The Permian ( / ˈ p ɜːr m i . ə n / PUR -mee-ən ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period 298.9 million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the sixth and last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison , who named it after the region of Perm in Russia .
99-712: The Batteriekopf is a peak in the Vosges Mountains, reaching an altitude of 1,311 meters. This mountain borders the Route des Crêtes and is located immediately south of the Rothenbachkopf , spanning the municipalities of Wildenstein and Metzeral . Part of the western slope of Batteriekopf is included in the Hautes-Chaumes du Rothenbach regional nature reserve which aims to preserve the summit lawns. This article related to geography of France
198-648: A Triassic rose sandstone) are embedded sometimes up to more than 500 m (1,600 ft) in thickness. The Lower Vosges in the north are dislocated plates of various sandstones, ranging from 300 to 600 m (1,000 to 2,000 ft) high. The Vosges are very similar to the corresponding range of the Black Forest across the Rhine since both lie within the same degrees of latitude, have similar geological formations and are characterized by forests on their lower slopes, above which are open pastures and rounded summits of
297-630: A city in Perm Krai. The stage was introduced by Alexandr Antonovich Stukenberg in 1890. The Kungurian currently lacks a defined GSSP. Recent proposals have suggested the appearance of Neostreptognathodus pnevi as the lower boundary. The Guadalupian Series is named after the Guadalupe Mountains in Texas and New Mexico, where extensive marine sequences of this age are exposed. It was named by George Herbert Girty in 1902. The Roadian
396-685: A coal swamp community, has an upper canopy consisting of lycopsid tree Sigillaria , with a lower canopy consisting of Marattialean tree ferns, and Noeggerathiales. Early conifers appeared in the Late Carboniferous, represented by primitive walchian conifers, but were replaced with more derived voltzialeans during the Permian. Permian conifers were very similar morphologically to their modern counterparts, and were adapted to stressed dry or seasonally dry climatic conditions. The increasing aridity, especially at low latitudes, facilitated
495-604: A dramatic increase in diversification during the Early Permian. Towards the end of the Permian, there was a substantial drop in both origination and extinction rates. The dominant insects during the Permian Period were early representatives of Paleoptera , Polyneoptera , and Paraneoptera . Palaeodictyopteroidea , which had represented the dominant group of insects during the Carboniferous, declined during
594-553: A forested region. Forests were cleared for agriculture, livestock and early industrial factories (such as charcoal works and glassworks ) and the water mills used water power . Concentrations of settlement and immigration took place and not only in areas where minerals were found. In the mining area of the Lièpvrette [ fr ] valley, for example, there was an influx of Saxon miners and mining specialists. From time to time, wars, plagues and religious conflicts saw
693-634: A geological point of view, a graben at the beginning of the Paleogene period caused the formation of Alsace and the uplift of the bedrock plates of the Vosges, in eastern France , and those in the Black Forest , in Germany . From a scientific view, the Vosges Mountains are not mountains as such, but rather the western edge of the unfinished Alsatian graben, stretching continuously as part of
792-540: A peak of diversity in the Cisuralian, with a substantial decline during the Guadalupian-Lopingian following Olson's extinction, with the family diversity dropping below Carboniferous levels. Embolomeres , a group of aquatic crocodile-like limbed vertebrates that are reptilliomorphs under some phylogenies. They previously had their last records in the Cisuralian, are now known to have persisted into
891-525: A protective cover, over plants such as ferns that disperse spores in a wetter environment. The first modern trees ( conifers , ginkgos and cycads ) appeared in the Permian. Three general areas are especially noted for their extensive Permian deposits—the Ural Mountains (where Perm itself is located), China, and the southwest of North America, including the Texas red beds. The Permian Basin in
990-524: A rather uniform altitude. Both areas exhibit steeper slopes towards the Rhine and a more gradual descent on the other side. Both the Vosges and the Black Forest were formed by isostatic uplift in response to the opening of the Rhine Graben , a major extensional basin. When such basins form, the thinning of the crust causes uplift immediately adjacent to the basin, decreasing with distance from
1089-588: A single formation (a stratotype ) identifying the lower boundary of the stage. The ages of the Permian, from youngest to oldest, are: For most of the 20th century, the Permian was divided into the Early and Late Permian, with the Kungurian being the last stage of the Early Permian. Glenister and colleagues in 1992 proposed a tripartite scheme, advocating that the Roadian-Capitanian was distinct from
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#17327901193011188-534: Is a stub . You can help Misplaced Pages by expanding it . Vosges The Vosges ( / v oʊ ʒ / VOHZH , French: [voʒ] ; German : Vogesen [voˈɡeːzn̩] ; Franconian and Alemannic German : Vogese ) are a range of medium mountains in Eastern France , near its border with Germany . Together with the Palatine Forest to the north on
