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Beech

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Genus ( / ˈ dʒ iː n ə s / ; pl. : genera / ˈ dʒ ɛ n ər ə / ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses . In binomial nomenclature , the genus name forms the first part of the binomial species name for each species within the genus.

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79-787: See text Beech ( Fagus ) is a genus of deciduous trees in the family Fagaceae , native to subtropical (accessory forest element) and temperate (as dominant element of mesophytic forests) Eurasia and North America . There are 14 accepted species in two distinct subgenera, Englerianae Denk & G.W.Grimm and Fagus . The subgenus Englerianae is found only in East Asia, distinctive for its low branches, often made up of several major trunks with yellowish bark. The better known species of subgenus Fagus are native to Europe , western and eastern Asia and eastern North America. They are high-branching trees with tall, stout trunks and smooth silver-grey bark. The European beech Fagus sylvatica

158-412: A phosphodiester bond and the bases are held together by hydrogen bonds . Nuclear DNA and mitochondrial DNA differ in many ways, starting with location and structure. Nuclear DNA is located within the nucleus of eukaryote cells and usually has two copies per cell while mitochondrial DNA is located in the mitochondria and contains 100–1,000 copies per cell. The structure of nuclear DNA chromosomes

237-530: A bitter taste (though not nearly as bitter as acorns ) and a high tannin content. The most recent classification system of the genus recognizes 14 species in two distinct subgenera, subgenus Englerianae and Fagus . Beech species can be diagnosed by phenotypical and/or genotypical traits. Species of subgenus Engleriana are found only in East Asia, and are notably distinct from species of subgenus Fagus in that these beeches are low-branching trees, often made up of several major trunks with yellowish bark and

316-649: A bitter, astringent, or in some cases, mild and nut-like taste. According to the Roman statesman Pliny the Elder in his work Natural History , beechnut was eaten by the people of Chios when the town was besieged, writing of the fruit: "that of the beech is the sweetest of all; so much so, that, according to Cornelius Alexander, the people of the city of Chios, when besieged, supported themselves wholly on mast". They can also be roasted and pulverized into an adequate coffee substitute . The leaves can be steeped in liquor to give

395-536: A case. Techniques used include polymerase chain reaction (PCR), which allows one to utilize very small amounts of DNA by making copies of targeted regions on the molecule, also known as short tandem repeats (STRs). Like mitosis , meiosis is a form of eukaryotic cell division . Meiosis gives rise to four unique daughter cells, each of which has half the number of chromosomes as the parent cell. Because meiosis creates cells that are destined to become gametes (or reproductive cells), this reduction in chromosome number

474-428: A cell goes through an interphase period in which it grows, replicates its chromosomes, and checks all of its systems to ensure that it is ready to divide. Like mitosis, meiosis also has distinct stages called prophase , metaphase , anaphase , and telophase . A key difference, however, is that during meiosis, each of these phases occurs twice — once during the first round of division, called meiosis I, and again during

553-426: A decorative wood. The timber can be used to build chalets, houses, and log cabins. Beech wood is used for the stocks of military rifles when traditionally preferred woods such as walnut are scarce or unavailable or as a lower-cost alternative. The edible fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from the tree in autumn. They are small, roughly triangular, and edible, with

632-480: A diverse set of DNA repair processes that remove nuclear DNA damages. These repair processes include base excision repair , nucleotide excision repair , homologous recombinational repair, non-homologous end joining and microhomology-mediated end joining . Such repair processes are essential for maintaining nuclear DNA stability. Failure of repair activity to keep up with the occurrence of damages has various negative consequences. Nuclear DNA damages, as well as

711-407: A genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

790-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

869-504: A letter of the alphabet) is буква ( bukva ), while Serbo-Croatian and Slovene use " bukva " to refer to the tree. The pigment bistre was made from beech wood soot . Beech litter raking as a replacement for straw in animal husbandry was an old non-timber practice in forest management that once occurred in parts of Switzerland in the 17th century. Beech has been listed as one of the 38 plants whose flowers are used to prepare Bach flower remedies . Genus The composition of

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948-457: A light green/yellow liqueur. In antiquity, the bark of the beech tree was used by Indo-European people for writing-related purposes, especially in a religious context. Beech wood tablets were a common writing material in Germanic societies before the development of paper . The Old English bōc has the primary sense of "beech" but also a secondary sense of "book", and it is from bōc that

