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63-607: Many, see text Gryllus is a genus of field cricket ( Orthoptera , Gryllidae , Gryllinae ). Members of the genus are typically 15–31 mm long and darkly coloured. The type species is Gryllus campestris L.: the European field cricket. Until the mid-1950s, native field crickets in eastern North America were all assigned to a single species, Acheta assimilis . Although regional variation in calling song and life history were noted, no morphological characters could be found to reliably distinguish these variants. Building upon

126-457: A food resource when in heat. Outside of the breeding season their efforts were not as persistent or successful. This shows that the food-for-sex hypothesis likely plays a role in the food sharing among canids and acts as a direct benefit for the females. Another study on free-ranging dogs found that social factors played a significant role in the determination of mating pairs. The study, done in 2014, looked at social regulation of reproduction in

189-414: A large role in the determination of mating pairs in dogs. Canids also show a wide range of parental care and in 2018 a study showed that sexual conflict plays a role in the determination of intersexual parental investment . The studied looked at coyote mating pairs and found that paternal investment was increased to match or near match the maternal investment. The amount of parental care provided by

252-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

315-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

378-705: A longer duration for a reduced energy requirement. By doing this the mothers increasing the likelihood of their pups surviving infancy and reaching adulthood and thereby increase their own fitness. A study done in 2017 found that aggression between male and female gray wolves varied and changed with age. Males were more likely to chase away rival packs and lone individuals than females and became increasingly aggressive with age. Alternatively, females were found to be less aggressive and constant in their level of aggression throughout their life. This requires further research but suggests that intersexual aggression levels in gray wolves relates to their mating system . Tooth breakage

441-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

504-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

567-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

630-592: A workshop hosted by the IUCN /SSC Canid Specialist Group recommends that because DNA evidence shows the side-striped jackal ( Canis adustus ) and black-backed jackal ( Canis mesomelas ) to form a monophyletic lineage that sits outside of the Canis / Cuon / Lycaon clade, that they should be placed in a distinct genus, Lupulella Hilzheimer, 1906 with the names Lupulella adusta and Lupulella mesomelas . The fossil record shows that feliforms and caniforms emerged within

693-409: Is a frequent result of carnivores' feeding behaviour. Carnivores include both pack hunters and solitary hunters. The solitary hunter depends on a powerful bite at the canine teeth to subdue their prey, and thus exhibits a strong mandibular symphysis . In contrast, a pack hunter, which delivers many shallower bites, has a comparably weaker mandibular symphysis. Thus, researchers can use the strength of

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756-595: Is also higher when taking and consuming large prey. In comparison to extant gray wolves, the extinct Beringian wolves included many more individuals with moderately to heavily worn teeth and with a significantly greater number of broken teeth. The frequencies of fracture ranged from a minimum of 2% found in the Northern Rocky Mountain wolf (Canis lupus irremotus) up to a maximum of 11% found in Beringian wolves. The distribution of fractures across

819-607: Is based on the DNA phylogeny of Lindblad-Toh et al . (2005), modified to incorporate recent findings on Canis species, Canis latrans (coyote) [REDACTED] Canis rufus (red wolf) [REDACTED] Canis lycaon (Algonquin wolf) [REDACTED] Canis lupus (gray wolf) [REDACTED] Canis familiaris (domestic dog) [REDACTED] Canis lupaster ( African golden wolf ) [REDACTED] Canis simensis ( Ethiopian wolf ) [REDACTED] Canis aureus ( golden jackal ) [REDACTED] In 2019,

882-445: Is commonly referred to as the wolf event . It is associated with the formation of the mammoth steppe and continental glaciation. Canis spread to Europe in the forms of C. arnensis , C. etruscus , and C. falconeri . However, a 2021 genetic study of the dire wolf ( Aenocyon dirus ), previously considered a member of Canis , found that it represented the last member of an ancient lineage of canines originally indigenous to

945-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

1008-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

1071-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

1134-399: Is used for slicing and the talonid is used for grinding. The ratio between the trigonid and the talonid indicates a carnivore's dietary habits, with a larger trigonid indicating a hypercarnivore and a larger talonid indicating a more omnivorous diet. Because of its low variability, the length of the lower carnassial is used to provide an estimate of a carnivore's body size. A study of

1197-555: The African wild dog . The gray wolf ranked between these two. The eating of bone increases the risk of accidental fracture due to the relatively high, unpredictable stresses that it creates. The most commonly broken teeth are the canines, followed by the premolars, carnassial molars, and incisors. Canines are the teeth most likely to break because of their shape and function, which subjects them to bending stresses that are unpredictable in direction and magnitude. The risk of tooth fracture

