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97-503: Gastropods ( / ˈ ɡ æ s t r ə p ɒ d z / ), commonly known as slugs and snails , belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ( / ɡ æ s ˈ t r ɒ p ə d ə / ). This class comprises snails and slugs from saltwater, freshwater, and from the land. There are many thousands of species of sea snails and slugs , as well as freshwater snails , freshwater limpets , land snails and slugs . The class Gastropoda

194-442: A taxonomic rank ; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain , kingdom , phylum ( division is sometimes used in botany in place of phylum ), class , order , family , genus , and species . The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed

291-529: A warning , when they are poisonous or contain stinging cells , or to camouflage them on the brightly colored hydroids , sponges , and seaweeds on which many of the species are found. Lateral outgrowths on the body of nudibranchs are called cerata . These contain an outpocketing of digestive glands called the diverticula . The sensory organs of gastropods include olfactory organs , eyes, statocysts and mechanoreceptors . Gastropods have no hearing. In terrestrial gastropods (land snails and slugs),

388-469: A basis of morphological and physiological facts as possible, and one in which "place is found for all observational and experimental data relating, even if indirectly, to the constitution, subdivision, origin, and behaviour of species and other taxonomic groups". Ideals can, it may be said, never be completely realized. They have, however, a great value of acting as permanent stimulants, and if we have some, even vague, ideal of an "omega" taxonomy we may progress

485-557: A chaotic and disorganized taxonomic literature. He not only introduced the standard of class, order, genus, and species, but also made it possible to identify plants and animals from his book, by using the smaller parts of the flower (known as the Linnaean system ). Plant and animal taxonomists regard Linnaeus' work as the "starting point" for valid names (at 1753 and 1758 respectively). Names published before these dates are referred to as "pre-Linnaean", and not considered valid (with

582-443: A different sense, to mean the delimitation of species (not subspecies or taxa of other ranks), using whatever investigative techniques are available, and including sophisticated computational or laboratory techniques. Thus, Ernst Mayr in 1968 defined " beta taxonomy " as the classification of ranks higher than species. An understanding of the biological meaning of variation and of the evolutionary origin of groups of related species

679-487: A few ciliary feeders, in which the radula is reduced or absent. Land-dwelling species can chew up leaves, bark, fruit and decomposing animals while marine species can scrape algae off the rocks on the seafloor. Certain species such as the Archaeogastropda maintain horizontal rows of slender marginal teeth. In some species that have evolved into endoparasites, such as the eulimid Thyonicola doglieli , many of

776-438: A fossil record. Gastropoda (previously known as univalves and sometimes spelled "Gasteropoda") are a major part of the phylum Mollusca, and are the most highly diversified class in the phylum , with 65,000 to 80,000 living snail and slug species . The anatomy , behavior, feeding, and reproductive adaptations of gastropods vary significantly from one clade or group to another, so stating many generalities for all gastropods

873-400: A gastropod is called a protoconch . Most shelled gastropods have a one piece shell (with exceptional bivalved gastropods ), typically coiled or spiraled, at least in the larval stage. This coiled shell usually opens on the right-hand side (as viewed with the shell apex pointing upward). Numerous species have an operculum , which in many species acts as a trapdoor to close the shell. This

970-446: A gastropod is usually adapted to the food that a species eats. The simplest gastropods are the limpets and abalones , herbivores that use their hard radula to rasp at seaweeds on rocks. Many marine gastropods are burrowers, and have a siphon that extends out from the mantle edge. Sometimes the shell has a siphonal canal to accommodate this structure. A siphon enables the animal to draw water into their mantle cavity and over

1067-455: A little way down the Greek alphabet. Some of us please ourselves by thinking we are now groping in a "beta" taxonomy. Turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy. Later authors have used the term in

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1164-491: A marine environment. Gastropods have a worldwide distribution, from the near Arctic and Antarctic zones to the tropics. They have become adapted to almost every kind of existence on earth, having colonized nearly every available medium. In habitats where not enough calcium carbonate is available to build a really solid shell, such as on some acidic soils on land, various species of slugs occur, and also some snails with thin, translucent shells, mostly or entirely composed of

1261-629: A natural defense against hurricane and tsunami damage in particular. The Sundarbans and Bhitarkanika Mangroves are two of the large mangrove forests in the world, both on the coast of the Bay of Bengal . Some seas and lakes are brackish. The Baltic Sea is a brackish sea adjoining the North Sea . Originally the Eridanos river system prior to the Pleistocene , since then it has been flooded by