1287-642: Is a global hiatus in the terrestrial fossil record during the late Kungurian and early Roadian , referred to as "Olson's Gap" that obscures the nature of the transition. Other proposals have suggested that the North American and Russian records overlap, with the latest terrestrial North American deposition occurring during the Roadian, suggesting that there was an extinction event, dubbed " Olson's Extinction ". The Middle Permian faunas of South Africa and Russia are dominated by therapsids, most abundantly by
1386-663: Is defined by the first appearance of the conodont Jinogondolella aserrata. The Capitanian is named after the Capitan Reef in the Guadalupe Mountains of Texas, named by George Burr Richardson in 1904, and first used in a chronostratigraphic sense by Glenister and Furnish in 1961 as a substage of the Guadalupian Stage. The Capitanian was ratified as an international stage by the ICS in 2001. The GSSP for
1485-773: Is the highest peak at 1,424 m (4,672 ft), followed by the Storkenkopf (1,366 m, 4,482 ft), and the Hohneck (1,364 m, 4,475 ft). Geographically , the Vosges Mountains are wholly in France , far above the Col de Saverne separating them from the Palatinate Forest in Germany . The latter area logically continues the same Vosges geologic structure but traditionally receives this different name for historical and political reasons. From 1871 to 1918
1584-452: Is thought to have existed during the Early Permian. Though the fossil record is fragmentary, lungfish appear to have undergone an evolutionary diversification and size increase in freshwater habitats during the Early Permian, but subsequently declined during the middle and late Permian. Conodonts experienced their lowest diversity of their entire evolutionary history during the Permian. Permian chondrichthyan faunas are poorly known. Members of
1683-914: The Capitanian Stage of the Permian was marked by the major Capitanian mass extinction event , associated with the eruption of the Emeishan Traps . The Permian (along with the Paleozoic) ended with the Permian–Triassic extinction event (colloquially known as the Great Dying), the largest mass extinction in Earth's history (which is the last of the three or four crises that occurred in the Permian), in which nearly 81% of marine species and 70% of terrestrial species died out, associated with
1782-569: The Celts . After the Roman era, Alemanni also settled in the east, and Franks in the northwest. Contrary to widespread belief, the main ridge of the Vosges coincided with the historical Roman-Germanic language boundary only in the southern Vosges. Old Romance ( Altromanisch ) is spoken east of the main ridge: in the valley of the Weiss around Lapoutroie , the valley of Lièpvrette (nowadays also called
1881-535: The First World War : Permian The Permian witnessed the diversification of the two groups of amniotes , the synapsids and the sauropsids ( reptiles ). The world at the time was dominated by the supercontinent Pangaea , which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa . The Carboniferous rainforest collapse left behind vast regions of desert within
1980-467: The Geological Society of London , after extensive Russian explorations undertaken with Édouard de Verneuil in the vicinity of the Ural Mountains in the years 1840 and 1841. Murchison identified "vast series of beds of marl , schist , limestone , sandstone and conglomerate" that succeeded Carboniferous strata in the region. Murchison, in collaboration with Russian geologists, named
2079-710: The Kastelberg to 1,350 m (4,429 ft); and the Ballon d'Alsace to 1,247 m (4,091 ft). The Col de Saales, between the Higher and Central Vosges, reaches nearly 579 m (1,900 ft), both lower and narrower than the Higher Vosges, with Mont Donon at 1,008 m (3,307 ft) being the highest point of this Nordic section. The highest mountains and peaks of the Vosges (with Alsatian or German names in brackets) are: Two nature parks lie within
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#17327901193012178-726: The North China Craton , the South China Block and Indochina fused to each other and Pangea by the end of the Permian. The Zechstein Sea , a hypersaline epicontinental sea , existed in what is now northwestern Europe. Large continental landmass interiors experience climates with extreme variations of heat and cold (" continental climate ") and monsoon conditions with highly seasonal rainfall patterns. Deserts seem to have been widespread on Pangaea. Such dry conditions favored gymnosperms , plants with seeds enclosed in
2277-704: The Paleo-Tethys Ocean , a large ocean that existed between Asia and Gondwana. The Cimmeria continent rifted away from Gondwana and drifted north to Laurasia , causing the Paleo-Tethys Ocean to shrink. A new ocean was growing on its southern end, the Neotethys Ocean , an ocean that would dominate much of the Mesozoic Era. A magmatic arc, containing Hainan on its southwesternmost end, began to form as Panthalassa subducted under
2376-621: The Parc naturel régional des Ballons des Vosges (established in 1989). Meteorologically , as a consequence of the Foehn effect the difference between the eastern and western mean slopes of the range is very marked. The main air streams come generally from the west and southwest, so the Alsatian central plains just under the Hautes-Vosges receive much less water than the south-west front of