1027-489: A long history of cultivation, Bulgaria for example, do not exhibit this pattern, so how much human activity has influenced the spread of beech trees is as yet unclear. The primeval beech forests of the Carpathians are also an example of a singular, complete, and comprehensive forest dominated by a single tree species - the beech tree. Forest dynamics here were allowed to proceed without interruption or interference since

1106-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

1185-687: A naturally growing forest tree, beech marks the important border between the European deciduous forest zone and the northern pine forest zone. This border is important for wildlife and fauna. In Denmark and Scania at the southernmost peak of the Scandinavian peninsula, southwest of the natural spruce boundary, it is the most common forest tree. It grows naturally in Denmark and southern Norway and Sweden up to about 57–59°N. The most northern known naturally growing (not planted) beech trees are found in

1264-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

1343-788: A separate family, the Nothofagaceae (which remains a member of the order Fagales ). They are found throughout the Southern Hemisphere in Australia , New Zealand , New Guinea , New Caledonia , as well as Argentina and Chile (principally Patagonia and Tierra del Fuego ). Species treated in Denk et al. (2024) and listed in Plants of the World Online (POWO) : Erroneously synonymized by some authors (e.g. POWO) with

1422-533: A series of short DNA segments called Okazaki fragments . Each Okazaki fragment requires a separate RNA primer. As the Okazaki fragments are synthesized, the RNA primers are replaced with DNA nucleotides and the fragments are bonded together in a continuous complementary strand. Damage of nuclear DNA is a persistent problem arising from a variety of disruptive endogenous and exogenous sources. Eukaryotes have evolved

1501-476: A short sequence of RNA nucleotides, complementary to a small, initial section of the DNA strand being prepared for replication. DNA polymerase is then able to add DNA nucleotides to the RNA primer and thus begin the process of constructing a new complementary strand of DNA. Later the RNA primer is enzymatically removed and replaced with the appropriate sequence of DNA nucleotides. Because the two complementary strands of

1580-560: A small grove north of Bergen on the west coast of Norway. Near the city of Larvik is the largest naturally occurring beech forest in Norway, Bøkeskogen . Some research suggests that early agriculture patterns supported the spread of beech in continental Europe. Research has linked the establishment of beech stands in Scandinavia and Germany with cultivation and fire disturbance, i.e. early agricultural practices. Other areas which have

1659-575: A substantially different nucleome (nuclear DNA) , especially in noncoding, highly variable gene regions such as the spacers of the nuclear-encoded ribosomal RNA genes (ribosomal DNA) . Further differentiating characteristics include the whitish bloom on the underside of the leaves, the visible tertiary leaf veins, and a long, smooth cupule-peduncle. Originally proposed but not formalized by botanist Chung-Fu Shen in 1992, this group comprised two Japanese species, F. japonica and F. okamotoi, and one Chinese species, F. engleriana . While

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1738-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

1817-421: A template to guide the synthesis of the new complementary polynucleotide of DNA. The DNA single-strand template serves to guide the synthesis of a complementary strand of DNA. DNA replication begins at a specific site in the DNA molecule called the origin of replication . The enzyme helicase unwinds and separates a portion of the DNA molecule after which single-strand binding proteins react with and stabilize

1896-437: A tone between those of maple and birch , the two most popular drum woods. The textile modal is a kind of rayon often made wholly from reconstituted cellulose of pulped beech wood. The European species Fagus sylvatica yields a tough, utility timber. It weighs about 720 kg per cubic metre and is widely used for furniture construction, flooring, and engineering purposes, in plywood and household items, but rarely as

1975-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

2054-462: Is a long polymer chain of repeating nucleotides . Each nucleotide is composed of a five-carbon sugar, a phosphate group, and an organic base. Nucleotides are distinguished by their bases: purines , large bases that include adenine and guanine ; and pyrimidines , small bases that include thymine and cytosine . Chargaff's rules state that adenine always pairs with thymine, and guanine always with cytosine. The phosphate groups are held together by

2133-439: Is also used to smoke Westphalian ham , andouille sausage, and some cheeses. Beeches are monoecious , bearing both male and female flowers on the same plant. The small flowers are unisexual, the female flowers borne in pairs, the male flowers wind-pollinating catkins . They are produced in spring shortly after the new leaves appear. The fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from