1358-504: A notable renaissance, principally with respect to theoretical content. Part of the theoretical material has to do with evolutionary areas (topics e and f above), the rest relates especially to the problem of classification. Taxonomy is that part of Systematics concerned with topics (a) to (d) above. A whole set of terms including taxonomy, systematic biology, systematics , scientific classification, biological classification, and phylogenetics have at times had overlapping meanings – sometimes

1455-516: A ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the evolutionary relationships among organisms, both living and extinct. The exact definition of taxonomy varies from source to source, but

1552-400: A reference to the fact that the animal's "foot" is positioned below its guts. The earlier name "univalve" means one valve (or shell), in contrast to bivalves , such as clams, which have two valves or shells. At all taxonomic levels, gastropods are second only to insects in terms of their diversity . Gastropods have the greatest numbers of named mollusk species. However, estimates of

1649-477: A single continuum, as per the scala naturae (the Natural Ladder). This, as well, was taken into consideration in the great chain of being. Advances were made by scholars such as Procopius , Timotheus of Gaza , Demetrios Pepagomenos , and Thomas Aquinas . Medieval thinkers used abstract philosophical and logical categorizations more suited to abstract philosophy than to pragmatic taxonomy. During

1746-535: A species only found in deep water in the Baltic, while pike are confined to the less saline surface waters. The Caspian Sea is the world's largest lake and contains brackish water with a salinity about one-third that of normal seawater. The Caspian is famous for its peculiar animal fauna, including one of the few non-marine seals (the Caspian seal ) and the great sturgeons , a major source of caviar . Hudson Bay

1843-613: A taxon involves five main requirements: However, often much more information is included, like the geographic range of the taxon, ecological notes, chemistry, behavior, etc. How researchers arrive at their taxa varies: depending on the available data, and resources, methods vary from simple quantitative or qualitative comparisons of striking features, to elaborate computer analyses of large amounts of DNA sequence data. Brackish water Technically, brackish water contains between 0.5 and 30 grams of salt per litre—more often expressed as 0.5 to 30 parts per thousand (‰), which

1940-524: A truly scientific attempt to classify organisms did not occur until the 18th century, with the possible exception of Aristotle, whose works hint at a taxonomy. Earlier works were primarily descriptive and focused on plants that were useful in agriculture or medicine. There are a number of stages in this scientific thinking. Early taxonomy was based on arbitrary criteria, the so-called "artificial systems", including Linnaeus 's system of sexual classification for plants (Linnaeus's 1735 classification of animals

2037-554: Is a specific gravity of between 1.0004 and 1.0226. Thus, brackish covers a range of salinity regimes and is not considered a precisely defined condition. It is characteristic of many brackish surface waters that their salinity can vary considerably over space or time. Water with a salt concentration greater than 30‰ is considered saline . See the salinity table from the Misplaced Pages salinity article . Brackish water condition commonly occurs when fresh water meets seawater. In fact,

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2134-555: Is a brackish marginal sea of the Arctic Ocean , it remains brackish due its limited connections to the open ocean, very high levels freshwater surface runoff input from the large Hudson Bay drainage basin , and low rate of evaporation due to being completely covered in ice for over half the year. In the Black Sea the surface water is brackish with an average salinity of about 17–18 parts per thousand compared to 30 to 40 for

2231-497: Is a critical component of the taxonomic process. As a result, it informs the user as to what the relatives of the taxon are hypothesized to be. Biological classification uses taxonomic ranks, including among others (in order from most inclusive to least inclusive): Domain , Kingdom , Phylum , Class , Order , Family , Genus , Species , and Strain . The "definition" of a taxon is encapsulated by its description or its diagnosis or by both combined. There are no set rules governing

2328-470: Is a diverse and highly successful class of mollusks within the phylum Mollusca. It contains a vast total of named species, second only to the insects in overall number. The fossil history of this class goes back to the Late Cambrian . As of 2017, 721 families of gastropods are known, of which 245 are extinct and appear only in the fossil record, while 476 are currently extant with or without

2425-400: Is a novel analysis of the variation patterns in a particular taxon . This analysis may be executed on the basis of any combination of the various available kinds of characters, such as morphological, anatomical , palynological , biochemical and genetic . A monograph or complete revision is a revision that is comprehensive for a taxon for the information given at a particular time, and for

2522-471: Is a part of the behavior of mating gastropods with some pulmonate families of land snails creating and utilizing love darts , the throwing of which have been identified as a form of sexual selection . The main aspects of the life cycle of gastropods include: The diet of gastropods differs according to the group considered. Marine gastropods include some that are herbivores , detritus feeders, predatory carnivores , scavengers , parasites , and also

