118-507: Triticum monococcum subsp. monococcum Triticum monococcum ssp. boeoticum Einkorn wheat (from German Einkorn , literally "single grain") is either a wild species of wheat ( Triticum ) or its domesticated form . The wild form is T. boeoticum (syn. T. m. subsp. boeoticum ), and the domesticated form is T. monococcum (syn. T. m. subsp. monococcum ). Einkorn is a diploid species (2 n = 14 chromosomes) of hulled wheat, with tough glumes ( husks ) that tightly enclose
236-479: A blacksmith god, and Batradz , a mighty hero. The folklore of the Caucasus shows ancient Iranian Zoroastrian influence, involve battles with ancient Goths , Huns and Khazars , and contain many connections with ancient Indian , Norse Scandinavian , and Greek cultures. Caucasian folklore contains many links with the myths of the ancient Greeks. There are resemblances between the mother goddess Satanaya and
354-438: A combination of artificial and natural selection . This complexity and diversity of status has led to much confusion in the naming of wheats. Hexaploid species (6N) Tetraploid species (4N) Diploid species (2N) The wild species of wheat, along with the domesticated varieties einkorn , emmer and spelt , have hulls. This more primitive morphology (in evolutionary terms) consists of toughened glumes that tightly enclose
472-535: A decline in climate resilience of wheat has been observed. Hunter-gatherers in West Asia harvested wild wheats for thousands of years before they were domesticated , perhaps as early as 21,000 BC, but they formed a minor component of their diets. In this phase of pre-domestication cultivation, early cultivars were spread around the region and slowly developed the traits that came to characterise their domesticated forms. Repeated harvesting and sowing of
590-409: A good crop of grain. The farmer may intervene while the crop is growing to add fertilizer , water by irrigation , or pesticides such as herbicides to kill broad-leaved weeds or insecticides to kill insect pests. The farmer may assess soil minerals, soil water, weed growth, or the arrival of pests to decide timely and cost-effective corrective actions, and crop ripeness and water content to select
708-470: A less reliable method based on grain size. Einkorn and emmer are considered two of the founder crops cultivated by the first farming societies in Neolithic West Asia. These communities also cultivated naked wheats ( T. aestivum and T. durum ) and a now-extinct domesticated form of Zanduri wheat ( T. timopheevii ), as well as a wide variety of other cereal and non-cereal crops. Wheat
826-681: A low quality for human nutrition, according to the DIAAS protein quality evaluation method. Though they contain adequate amounts of the other essential amino acids, at least for adults, wheat proteins are deficient in the essential amino acid lysine . Because the proteins present in the wheat endosperm (gluten proteins) are particularly poor in lysine, white flours are more deficient in lysine compared with whole grains. Significant efforts in plant breeding are made to develop lysine-rich wheat varieties, without success, as of 2017 . Supplementation with proteins from other food sources (mainly legumes )
944-450: A major contribution to the nutritional welfare of the poor. Some wheat species are diploid , with two sets of chromosomes , but many are stable polyploids , with four sets of chromosomes ( tetraploid ) or six ( hexaploid ). Einkorn wheat ( Triticum monococcum ) is diploid (AA, two complements of seven chromosomes, 2n=14). Most tetraploid wheats (e.g. emmer and durum wheat) are derived from wild emmer , T. dicoccoides . Wild emmer
1062-475: A major contributor to increased grain output. More widely over the last 40 years, a massive increase in fertilizer use together with the increased availability of semi-dwarf varieties in developing countries, has greatly increased yields per hectare. In developing countries, use of (mainly nitrogenous) fertilizer increased 25-fold in this period. However, farming systems rely on much more than fertilizer and breeding to improve productivity. A good illustration of this
1180-470: A source of straw , which could be used for fuel, wicker-making , or wattle and daub construction. Domestic wheat was quickly spread to regions where its wild ancestors did not grow naturally. Emmer was introduced to Cyprus as early as 8600 BC and einkorn c. 7500 BC ; emmer reached Greece by 6500 BC, Egypt shortly after 6000 BC, and Germany and Spain by 5000 BC. "The early Egyptians were developers of bread and
1298-510: A spit, heat it up, stab it into the giant's eye, and escape. There are also links with the ancient Greek myth of Prometheus . Many legends, widespread in the Caucasus, contain motifs shared with the Prometheus story. These motifs include a giant hero, his conflict with God or gods, the stealing of fire and giving it to men, being chained, and being tormented by a bird who pecks at his liver (or heart). The Adyge / Circassian Nart Nasran,
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#17327908462591416-626: A tight clump which inhibits germination and makes the mutant seedlings susceptible to disease. But harvesting einkorn with intact seed heads was easier for early human harvesters, who could then manually break apart the seed heads and scatter any kernels not eaten. Over time and through selection, conscious or unconscious, the human preference for intact seed heads created the domestic variety, which has slightly larger kernels than wild einkorn. Domesticated einkorn thus requires human planting and harvesting for its continuing existence. This process of domestication may have taken only 20 to 200 years, resulting in
1534-426: A viable crop. When the use of seed drills replaced broadcasting sowing of seed in the 18th century, another great increase in productivity occurred. Yields of pure wheat per unit area increased as methods of crop rotation were applied to land that had long been in cultivation, and the use of fertilizers became widespread. Improved agricultural husbandry has more recently included pervasive automation , starting with
1652-418: A wheat plant is known as the flag leaf. It is denser and has a higher photosynthetic rate than other leaves, to supply carbohydrate to the developing ear. In temperate countries the flag leaf, along with the second and third highest leaf on the plant, supply the majority of carbohydrate in the grain and their condition is paramount to yield formation. Wheat is unusual among plants in having more stomata on