2212-523: Is critical — without it, the union of two gametes during fertilization would result in offspring with twice the normal number of chromosomes. Meiosis creates new combinations of genetic material in each of the four daughter cells. These new combinations result from the exchange of DNA between paired chromosomes. Such an exchange means that the gametes produced through meiosis often exhibit considerable genetic variation. Meiosis involves two rounds of nuclear division, not just one. Prior to undergoing meiosis,

2291-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

2370-601: Is less than 0.3% while that of mitochondrial DNA is generally higher. Nuclear DNA is known as the molecule of life and contains the genetic instructions for the development of all eukaryotic organisms. It is found in almost every cell in the human body, with exceptions such as red blood cells . Everyone has a unique genetic blueprint, even identical twins. Forensic departments such as the Bureau of Criminal Apprehension (BCA) and Federal Bureau of Investigation (FBI) are able to use techniques involving nuclear DNA to compare samples in

2449-434: Is linear with open ends and includes 46 chromosomes and contains for example 3 billion nucleotides in humans while the structure of Mitochondrial DNA chromosome is usually closed, circular, and contains for example 16,569 nucleotides in humans. Nuclear DNA in animals is diploid , ordinarily inheriting the DNA from two parents, while mitochondrial DNA is haploid , coming only from the mother. The mutation rate for nuclear DNA

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2528-428: Is not planted much as an ornamental due to slower growth and less resistance to urban pollution. It most commonly occurs as an overstory component in the northern part of its range with sugar maple, transitioning to other forest types further south such as beech-magnolia. American beech is rarely encountered in developed areas except as a remnant of a forest that was cut down for land development. The dead brown leaves of

2607-474: Is smooth and light gray. The fruit is a small, sharply three-angled nut 10–15 mm ( 3 ⁄ 8 – 5 ⁄ 8  in) long, borne singly or in pairs in soft-spined husks 1.5–2.5 cm ( 5 ⁄ 8 –1 in) long, known as cupules. The husk can have a variety of spine- to scale-like appendages, the character of which is, in addition to leaf shape, one of the primary ways beeches are differentiated. The nuts are called beechnuts or beech mast and have

2686-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

2765-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

2844-426: Is the most commonly cultivated species, yielding a utility timber used for furniture construction, flooring and engineering purposes, in plywood, and household items. The timber can be used to build homes. Beechwood makes excellent firewood . Slats of washed beech wood are spread around the bottom of fermentation tanks for Budweiser beer. Beech logs are burned to dry the malt used in some German smoked beers . Beech

2923-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

3002-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

3081-652: The cryptic sister species F. hayatae and F. pashanica ) , two pseudo-cryptic species in eastern North America ( F. grandifolia , F. mexicana ), and a species complex of at least four species ( F. caspica , F. hohenackeriana , F. orientalis , F. sylvatica ) in Western Eurasia . Their genetics are highly complex and include both species-unique alleles as well as alleles and ribosomal DNA spacers that are shared between two or more species. The western Eurasian species are characterized by morphological and genetical gradients. Research suggests that

3160-575: The mutations and epigenetic alterations that such damages cause, are considered to be a major cause of cancer . Nuclear DNA damages are also implicated in aging and neurodegenerative diseases . Nuclear DNA is subject to mutation . A major cause of mutation is inaccurate DNA replication , often by specialized DNA polymerases that synthesize past DNA damages in the template strand (error-prone trans-lesion synthesis ). Mutations also arise by inaccurate DNA repair. The microhomology-mediated end joining pathway for repair of double-strand breaks

3239-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

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3318-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

3397-636: The American beech remain on the branches until well into the following spring, when the new buds finally push them off. East Asia is home to eight species of Fagus , only one of which ( F. crenata ) is occasionally planted in Western countries. Smaller than F. sylvatica and F. grandifolia , this beech is one of the most common hardwoods in its native range. Beech grows on a wide range of soil types, acidic or basic, provided they are not waterlogged. The tree canopy casts dense shade and thickens

3476-721: The Crimean F. × taurica, from which it differs morphologically and genetically. Numerous species have been named globally from the fossil record spanning from the Cretaceous to the Pleistocene . Fossil species formerly placed in Fagus include: The name of the tree in Latin, fagus (from whence the generic epithet ), is cognate with English "beech" and of Indo-European origin, and played an important role in early debates on