2619-458: Is a resource for fossils. Biological taxonomy is a sub-discipline of biology , and is generally practiced by biologists known as "taxonomists", though enthusiastic naturalists are also frequently involved in the publication of new taxa. Because taxonomy aims to describe and organize life , the work conducted by taxonomists is essential for the study of biodiversity and the resulting field of conservation biology . Biological classification

2716-454: Is considered a part of taxonomy (definitions 1 and 2), or a part of systematics outside taxonomy. For example, definition 6 is paired with the following definition of systematics that places nomenclature outside taxonomy: In 1970, Michener et al. defined "systematic biology" and "taxonomy" (terms that are often confused and used interchangeably) in relation to one another as follows: Systematic biology (hereafter called simply systematics)

2813-422: Is difficult. The class Gastropoda has an extraordinary diversification of habitats . Representatives live in gardens, woodland, deserts, and on mountains; in small ditches, great rivers, and lakes; in estuaries , mudflats , the rocky intertidal , the sandy subtidal, the abyssal depths of the oceans, including the hydrothermal vents , and numerous other ecological niches, including parasitic ones. Although

2910-419: Is even more important for the second stage of taxonomic activity, the sorting of species into groups of relatives ("taxa") and their arrangement in a hierarchy of higher categories. This activity is what the term classification denotes; it is also referred to as "beta taxonomy". How species should be defined in a particular group of organisms gives rise to practical and theoretical problems that are referred to as

3007-433: Is muscular, and the second is mutagenetic . The effects of torsion are primarily physiological. The organism develops by asymmetrical growth, with the majority of growth occurring on the left side. This leads to the loss of right-side anatomy that in most bilaterians is a duplicate of the left side anatomy. The essential feature of this asymmetry is that the anus generally lies to one side of the median plane. The gill-combs ,

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3104-468: Is not the most important sense, because they are mainly nocturnal animals. The nervous system of gastropods includes the peripheral nervous system and the central nervous system . The central nervous system consists of ganglia connected by nerve cells. It includes paired ganglia: the cerebral ganglia, pedal ganglia, osphradial ganglia, pleural ganglia, parietal ganglia and the visceral ganglia. There are sometimes also buccal ganglia. The radula of

3201-638: Is probably not even a mollusk. As such, it's not until the Ordovician that the first crown-group members arise. By the Ordovician period the gastropods were a varied group present in a range of aquatic habitats. Commonly, fossil gastropods from the rocks of the early Palaeozoic era are too poorly preserved for accurate identification. Still, the Silurian genus Poleumita contains fifteen identified species. Fossil gastropods were less common during

3298-400: Is the field that (a) provides scientific names for organisms, (b) describes them, (c) preserves collections of them, (d) provides classifications for the organisms, keys for their identification, and data on their distributions, (e) investigates their evolutionary histories, and (f) considers their environmental adaptations. This is a field with a long history that in recent years has experienced

3395-459: Is usually made of a horn-like material, but in some molluscs it is calcareous. In the land slugs, the shell is reduced or absent, and the body is streamlined. Some gastropods have adult shells which are bottom heavy due to the presence of a thick, often broad, convex ventral callus deposit on the inner lip and adapical to the aperture which may be important for gravitational stability. Some sea slugs are very brightly colored. This serves either as

3492-422: Is very low and the fish fauna consists predominantly of freshwater species such as roach , dace , carp , perch , and pike . The Thames Estuary becomes brackish between Battersea and Gravesend , and the diversity of freshwater fish species present is smaller, primarily roach and dace; euryhaline marine species such as flounder , European seabass , mullet , and smelt become much more common. Further east,

3589-529: The Aristotelian system , with additions concerning the philosophical and existential order of creatures. This included concepts such as the great chain of being in the Western scholastic tradition, again deriving ultimately from Aristotle. The Aristotelian system did not classify plants or fungi , due to the lack of microscopes at the time, as his ideas were based on arranging the complete world in

3686-585: The Neomura , the clade that groups together the Archaea and Eucarya , would have evolved from Bacteria, more precisely from Actinomycetota . His 2004 classification treated the archaeobacteria as part of a subkingdom of the kingdom Bacteria, i.e., he rejected the three-domain system entirely. Stefan Luketa in 2012 proposed a five "dominion" system, adding Prionobiota ( acellular and without nucleic acid ) and Virusobiota (acellular but with nucleic acid) to