2475-549: The Sydney Basin , and palaeoclimatic models of the Earth's climate based on the behaviour of modern weather patterns showing that such a megamonsoon would occur given the continental arrangement of the Permian. The aforementioned increasing equatorial aridity was likely driven by the development and intensification of this Pangaean megamonsoon. Permian marine deposits are rich in fossil mollusks , brachiopods , and echinoderms . Brachiopods were highly diverse during
2574-447: The U.S. states of Texas and New Mexico is so named because it has one of the thickest deposits of Permian rocks in the world. Sea levels dropped slightly during the earliest Permian (Asselian). The sea level was stable at several tens of metres above present during the Early Permian, but there was a sharp drop beginning during the Roadian, culminating in the lowest sea level of the entire Palaeozoic at around present sea level during
2673-680: The Val d'Argent ; "Valley of Silver"), parts of the canton of Villé valley [ fr ] ( Vallée de Villé ) and parts of the Bruche valley ( Vallée de la Bruche ). By contrast, those parts of the northern Vosges and the whole of the Wasgau , which lie north of the Breusch valley, fall within the Germanic-speaking area because, from Schirmeck the historical linguistic boundary turns to
2772-647: The first appearance datum of specific species of conodont , an enigmatic group of jawless chordates with hard tooth-like oral elements. Conodonts are used as index fossils for most of the Palaeozoic and the Triassic. The Cisuralian Series is named after the strata exposed on the western slopes of the Ural Mountains in Russia and Kazakhstan. The name was proposed by J. B. Waterhouse in 1982 to comprise
2871-577: The pseudosuchians , dinosaurs , and pterosaurs in the following Triassic, first appeared and diversified during the Late Permian, including the first appearance of the Archosauriformes during the latest Permian. Cynodonts , the group of therapsids ancestral to modern mammals , first appeared and gained a worldwide distribution during the Late Permian. Another group of therapsids, the therocephalians (such as Lycosuchus ), arose in
2970-605: The Asselian, Sakmarian, and Artinskian stages. The Kungurian was later added to conform to the Russian "Lower Permian". Albert Auguste Cochon de Lapparent in 1900 had proposed the "Uralian Series", but the subsequent inconsistent usage of this term meant that it was later abandoned. The Asselian was named by the Russian stratigrapher V.E. Ruzhenchev in 1954, after the Assel River in the southern Ural Mountains. The GSSP for
3069-514: The Cisuralian, Guadalupian, and Lopingian. Geologists divide the rocks of the Permian into a stratigraphic set of smaller units called stages , each formed during corresponding time intervals called ages. Stages can be defined globally or regionally. For global stratigraphic correlation, the International Commission on Stratigraphy (ICS) ratify global stages based on a Global Boundary Stratotype Section and Point (GSSP) from
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3168-628: The Cisuralian. Another cool period began around the middle Capitanian. This cool period, lasting for 3-4 Myr, was known as the Kamura Event. It was interrupted by the Emeishan Thermal Excursion in the late part of the Capitanian, around 260 million years ago, corresponding to the eruption of the Emeishan Traps . This interval of rapid climate change was responsible for the Capitanian mass extinction event. During
3267-450: The Cisuralian. Permian synapsids included some large members such as Dimetrodon . The special adaptations of synapsids enabled them to flourish in the drier climate of the Permian and they grew to dominate the vertebrates. A faunal turnover occurred around the transition between the Cisuralian and Guadalupian, with the decline of amphibians and the replacement of pelycosaurs (a paraphyletic group) with more advanced therapsids , although
3366-500: The Early Permian Chemnitz petrified forest of Germany demonstrates that they had complex branching patterns similar to modern angiosperm trees. By the Late Permian, high thin forests had become widespread across the globe, as evidenced by the global distribution of weigeltisaurids. The oldest likely record of Ginkgoales (the group containing Ginkgo and its close relatives) is Trichopitys heteromorpha from
3465-568: The Early Triassic, but a Permian origin is suspected. The diversity of coelacanths is relatively low throughout the Permian in comparison to other marine fishes, though there is an increase in diversity during the terminal Permian (Changhsingian), corresponding with the highest diversity in their evolutionary history during the Early Triassic. Diversity of freshwater fish faunas was generally low and dominated by lungfish and "Paleopterygians". The last common ancestor of all living lungfish
3564-770: The German side of the border, they form a single geomorphological unit and low mountain range of around 8,000 km (3,100 sq mi) in area. It runs in a north-northeast direction from the Burgundian Gate (the Belfort – Ronchamp – Lure line) to the Börrstadt Basin (the Winnweiler – Börrstadt – Göllheim line), and forms the western boundary of the Upper Rhine Plain . The Grand Ballon
3663-472: The Late Permian. Members of the modern orders Archostemata and Adephaga are known from the Late Permian. Complex wood boring traces found in the Late Permian of China suggest that members of Polyphaga , the most diverse group of modern beetles, were also present by the Late Permian. The terrestrial fossil record of the Permian is patchy and temporally discontinuous. Early Permian records are dominated by equatorial Europe and North America, while those of
