55-401: Ciconia abdimii Ciconia boyciana Ciconia ciconia Ciconia episcopus Ciconia maguari Ciconia microscelis Ciconia nigra Ciconia stormi Dissoura Euxenura Sphenorhynchus Ciconia ( / s ɪ ˈ k oʊ . n i . ə / sih- KOH -nee-uh ; Classical Latin : [kiˈkoː.ni.a] ) is a genus of birds in the stork family. Six of
110-473: A monotreme , appeared. The predatory South American phorusrhacids were rare in this time; among the last was Titanis , a large phorusrhacid that migrated to North America and rivaled mammals as top predator. Other birds probably evolved at this time, some modern (such as the genera Cygnus , Bubo , Struthio and Corvus ), some now extinct. Alligators and crocodiles died out in Europe as
165-614: A rock shelter on Réunion was probably of a bird taken there as food by early settlers; no known account mentions the presence of storks on the Mascarenes , and while this subfossil was initially believed to be from a stork, it is today assigned to the Réunion ibis ( Threskiornis solitarius ) which is quite similar to storks osteologically and was not yet described when the bone was discovered (Cowles, 1994). Ciconia abdimii Abdim's stork ( Ciconia abdimii ), also known as
220-462: A duller, browner version of the adult. Depending on species, breeding can be in solitary pairs or colonies . Pairs usually stay together for life. They typically build large stick nests in trees, although the Abdim's stork sometimes will nest on cliffs, the maguari stork will nest on the ground and at least three species will construct their nests on human habitations. One of these, the white stork ,
275-406: A large neocortex relative to body mass and stone tool manufacture. Improvements in dating methods and in the use of climate proxies have provided scientists with the means to test hypotheses of the evolution of human ancestors. Early hypotheses of the evolution of human traits emphasized the selective pressures produced by particular habitats. For example, many scientists have long favored
330-429: A particular habitat. The change to a cooler, drier, more seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide. Deciduous forests proliferated, coniferous forests and tundra covered much of the north, and grasslands spread on all continents (except Antarctica). Eastern Africa in particular saw a huge expansion of C 4 grasslands. Tropical forests were limited to
385-968: A tight band around the equator, and in addition to dry savannahs , deserts appeared in Asia and Africa. Both marine and continental faunas were essentially modern, although continental faunas were a bit more primitive than today. The land mass collisions meant great migration and mixing of previously isolated species, such as in the Great American Interchange . Herbivores got bigger, as did specialized predators. In North America, rodents , large mastodons and gomphotheres , and opossums continued successfully, while hoofed animals ( ungulates ) declined, with camel , deer , and horse all seeing populations recede. Three-toed horses ( Nannippus ), oreodonts , protoceratids , and chalicotheres became extinct. Borophagine dogs and Agriotherium became extinct, but other carnivores including
440-552: Is a key finding of research into a lake-sediment core obtained in Eastern Siberia, which is of exceptional importance because it has provided the longest continuous late Cenozoic land-based sedimentary record thus far. During the late Zanclean, Italy remained relatively warm and humid. Central Asia became more seasonal during the Pliocene, with colder, drier winters and wetter summers, which contributed to an increase in
495-589: Is administered by the Association of Zoos and Aquariums and in Europe by the European Association of Zoos and Aquaria . Ciconia abdimii is a black stork with grey legs, red knees and feet, grey bill and white underparts. It has red facial skin in front of the eye and blue skin near the bill in breeding season. It is the smallest species of stork, at 73 cm (29 in) and a weight of just over 1 kg (2.2 lb). The Abdim's stork
550-400: Is mostly insectivorous , feeding on locusts , caterpillars and other large insects, although these birds will also eat small reptiles, amphibians, mice, crabs and eggs. The female lays two to three eggs and is slightly smaller than the male. Pliocene The Pliocene ( / ˈ p l aɪ . ə s iː n , ˈ p l aɪ . oʊ -/ PLY -ə-seen, PLY -oh- ; also Pleiocene ) is
605-627: Is no evidence that the population is breeding and may only persist due to continuing releases or escapes. [1] Archived 2018-12-13 at the Wayback Machine Widespread and common throughout its large range, Abdim's stork is evaluated as Least Concern on the IUCN Red List of Threatened Species . It is the subject of several nationally coordinated breeding programs: in the United States, the plan for this species