1260-663: The Ethiopian wolf ( C. simensis ), eastern wolf ( C. lycaon ), and the African golden wolf ( C. lupaster ) are four of the many Canis species referred to as "wolves". Species that are too small to attract the word "wolf" are called coyotes in the Americas and jackals elsewhere. Although these may not be more closely related to each other than they are to C. lupus , they are, as fellow Canis species, more closely related to wolves and domestic dogs than they are to foxes , maned wolves , or other canids which do not belong to

1323-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

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1386-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

1449-576: The Middle Pleistocene and was limited in Eurasia to the small wolves of the Canis mosbachensis –Canis variabilis group and the large hypercarnivorous Canis (Xenocyon) lycaonoides . The hypercarnivore Xenocyon gave rise to the modern dhole and the African wild dog . Dentition relates to the arrangement of teeth in the mouth, with the dental notation for the upper-jaw teeth using

1512-595: The Pliocene and Pleistocene , which is consistent with fossil evidence suggesting that much of the African canid fauna diversity resulted from the immigration of Eurasian ancestors, likely coincident with Plio-Pleistocene climatic oscillations between arid and humid conditions. In 2017, the fossil remains of a new Canis species, named Canis othmanii , was discovered among remains found at Wadi Sarrat, Tunisia, from deposits that date 700,000 years ago. This canine shows

1575-659: The gray wolf and coyote , likely only arrived in the New World during the Late Pleistocene , where their dietary flexibility and/or ability to hybridize with other canids allowed them to survive the Quaternary extinction event , unlike the dire wolf. Xenocyon (strange wolf) is an extinct subgenus of Canis . The diversity of the Canis group decreased by the end of the Early Pleistocene to

1638-419: The nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

1701-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

1764-533: The African hunting dog (142), the gray wolf (136), the dhole (112), and the dingo (108). The bite force at the carnassials showed a similar trend to the canines. A predator's largest prey size is strongly influenced by its biomechanical limits. There is little variance among male and female canids. Canids tend to live as monogamous pairs. Wolves, dholes , coyotes , and jackals live in groups that include breeding pairs and their offspring. Wolves may live in extended family groups. To take prey larger than themselves,

1827-404: The African wild dog, the dhole, and the gray wolf depend on their jaws as they cannot use their forelimbs to grapple with prey. They work together as a pack consisting of an alpha pair and their offspring from the current and previous years. Social mammal predators prey on herbivores with a body mass similar to that of the combined mass of the predator pack. The gray wolf specializes in preying on

1890-518: The ICZN's Direction 22 added Canis familiaris as the type species for genus Canis to the official list. Canis is primitive relative to Cuon , Lycaon , and Xenocyon in its relatively larger canines and lack of such dental adaptations for hypercarnivory as m1–m2 metaconid and entoconid small or absent; M1–M2 hypocone small; M1–M2 lingual cingulum weak; M2 and m2 small, may be single-rooted; m3 small or absent; and wide palate. The cladogram below

1953-620: The Late Miocene and Early Pliocene but they were not the top predators. For Canis populations in the New World, Eucyon in North America gave rise to early North American Canis which first appeared in the Miocene (6 million YBP) in south-western United States and Mexico. By 5 million YBP the larger Canis lepophagus , ancestor of wolves and coyotes, appeared in the same region. Around 5 million years ago, some of

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2016-563: The New World that had diverged prior to the appearance of Canis , and that its lineage had been distinct since the Miocene with no evidence of introgression with Canis . The study hypothesized that the Neogene canids in the New World, Canis armbrusteri and Canis edwardii , were possibly members of the distinct dire wolf lineage that had convergently evolved a very similar appearance to members of Canis . True members of Canis , namely

2079-660: The Old World Eucyon evolved into the first members of Canis , and the position of the canids would change to become a dominant predator across the Palearctic . The wolf-sized C. chihliensis appeared in northern China in the Mid-Pliocene around 4-3 million YBP. This was followed by an explosion of Canis evolution across Eurasia in the Early Pleistocene around 1.8 million YBP in what

2142-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

2205-646: The clade Carnivoramorpha 43 million YBP . The caniforms included the fox-like genus Leptocyon , whose various species existed from 24 million YBP before branching 11.9 million YBP into Vulpes (foxes) and Canini (canines). The jackal-sized Eucyon existed in North America from 10 million YBP and by the Early Pliocene about 6-5 million YBP the coyote-like Eucyon davisi invaded Eurasia. The canids that had emigrated from North America to Eurasia – Eucyon , Vulpes , and Nyctereutes – were small to medium-sized predators during

2268-449: The dogs. They found that females in heat searched out dominant males and were more likely to mate with a dominant male who appeared to be a quality leader. The females were more likely to reject submissive males. Furthermore, cases of male-male competition were more aggressive in the presence of high ranking females. This suggests that females prefer dominant males and males prefer high ranking females meaning social cues and status play

2331-458: The estimated bite force at the canine teeth of a large sample of living and fossil mammalian predators, when adjusted for their body mass, found that for placental mammals the bite force at the canines (in Newtons /kilogram of body weight) was greatest in the extinct dire wolf (163), followed among the modern canids by the four hypercarnivores that often prey on animals larger than themselves:

2394-424: The extinct dire wolf. This indicates that these are both better adapted for cracking bone than other canids. A study of nine modern carnivores indicate that one in four adults had suffered tooth breakage and that half of these breakages were of the canine teeth. The highest frequency of breakage occurred in the spotted hyena, which is known to consume all of its prey including the bone. The least breakage occurred in

2457-500: The fathers also was shown to fluctuated depending on the level of care provided by the mother. Another study on parental investment showed that in free-ranging dogs, mothers modify their energy and time investment into their pups as they age. Due to the high mortality of free-range dogs at a young age a mother's fitness can be drastically reduced. This study found that as the pups aged the mother shifted from high-energy care to lower-energy care so that they can care for their offspring for

2520-484: The following species as of March 2023: Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of

2583-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

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2646-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

2709-629: The genus Canis . The word "jackal" is applied to the golden jackal ( C. aureus ), found across southwestern and south-central Asia, and the Balkans in Europe. The first record of Canis on the African continent is Canis sp. A from South Turkwel, Kenya, dated 3.58–3.2 million years ago. In 2015, a study of mitochondrial genome sequences and whole genome nuclear sequences of African and Eurasian canids indicated that extant wolf-like canids have colonised Africa from Eurasia at least 5 times throughout

2772-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

2835-626: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Canis Extant: Extinct: Canis is a genus of the Caninae which includes multiple extant species, such as wolves , dogs , coyotes , and golden jackals . Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails. The genus Canis ( Carl Linnaeus , 1758)

2898-471: The mandibular symphysis in fossil carnivore specimens to determine what kind of hunter it was – a pack hunter or a solitary hunter – and even how it consumed its prey. The mandibles of canids are buttressed behind the carnassial teeth to crack bones with their post-carnassial teeth (molars M2 and M3). A study found that the modern gray wolf and the red wolf ( C.   rufus ) possess greater buttressing than all other extant canids and

2961-546: The middle one being longer and more slender than the rest, in American and European species. African species, however, show more variability, making it more difficult to assign species to a genus with the same degree of certainty as there is more overlap with closely related genera. Gryllus is one of the most widespread cricket genera. It is found in Africa , North and South America , Europe and Asia . The genus contains

3024-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

3087-407: The muscle and tendon of prey. Canids use their premolars for cutting and crushing except for the upper fourth premolar P4 (the upper carnassial) that is only used for cutting. They use their molars for grinding except for the lower first molar m1 (the lower carnassial) that has evolved for both cutting and grinding depending on the candid's dietary adaptation. On the lower carnassial the trigonid

3150-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

3213-464: The pioneering work of Fulton, Alexander used male calling song, life history and crosses between putative species to revise the taxonomy of gryllines in the eastern United States, and recognized five species, although at the time they were still classified in the genus Acheta . Species in this genus often look similar to species from other genera. They can often only be distinguished by the male genitalia. The epithallus typically consists of three lobes,

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3276-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

3339-470: The same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but the French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or

3402-408: The scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of a species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in

3465-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

3528-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

3591-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

3654-400: The tooth row also differs, with Beringian wolves having much higher frequencies of fracture for incisors, carnassials, and molars. A similar pattern was observed in spotted hyenas, suggesting that increased incisor and carnassial fracture reflects habitual bone consumption because bones are gnawed with the incisors and then cracked with the carnassials and molars. The gray wolf ( C. lupus ),

3717-409: The upper-case letters I to denote incisors , C for canines , P for premolars , and M for molars , and the lower-case letters i, c, p and m to denote the mandible teeth . Teeth are numbered using one side of the mouth and from the front of the mouth to the back. In carnivores , the upper premolar P4 and the lower molar m1 form the carnassials that are used together in a scissor-like action to shear

3780-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

3843-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

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3906-709: The vulnerable individuals of large prey, and a pack of timber wolves can bring down a 500 kg (1,100 lb) moose. The genus Canis contains many different species and has a wide range of different mating systems that varies depending on the type of canine and the species. In a study done in 2017, it was found that in some species of canids females use their sexual status to gain food resources. The study looked at wolves and dogs. Wolves are typically monogamous and form pair-bonds ; whereas dogs are promiscuous when free-range and mate with multiple individuals. The study found that in both species females tried to gain access to food more and were more successful in monopolizing

3969-627: Was published in the 10th edition of Systema Naturae and included the dog-like carnivores: the domestic dog, wolves, coyotes and jackals. All species within Canis are phylogenetically closely related with 78 chromosomes and can potentially interbreed . In 1926, the International Commission on Zoological Nomenclature (ICZN) in Opinion 91 included Genus Canis on its Official Lists and Indexes of Names in Zoology . In 1955,

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