3762-617: The Late Permian. By the Changhsingian, only a handful (4-6) genera remained. Corals exhibited a decline in diversity over the course of the Middle and Late Permian. Terrestrial life in the Permian included diverse plants, fungi , arthropods , and various types of tetrapods . The period saw a massive desert covering the interior of Pangaea . The warm zone spread in the northern hemisphere, where extensive dry desert appeared. The rocks formed at that time were stained red by iron oxides,
3861-623: The Lopingian in China. Modern amphibians ( lissamphibians ) are suggested to have originated during Permian, descending from a lineage of dissorophoid temnospondyls or lepospondyls . The diversity of fish during the Permian is relatively low compared to the following Triassic. The dominant group of bony fishes during the Permian were the " Paleopterygii " a paraphyletic grouping of Actinopterygii that lie outside of Neopterygii . The earliest unequivocal members of Neopterygii appear during
3960-605: The Lopingian series. The GSSP for the base of the Wuchiapingian is located at Penglaitan, Guangxi , China and was ratified in 2004. The boundary is defined by the first appearance of Clarkina postbitteri postbitteri The Changhsingian was originally derived from the Changxing Limestone, a geological unit first named by the Grabau in 1923, ultimately deriving from Changxing County , Zhejiang .The GSSP for
4059-408: The Lopingian to a series, including all Permian deposits in South China that overlie the Maokou Limestone. In 1995, a vote by the Subcommission on Permian Stratigraphy of the ICS adopted the Lopingian as an international standard chronostratigraphic unit. The Wuchiapinginan and Changhsingian were first introduced in 1962, by J. Z. Sheng as the "Wuchiaping Formation" and "Changhsing Formation" within
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4158-450: The Middle Permian. There were no flying vertebrates, though the extinct lizard-like reptile family Weigeltisauridae from the Late Permian had extendable wings like modern gliding lizards , and are the oldest known gliding vertebrates. Permian stem-amniotes consisted of lepospondyli and batrachosaurs , according to some phylogenies; according to others, stem-amniotes are represented only by diadectomorphs . Temnospondyls reached
4257-547: The Middle and Late Permian are dominated by temperate Karoo Supergroup sediments of South Africa and the Ural region of European Russia. Early Permian terrestrial faunas of North America and Europe were dominated by primitive pelycosaur synapsids including the herbivorous edaphosaurids , and carnivorous sphenacodontids , diadectids and amphibians . Early Permian reptiles, such as acleistorhinids , were mostly small insectivores. Synapsids (the group that would later include mammals) thrived and diversified greatly during
4356-410: The Ochoan, corresponding to the Lopingian. During the Permian, all the Earth 's major landmasses were collected into a single supercontinent known as Pangaea , with the microcontinental terranes of Cathaysia to the east. Pangaea straddled the equator and extended toward the poles, with a corresponding effect on ocean currents in the single great ocean (" Panthalassa ", the "universal sea"), and
4455-582: The Permian, representing up to a third of all insects at some localities. Mecoptera (sometimes known as scorpionflies) first appeared during the Early Permian, going on to become diverse during the Late Permian. Some Permian mecopterans, like Mesopsychidae have long proboscis that suggest they may have pollinated gymnosperms. The earliest known beetles appeared at the beginning of the Permian. Early beetles such as members of Permocupedidae were likely xylophagous , feeding on decaying wood. Several lineages such as Schizophoridae expanded into aquatic habitats by
4554-473: The Permian-Triassic boundary, corresponding to the eruption of the Siberian Traps , which released more than 5 teratonnes of CO 2 , more than doubling the atmospheric carbon dioxide concentration. A -2% δ O excursion signifies the extreme magnitude of this climatic shift. This extremely rapid interval of greenhouse gas release caused the Permian-Triassic mass extinction, as well as ushering in an extreme hothouse that persisted for several million years into
4653-454: The Permian. Xenacanthiformes , another extinct group of shark-like chondrichthyans, were common in freshwater habitats, and represented the apex predators of freshwater ecosystems. Four floristic provinces in the Permian are recognised, the Angaran , Euramerican, Gondwanan, and Cathaysian realms. The Carboniferous Rainforest Collapse would result in the replacement of lycopsid -dominated forests with tree-fern dominated ones during
4752-456: The Permian. The extinct order Productida was the predominant group of Permian brachiopods, accounting for up to about half of all Permian brachiopod genera. Brachiopods also served as important ecosystem engineers in Permian reef complexes. Amongst ammonoids , Goniatitida were a major group during the Early-Mid Permian, but declined during the Late Permian. Members of the order Prolecanitida were less diverse. The Ceratitida originated from