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#1732780182819660-406: Is probably the best known of all storks, with a wealth of legend and folklore associated with this familiar summer visitor to Europe. These storks feed on frogs , insects , fish , crustaceans , small birds, lizards and rodents . They fly with the neck outstretched, like most other storks, but unlike herons which retract their neck in flight. The migratory species like the white stork and
715-588: The Hardangervidda plateau to 1200 m in the Early Pliocene. In Southern Sweden similar movements elevated the South Swedish highlands leading to a deflection of the ancient Eridanos river from its original path across south-central Sweden into a course south of Sweden. The Pliocene is bookended by two significant events in the evolution of human ancestors. The first is the appearance of
770-686: The Isthmus of Panama during the Pliocene, making possible the Great American Interchange and bringing a nearly complete end to South America's distinctive native ungulate fauna, though other South American lineages like its predatory mammals were already extinct by this point and others like xenarthrans continued to do well afterwards. The formation of the Isthmus had major consequences on global temperatures, since warm equatorial ocean currents were cut off and an Atlantic cooling cycle began, with cold Arctic and Antarctic waters decreasing temperatures in
825-487: The Latin word for "stork", and was originally recorded in the works of Horace and Ovid . The Abdim's stork is the smallest of the family, but other species in the genus are generally medium-sized storks, with long legs and a long thick bill. The members of this genus are more variable in plumage than other stork genera, but all species are black (at least to the wings) and white (at least underparts or neck). Juveniles are
880-571: The Netherlands the Pliocene is divided into these stages (old to young): Brunssumian C, Reuverian A, Reuverian B, Reuverian C, Praetiglian , Tiglian A, Tiglian B, Tiglian C1-4b, Tiglian C4c, Tiglian C5, Tiglian C6 and Eburonian . The exact correlations between these local stages and the International Commission on Stratigraphy (ICS) stages is not established. During the Pliocene epoch (5.3 to 2.6 million years ago (Ma)),
935-478: The North Atlantic and North Pacific Ocean beds. Mid-latitude glaciation was probably underway before the end of the epoch. The global cooling that occurred during the Pliocene may have accelerated on the disappearance of forests and the spread of grasslands and savannas. During the Pliocene the earth climate system response shifted from a period of high frequency-low amplitude oscillation dominated by
990-648: The Northern Hemisphere . Fossil members of the genus include: A distal radius in Late Pleistocene deposits of San Josecito Cavern (Mexico) may belong in this genus or in Mycteria ; it is smaller than that of any known American stork, Ciconia or otherwise. The proposed fossil genus Prociconia from Brazil , also of Late Pleistocene age, may be a junior synonym of either this genus or Jabiru . A distal tarsometatarsus found in
1045-604: The Sahel to South Africa, being absent mainly from forests, dense woodlands and deserts. A smaller populations occurs in Yemen. It breeds colonially in trees, on cliffs or rooftops in the northern half of its range (north of the Equator ) during the wet season from May to August, migrating to eastern and southern Africa for the remainder of the year. This stork has escaped or been deliberately released into Florida , U.S., but there
1100-790: The black stork soar on broad wings and rely on thermals of hot air for sustained long distance flight. Since thermals only form over land, these storks, like large raptors , must cross the Mediterranean at the narrowest points, and many of these birds can be seen going through the Straits of Gibraltar and the Bosphorus on migration. The genus contains eight extant species: [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] [REDACTED] The fossil record of
1155-719: The epoch in the geologic time scale that extends from 5.33 to 2.58 million years ago (Ma). It is the second and most recent epoch of the Neogene Period in the Cenozoic Era . The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene,
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#17327801828191210-497: The hominin Australopithecus anamensis in the early Pliocene, around 4.2 million years ago. The second is the appearance of Homo , the genus that includes modern humans and their closest extinct relatives, near the end of the Pliocene at 2.6 million years ago. Key traits that evolved among hominins during the Pliocene include terrestrial bipedality and, by the end of the Pliocene, encephalized brains (brains with