3783-647: The Palaeozoic era than bivalves . Most of the gastropods of the Palaeozoic era belong to primitive groups, a few of which still survive. By the Carboniferous period many of the shapes seen in living gastropods can be matched in the fossil record, but despite these similarities in appearance the majority of these older forms are not directly related to living forms. It was during the Mesozoic era that

3880-479: The Planorbidae , have hemoglobin as the respiratory protein. In one large group of sea slugs, the gills are arranged as a rosette of feathery plumes on their backs, which gives rise to their other name, nudibranchs . Some nudibranchs have smooth or warty backs with no visible gill mechanism, such that respiration may likely take place directly through the skin. Gastropods have open circulatory system and

3977-512: The Renaissance and the Age of Enlightenment , categorizing organisms became more prevalent, and taxonomic works became ambitious enough to replace the ancient texts. This is sometimes credited to the development of sophisticated optical lenses, which allowed the morphology of organisms to be studied in much greater detail. One of the earliest authors to take advantage of this leap in technology

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4074-473: The continental slope and the continental rise are home to the highest diversity, while the continental shelf and abyssal depths have a low diversity of marine gastropods. Gastropods are found in a wide range of aquatic and terrestrial habitats, from deep ocean trenches to deserts. Some of the more familiar and better-known gastropods are terrestrial gastropods (the land snails and slugs). Some live in fresh water, but most named species of gastropods live in

4171-406: The crab-eating frog , Fejervarya cancrivora (formerly Rana cancrivora ). Mangroves represent important nesting sites for numerous birds groups such as herons, storks, spoonbills, ibises, kingfishers, shorebirds and seabirds. Although often plagued with mosquitoes and other insects that make them unpleasant for humans, mangrove swamps are very important buffer zones between land and sea, and are

4268-445: The olfactory organs , the foot slime-gland , nephridia , and the auricle of the heart are single or at least are more developed on one side of the body than the other. Furthermore, there is only one genital orifice, which lies on the same side of the body as the anus. Furthermore, the anus becomes redirected to the same space as the head. This is speculated to have some evolutionary function, as prior to torsion, when retracting into

4365-439: The species problem . The scientific work of deciding how to define species has been called microtaxonomy. By extension, macrotaxonomy is the study of groups at the higher taxonomic ranks subgenus and above, or simply in clades that include more than one taxon considered a species, expressed in terms of phylogenetic nomenclature . While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations,

4462-467: The vertebrates ), as well as groups like the sharks and cetaceans , are commonly used. His student Theophrastus (Greece, 370–285 BC) carried on this tradition, mentioning some 500 plants and their uses in his Historia Plantarum . Several plant genera can be traced back to Theophrastus, such as Cornus , Crocus , and Narcissus . Taxonomy in the Middle Ages was largely based on

4559-488: The 1960s. In 1958, Julian Huxley used the term clade . Later, in 1960, Cain and Harrison introduced the term cladistic . The salient feature is arranging taxa in a hierarchical evolutionary tree , with the desideratum that all named taxa are monophyletic. A taxon is called monophyletic if it includes all the descendants of an ancestral form. Groups that have descendant groups removed from them are termed paraphyletic , while groups representing more than one branch from

4656-484: The North Sea but still receives so much freshwater from the adjacent lands that the water is brackish. As seawater is denser, the water in the Baltic is stratified, with seawater at the bottom and freshwater at the top. Limited mixing occurs because of the lack of tides and storms, with the result that the fish fauna at the surface is freshwater in composition while that lower down is more marine. Cod are an example of

4753-491: The Origin of Species (1859) led to a new explanation for classifications, based on evolutionary relationships. This was the concept of phyletic systems, from 1883 onwards. This approach was typified by those of Eichler (1883) and Engler (1886–1892). The advent of cladistic methodology in the 1970s led to classifications based on the sole criterion of monophyly , supported by the presence of synapomorphies . Since then,

4850-475: The adult stage—though in some, the coiling may not be very visible, for example in cowries . In a number of families of species, such as all the various limpets , the shell is coiled only in the larval stage, and is a simple conical structure after that. In the scientific literature, gastropods were described as "gasteropodes" by Georges Cuvier in 1795. The word gastropod comes from Greek γαστήρ ( gastḗr 'stomach') and πούς ( poús 'foot'),

4947-466: The ancestors of many of the living gastropods evolved. Taxonomy (biology) In biology , taxonomy (from Ancient Greek τάξις ( taxis )  'arrangement' and -νομία ( -nomia )  ' method ') is the scientific study of naming, defining ( circumscribing ) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given