4851-515: The Permian. This is likely due to competition by Hemiptera , due to their similar mouthparts and therefore ecology. Primitive relatives of damselflies and dragonflies ( Meganisoptera ), which include the largest flying insects of all time, also declined during the Permian. Holometabola , the largest group of modern insects, also diversified during this time. " Grylloblattidans ", an extinct group of winged insects thought to be related to modern ice crawlers , reached their apex of diversity during
4950-399: The Rhine valley, the Black Forest and the distant, snow-covered Swiss mountains. The massif known in Latin as Vosago mons or Vosego silva , sometimes Vogesus mons , was extended to the vast woods covering the region. Later, German speakers referred to the same region as Vogesen or Wasgenwald . Over the centuries, settlement population density grew gradually, as was typical for
5049-475: The Usolka section in the southern Urals, which was ratified in 2018. The GSSP is defined by the first appearance of Sweetognathus binodosus . The Artinskian was named after the city of Arti in Sverdlovsk Oblast , Russia. It was named by Karpinsky in 1874. The Artinskian currently lacks a defined GSSP. The proposed definition for the base of the Artinskian is the first appearance of Sweetognathus aff. S. whitei. The Kungurian takes its name after Kungur ,
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#17327901193015148-400: The Vosges Mountains. The highlands of the arrondissement of Remiremont receive as annual rainfall or snowfall more than 2,000 mm (79 in) of precipitation yearly, whereas some dry countryside near Colmar receives less than 500 mm (20 in) of water in the event of insufficient storms. The temperature is much lower in the west front of the mountains than in the low plains behind
5247-403: The Vosges marked for the most part the border between Germany and France, due to the Franco-Prussian War . The elongated massif is divided south to north into three sections: In addition, the term "Central Vosges" is used to designate the various lines of summits, especially those above 1,000 m (3,300 ft) in elevation. The French department of Vosges is named after the range. From
5346-421: The Vosges were the scene of the Battle of Trippstadt . From 1871 to 1918, they formed the main border line between France and the German Empire . The demarcation line stretched from the Ballon d'Alsace in the south to Mont Donon in the north with the lands east of it being incorporated into Germany as part of Alsace-Lorraine . The Vosges saw extensive fighting during the world wars. During World War I, there
5445-463: The Vosges: the Ballons des Vosges Nature Park and the Northern Vosges Regional Nature Park . The Northern Vosges Nature Park and the Palatinate Forest Nature Park on the German side of the border form the cross-border UNESCO -designated Palatinate Forest-North Vosges Biosphere Reserve . In the late 20th century, a wide area of the massif was included in two protected areas , the Parc naturel régional des Vosges du Nord (established in 1976) and
5544-401: The Word Formation by Johan August Udden in 1916, Glenister and Furnish in 1961 was the first publication to use it as a chronostratigraphic term as a substage of the Guadalupian Stage. The GSSP for the base of the Wordian is located in Guadalupe Pass, Texas, within the sediments of the Getaway Limestone Member of the Cherry Canyon Formation , which was ratified in 2001. The base of the Wordian
5643-413: The Wuchiapingian, followed by a slight rise during the Changhsingian. The Permian was cool in comparison to most other geologic time periods, with modest pole to Equator temperature gradients. At the start of the Permian, the Earth was still in the Late Paleozoic icehouse (LPIA), which began in the latest Devonian and spanned the entire Carboniferous period, with its most intense phase occurring during
5742-506: The amount of lava estimated to have been produced during this period, the worst-case scenario is the release of enough carbon dioxide from the eruptions to raise world temperatures five degrees Celsius. Another hypothesis involves ocean venting of hydrogen sulfide gas. Portions of the deep ocean will periodically lose all of its dissolved oxygen allowing bacteria that live without oxygen to flourish and produce hydrogen sulfide gas. If enough hydrogen sulfide accumulates in an anoxic zone ,
5841-411: The ancestors of many present-day families. Rich forests were present in many areas, with a diverse mix of plant groups. The southern continent saw extensive seed fern forests of the Glossopteris flora. Oxygen levels were probably high there. The ginkgos and cycads also appeared during this period. Insects, which had first appeared and become abundant during the preceding Carboniferous, experienced
5940-405: The base of the Asselian is located in the Aidaralash River valley near Aqtöbe , Kazakhstan, which was ratified in 1996. The beginning of the stage is defined by the first appearance of Streptognathodus postfusus . The Sakmarian is named in reference to the Sakmara River in the southern Urals, and was coined by Alexander Karpinsky in 1874. The GSSP for the base of the Sakmarian is located at
6039-418: The base of the Capitanian is located at Nipple Hill in the southeast Guadalupe Mountains of Texas, and was ratified in 2001, the beginning of the stage is defined by the first appearance of Jinogondolella postserrata. The Lopingian was first introduced by Amadeus William Grabau in 1923 as the "Loping Series" after Leping , Jiangxi , China. Originally used as a lithostraphic unit, T.K. Huang in 1932 raised