1770-499: A wheat that was easier to harvest. Einkorn wheat is one of the earliest cultivated forms of wheat, alongside emmer wheat ( T. dicoccum ). Hunter gatherers in the Fertile Crescent may have started harvesting einkorn as early as 30,000 years ago, according to archaeological evidence from Syria . Although gathered from the wild for thousands of years, einkorn wheat was first domesticated approximately 10,000 years BP in
1888-506: A year's supply. Wheat grain was stored after threshing , with the chaff removed. It was then processed into flour using ground stone mortars . Bread made from ground einkorn and the tubers of a form of club rush ( Bolboschoenus glaucus ) was made as early as 12,400 BC. At Çatalhöyük ( c. 7100–6000 BC ), both wholegrain wheat and flour was used to prepare bread, porridge and gruel . Apart from food, wheat may also have been important to Neolithic societies as
2006-452: Is Australian wheat growing in the southern winter cropping zone, where, despite low rainfall (300 mm), wheat cropping is successful even with relatively little use of nitrogenous fertilizer. This is achieved by crop rotation with leguminous pastures. The inclusion of a canola crop in the rotations has boosted wheat yields by a further 25%. In these low rainfall areas, better use of available soil-water (and better control of soil erosion)
2124-399: Is a "covered wheat" as its kernels do not break free from its seed coat ( glume ) with threshing. This makes it difficult to separate the husk from the seed. Einkorn is a common food in northern Provence (France). It is used for bulgur or as animal feed in mountainous areas of countries including France, India, Italy, Morocco, the former Yugoslavia , and Turkey. It contains gluten (so
2242-678: Is a region spanning Eastern Europe and Western Asia . It is situated between the Black Sea and the Caspian Sea , mainly comprising Armenia , Azerbaijan , Georgia , and parts of Southern Russia . The Caucasus Mountains , including the Greater Caucasus range, have conventionally been considered as a natural barrier between Europe and Asia , bisecting the Eurasian landmass. Mount Elbrus , Europe 's highest mountain,
2360-574: Is a rich source (20% or more of the Daily Value , DV) of multiple dietary minerals , such as manganese , phosphorus , magnesium , zinc , and iron (table). The B vitamins , niacin (36% DV), thiamine (33% DV), and vitamin B6 (23% DV), are present in significant amounts (table). Wheat is a significant source of vegetable proteins in human food, having a relatively high protein content compared to other major cereals. However, wheat proteins have
2478-400: Is achieved by retaining the stubble after harvesting and by minimizing tillage. Pests and diseases consume 21.47% of the world's wheat crop annually. There are many wheat diseases, mainly caused by fungi, bacteria, and viruses . Plant breeding to develop new disease-resistant varieties, and sound crop management practices are important for preventing disease. Fungicides, used to prevent
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#17327908462592596-459: Is also important to identify periods of higher risk from the climate. Farmers benefit from knowing when the 'flag leaf' (last leaf) appears, as this leaf represents about 75% of photosynthesis reactions during the grain filling period, and so should be preserved from disease or insect attacks to ensure a good yield. Several systems exist to identify crop stages, with the Feekes and Zadoks scales being
2714-402: Is commonly used to compensate for this deficiency, since the limitation of a single essential amino acid causes the others to break down and become excreted, which is especially important during growth. Consumed worldwide by billions of people, wheat is a significant food for human nutrition, particularly in the least developed countries where wheat products are primary foods. When eaten as
2832-609: Is considered by some sources to be the dividing line between Europe and Southwest Asia . According to that, the highest peak in the Caucasus, Mount Elbrus (5,642 meters) located in western Ciscaucasus, is considered the highest point in Europe. The Kuma-Manych Depression , the geologic depression that divides the Russian Plain from the North Caucasus foreland is often regarded by classical and non-British sources as
2950-418: Is greater than for all other crops combined. Global demand for wheat is increasing due to the unique viscoelastic and adhesive properties of gluten proteins, which facilitate the production of processed foods, whose consumption is increasing as a result of the worldwide industrialization process and westernization of diets . Wheat became a central agriculture endeavor in the worldwide British Empire in
3068-477: Is greater than for all other crops combined. In 2021, world wheat production was 771 million tonnes (850 million short tons), making it the second most-produced cereal after maize (known as corn in North America and Australia; wheat is often called corn in countries including Britain). Since 1960, world production of wheat and other grain crops has tripled and is expected to grow further through
3186-473: Is itself the result of a hybridization between two diploid wild grasses, T. urartu and a wild goatgrass such as Ae. speltoides . The hybridization that formed wild emmer (AABB, four complements of seven chromosomes in two groups, 4n=28) occurred in the wild, long before domestication, and was driven by natural selection . Hexaploid wheats evolved in farmers' fields as wild emmer hybridized with another goatgrass, Ae. squarrosa or Ae. tauschii , to make
3304-629: Is likely to be prioritised in drought-adapted crops, while stem non-structural carbohydrate is prioritised in varieties developed for countries where disease is a bigger issue. Depending on variety, wheat may be awned or not awned. Producing awns incurs a cost in grain number, but wheat awns photosynthesise more efficiently than their leaves with regards to water usage, so awns are much more frequent in varieties of wheat grown in hot drought-prone countries than those generally seen in temperate countries. For this reason, awned varieties could become more widely grown due to climate change . In Europe, however,
3422-558: Is likewise called the Caucasus by many." In the Tale of Past Years (1113 AD), it is stated that Old East Slavic Кавкасийскыѣ горы ( Kavkasijskyě gory ) came from Ancient Greek Καύκασος ( Kaúkasos ), which, according to M. A. Yuyukin, is a compound word that can be interpreted as the 'mountain of the seagull(s)' (καύ-: καύαξ, καύηξ, -ηκος, κήξ, κηϋξ 'a kind of seagull' + the reconstructed *κάσος 'mountain' or 'rock' richly attested both in place and personal names). In Georgian tradition,