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5044-522: The animal and plant kingdoms toward the end of the 18th century, well before Charles Darwin's On the Origin of Species was published. The pattern of the "Natural System" did not entail a generating process, such as evolution, but may have implied it, inspiring early transmutationist thinkers. Among early works exploring the idea of a transmutation of species were Zoonomia in 1796 by Erasmus Darwin (Charles Darwin's grandfather), and Jean-Baptiste Lamarck 's Philosophie zoologique of 1809. The idea

5141-540: The body cavity, and digestive glands opening into the stomach . Courtship is a part of mating behavior in some gastropods, including some of the Helicidae . Again, in some land snails, an unusual feature of the reproductive system of gastropods is the presence and utilization of love darts . In many marine gastropods other than the opisthobranchs , there are separate sexes ( dioecious / gonochoric ); most land gastropods, however, are hermaphrodites . Courtship

5238-426: The core of the discipline remains: the conception, naming, and classification of groups of organisms. As points of reference, recent definitions of taxonomy are presented below: The varied definitions either place taxonomy as a sub-area of systematics (definition 2), invert that relationship (definition 6), or appear to consider the two terms synonymous. There is some disagreement as to whether biological nomenclature

5335-418: The deep-sea floor has been studied biologically. The total number of living species of freshwater snails is about 4,000. Recently extinct species of gastropods (extinct since 1500) number 444, 18 species are now extinct in the wild (but still exist in captivity), and 69 species are "possibly extinct". The number of prehistoric (fossil) species of gastropods is at least 15,000 species. In marine habitats,

5432-550: The definition of taxa, but the naming and publication of new taxa is governed by sets of rules. In zoology , the nomenclature for the more commonly used ranks ( superfamily to subspecies ), is regulated by the International Code of Zoological Nomenclature ( ICZN Code ). In the fields of phycology , mycology , and botany , the naming of taxa is governed by the International Code of Nomenclature for algae, fungi, and plants ( ICN ). The initial description of

5529-422: The earliest representatives of the group appearing in the Late Cambrian ( Chippewaella , Strepsodiscus ), though their only gastropod character is a coiled shell, so they could lie in the stem lineage, if they are gastropods at all. Earliest Cambrian organisms like Helcionella , Barskovia and Scenella are no longer considered gastropods, and the tiny coiled Aldanella of earliest Cambrian time

5626-399: The entire world. Other (partial) revisions may be restricted in the sense that they may only use some of the available character sets or have a limited spatial scope. A revision results in a conformation of or new insights in the relationships between the subtaxa within the taxon under study, which may lead to a change in the classification of these subtaxa, the identification of new subtaxa, or

5723-494: The evidentiary basis has been expanded with data from molecular genetics that for the most part complements traditional morphology . Naming and classifying human surroundings likely began with the onset of language. Distinguishing poisonous plants from edible plants is integral to the survival of human communities. Medicinal plant illustrations show up in Egyptian wall paintings from c.  1500 BC , indicating that

5820-524: The exception of spiders published in Svenska Spindlar ). Even taxonomic names published by Linnaeus himself before these dates are considered pre-Linnaean. Modern taxonomy is heavily influenced by technology such as DNA sequencing , bioinformatics , databases , and imaging . A pattern of groups nested within groups was specified by Linnaeus' classifications of plants and animals, and these patterns began to be represented as dendrograms of

5917-486: The first modern groups tied to fossil ancestors was birds. Using the then newly discovered fossils of Archaeopteryx and Hesperornis , Thomas Henry Huxley pronounced that they had evolved from dinosaurs, a group formally named by Richard Owen in 1842. The resulting description, that of dinosaurs "giving rise to" or being "the ancestors of" birds, is the essential hallmark of evolutionary taxonomic thinking. As more and more fossil groups were found and recognized in

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6014-687: The formal naming of clades. Linnaean ranks are optional and have no formal standing under the PhyloCode , which is intended to coexist with the current, rank-based codes. While popularity of phylogenetic nomenclature has grown steadily in the last few decades, it remains to be seen whether a majority of systematists will eventually adopt the PhyloCode or continue using the current systems of nomenclature that have been employed (and modified, but arguably not as much as some systematists wish) for over 250 years. Well before Linnaeus, plants and animals were considered separate Kingdoms. Linnaeus used this as