1265-634: The savannah hypothesis . This proposes that the evolution of terrestrial bipedality and other traits was an adaptive response to Pliocene climate change that transformed forests into more open savannah . This was championed by Grafton Elliot Smith in his 1924 book, The Evolution of Man , as "the unknown world beyond the trees", and was further elaborated by Raymond Dart as the killer ape theory . Other scientists, such as Sherwood L. Washburn , emphasized an intrinsic model of hominin evolution. According to this model, early evolutionary developments triggered later developments. The model placed little emphasis on
1320-493: The weasel family diversified, and dogs and short-faced bears did well. Ground sloths , huge glyptodonts , and armadillos came north with the formation of the Isthmus of Panama. The latitudinal diversity gradient among terrestrial North American mammals became established during this epoch some time after 4 Ma. In Eurasia rodents did well, while primate distribution declined. Elephants , gomphotheres and stegodonts were successful in Asia (the largest land mammals of
1375-652: The white-bellied stork , is a stork belonging to the family Ciconiidae . It is the smallest species of stork, feeds mostly on insects, and is found widely in open habitats in Sub-Saharan Africa and in Yemen . The common name commemorates the Turkish Governor of Wadi Halfa in Sudan , Bey El-Arnaut Abdim (1780–1827). Abdim's stork is found widely in open habitats throughout Sub-Saharan Africa, from
1430-469: The 41,000-year period of Earth's obliquity to one of low-frequency, high-amplitude oscillation dominated by the 100,000-year period of the orbital eccentricity characteristic of the Pleistocene glacial-interglacial cycles. During the late Pliocene and early Pleistocene, 3.6 to 2.6 Ma, the Arctic was much warmer than it is at the present day (with summer temperatures some 8 °C warmer than today). That
1485-595: The Earth's climate became cooler and drier, as well as more seasonal, marking a transition between the relatively warm Miocene to the cooler Pleistocene . However, the beginning of the Pliocene was marked by an increase in global temperatures relative to the cooler Messinian . This increase was related to the 1.2 million year obliquity amplitude modulation cycle . By 3.3-3.0 Ma, during the Mid-Piacenzian Warm Period (mPWP), global average temperature
1540-510: The Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in the Pleistocene. As with other older geologic periods, the geological strata that define the start and end are well-identified but the exact dates of the start and end of the epoch are slightly uncertain. The boundaries defining the Pliocene are not set at an easily identified worldwide event but rather at regional boundaries between
1595-628: The Pliocene contains the Dacian (roughly equal to the Zanclean) and Romanian (roughly equal to the Piacenzian and Gelasian together) stages. As usual in stratigraphy, there are many other regional and local subdivisions in use. In Britain , the Pliocene is divided into the following stages (old to young): Gedgravian, Waltonian , Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian, Pre-Pastonian or Baventian, Pastonian and Beestonian . In
1650-566: The Pliocene were such proboscideans as Deinotherium , Anancus , and Mammut borsoni , ) though proboscidean diversity declined significantly during the Late Pliocene. Hyraxes migrated north from Africa. Horse diversity declined, while tapirs and rhinos did fairly well. Bovines and antelopes were successful; some camel species crossed into Asia from North America. Hyenas and early saber-toothed cats appeared, joining other predators including dogs, bears, and weasels. Africa
1705-546: The Western Atlantic, assemblages of bivalves exhibited remarkable stasis with regards to their basal metabolic rates throughout the various climatic changes of the Pliocene. The Pliocene was a high water mark for species diversity among Caribbean corals. From 5 to 2 Ma, coral species origination rates were relatively high in the Caribbean, although a noticeable extinction event and drop in diversity occurred at
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1760-544: The abundance of C 4 plants across the region. In the Loess Plateau , δ13C values of occluded organic matter increased by 2.5% while those of pedogenic carbonate increased by 5% over the course of the Late Miocene and Pliocene, indicating increased aridification. Further aridification of Central Asia was caused by the development of Northern Hemisphere glaciation during the Late Pliocene. A sediment core from
1815-735: The climate cooled. Venomous snake genera continued to increase as more rodents and birds evolved. Rattlesnakes first appeared in the Pliocene. The modern species Alligator mississippiensis , having evolved in the Miocene, continued into the Pliocene, except with a more northern range; specimens have been found in very late Miocene deposits of Tennessee . Giant tortoises still thrived in North America, with genera like Hesperotestudo . Madtsoid snakes were still present in Australia. The amphibian order Allocaudata became extinct. In
