally a thickening. The seed coat forms from the two integuments or outer layers of cells of the ovule, which derive from tissue from the mother plant, the inner integument forms the tegmen and the outer forms the testa . (The seed coats of some monocotyledon plants, such as
15196-523: The potential increase in productivity with mixed stands and plantations, compared to plots of only one species. The relative yield total values of the mixed plantings steadily increase with time. C. sativa responds well to competitive pressure from Pseudotsuga menziesii , the latter also showing a higher productivity. C. dentata seedlings in Ohio reforestation efforts are best achieved by planting them in places with little or no arboreous land cover, because of
15327-466: The preferred range is from pH 5.5-6.0. It does not grow well on alkaline soils , such as chalk , but thrives on soils such as those derived from granite , sandstone , or schist . On alkaline soils, chestnut trees can be grown by grafting them onto oak rootstocks. Recently cleared land is best avoided to help resist the root rot , Armillaria mellia . Castanea likes a full sun position. An experiment with C. dentata seedlings in Ohio confirmed
15458-405: The rate of germination. This is given as a percent of germination over a certain amount of time, 90% germination in 20 days, for example. 'Dormancy' is covered above; many plants produce seeds with varying degrees of dormancy, and different seeds from the same fruit can have different degrees of dormancy. It's possible to have seeds with no dormancy if they are dispersed right away and do not dry (if
15589-524: The same name of "Great Chestnut of Tortworth" in the days of Stephen . This tree measured over 50 ft (15 m) in circumference at 5 ft (1.5 m) from the ground in 1720. The Hundred Horse Chestnut in the chestnut forests on Mount Etna is the oldest living chestnut tree and is said to be even larger. Chestnut trees particularly flourish in the Mediterranean basin . In 1584, the governor of Genoa , which dominated Corsica , ordered all
15720-415: The scar forming an oval depression, the hilum . Anatropous ovules have a portion of the funiculus that is adnate (fused to the seed coat), and which forms a longitudinal ridge, or raphe , just above the hilum. In bitegmic ovules (e.g. Gossypium described here) both inner and outer integuments contribute to the seed coat formation. With continuing maturation the cells enlarge in the outer integument. While
15851-413: The seed affects its health and germination ability: since the seed has a living embryo, over time cells die and cannot be replaced. Some seeds can live for a long time before germination, while others can only survive for a short period after dispersal before they die. Seed vigor is a measure of the quality of seed, and involves the viability of the seed, the germination percentage, germination rate, and
15982-405: The seeds. The ovule consists of a number of components: The shape of the ovules as they develop often affects the final shape of the seeds. Plants generally produce ovules of four shapes: the most common shape is called anatropous , with a curved shape. Orthotropous ovules are straight with all the parts of the ovule lined up in a long row producing an uncurved seed. Campylotropous ovules have
16113-507: The seeds. They arose during the late Devonian period (416 million to 358 million years ago). From these early gymnosperms, seed ferns evolved during the Carboniferous period (359 to 299 million years ago); they had ovules that were borne in a cupule, which consisted of groups of enclosing branches likely used to protect the developing seed. Published literature about seed storage, viability and its hygrometric dependence began in
16244-430: The sides and develop broad, rounded, dense crowns at maturity. The foliage of the European and American species has striking yellow autumn coloring. Its bark is smooth when young, of a vinous maroon or red-brown color for the American chestnut, grey for the European chestnut. With age, American species' bark becomes grey and darker, thick, and deeply furrowed ; the furrows run longitudinally, and tend to twist around
16375-693: The smallest species of chinkapins, often shrubby, to the giant of past American forests, C. dentata that could reach 60 metres (200 feet). Between these extremes are found the Japanese chestnut ( C. crenata ) at 10 m average; followed by the Chinese chestnut ( C. mollissima ) at about 15 m , then the European chestnut ( C. sativa ) around 30 m . The Chinese and more so the Japanese chestnuts are both often multileadered and wide-spreading, whereas European and especially American species tend to grow very erect when planted among others, with little tapering of their columnar trunks , which are firmly set and massive. When standing on their own, they spread on
16506-420: The soil surface is the next phase of the plant's growth and is called seedling establishment. Three fundamental conditions must exist before germination can occur. (1) The embryo must be alive, called seed viability. (2) Any dormancy requirements that prevent germination must be overcome. (3) The proper environmental conditions must exist for germination. Far red light can prevent germination. Seed viability
16637-507: The strength of the seedlings produced. The germination percentage is simply the proportion of seeds that germinate from all seeds subject to the right conditions for growth. The germination rate is the length of time it takes for the seeds to germinate. Germination percentages and rates are affected by seed viability, dormancy and environmental effects that impact on the seed and seedling. In agriculture and horticulture quality seeds have high viability, measured by germination percentage plus
16768-597: The trunk as the tree ages, sometimes reminiscent of a large cable with twisted strands. The leaves are simple, ovate or lanceolate , 10–30 cm long and 4–10 cm wide, with sharply pointed, widely spaced teeth, with shallow rounded sinuates between. The flowers follow the leaves, appearing in late spring or early summer or into July. They are arranged in long catkins of two kinds, with both kinds being borne on every tree. Some catkins are made of only male flowers, which mature first. Each flower has eight stamens , or 10 to 12 for C. mollissima . The ripe pollen carries
16899-464: The uterine lining that sheds off after birth. In botany and horticulture , deciduous plants , including trees , shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year. This process is called abscission . In some cases leaf loss coincides with winter—namely in temperate or polar climates . In other parts of the world, including tropical, subtropical, and arid regions, plants lose their leaves during
17030-442: The varieties, although usually smaller than the Japanese chestnut. The unrelated horse chestnuts (genus Aesculus ) are not true chestnuts, but are named for producing nuts of similar appearance that are mildly poisonous to humans. True chestnuts should also not be confused with water chestnuts , which are tubers of an aquatic herbaceous plant in the sedge family Cyperaceae . Other species commonly mistaken for chestnut trees are
17161-426: Was referred to as albumen , and the inner endosperm layer as vitellus. Although misleading, the term began to be applied to all the nutrient matter. This terminology persists in referring to endospermic seeds as "albuminous". The nature of this material is used in both describing and classifying seeds, in addition to the embryo to endosperm size ratio. The endosperm may be considered to be farinaceous (or mealy) in which
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