3540-458: Is more diverse, with domesticated varieties falling into two major groups: hulled or non-shattering, in which threshing separates the whole spikelet ; and free-threshing, where the individual grains are separated. Both varieties probably existed in prehistory, but over time free-threshing cultivars became more common. Wild emmer was first cultivated in the southern Levant , as early as 9600 BC. Genetic studies have found that, like einkorn, it
3658-399: Is normally planted after tilling the soil by ploughing and then harrowing to kill weeds and create an even surface. The seeds are then scattered on the surface, or drilled into the soil in rows. Winter wheat lies dormant during a winter freeze. It needs to develop to a height of 10 to 15 cm before the cold intervenes, so as to be able to survive the winter; it requires a period with
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3776-411: Is not suitable for people with gluten-related disorders ) and has a higher percentage of protein than modern red wheats. It is considered more nutritious because it has higher levels of fat , phosphorus , potassium , pyridoxine , and beta-carotene . Einkorn is the source of many potential introgressions for immunity – Nikolai Vavilov called it an "accumulator of complex immunities." T. monococcum
3894-511: Is not the same as a wheat allergy . Other diseases triggered by eating wheat are non-coeliac gluten sensitivity (estimated to affect 0.5% to 13% of the general population ), gluten ataxia , and dermatitis herpetiformis . Certain short-chain carbohydrates present in wheat, known as FODMAPs (mainly fructose polymers ), may be the cause of non-coeliac gluten sensitivity. As of 2019 , reviews have concluded that FODMAPs only explain certain gastrointestinal symptoms, such as bloating , but not
4012-840: Is situated in the Western Caucasus area of Russia . On the southern side, the Lesser Caucasus includes the Javakheti Plateau and the Armenian highlands , part of which is in Turkey. The Caucasus is divided into the North Caucasus and South Caucasus , although the Western Caucasus also exists as a distinct geographic space within the North Caucasus. The Greater Caucasus mountain range in
4130-499: Is speculative given that Pelasgian is so poorly known. The term Caucasus is not only used for the mountains themselves but also includes Ciscaucasia (which is part of the Russian Federation ) and Transcaucasia . According to Alexander Mikaberidze , Transcaucasia is a "Russo-centric" term. The Transcaucasus region and Dagestan were the furthest points of Parthian and later Sasanian expansions, with areas to
4248-445: Is the source of Sr21 , a stem rust resistance gene which has been introgressed into hexaploid worldwide. It is also the source of Yr34 , a resistance gene for yellow rust . The salt-tolerance feature of T. monococcum has been bred into durum wheat. Wheat Wheat is a grass widely cultivated for its seed , a cereal grain that is a staple food around the world. The many species of wheat together make up
4366-493: Is valuable to impart viscoelastic functional qualities in dough , enabling the preparation of diverse processed foods such as breads, noodles, and pasta that facilitate wheat consumption. Raw red winter wheat is 13% water, 71% carbohydrates including 12% dietary fiber , 13% protein , and 2% fat (table). Some 75–80% of the protein content is as gluten . In a reference amount of 100 grams (3.5 oz), wheat provides 1,368 kilojoules (327 kilocalories) of food energy and
4484-502: Is variously a one-eyed rock-throwing cannibal, who lives in a cave (the exit of which is often blocked by a stone), kills the hero's companions, is blinded by a hot stake, and whose flock of animals is stolen by the hero and his men, all motifs which (along with still others) are also found in the Polyphemus story. In one example from Georgia , two brothers, who are being held prisoner by a giant one-eyed shepherd called "One-eye", take
4602-694: The Achaemenid Empire , Parthia , and the Sassanid Empire , who would altogether rule the Caucasus for many hundreds of years. In 95–55 BC, under the reign of the Armenian king Tigranes the Great , the Kingdom of Armenia included Kingdom of Armenia, vassals Iberia, Albania, Parthia, Atropatene , Mesopotamia , Cappadocia , Cilicia , Syria , Nabataean kingdom , and Judea . By the time of
4720-642: The Caucasus , the Balkans, and central Europe. Einkorn wheat was more commonly grown in cooler climates than emmer wheat , the other domesticated wheat. Cultivation of einkorn in the Middle East began to decline in favor of emmer wheat around 2000 BC. Cultivation of einkorn was never extensive in Italy , southern France , and Spain . Einkorn continued to be cultivated in some areas of northern Europe throughout
4838-741: The Georgian Amirani , the Chechen Pkharmat , and the Abkhazian Abrskil , are examples of such Prometheus-like figures. The Caucasus is an area of great ecological importance. The region is included in the list of 34 world biodiversity hotspots . It harbors some 6400 species of higher plants, 1600 of which are endemic to the region. Its wildlife includes Persian leopards , brown bears , wolves , bison , marals , golden eagles and hooded crows . Among invertebrates , some 1000 spider species are recorded in
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4956-773: The Greater Caucasus mountain range. It consists of Southern Russia , mainly the North Caucasian Federal District 's autonomous republics and the Krais in Southern Russia, and the northernmost parts of Georgia and Azerbaijan . The North Caucasus lies between the Black Sea to its west, the Caspian Sea to its east, and borders the Southern Federal District to its north. The two Federal Districts are collectively referred to as "Southern Russia". The South Caucasus borders
5074-622: The Kartvelian , Northwest Caucasian , and Northeast Caucasian language families are indigenous to the area. Pliny the Elder 's Natural History (77–79 AD) derives the name of the Caucasus from a Scythian name, Croucasis , which supposedly means 'shimmering with snow'. German linguist Paul Kretschmer notes that the Latvian word kruvesis also means 'frozen mud'. Isidore of Seville 's Etymologies ( c. 625 AD) also says
5192-549: The Middle Ages and until the early part of the 20th century. Wild and domesticated einkorns are diploid wheats. Unlike emmer and bread wheat , which were formed by hybridisation with Aegilops goatgrasses, einkorn is not a hybrid. Einkorn wheat is low-yielding but can survive on poor, dry, marginal soils where other varieties of wheat will not. It is primarily eaten boiled in whole grains or in porridge. As with other ancient varieties of wheat such as emmer , Einkorn