6111-411: The gill. They use the siphon primarily to "taste" the water to detect prey from a distance. Gastropods with siphons tend to be either predators or scavengers. Almost all marine gastropods breathe with a gill , but many freshwater species, and the majority of terrestrial species, have a pallial lung . The respiratory protein in almost all gastropods is hemocyanin , but one freshwater pulmonate family,

6208-466: The late 19th and early 20th centuries, palaeontologists worked to understand the history of animals through the ages by linking together known groups. With the modern evolutionary synthesis of the early 1940s, an essentially modern understanding of the evolution of the major groups was in place. As evolutionary taxonomy is based on Linnaean taxonomic ranks, the two terms are largely interchangeable in modern use. The cladistic method has emerged since

6305-401: The merger of previous subtaxa. Taxonomic characters are the taxonomic attributes that can be used to provide the evidence from which relationships (the phylogeny ) between taxa are inferred. Kinds of taxonomic characters include: The term " alpha taxonomy " is primarily used to refer to the discipline of finding, describing, and naming taxa , particularly species. In earlier literature,

6402-467: The most extensive brackish water habitats worldwide are estuaries , where a river meets the sea. The River Thames flowing through London is a classic river estuary. The town of Teddington a few miles west of London marks the boundary between the tidal and non-tidal parts of the Thames, although it is still considered a freshwater river about as far east as Battersea insofar as the average salinity

6499-572: The name "snail" can be, and often is, applied to all the members of this class, commonly this word means only those species with an external shell big enough that the soft parts can withdraw completely into it. Slugs are gastropods that have no shell or a very small, internal shell; semislugs are gastropods that have a shell that they can partially retreat into but not entirely. The marine shelled species of gastropods include species such as abalone , conches , periwinkles , whelks , and numerous other sea snails that produce seashells that are coiled in

6596-549: The oceans. The deep, anoxic water of the Black Sea originates from warm, salty water of the Mediterranean . Lake Texoma , a reservoir on the border between the U.S. states of Texas and Oklahoma , is a rare example of a brackish lake that is neither part of an endorheic basin nor a direct arm of the ocean, though its salinity is considerably lower than that of the other bodies of water mentioned here. The reservoir

6693-484: The olfactory organs, located on the tips of the four tentacles , are the most important sensory organ. The chemosensory organs of opisthobranch marine gastropods are called rhinophores . The majority of gastropods have simple visual organs, eye spots either at the tip or base of the tentacles. However, "eyes" in gastropods range from simple ocelli that only distinguish light and dark, to more complex pit eyes , and even to lens eyes . In land snails and slugs, vision

6790-507: The origin of Patellogastropoda and Heterobranchia , whereas fewer changes occurred between the ancestors of Vetigastropoda (only tRNAs D, C and N) and Caenogastropoda (a large single inversion , and translocations of the tRNAs D and N). Within Heterobranchia, gene order seems relatively conserved, and gene rearrangements are mostly related with transposition of tRNA genes. The first gastropods were exclusively marine, with

6887-431: The pens the farmed fish are kept in. Another important brackish water habitat is the mangrove swamp or mangal . Many, though not all, mangrove swamps fringe estuaries and lagoons where the salinity changes with each tide. Among the most specialised residents of mangrove forests are mudskippers , fish that forage for food on land, and archer fish , perch-like fish that "spit" at insects and other small animals living in

6984-434: The possibilities of closer co-operation with their cytological, ecological and genetics colleagues and to acknowledge that some revision or expansion, perhaps of a drastic nature, of their aims and methods, may be desirable ... Turrill (1935) has suggested that while accepting the older invaluable taxonomy, based on structure, and conveniently designated "alpha", it is possible to glimpse a far-distant taxonomy built upon as wide

7081-553: The protein conchiolin . Snails such as Sphincterochila boissieri and Xerocrassa seetzeni have adapted to desert conditions. Other snails have adapted to an existence in ditches, near deepwater hydrothermal vents , in oceanic trenches 10,000 meters (6 miles) below the surface, the pounding surf of rocky shores , caves , and many other diverse areas. Gastropods can be accidentally transferred from one habitat to another by other animals, e.g. by birds . Snails are distinguished by an anatomical process known as torsion , where

7178-818: The rank of Order, although both exclude fossil representatives. A separate compilation (Ruggiero, 2014) covers extant taxa to the rank of Family. Other, database-driven treatments include the Encyclopedia of Life , the Global Biodiversity Information Facility , the NCBI taxonomy database , the Interim Register of Marine and Nonmarine Genera , the Open Tree of Life , and the Catalogue of Life . The Paleobiology Database