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#17327901193016138-404: The base of the Changhsingian is located 88 cm above the base of the Changxing Limestone in the Meishan D section, Zhejiang, China and was ratified in 2005, the boundary is defined by the first appearance of Clarkina wangi. The GSSP for the base of the Triassic is located at the base of Bed 27c at the Meishan D section, and was ratified in 2001. The GSSP is defined by the first appearance of
6237-424: The basin. Thus, the highest range of peaks rises immediately adjacent to the basin and increasingly lower mountains rise further from the basin. The highest points are in the Hautes Vosges : the Grand Ballon , in ancient times called Ballon de Guebwiller or Ballon de Murbach, rises to 1,424 m (4,672 ft); the Storckenkopf to 1,366 m (4,482 ft); the Hohneck to 1,364 m (4,475 ft);
6336-429: The chondrichthyan clade Holocephali , which contains living chimaeras , reached their apex of diversity during the Carboniferous-Permian, the most famous Permian representative being the "buzz-saw shark" Helicoprion , known for its unusual spiral shaped spiral tooth whorl in the lower jaw. Hybodonts , a group of shark-like chondrichthyans, were widespread and abundant members of marine and freshwater faunas throughout
6435-449: The climate became notably more arid at the end of the Carboniferous and beginning of the Permian. Nonetheless, temperatures continued to cool during most of the Asselian and Sakmarian, during which the LPIA peaked. By 287 million years ago, temperatures warmed and the South Pole ice cap retreated in what was known as the Artinskian Warming Event (AWE), though glaciers remained present in the uplands of eastern Australia, and perhaps also
6534-399: The conodont Hindeodus parvus . The Russian Tatarian Stage includes the Lopingian, Capitanian and part of the Wordian, while the underlying Kazanian includes the rest of the Wordian as well as the Roadian. In North America, the Permian is divided into the Wolfcampian (which includes the Nealian and the Lenoxian stages); the Leonardian (Hessian and Cathedralian stages); the Guadalupian; and
6633-434: The continental interior by the more advanced seed ferns and early conifers as a result of the Carboniferous rainforest collapse . At the close of the Permian, lycopod and equisete swamps reminiscent of Carboniferous flora survived only in Cathaysia , a series of equatorial islands in the Paleo-Tethys Ocean that later would become South China . The Permian saw the radiation of many important conifer groups, including
6732-449: The continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibian ancestors. Various authors recognise at least three, and possibly four extinction events in the Permian. The end of the Early Permian ( Cisuralian ) saw a major faunal turnover, with most lineages of primitive " pelycosaur " synapsids becoming extinct, being replaced by more advanced therapsids . The end of
6831-458: The decline of early synapsid clades was apparently a slow event that lasted about 20 Ma, from the Sakmarian to the end of the Kungurian . Predator-prey interactions among terrestrial synapsids became more dynamic. If terrestrial deposition ended around the end of the Cisuralian in North America and began in Russia during the early Guadalupian, a continuous record of the transition is not preserved. Uncertain dating has led to suggestions that there
6930-457: The depopulation of territories – in their wake it was not uncommon for people to be relocated there from other areas. On the lower heights and buttresses of the main chain on the Alsatian side are numerous castles, generally in ruins, testifying to the importance of this crucial crossroads of Europe, violently contested for centuries. At several points on the main ridge, especially at Sainte Odile above Ribeauvillé (German: Rappoltsweiler), are
7029-412: The diverse Dinocephalia . Dinocephalians become extinct at the end of the Middle Permian, during the Capitanian mass extinction event . Late Permian faunas are dominated by advanced therapsids such as the predatory sabertoothed gorgonopsians and herbivorous beaked dicynodonts , alongside large herbivorous pareiasaur parareptiles . The Archosauromorpha , the group of reptiles that would give rise to
7128-465: The earliest Permian of France. The oldest known fossils definitively assignable to modern cycads are known from the Late Permian. In Cathaysia, where a wet tropical frost-free climate prevailed, the Noeggerathiales , an extinct group of tree fern-like progymnosperms were a common component of the flora The earliest Permian (~ 298 million years ago) Cathyasian Wuda Tuff flora, representing
7227-573: The early Wuchiapingian, following the emplacement of the Emeishan Traps, global temperatures declined as carbon dioxide was weathered out of the atmosphere by the large igneous province's emplaced basalts. The late Wuchiapingian saw the finale of the Late Palaeozoic Ice Age, when the last Australian glaciers melted. The end of the Permian is marked by a temperature excursion, much larger than the Emeishan Thermal Excursion, at
7326-494: The end of the Permian. Nautiloids , a subclass of cephalopods, surprisingly survived this occurrence. There is evidence that magma , in the form of flood basalt , poured onto the Earth's surface in what is now called the Siberian Traps , for thousands of years, contributing to the environmental stress that led to mass extinction. The reduced coastal habitat and highly increased aridity probably also contributed. Based on