5310-745: The Muslim conquest of Persia , large parts of the region came under the rule of the Arabs , and Islam penetrated the region. In the 10th century, the Alans (proto- Ossetians ) founded the Kingdom of Alania , that flourished in the Northern Caucasus , roughly in the location of latter-day Circassia and modern North Ossetia–Alania , until its destruction by the Mongol invasion in 1238–39. During
5428-755: The Ottoman Empire . In the second half of the 19th century, the Russian Empire also conquered the North Caucasus. In the aftermath of the Caucasian Wars , the Russian military perpetrated an ethnic cleansing of Circassians , expelling this indigenous population from its homeland. Between the 1850s and World War I, about a million North Caucasian Muslims arrived in the Ottoman Empire as refugees. Having killed and deported most of
5546-512: The Pre-Pottery Neolithic A (PPNA) or B (PPNB) periods. Evidence from DNA fingerprinting suggests einkorn was first domesticated near Karaca Dağ in southeast Turkey, an area in which a number of PPNB farming villages have been found. One theory by Yuval Noah Harari suggests that the domestication of einkorn was linked to intensive agriculture to support the nearby Göbekli Tepe site. An important characteristic facilitating
5664-606: The Simurgh . The Roman poet Ovid placed the Caucasus in Scythia and depicted it as a cold and stony mountain which was the abode of personified hunger. The Greek hero Jason sailed to the west coast of the Caucasus in pursuit of the Golden Fleece , and there met Medea , a daughter of King Aeëtes of Colchis . The Caucasus has a rich folklore tradition. This tradition has been preserved orally—necessitated by
5782-436: The extra-digestive symptoms that people with non-coeliac gluten sensitivity may develop. Other wheat proteins, amylase-trypsin inhibitors, have been identified as the possible activator of the innate immune system in coeliac disease and non-coeliac gluten sensitivity. These proteins are part of the plant's natural defense against insects and may cause intestinal inflammation in humans. In 2022, world wheat production
5900-678: The grains . The cultivated form is similar to the wild, except that the ear stays intact when ripe and the seeds are larger. The domestic form is known as petit épeautre in French, Einkorn in German, "einkorn" or "littlespelt" in English, piccolo farro in Italian and escanda menor in Spanish. The name refers to the fact that each spikelet contains only one grain. Einkorn wheat
6018-853: The hexaploid wheats including bread wheat . A 2007 molecular phylogeny of the wheats gives the following not fully-resolved cladogram of major cultivated species; the large amount of hybridisation makes resolution difficult. Markings like "6N" indicate the degree of polyploidy of each species: Barley 2N, rye 2N/4N, and other cereals Triticum monococcum ( einkorn ) 2N × Aegilotriticum hybrids ( Aegilops x Triticum ) 6N Triticum timopheevii (zanduri wheat) and others 4N Triticum aestivum ( common or bread wheat ) 6N Triticum durum/turgidum ( durum wheat) 4N Triticum spelta ( spelt ) 6N Triticum turanicum ( khorasan wheat ) 4N Triticum dicoccum ( emmer ) 4N many other species During 10,000 years of cultivation, numerous forms of wheat, many of them hybrids , have developed under
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#17327908462596136-488: The whole grain , wheat is a source of multiple nutrients and dietary fiber . In a small part of the general population, gluten – which comprises most of the protein in wheat – can trigger coeliac disease , noncoeliac gluten sensitivity , gluten ataxia , and dermatitis herpetiformis . Wheat is a stout grass of medium to tall height. Its stem is jointed and usually hollow, forming a straw. There can be many stems on one plant. It has long narrow leaves, their bases sheathing
6254-427: The whole grain , wheat supplies multiple nutrients and dietary fiber recommended for children and adults. In genetically susceptible people, wheat gluten can trigger coeliac disease . Coeliac disease affects about 1% of the general population in developed countries . The only known effective treatment is a strict lifelong gluten-free diet . While coeliac disease is caused by a reaction to wheat proteins, it
6372-690: The "inhabitants of the southern coast of the Black Sea ". It was also noted that in Nakh Ков гас ( Kov gas ) means "gateway to steppe". The modern endonym for the region is usually similar in many languages, and is generally between Kavkaz and Kaukaz . The North Caucasus region is also known as the Ciscaucasus , whereas the South Caucasus region is alternatively known as the Transcaucasus . The North Caucasus contains most of
6490-617: The 19th century, and remains of great importance in Australia, Canada and India. In Australia, with vast lands and a limited work force, expanded production depended on technological advances, especially regarding irrigation and machinery. By the 1840s there were 900 growers in South Australia . They used "Ridley's Stripper", a reaper-harvester perfected by John Ridley in 1843, to remove the heads of grain. In Canada, modern farm implements made large scale wheat farming possible from
6608-513: The 20th century, global wheat output expanded by about 5-fold, but until about 1955 most of this reflected increases in wheat crop area, with lesser (about 20%) increases in crop yields per unit area. After 1955 however, there was a ten-fold increase in the rate of wheat yield improvement per year, and this became the major factor allowing global wheat production to increase. Thus technological innovation and scientific crop management with synthetic nitrogen fertilizer , irrigation and wheat breeding were
6726-742: The Armenians of Western Armenia during the Armenian genocide , the Turks intended to eliminate the Armenian population of Eastern Armenia . During the 1920 Turkish–Armenian War , 60,000 to 98,000 Armenian civilians were estimated to have been killed by the Turkish army. In the 1940s, around 480,000 Chechens and Ingush , 120,000 Karachay – Balkars and Meskhetian Turks , thousands of Kalmyks , and 200,000 Kurds in Nakchivan and Caucasus Germans were deported en masse to Central Asia and Siberia by
6844-578: The Caucasus region. As a result of her military campaigns and the temporary fall of the Byzantine Empire in 1204, Georgia became the strongest Christian state in the whole Near East area, encompassing most of the Caucasus stretching from Northern Iran and Northeastern Turkey to the North Caucasus. The Caucasus region was conquered by the Ottomans , Turco-Mongols , local kingdoms and khanates, as well as, once again, Iran . Up to and including