7275-500: The river. This type of ecological succession from freshwater to marine ecosystem is typical of river estuaries. River estuaries form important staging points during the migration of anadromous and catadromous fish species, such as salmon , shad and eels , giving them time to form social groups and to adjust to the changes in salinity. Salmon are anadromous, meaning they live in the sea but ascend rivers to spawn; eels are catadromous, living in rivers and streams, but returning to

7372-412: The salinity increases and the freshwater fish species are completely replaced by euryhaline marine ones, until the river reaches Gravesend, at which point conditions become fully marine and the fish fauna resembles that of the adjacent North Sea and includes both euryhaline and stenohaline marine species. A similar pattern of replacement can be observed with the aquatic plants and invertebrates living in

7469-407: The same, sometimes slightly different, but always related and intersecting. The broadest meaning of "taxonomy" is used here. The term itself was introduced in 1813 by de Candolle , in his Théorie élémentaire de la botanique . John Lindley provided an early definition of systematics in 1830, although he wrote of "systematic botany" rather than using the term "systematics". Europeans tend to use

7566-727: The sea to breed. Besides the species that migrate through estuaries, there are many other fish that use them as "nursery grounds" for spawning or as places young fish can feed and grow before moving elsewhere. Herring and plaice are two commercially important species that use the Thames Estuary for this purpose. Estuaries are also commonly used as fishing grounds and as places for fish farming or ranching. For example, Atlantic salmon farms are often located in estuaries, although this has caused controversy, because in doing so, fish farmers expose migrating wild fish to large numbers of external parasites such as sea lice that escape from

7663-419: The shell, first the posterior end would get pulled in, and then the anterior. Now, the front can be retracted more easily, perhaps suggesting a defensive purpose. Gastropods typically have a well-defined head with two or four sensory tentacles with eyes, and a ventral foot. The foremost division of the foot is called the propodium. Its function is to push away sediment as the snail crawls. The larval shell of

7760-592: The standard gastropod features are strongly reduced or absent. A few sea slugs are herbivores and some are carnivores. The carnivorous habit is due to specialisation. Many gastropods have distinct dietary preferences and regularly occur in close association with their food species. Some predatory carnivorous gastropods include: cone shells , Testacella , Daudebardia , turrids , ghost slugs and others. Gastropods exhibit an important degree of variation in mitochondrial gene organization when compared to other animals. Main events of gene rearrangement occurred at

7857-472: The term had a different meaning, referring to morphological taxonomy, and the products of research through the end of the 19th century. William Bertram Turrill introduced the term "alpha taxonomy" in a series of papers published in 1935 and 1937 in which he discussed the philosophy and possible future directions of the discipline of taxonomy. ... there is an increasing desire amongst taxonomists to consider their problems from wider viewpoints, to investigate

7954-482: The terms "systematics" and "biosystematics" for the study of biodiversity as a whole, whereas North Americans tend to use "taxonomy" more frequently. However, taxonomy, and in particular alpha taxonomy , is more specifically the identification, description, and naming (i.e., nomenclature) of organisms, while "classification" focuses on placing organisms within hierarchical groups that show their relationships to other organisms. A taxonomic revision or taxonomic review

8051-510: The three-domain method is the separation of Archaea and Bacteria , previously grouped into the single kingdom Bacteria (a kingdom also sometimes called Monera ), with the Eukaryota for all organisms whose cells contain a nucleus . A small number of scientists include a sixth kingdom, Archaea, but do not accept the domain method. Thomas Cavalier-Smith , who published extensively on the classification of protists , in 2002 proposed that

8148-427: The top rank, dividing the physical world into the vegetable, animal and mineral kingdoms. As advances in microscopy made the classification of microorganisms possible, the number of kingdoms increased, five- and six-kingdom systems being the most common. Domains are a relatively new grouping. First proposed in 1977, Carl Woese 's three-domain system was not generally accepted until later. One main characteristic of

8245-712: The total number of gastropod species vary widely, depending on cited sources. The number of gastropod species can be ascertained from estimates of the number of described species of Mollusca with accepted names: about 85,000 (minimum 50,000, maximum 120,000). But an estimate of the total number of Mollusca, including undescribed species, is about 240,000 species. The estimate of 85,000 mollusks includes 24,000 described species of terrestrial gastropods. Different estimates for aquatic gastropods (based on different sources) give about 30,000 species of marine gastropods, and about 5,000 species of freshwater and brackish gastropods. Many deep-sea species remain to be discovered, as only 0.0001% of