1870-475: The early-mid Pliocene (5 Ma – 3 Ma), after three-million-year intervals with modern or glacial ice volume became longer and collapse occurs only at times when warmer global temperature coincide with strong austral summer insolation anomalies. Continents continued to drift , moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations. South America became linked to North America through
1925-570: The end of this interval. Oceans continued to be relatively warm during the Pliocene, though they continued cooling. The Arctic ice cap formed , drying the climate and increasing cool shallow currents in the North Atlantic. Deep cold currents flowed from the Antarctic. The formation of the Isthmus of Panama about 3.5 million years ago cut off the final remnant of what was once essentially a circum-equatorial current that had existed since
1980-549: The first time since the Cretaceous , with North American rodents and primates mixing with southern forms. Litopterns and the notoungulates , South American natives, were mostly wiped out, except for the macrauchenids and toxodonts , which managed to survive. Small weasel-like carnivorous mustelids , coatis and short-faced bears migrated from the north. Grazing glyptodonts , browsing giant ground sloths and smaller caviomorph rodents , pampatheres , and armadillos did
2035-542: The genus is extensive, indicating that Ciconia storks were once more widespread than they are today. Although the known material tends to suggest that the genus evolved around the Atlantic, possibly in western Europe or Africa, the comparative lack of fossil sites in Asia makes this assumption not well-founded presently. All that can be said is that by the Early Pliocene , Ciconia was widespread at least all over
2090-413: The influence of seasonal variation. More recent research has emphasized the variability selection hypothesis, which proposes that variability in climate fostered development of hominin traits. Improved climate proxies show that the Pliocene climate of east Africa was highly variable, suggesting that adaptability to varying conditions was more important in driving hominin evolution than the steady pressure of
2145-555: The northern South China Sea shows an increase in dust storm activity during the middle Pliocene. The South Asian Summer Monsoon (SASM) increased in intensity after 2.95 Ma, likely because of enhanced cross-equatorial pressure caused by the reorganisation of the Indonesian Throughflow. In the south-central Andes , an arid period occurred from 6.1 to 5.2 Ma, with another occurring from 3.6 to 3.3 Ma. These arid periods are coincident with global cold periods, during which
2200-754: The now-separated Atlantic Ocean. Africa 's collision with Europe formed the Mediterranean Sea , cutting off the remnants of the Tethys Ocean . The border between the Miocene and the Pliocene is also the time of the Messinian salinity crisis . During the Late Pliocene, the Himalayas became less active in their uplift, as evidenced by sedimentation changes in the Bengal Fan . The land bridge between Alaska and Siberia ( Beringia )
2255-512: The onset of continental glaciation in the Arctic, suggesting that vegetation changes in Australia during the Pliocene behaved similarly to during the Late Pleistocene and were likely characterised by comparable cycles of aridity and humidity. The equatorial Pacific Ocean sea surface temperature gradient was considerably lower than it is today. Mean sea surface temperatures in the east were substantially warmer than today but similar in
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2310-400: The opposite, migrating to the north and thriving there. The marsupials remained the dominant Australian mammals, with herbivore forms including wombats and kangaroos , and the huge Diprotodon . Carnivorous marsupials continued hunting in the Pliocene, including dasyurids , the dog-like thylacine and cat-like Thylacoleo . The first rodents arrived in Australia. The modern platypus ,
2365-628: The position of the Southern Hemisphere westerlies shifted northward and disrupted the South American Low Level Jet, which brings moisture to southeastern South America. From around 3.8 Ma to about 3.3 Ma, North Africa experienced an extended humid period. In northwestern Africa, tropical forests extended up to Cape Blanc during the Zanclean until around 3.5 Ma. During the Piacenzian, from about 3.5 to 2.6 Ma,
2420-514: The recent", referring to the essentially modern marine mollusc fauna. In the official timescale of the ICS , the Pliocene is subdivided into two stages . From youngest to oldest they are: The Piacenzian is sometimes referred to as the Late Pliocene, whereas the Zanclean is referred to as the Early Pliocene. In the system of In the Paratethys area (central Europe and parts of western Asia)