6962-712: The Caucasus was usually incorporated into the Iranian world . At the beginning of the 19th century, the Russian Empire conquered the territory from Qajar Iran . The territory of the Caucasus region was inhabited by Homo erectus since the Paleolithic Era . In 1991, early Hominini fossils dating back 1.8 million years were found at the Dmanisi archaeological site in Georgia . Scientists now classify
7080-660: The Caucasus. Most of arthropod biodiversity is concentrated on Great and Lesser Caucasus ranges. The region has a high level of endemism and several relict animals and plants, the fact reflecting the presence of refugial forests, which survived the Ice Age in the Caucasus Mountains. The Caucasus forest refugium is the largest throughout the Western Asian (near Eastern) region. The area has multiple representatives of disjunct relict groups of plants with
7198-669: The Greater Caucasus range and Southern Russia to its north, the Black Sea and Turkey to its west, the Caspian Sea to its east, and Iran to its south. It contains the Lesser Caucasus mountain range and surrounding lowlands. All of Armenia , Azerbaijan (excluding the northernmost parts), and Georgia (excluding the northernmost parts) are in the South Caucasus. The watershed along the Greater Caucasus range
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#17327908462597316-460: The Greek goddess of love Aphrodite . The story of how the trickster Nart Sosruquo, became invulnerable parallels that of the Greek hero Achilles . The ancient Greek Amazons may be connected to a Caucasian "warrior Forest-Mother, Amaz-an". Caucasian legends include stories involving giants similar to Homer 's Polyphemus story. In these stories, the giant is almost always a shepherd , and he
7434-642: The Middle Ages, Bagratid Armenia , Kingdom of Tashir-Dzoraget , Kingdom of Syunik and Principality of Khachen organized local Armenian population facing multiple threats after the fall of antique Kingdom of Armenia . Caucasian Albania maintained close ties with Armenia and the Church of Caucasian Albania shared the same Christian dogmas with the Armenian Apostolic Church and had a tradition of their Catholicos being ordained through
7552-662: The Patriarch of Armenia. In the 12th century, the Georgian king David the Builder drove the Muslims out of the Caucasus and made the Kingdom of Georgia a strong regional power. In 1194–1204 Georgian Queen Tamar 's armies crushed new Seljuk Turkish invasions from the southeast and south and launched several successful campaigns into Seljuk Turkish-controlled Southern Armenia. The Georgian Kingdom continued military campaigns in
7670-502: The Russian Federation. The Russian divisions include Dagestan , Chechnya , Ingushetia , North Ossetia–Alania , Kabardino–Balkaria , Karachay–Cherkessia , Adygea , Krasnodar Krai , and Stavropol Krai , in clockwise order. Two territories in the region claim independence but are recognized as such by only a handful of entities: Abkhazia , and South Ossetia . Abkhazia and South Ossetia are largely recognized by
7788-445: The Southern Caucasus (except western Georgia), northwestern Iran, the northeastern Caucasus, eastern Turkey, and as far as Syria. Under Ashurbanipal (669–627 BC), the boundaries of the Assyrian Empire reached as far as the Caucasus Mountains. Later ancient kingdoms of the region included Armenia , Albania , Colchis and Iberia , among others. These kingdoms were later incorporated into various Iranian empires, including Media ,
7906-543: The Soviet security apparatus. About a quarter of them died. The Southern Caucasus region was unified as a single political entity twice – during the Russian Civil War ( Transcaucasian Democratic Federative Republic ) from 9 April 1918 to 26 May 1918, and under the Soviet rule ( Transcaucasian SFSR ) from 12 March 1922 to 5 December 1936. Following the dissolution of the Soviet Union in 1991, Georgia , Azerbaijan and Armenia became independent nations. The region has been subject to various territorial disputes since
8024-446: The area. Russian is used as a lingua franca most notably in the North Caucasus. The peoples of the northern and southern Caucasus mostly are Shia Muslims , Sunni Muslims , Eastern Orthodox Christians or Armenian Christians . Located on the peripheries of Turkey , Iran , and Russia , the region has been an arena for political, military, religious, and cultural rivalries and expansionism for centuries. Throughout its history,
8142-464: The assemblage of fossil skeletons as the subspecies Homo erectus georgicus . The site yields the earliest unequivocal evidence for the presence of early humans outside the African continent; and the Dmanisi skulls are the five oldest hominins ever found outside Africa . Kura–Araxes culture from about 4000 BC until about 2000 BC enveloped a vast area of approximately 1,000 km by 500 km, and mostly encompassed, on modern-day territories,
8260-427: The closest relatives in Eastern Asia, southern Europe, and even North America. Over 70 species of forest snails of the region are endemic. Some relict species of vertebrates are Caucasian parsley frog , Caucasian salamander , Robert's snow vole , and Caucasian grouse , and there are almost entirely endemic groups of animals such as lizards of genus Darevskia . In general, the species composition of this refugium
8378-426: The collapse of the Soviet Union, leading to the First Nagorno-Karabakh War (1988–1994), the East Prigorodny Conflict (1989–1991), the War in Abkhazia (1992–93) , the First Chechen War (1994–1996), the Second Chechen War (1999–2009), Russo-Georgian War (2008), and the Second Nagorno-Karabakh War (2020). In Greek mythology , the Caucasus was one of the pillars supporting the world. After presenting man with
8496-414: The deepest of arable crops, extending as far down as 2 metres (6 ft 7 in). While the roots of a wheat plant are growing, the plant also accumulates an energy store in its stem, in the form of fructans , which helps the plant to yield under drought and disease pressure, but it has been observed that there is a trade-off between root growth and stem non-structural carbohydrate reserves. Root growth
8614-485: The domestic form comes after c. 8800 BC in southern Turkey, at Çayönü , Cafer Höyük , and possibly Nevalı Çori . Genetic evidence indicates that it was domesticated in multiple places independently. Wild emmer wheat ( T. turgidum subsp. dicoccoides ) is less widespread than einkorn, favouring the rocky basaltic and limestone soils found in the hilly flanks of the Fertile Crescent. It