8342-436: The traditional three domains. Partial classifications exist for many individual groups of organisms and are revised and replaced as new information becomes available; however, comprehensive, published treatments of most or all life are rarer; recent examples are that of Adl et al., 2012 and 2019, which covers eukaryotes only with an emphasis on protists, and Ruggiero et al., 2015, covering both eukaryotes and prokaryotes to

8439-437: The transport fluid is hemolymph . Hemocyanin is present in the hemolymph as the respiratory pigment. The primary organs of excretion in gastropods are nephridia , which produce either ammonia or uric acid as a waste product. The nephridium also plays an important role in maintaining water balance in freshwater and terrestrial species. Additional organs of excretion, at least in some species, include pericardial glands in

8536-518: The tree of life are called polyphyletic . Monophyletic groups are recognized and diagnosed on the basis of synapomorphies , shared derived character states. Cladistic classifications are compatible with traditional Linnean taxonomy and the Codes of Zoological and Botanical nomenclature , to a certain extent. An alternative system of nomenclature, the International Code of Phylogenetic Nomenclature or PhyloCode has been proposed, which regulates

8633-420: The trees, knocking them into the water where they can be eaten. Like estuaries, mangrove swamps are extremely important breeding grounds for many fish, with species such as snappers , halfbeaks , and tarpon spawning or maturing among them. Besides fish, numerous other animals use mangroves, including such species as the saltwater crocodile , American crocodile , proboscis monkey , diamondback terrapin , and

8730-642: The uses of different species were understood and that a basic taxonomy was in place. Organisms were first classified by Aristotle ( Greece , 384–322 BC) during his stay on the Island of Lesbos . He classified beings by their parts, or in modern terms attributes , such as having live birth, having four legs, laying eggs, having blood, or being warm-bodied. He divided all living things into two groups: plants and animals . Some of his groups of animals, such as Anhaima (animals without blood, translated as invertebrates ) and Enhaima (animals with blood, roughly

8827-404: The visceral mass of the animal rotates 180° to one side during development, such that the anus is situated more or less above the head. This process is unrelated to the coiling of the shell, which is a separate phenomenon. Torsion is present in all gastropods, but the opisthobranch gastropods are secondarily untorted to various degrees. Torsion occurs in two stages. The first, mechanistic stage

8924-486: Was Methodus Plantarum Nova (1682), in which he published details of over 18,000 plant species. At the time, his classifications were perhaps the most complex yet produced by any taxonomist, as he based his taxa on many combined characters. The next major taxonomic works were produced by Joseph Pitton de Tournefort (France, 1656–1708). His work from 1700, Institutiones Rei Herbariae , included more than 9000 species in 698 genera, which directly influenced Linnaeus, as it

9021-519: Was created by the damming of the Red River of the South , which (along with several of its tributaries) receives large amounts of salt from natural seepage from buried deposits in the upstream region. The salinity is high enough that striped bass , a fish normally found only in salt water, has self-sustaining populations in the lake. Brackish water is being used by humans in many different sectors. It

9118-555: Was entitled " Systema Naturae " ("the System of Nature"), implying that he, at least, believed that it was more than an "artificial system"). Later came systems based on a more complete consideration of the characteristics of taxa, referred to as "natural systems", such as those of de Jussieu (1789), de Candolle (1813) and Bentham and Hooker (1862–1863). These classifications described empirical patterns and were pre- evolutionary in thinking. The publication of Charles Darwin 's On

9215-615: Was popularized in the Anglophone world by the speculative but widely read Vestiges of the Natural History of Creation , published anonymously by Robert Chambers in 1844. With Darwin's theory, a general acceptance quickly appeared that a classification should reflect the Darwinian principle of common descent . Tree of life representations became popular in scientific works, with known fossil groups incorporated. One of

9312-543: Was the Italian physician Andrea Cesalpino (1519–1603), who has been called "the first taxonomist". His magnum opus De Plantis came out in 1583, and described more than 1500 plant species. Two large plant families that he first recognized are in use: the Asteraceae and Brassicaceae . In the 17th century John Ray ( England , 1627–1705) wrote many important taxonomic works. Arguably his greatest accomplishment

9409-429: Was the text he used as a young student. The Swedish botanist Carl Linnaeus (1707–1778) ushered in a new era of taxonomy. With his major works Systema Naturae 1st Edition in 1735, Species Plantarum in 1753, and Systema Naturae 10th Edition , he revolutionized modern taxonomy. His works implemented a standardized binomial naming system for animal and plant species, which proved to be an elegant solution to

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