3555-407: The DNA molecule are oriented in opposite directions and the DNA polymerase can only accommodate replication in one direction, two different mechanisms for copying the strands of DNA are employed. One strand is replicated continuously towards unwinding, separating the portion of the original DNA molecule; while the other strand is replicated discontinuously in the opposite direction with the formation of

3634-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

3713-517: The bottom of fermentation tanks for Budweiser beer. This provides a complex surface on which the yeast can settle, so that it does not pile up, preventing yeast autolysis which would contribute off-flavours to the beer. Beech logs are burned to dry the malt used in German smoked beers . Beech is also used to smoke Westphalian ham , traditional andouille (an offal sausage) from Normandy, and some cheeses. Some drums are made from beech, which has

3792-577: The disease has become widespread in Connecticut, Massachusetts and Rhode Island, and in portions of coastal New Hampshire and coastal and central Maine. The beech most commonly grown as an ornamental tree is the European beech ( Fagus sylvatica ), widely cultivated in North America as well as its native Europe. Many varieties are in cultivation, notably the weeping beech F. sylvatica 'Pendula', several varieties of copper or purple beech,

3871-821: The eastern United States and southeastern Canada, with a disjunct sister species in Mexico ( F. mexicana ). There are the only extant (surviving) Fagus species in the Western Hemisphere. Before the Pleistocene Ice Age, it is believed to have spanned the entire width of the continent from the Atlantic Ocean to the Pacific but now is confined to the east of the Great Plains. F. grandifolia tolerates hotter climates than European species but

3950-657: The exception of the Western Eurasian and North American species. Fagus is the first diverging lineage in the evolution of the Fagaceae family, which also includes oaks and chestnuts . The oldest fossils that can be assigned to the beech lineage are 81–82 million years old pollen from the Late Cretaceous of Wyoming , United States. The southern beeches (genus Nothofagus ) historically thought closely related to beeches, are treated as members of

4029-728: The fern-leaved beech F. sylvatica 'Asplenifolia', and the tricolour beech F. sylvatica 'Roseomarginata'. The columnar Dawyck beech ( F. sylvatica 'Dawyck') occurs in green, gold, and purple forms, named after Dawyck Botanic Garden in the Scottish Borders, one of the four garden sites of the Royal Botanic Garden Edinburgh . Beech wood is an excellent firewood , easily split and burning for many hours with bright but calm flames. Slats of beech wood are washed in caustic soda to leach out any flavour or aroma characteristics and are spread around

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4108-752: The first representatives of the modern-day genus were already present in the Paleocene of Arctic North America (western Greenland ) and quickly radiated across the high latitudes of the Northern Hemisphere , with a first diversity peak in the Miocene of northeastern Asia . The contemporary species are the product of past, repeated reticulate evolutionary processes ( outbreeding , introgression , hybridization ). As far as studied, heterozygosity and intragenomic variation are common in beech species, and their chloroplast genomes are nonspecific with

4187-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

4266-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

4345-490: The geographical origins of the Indo-European people , the beech argument . Greek φηγός (figós) is from the same root, but the word was transferred to the oak tree (e.g. Iliad 16.767) as a result of the absence of beech trees in southern Greece . Fagus sylvatica was a late entrant to Great Britain after the last glaciation, and may have been restricted to basic soils in the south of England. Some suggest that it

4424-479: The ground with leaf litter . In North America, they can form beech-maple climax forests by partnering with the sugar maple . The beech blight aphid ( Grylloprociphilus imbricator ) is a common pest of American beech trees. Beeches are also used as food plants by some species of Lepidoptera . Beech bark is extremely thin and scars easily. Since the beech tree has such delicate bark, carvings, such as lovers' initials and other forms of graffiti, remain because

4503-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

4582-571: The last ice age. Nowadays, they are amongst the last pure beech forests in Europe to document the undisturbed postglacial repopulation of the species, which also includes the unbroken existence of typical animals and plants. These virgin beech forests and similar forests across 12 countries in continental Europe were inscribed on the UNESCO World Heritage List in 2007. The American beech ( Fagus grandifolia ) occurs across much of

4661-490: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Nuclear DNA Nuclear DNA ( nDNA ), or nuclear deoxyribonucleic acid , is the DNA contained within each cell nucleus of a eukaryotic organism . It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for