7425-704: The eruption of the Siberian Traps . It took well into the Triassic for life to recover from this catastrophe; on land, ecosystems took 30 million years to recover. Prior to the introduction of the term Permian , rocks of equivalent age in Germany had been named the Rotliegend and Zechstein , and in Great Britain as the New Red Sandstone . The term Permian was introduced into geology in 1841 by Sir Roderick Impey Murchison , president of
7524-580: The family Daraelitidae within Prolecanitida during the mid-Permian, and extensively diversified during the Late Permian. Only three families of trilobite are known from the Permian, Proetidae , Brachymetopidae and Phillipsiidae . Diversity, origination and extinction rates during the Early Permian were low. Trilobites underwent a diversification during the Kungurian-Wordian, the last in their evolutionary history, before declining during
7623-429: The gas can rise into the atmosphere. Oxidizing gases in the atmosphere would destroy the toxic gas, but the hydrogen sulfide would soon consume all of the atmospheric gas available. Hydrogen sulfide levels might have increased dramatically over a few hundred years. Models of such an event indicate that the gas would destroy ozone in the upper atmosphere allowing ultraviolet radiation to kill off species that had survived
7722-692: The larger Tertiary formations. Erosive glacial action was the primary catalyst for development of the highland massif feature. The Vosges in their southern and central parts are called the Hautes Vosges . These consist of a large Carboniferous mountain eroded just before the Permian Period with gneiss , granites , porphyritic masses or other volcanic intrusions. The north, south and west parts are less eroded by glaciers, and here Vosges Triassic and Permian red sandstone remains are found in large beds. The grès vosgien (a French name for
7821-615: The last two from the Vosges Mountains. The rivers Moselle , Meurthe and Sarre and their numerous affluents all rise on the Lorraine side. In the High Moselle and Meurthe basins, moraines , boulders and polished rocks testify to the former existence of glaciers which once covered the top of the Vosges. The mountain lakes caused by the original glacial phenomena are surrounded by pines, beeches and maples , and green meadows provide pasture for large herds of cattle, with views of
7920-748: The late Carboniferous in Euramerica, and result in the differentiation of the Cathaysian floras from those of Euramerica. The Gondwanan floristic region was dominated by Glossopteridales , a group of woody gymnosperm plants, for most of the Permian, extending to high southern latitudes. The ecology of the most prominent glossopterid, Glossopteris , has been compared to that of bald cypress , living in mires with waterlogged soils. The tree-like calamites , distant relatives of modern horsetails , lived in coal swamps and grew in bamboo -like vertical thickets. A mostly complete specimen of Arthropitys from
8019-664: The latter part of the Pennsylvanian epoch. A significant trend of increasing aridification can be observed over the course of the Cisuralian. Early Permian aridification was most notable in Pangaean localities at near-equatorial latitudes. Sea levels also rose notably in the Early Permian as the LPIA slowly waned. At the Carboniferous-Permian boundary, a warming event occurred. In addition to becoming warmer,
8118-555: The massif, especially in summer. On the eastern slope economic vineyards reach to a height of 400 m (1,300 ft); on the other hand, in the mountains, it is a land of pasture and forest. The only rivers in Alsace are the Ill coming from south Alsace (or Sundgau), and the Bruche d'Andlau and the Bruche which have as tributaries other, shorter but sometimes powerful streams coming like
8217-514: The most extensive extinction event recorded in paleontology : the Permian–Triassic extinction event . 90 to 95% of marine species became extinct , as well as 70% of all land organisms. It is also the only known mass extinction of insects. Recovery from the Permian–Triassic extinction event was protracted; on land, ecosystems took 30 million years to recover. Trilobites , which had thrived since Cambrian times, finally became extinct before
8316-473: The most potent greenhouse gases ) into the atmosphere to raise world temperatures an additional five degrees Celsius. The frozen methane hypothesis helps explain the increase in carbon-12 levels found midway in the Permian–Triassic boundary layer. It also helps explain why the first phase of the layer's extinctions was land-based, the second was marine-based (and starting right after the increase in C-12 levels), and
8415-414: The mountainous regions of far northern Siberia. Southern Africa also retained glaciers during the late Cisuralian in upland environments. The AWE also witnessed aridification of a particularly great magnitude. In the late Kungurian, cooling resumed, resulting in a cool glacial interval that lasted into the early Capitanian, though average temperatures were still much higher than during the beginning of
8514-520: The mountainside under heavy fire from the Germans. Two previous rescues failed. The 442nd suffered 800 casualties, rescued the Texans, and took the mountain. On 20 January 1992 Air Inter Flight 148 crashed into the Vosges Mountains while circling to land at Strasbourg International Airport, killing 87 people. In pre-Roman times, the Vosges was empty of settlements or was colonized and dominated by