2475-499: The region was forested at irregular intervals and contained a significant Saharan palaeoriver until 3.35 Ma, when trade winds began to dominate over fluvial transport of pollen. Around 3.26 Ma, a strong aridification event that was followed by a return to more humid conditions, which was itself followed by another aridification around 2.7 Ma. From 2.6 to 2.4 Ma, vegetation zones began repeatedly shifting latitudinally in response to glacial-interglacial cycles. The climate of eastern Africa
2530-527: The scene. Bears, dogs and weasels (originally from North America) joined cats, hyenas and civets as the African predators, forcing hyenas to adapt as specialized scavengers. Most mustelids in Africa declined as a result of increased competition from the new predators, although Enhydriodon omoensis remained an unusually successful terrestrial predator. South America was invaded by North American species for
2585-573: The seven living species occur in the Old World , but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times. The genus was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the white stork ( Ciconia ciconia ) as the type species . The genus name is
2640-400: The surrounding environment. Anthropologists tended to focus on intrinsic models while geologists and vertebrate paleontologists tended to put greater emphasis on habitats. Alternatives to the savanna hypothesis include the woodland/forest hypothesis, which emphasizes the evolution of hominins in closed habitats, or hypotheses emphasizing the influence of colder habitats at higher latitudes or
2695-514: The warmer Miocene and the relatively cooler Pleistocene. The upper boundary was set at the start of the Pleistocene glaciations. Charles Lyell (later Sir Charles) gave the Pliocene its name in Principles of Geology (volume 3, 1833). The word pliocene comes from the Greek words πλεῖον ( pleion , "more") and καινός ( kainos , "new" or "recent") and means roughly "continuation of
2750-505: The west. This condition has been described as a permanent El Niño state, or “El Padre.” Several mechanisms have been proposed for this pattern, including increased tropical cyclone activity . The extent of the West Antarctic Ice Sheet oscillated at the 40 kyr period of Earth's obliquity. Ice sheet collapse occurred when the global average temperature was 3 °C warmer than today and carbon dioxide concentration
2805-470: Was 2–3 °C higher than today, while carbon dioxide levels were the same as today (400 ppm). Global sea level was about 25 m higher, though its exact value is uncertain. The northern hemisphere ice sheet was ephemeral before the onset of extensive glaciation over Greenland that occurred in the late Pliocene around 3 Ma. The formation of an Arctic ice cap is signaled by an abrupt shift in oxygen isotope ratios and ice-rafted cobbles in
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#17327801828192860-511: Was at 400 ppmv. This resulted in open waters in the Ross Sea . Global sea-level fluctuation associated with ice-sheet collapse was probably up to 7 meters for the west Antarctic and 3 meters for the east Antarctic. Model simulations are consistent with reconstructed ice-sheet oscillations and suggest a progression from a smaller to a larger West Antarctic ice sheet in the last 5 million years. Intervals of ice sheet collapse were much more common in
2915-486: Was dominated by hoofed animals, and primates continued their evolution, with australopithecines (some of the first hominins ) and baboon-like monkeys such as the Dinopithecus appearing in the late Pliocene. Rodents were successful, and elephant populations increased. Cows and antelopes continued diversification and overtook pigs in numbers of species. Early giraffes appeared. Horses and modern rhinos came onto
2970-452: Was first flooded near the start of the Pliocene, allowing marine organisms to spread between the Arctic and Pacific Oceans. The bridge would continue to be periodically flooded and restored thereafter. Pliocene marine formations are exposed in northeast Spain , southern California , New Zealand , and Italy . During the Pliocene parts of southern Norway and southern Sweden that had been near sea level rose. In Norway this rise elevated
3025-545: Was very similar to what it is today. Unexpectedly, the expansion of grasslands in eastern Africa during this epoch appears to have been decoupled from aridification and not caused by it, as evidenced by their asynchrony. Southwestern Australia hosted heathlands , shrublands , and woodlands with a greater species diversity compared to today during the Middle and Late Pliocene. Three different aridification events occurred around 2.90, 2.59, and 2.56 Ma, and may have been linked to
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