4740-428: The modern word derives. In modern German, the word for "book" is Buch, with Buche meaning "beech tree". In modern Dutch, the word for "book" is boek, with beuk meaning "beech tree". In Swedish, these words are the same, bok meaning both "beech tree" and "book". There is a similar relationship in some Slavic languages. In Russian and Bulgarian , the word for beech is бук ( buk ), while that for "letter" (as in

4819-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

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4898-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

4977-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

5056-521: The rest. It adheres to Mendelian inheritance , with information coming from two parents, one male and one female—rather than matrilineally (through the mother) as in mitochondrial DNA. Nuclear DNA is a nucleic acid , a polymeric biomolecule or biopolymer , found in the nucleus of eukaryotic cells. Its structure is a double helix , with two strands wound around each other, a structure first described by Francis Crick and James D. Watson (1953) using data collected by Rosalind Franklin . Each strand

5135-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

5214-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

5293-477: The second round of division, called meiosis II. Prior to cell division, the DNA material in the original cell must be duplicated so that after cell division, each new cell contains the full amount of DNA material. The process of DNA duplication is usually called replication . The replication is termed semiconservative since each new cell contains one strand of original DNA and one newly synthesized strand of DNA. The original polynucleotide strand of DNA serves as

5372-419: The separated, single-stranded sections of the DNA molecule. The enzyme complex DNA polymerase engages the separated portion of the molecule and initiates the process of replication. DNA polymerase can only connect new DNA nucleotides to a pre-existing chain of nucleotides. Therefore, replication begins as an enzyme called primase assembles an RNA primer at the origin of replication. The RNA primer consists of

5451-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

5530-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

5609-786: The status of F. okamotoi remains uncertain, the most recent systematic treatment based on morphological and genetic data confirmed a third species, F. multinervis, endemic to Ulleungdo , a South Korean island in the Sea of Japan . The beeches of Ulleungdo have been traditionally treated as a subspecies of F. engleriana, to which they are phenotypically identical, or as a variety of F. japonica. The differ from their siblings by their unique nuclear and plastid genotypes. The better known subgenus Fagus beeches are high-branching with tall, stout trunks and smooth silver-gray bark. This group includes five extant species in continental and insular East Asia ( F. crenata , F. longipetiolata , F. lucida , and

5688-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

5767-453: The tree in autumn. They are small, roughly triangular, and edible, with a bitter, astringent, or mild and nut-like taste. The European beech ( Fagus sylvatica ) is the most commonly cultivated, although few important differences are seen between species aside from detail elements such as leaf shape. The leaves of beech trees are entire or sparsely toothed, from 5–15 centimetres (2–6 inches) long and 4–10 cm (2–4 in) broad. The bark

5846-618: The tree is unable to heal itself. Beech bark disease is a fungal infection that attacks the American beech through damage caused by scale insects. Infection can lead to the death of the tree. Beech leaf disease is a disease that affects American beeches spread by the newly discovered nematode, Litylenchus crenatae mccannii . This disease was first discovered in Lake County, Ohio, in 2012 and has now spread to over 41 counties in Ohio, Pennsylvania, New York, and Ontario, Canada. As of 2024,

5925-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

6004-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

6083-583: The world (according to Guinness World Records ) is the Meikleour Beech Hedge in Meikleour , Perth and Kinross , Scotland. Fagus sylvatica is one of the most common hardwood trees in north-central Europe, in France constituting alone about 15% of all nonconifers. The Balkans are also home to the lesser-known oriental beech ( F. orientalis ) and Crimean beech ( F. taurica ). As

6162-511: Was designated for its beech woodlands, which are believed to be on the western edge of their natural range in this steep limestone gorge. Beech is not native to Ireland; however, it was widely planted in the 18th century and can become a problem shading out the native woodland understory. Beech is widely planted for hedging and in deciduous woodlands, and mature, regenerating stands occur throughout mainland Britain at elevations below about 650 m (2,100 ft). The tallest and longest hedge in

6241-457: Was introduced by Neolithic tribes who planted the trees for their edible nuts. The beech is classified as a native in the south of England and as a non-native in the north where it is often removed from 'native' woods. Large areas of the Chilterns are covered with beech woods, which are habitat to the common bluebell and other flora. The Cwm Clydach National Nature Reserve in southeast Wales

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