8732-439: The domestication of einkorn and other annual grains is that the plants are largely self-pollinating. Thus, the desirable (for human management) traits of einkorn could be perpetuated at less risk of cross-fertilization with wild plants which might have traits – e.g. smaller seeds, shattering seed heads, as less desirable for human management. From the northern part of the Fertile Crescent, the cultivation of einkorn wheat spread to
8850-766: The earliest known wheat with sufficient gluten for yeasted breads was found in a granary at Assiros in Macedonia dated to 1350 BC. From the Middle East , wheat continued to spread across Europe and to the Americas in the Columbian exchange . In the British Isles, wheat straw ( thatch ) was used for roofing in the Bronze Age , and remained in common use until the late 19th century. White wheat bread
8968-834: The early 19th century, most of the Southern Caucasus and southern Dagestan all formed part of the Persian Empire . In 1813 and 1828 by the Treaty of Gulistan and the Treaty of Turkmenchay respectively, the Persians were forced to irrevocably cede the Southern Caucasus and Dagestan to Imperial Russia . In the ensuing years after these gains, the Russians took the remaining part of the Southern Caucasus , comprising western Georgia, through several wars from
9086-521: The fact that for most of the languages involved, there was no alphabet until the early twentieth century—and only began to be written down in the late nineteenth century. One important tradition is that of the Nart sagas , which tell stories of a race of ancient heroes called the Narts. These sagas include such figures as Satanaya , the mother of the Narts, Sosruquo a shape changer and trickster, Tlepsh
9204-567: The farm and because of poor roads, inadequate storage technologies, inefficient supply chains and farmers' inability to bring the produce into retail markets dominated by small shopkeepers. Some 10% of total wheat production is lost at farm level, another 10% is lost because of poor storage and road networks, and additional amounts are lost at the retail level. In the Punjab region of the Indian subcontinent, as well as North China, irrigation has been
9322-456: The farmer have a detailed understanding of each stage of development in the growing plants. In particular, spring fertilizers , herbicides , fungicides , and growth regulators are typically applied only at specific stages of plant development. For example, it is currently recommended that the second application of nitrogen is best done when the ear (not visible at this stage) is about 1 cm in size (Z31 on Zadoks scale ). Knowledge of stages
9440-421: The first century BC, Zoroastrianism had become the dominant religion of the region; however, the region would go through two other religious transformations. Owing to the strong rivalry between Persia and Rome , and later Byzantium . The Romans first arrived in the region in the 1st century BC with the annexation of the kingdom of Colchis, which was later turned into the province of Lazicum . The next 600 years
9558-466: The genus Triticum ( / ˈ t r ɪ t ɪ k ə m / ); the most widely grown is common wheat ( T. aestivum ). The archaeological record suggests that wheat was first cultivated in the regions of the Fertile Crescent around 9600 BC. Botanically, the wheat kernel is a caryopsis , a type of fruit . Wheat is grown on a larger area of land than any other food crop (220.7 million hectares or 545 million acres in 2021). World trade in wheat
9676-464: The gift of fire, Prometheus (or Amirani in the Georgian version ) was chained there by Zeus , to have his liver eaten daily by an eagle as punishment for defying Zeus's wish to keep the "secret of fire" from humans. In Persian mythology , the Caucasus might be associated with the mythic Mount Qaf which is believed to surround the known world. It is the battlefield of Saoshyant and the nest of
9794-601: The glumes are fragile and the rachis tough. On threshing, the chaff breaks up, releasing the grains. Wheat grain classes are named by colour, season, and hardness. The classes used in the United States are: Wheat is a staple cereal worldwide. Raw wheat berries can be ground into flour or, using hard durum wheat only, can be ground into semolina ; germinated and dried creating malt ; crushed or cut into cracked wheat; parboiled (or steamed), dried, crushed and de-branned into bulgur also known as groats . If
9912-431: The glumes. The flowers are grouped into spikelets , each with between two and six flowers. Each fertilised carpel develops into a wheat grain or berry; botanically a fruit , it is often called a seed. The grains ripen to a golden yellow; a head of grain is called an ear. Leaves emerge from the shoot apical meristem in a telescoping fashion until the transition to reproduction i.e. flowering. The last leaf produced by
10030-562: The grains of wild grasses led to the creation of domestic strains, as mutant forms ('sports') of wheat were more amenable to cultivation. In domesticated wheat, grains are larger, and the seeds (inside the spikelets ) remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to shatter easily, dispersing the spikelets. Selection for larger grains and non-shattering heads by farmers might not have been deliberately intended, but simply have occurred because these traits made gathering
10148-479: The grains, and (in domesticated wheats) a semi-brittle rachis that breaks easily on threshing. The result is that when threshed, the wheat ear breaks up into spikelets. To obtain the grain, further processing, such as milling or pounding, is needed to remove the hulls or husks. Hulled wheats are often stored as spikelets because the toughened glumes give good protection against pests of stored grain. In free-threshing (or naked) forms, such as durum wheat and common wheat,
10266-535: The hill country in the northern part of the Fertile Crescent and Anatolia , although it has a wider distribution reaching into the Balkans and south to Jordan near the Dead Sea . It is a short variety of wild wheat, usually less than 70 centimetres (28 in) tall and is not very productive of edible seeds. The principal difference between wild einkorn and cultivated einkorn is the method of seed dispersal. In
10384-519: The late 1840s. By 1879, Saskatchewan was the center, followed by Alberta , Manitoba and Ontario , as the spread of railway lines allowed easy exports to Britain. By 1910, wheat made up 22% of Canada's exports, rising to 25% in 1930 despite the sharp decline in prices during the worldwide Great Depression . Efforts to expand wheat production in South Africa, Kenya and India were stymied by low yields and disease. However, by 2000 India had become