8613-561: The name Dyassic , from Dyas and Trias , though Murchison rejected this in 1871. The Permian system was controversial for over a century after its original naming, with the United States Geological Survey until 1941 considering the Permian a subsystem of the Carboniferous equivalent to the Mississippian and Pennsylvanian . The Permian Period is divided into three epochs , from oldest to youngest,
8712-713: The next geologic epoch, the Triassic. The Permian climate was also extremely seasonal and characterised by megamonsoons , which produced high aridity and extreme seasonality in Pangaea's interiors. Precipitation along the western margins of the Palaeo-Tethys Ocean was very high. Evidence for the megamonsoon includes the presence of megamonsoonal rainforests in the Qiangtang Basin of Tibet, enormous seasonal variation in sedimentation, bioturbation, and ichnofossil deposition recorded in sedimentary facies in
8811-633: The northwest and runs between Donon and Mutzigfelsen heading for Sarrebourg ( Saarburg ). The Germanic areas of the Vosges mountains are part of the Alemannic dialect region and cultural area and, in the north, also part of the Frankish dialect region and cultural area. The Romance-speaking areas are traditionally part of the Lorrain language region in the west and the Frainc-Comtou region in
8910-585: The period after the surrounding Russian region of Perm, which takes its name from the medieval kingdom of Permia that occupied the same area hundreds of years prior, and which is now located in the Perm Krai administrative region. Between 1853 and 1867, Jules Marcou recognised Permian strata in a large area of North America from the Mississippi River to the Colorado River and proposed
9009-646: The remains of a wall of unmortared stone with tenons of wood, about 1.8 to 2.2 m (6 to 7 ft) thick and 1.3 to 1.7 m (4 to 6 ft) high, called the Mur Païen (Pagan Wall). It was used for defence in the Middle Ages and archaeologists are divided as to whether it was built by the Romans , or before their arrival . During the French Revolutionary Wars , on 13 July 1794,
9108-419: The rest of the Late Permian, and should be regarded as a separate epoch. The tripartite split was adopted after a formal proposal by Glenister et al. (1999). Historically, most marine biostratigraphy of the Permian was based on ammonoids ; however, ammonoid localities are rare in Permian stratigraphic sections, and species characterise relatively long periods of time. All GSSPs for the Permian are based around
9207-439: The result of intense heating by the sun of a surface devoid of vegetation cover. A number of older types of plants and animals died out or became marginal elements. The Permian began with the Carboniferous flora still flourishing. About the middle of the Permian a major transition in vegetation began. The swamp -loving lycopod trees of the Carboniferous, such as Lepidodendron and Sigillaria , were progressively replaced in
9306-443: The south. For a long time the distribution of languages and dialects basically correlated with the pattern of settlement movements. However, the switch from German to French as the lingua franca which took place between the 17th and the 20th century across the whole of Alsace was not accompanied by any further significant movements of population. General texts: List of majors periodicals concerning Lorraine and South Lorraine: On
9405-408: The southeastern South China. The Central Pangean Mountains , which began forming due to the collision of Laurasia and Gondwana during the Carboniferous, reached their maximum height during the early Permian around 295 million years ago, comparable to the present Himalayas , but became heavily eroded as the Permian progressed. The Kazakhstania block collided with Baltica during the Cisuralian, while
9504-532: The spread of conifers and their increasing prevalence throughout terrestrial ecosystems. Bennettitales , which would go on to become in widespread the Mesozoic, first appeared during the Cisuralian in China. Lyginopterids , which had declined in the late Pennsylvanian and subsequently have a patchy fossil record, survived into the Late Permian in Cathaysia and equatorial east Gondwana. The Permian ended with
9603-413: The toxic gas. There are species that can metabolize hydrogen sulfide. Another hypothesis builds on the flood basalt eruption theory. An increase in temperature of five degrees Celsius would not be enough to explain the death of 95% of life. But such warming could slowly raise ocean temperatures until frozen methane reservoirs below the ocean floor near coastlines melted, expelling enough methane (among
9702-667: Was named in 1968 in reference to the Road Canyon Member of the Word Formation in Texas. The GSSP for the base of the Roadian is located 42.7m above the base of the Cutoff Formation in Stratotype Canyon, Guadalupe Mountains, Texas, and was ratified in 2001. The beginning of the stage is defined by the first appearance of Jinogondolella nankingensis . The Wordian was named in reference to
9801-563: Was severe and almost continuous fighting in the mountains. During World War II in October 1944, there was a fierce battle between German forces and the U.S. 442nd Regiment, a segregated unit composed of second-generation Japanese Americans (Nisei), during which the 442nd charged straight up the mountain to rescue the 1st Battalion of the 36th Infantry, formerly the Texas Guard—also known as the "Lost Battalion"—who were cut off and stranded on
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