10502-420: The main drivers of wheat output growth in the second half of the century. There were some significant decreases in wheat crop area, for instance in North America. Better seed storage and germination ability (and hence a smaller requirement to retain harvested crop for next year's seed) is another 20th-century technological innovation. In Medieval England, farmers saved one-quarter of their wheat harvest as seed for
10620-442: The middle of the 21st century. Global demand for wheat is increasing because of the usefulness of gluten to the food industry. Wheat is an important source of carbohydrates . Globally, it is the leading source of vegetable proteins in human food, having a protein content of about 13%, which is relatively high compared to other major cereals but relatively low in protein quality (supplying essential amino acids ). When eaten as
10738-568: The most widely used. Each scale is a standard system which describes successive stages reached by the crop during the agricultural season. For example, the stage of pollen formation from the mother cell, and the stages between anthesis and maturity, are susceptible to high temperatures, and this adverse effect is made worse by water stress. Technological advances in soil preparation and seed placement at planting time, use of crop rotation and fertilizers to improve plant growth, and advances in harvesting methods have all combined to promote wheat as
10856-487: The name means shining white like snow: "Thus, toward the east, where it rises to a greater height, it is called the Caucasus, due to the whiteness of its snow, for in an eastern language, caucasus means “white,” that is, shining white with a very thick snow cover. For the same reason the Scythians, who live next to this mountain range, call it Croacasim, for among them whiteness or snow is called casim. 3. The Taurus range
10974-429: The natural and historical boundary between Europe and Asia. Another opinion is that the rivers Kura and Rioni mark this border, or even that of the river Aras . The Caucasus is a linguistically , culturally and geographically diverse region. The nation states that compose the Caucasus today are the post-Soviet states Georgia (including Adjara and Abkhazia ), Azerbaijan (including Nakhchivan ), Armenia , and
11092-488: The need to systematically remove vegetation or till the soil. They may also have exploited natural wetlands and floodplains to practice décrue farming , sowing seeds in the soil left behind by receding floodwater. It was harvested with stone-bladed sickles . The ease of storing wheat and other cereals led farming households to become gradually more reliant on it over time, especially after they developed individual storage facilities that were large enough to hold more than
11210-473: The next crop, leaving only three-quarters for food and feed consumption. By 1999, the global average seed use of wheat was about 6% of output. In the 21st century, rising temperatures associated with global warming are reducing wheat yield in several locations. Wheat is an annual crop. It can be planted in autumn and harvested in early summer as winter wheat in climates that are not too severe, or planted in spring and harvested in autumn as spring wheat. It
11328-417: The north is mostly shared by Russia and Georgia as well as the northernmost parts of Azerbaijan . The Lesser Caucasus mountain range in the south is occupied by several independent states, mostly by Armenia, Azerbaijan, and Georgia, but also extends to parts of northeastern Turkey , and northern Iran . The region is known for its linguistic diversity : aside from Indo-European and Turkic languages,
11446-526: The north of the Greater Caucasus range practically impregnable. The mythological Mount Qaf , the world's highest mountain that ancient Iranian lore shrouded in mystery, was said to be situated in this region. The region is also one of the candidates for the location of Airyanem Vaejah , the apparent homeland of the Iranians of Zoroaster . In Middle Persian sources of the Sasanian era, the Caucasus range
11564-461: The raw wheat is broken into parts at the mill, as is usually done, the outer husk or bran can be used in several ways. Wheat is a major ingredient in such foods as bread , porridge , crackers , biscuits , muesli , pancakes , pasta , pies , pastries , pizza , semolina , cakes , cookies , muffins , rolls , doughnuts , gravy , beer , vodka , boza (a fermented beverage ), and breakfast cereals . In manufacturing wheat products, gluten
11682-452: The right moment to harvest. Harvesting involves reaping , cutting the stems to gather the crop; and threshing , breaking the ears to release the grain; both steps are carried out by a combine harvester . The grain is then dried so that it can be stored safe from mould fungi. Wheat normally needs between 110 and 130 days between sowing and harvest, depending upon climate, seed type, and soil conditions. Optimal crop management requires that
11800-517: The second largest producer of wheat in the world. In the 19th century the American wheat frontier moved rapidly westward. By the 1880s 70% of American exports went to British ports. The first successful grain elevator was built in Buffalo in 1842. The cost of transport fell rapidly. In 1869 it cost 37 cents to transport a bushel of wheat from Chicago to Liverpool . In 1905 it was 10 cents. In
11918-513: The seeds easier; nevertheless such 'incidental' selection was an important part of crop domestication . As the traits that improve wheat as a food source involve the loss of the plant's natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild. Wild einkorn wheat ( T. monococcum subsp. boeoticum ) grows across Southwest Asia in open parkland and steppe environments. It comprises three distinct races , only one of which, native to Southeast Anatolia ,
12036-485: The significant crop losses from fungal disease, can be a significant variable cost in wheat production. Estimates of the amount of wheat production lost owing to plant diseases vary between 10 and 25% in Missouri. A wide range of organisms infect wheat, of which the most important are viruses and fungi. Caucasus The Caucasus ( / ˈ k ɔː k ə s ə s / ) or Caucasia ( / k ɔː ˈ k eɪ ʒ ə / ),
12154-417: The stem, one above each joint. At the top of the stem is the flower head, containing some 20 to 100 flowers. Each flower contains both male and female parts. The flowers are wind-pollinated , with over 99% of pollination events being self-pollinations and the rest cross-pollinations . The flower is housed in a pair of small leaflike glumes . The two (male) stamens and (female) stigmas protrude outside
12272-405: The temperature at or near freezing, its dormancy then being broken by the thaw or rise in temperature. Spring wheat does not undergo dormancy. Wheat requires a deep soil , preferably a loam with organic matter, and available minerals including soil nitrogen, phosphorus, and potassium. An acid and peaty soil is not suitable. Wheat needs some 30 to 38 cm of rain in the growing season to form
12390-718: The term Caucasus is derived from Caucas ( Georgian : კავკასოსი Ḳavḳasosi ), the son of the Biblical Togarmah and legendary forefather of the Nakh peoples . According to German philologists Otto Schrader and Alfons A. Nehring, the Ancient Greek word Καύκασος ( Kaukasos ) is connected to Gothic hauhs 'high' as well as Lithuanian kaũkas 'hillock' and kaukarà 'hill, top', Russian куча 'heap'. British linguist Adrian Room claims that * kau- also means 'mountain' in Pelasgian , though this
12508-407: The upper ( adaxial ) side of the leaf, than on the under ( abaxial ) side. It has been theorised that this might be an effect of it having been domesticated and cultivated longer than any other plant. Winter wheat generally produces up to 15 leaves per shoot and spring wheat up to 9 and winter crops may have up to 35 tillers (shoots) per plant (depending on cultivar). Wheat roots are among
12626-614: The use of threshing machines , and progressing to large and costly machines like the combine harvester which greatly increased productivity. At the same time, better varieties such as Norin 10 wheat , developed in Japan in the 1930s, or the dwarf wheat developed by Norman Borlaug in the Green Revolution , greatly increased yields. In addition to gaps in farming system technology and knowledge, some large wheat grain-producing countries have significant losses after harvest at
12744-542: The use of the oven and developed baking into one of the first large-scale food production industries." By 4000 BC, wheat had reached the British Isles and Scandinavia . Wheat likely appeared in China 's lower Yellow River around 2600 BC. The oldest evidence for hexaploid wheat has been confirmed through DNA analysis of wheat seeds, dating to around 6400–6200 BC, recovered from Çatalhöyük . As of 2023,
12862-410: The wild variety the seed head usually shatters and drops the kernels (seeds) of wheat onto the ground. This facilitates a new crop of wheat. In the domestic variety, the seed head remains intact. While such a mutation may occasionally occur in the wild, it is not viable there in the long term: the intact seed head will only drop to the ground when the stalk rots, and the kernels will not scatter but form
12980-417: The world community as part of Georgia. The region has many different languages and language families. There are more than 50 ethnic groups living in the region. No fewer than three language families are unique to the area. In addition, Indo-European languages, such as East Slavic , Armenian and Ossetian , and Turkic languages , such as Azerbaijani , Kumyk language and Karachay–Balkar , are spoken in
13098-486: Was 808.4 million tonnes, led by China, India, and Russia which collectively provided 43.22% of the world total. As of 2019 , the largest exporters were Russia (32 million tonnes), United States (27), Canada (23) and France (20), while the largest importers were Indonesia (11 million tonnes), Egypt (10.4) and Turkey (10.0). In 2021, wheat was grown on 220.7 million hectares or 545 million acres worldwide, more than any other food crop. World trade in wheat
13216-462: Was domesticated in southeastern Anatolia, but only once. The earliest secure archaeological evidence for domestic emmer comes from Çayönü, c. 8300–7600 BC , where distinctive scars on the spikelets indicated that they came from a hulled domestic variety. Slightly earlier finds have been reported from Tell Aswad in Syria, c. 8500–8200 BC , but these were identified using
13334-429: Was domesticated. The main feature that distinguishes domestic einkorn from wild is that its ears do not shatter without pressure, making it dependent on humans for dispersal and reproduction. It also tends to have wider grains. Wild einkorn was collected at sites such as Tell Abu Hureyra ( c. 10,700–9000 BC ) and Mureybet ( c. 9800–9300 BC ), but the earliest archaeological evidence for
13452-601: Was historically a high status food, but during the nineteenth century it became in Britain an item of mass consumption, displacing oats , barley and rye from diets in the North of the country. It became "a sign of a high degree of culture". After 1860, the enormous expansion of wheat production in the United States flooded the world market, lowering prices by 40%, and (along with the expansion of potato growing) made
13570-534: Was marked by a conflict between Rome and Sassanid Empire for the control of the region. In western Georgia the eastern Roman rule lasted until the Middle Ages. As the Arsacid dynasty of Armenia (an eponymous branch of the Arsacid dynasty of Parthia ) was the first nation to adopt Christianity as state religion (in 301 AD), and Caucasian Albania and Georgia had become Christian entities, Christianity began to overtake Zoroastrianism and pagan beliefs. With
13688-445: Was one of the first plants to be domesticated and cultivated . The earliest clear evidence of the domestication of einkorn dates from 10,600 to 9,900 years before present (8650 BCE to 7950 BCE) from Çayönü and Cafer Höyük , two Early Pre-Pottery Neolithic B archaeological sites in southern Turkey . Remnants of einkorn were found with the iceman mummy Ötzi , dated the late 4th millenium BCE. Einkorn wheat commonly grows wild in
13806-666: Was referred to as Kaf Kof . The term resurfaced in Iranian tradition later on in a variant form when Ferdowsi , in his Shahnameh , referred to the Caucasus mountains as Kōh-i Kāf . "Most of the modern names of the Caucasus originate from the Greek Kaukasos (Lat., Caucasus ) and the Middle Persian Kaf Kof ". "The earliest etymon" of the name Caucasus comes from Kaz-kaz , the Hittite designation of
13924-456: Was relatively uncommon for the first thousand years of the Neolithic (when barley predominated), but became a staple after around 8500 BC. Early wheat cultivation did not demand much labour. Initially, farmers took advantage of wheat's ability to establish itself in annual grasslands by enclosing fields against grazing animals and re-sowing stands after they had been harvested, without
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