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67-488: Foothills primarily border mountains, especially those which are reached through low ridges that increase in size closer and closer to the mountain, but can also border uplands and higher hills. Areas where foothills exist, or areas commonly referred to as the foothills, include the: Another word for a foothill region is "piedmont", derived from "foot of the mount" in Romance languages . The Piedmont region of Italy lies in

134-498: A dialect continuum , where the speech variety of a location differs only slightly from that of a neighboring location, but over a longer distance these differences can accumulate to the point where two remote locations speak what may be unambiguously characterized as separate languages. This makes drawing language boundaries difficult, and as such there is no unambiguous way to divide the Romance varieties into individual languages. Even

201-417: A ' grade ', which are fruitless to precisely delineate, especially when including extinct species. Radiation results in the generation of new subclades by bifurcation, but in practice sexual hybridization may blur very closely related groupings. As a hypothesis, a clade can be rejected only if some groupings were explicitly excluded. It may then be found that the excluded group did actually descend from

268-471: A cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms worms or fishes were used within a strict cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically , outside of cladistics, e.g. as

335-422: A coarse impression of the complexity. A more detailed account will give details about fractions of introgressions between groupings, and even geographic variations thereof. This has been used as an argument for the use of paraphyletic groupings, but typically other reasons are quoted. Horizontal gene transfer is the mobility of genetic info between different organisms that can have immediate or delayed effects for

402-479: A difficulty for taxonomy , where the rank and (genus-)naming of established groupings may turn out to be inconsistent. Cladistics is now the most commonly used method to classify organisms. The original methods used in cladistic analysis and the school of taxonomy derived from the work of the German entomologist Willi Hennig , who referred to it as phylogenetic systematics (also the title of his 1966 book); but

469-489: A large number and variety of different kinds of characters are viewed as more robust than those based on more limited evidence. Mono-, para- and polyphyletic taxa can be understood based on the shape of the tree (as done above), as well as based on their character states. These are compared in the table below. Cladistics, either generally or in specific applications, has been criticized from its beginnings. Decisions as to whether particular character states are homologous ,

536-473: A lot of possible trees. Assigning names to each possible clade may not be prudent. Furthermore, established names are discarded in cladistics, or alternatively carry connotations which may no longer hold, such as when additional groups are found to have emerged in them. Naming changes are the direct result of changes in the recognition of mutual relationships, which often is still in flux, especially for extinct species. Hanging on to older naming and/or connotations

603-457: A period, many branches may have radiated, and it may take hundreds of millions of years for them to have whittled down to just two. Only then one can theoretically assign proper last common ancestors of groupings which do not inadvertently include earlier branches. The process of true cladistic bifurcation can thus take a much more extended time than one is usually aware of. In practice, for recent radiations, cladistically guided findings only give

670-444: A phylogenetic tree are used to justify decisions about character states, which are then used as evidence for the shape of the tree. Phylogenetics uses various forms of parsimony to decide such questions; the conclusions reached often depend on the dataset and the methods. Such is the nature of empirical science, and for this reason, most cladists refer to their cladograms as hypotheses of relationship. Cladograms that are supported by

737-447: A potential piece of evidence for grouping. Synapomorphies (shared, derived character states) are viewed as evidence of grouping, while symplesiomorphies (shared ancestral character states) are not. The outcome of a cladistic analysis is a cladogram – a tree -shaped diagram ( dendrogram ) that is interpreted to represent the best hypothesis of phylogenetic relationships. Although traditionally such cladograms were generated largely on

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804-598: A potential source of separatist movements; therefore, they have generally fought to eliminate it, by extensively promoting the use of the official language, restricting the use of the other languages in the media, recognizing them as mere "dialects", or even persecuting them. As a result, all of these languages are considered endangered to varying degrees according to the UNESCO Red Book of Endangered Languages , ranging from "vulnerable" (e.g. Sicilian and Venetian ) to "severely endangered" ( Franco-Provençal , most of

871-553: A powerful way to test hypotheses about cross-cultural relationships among folktales. Literature : Cladistic methods have been used in the classification of the surviving manuscripts of the Canterbury Tales , and the manuscripts of the Sanskrit Charaka Samhita . Historical linguistics : Cladistic methods have been used to reconstruct the phylogeny of languages using linguistic features. This

938-400: A precondition of their being synapomorphies, have been challenged as involving circular reasoning and subjective judgements. Of course, the potential unreliability of evidence is a problem for any systematic method, or for that matter, for any empirical scientific endeavor at all. Transformed cladistics arose in the late 1970s in an attempt to resolve some of these problems by removing

1005-406: A priori assumptions about phylogeny from cladistic analysis, but it has remained unpopular. The cladistic method does not identify fossil species as actual ancestors of a clade. Instead, fossil taxa are identified as belonging to separate extinct branches. While a fossil species could be the actual ancestor of a clade, there is no way to know that. Therefore, a more conservative hypothesis is that

1072-846: A single country. Portuguese is the official language of six African countries ( Angola , Cape Verde , Guinea-Bissau , Mozambique , Equatorial Guinea , and São Tomé and Príncipe ), and is spoken as a native language by perhaps 16 million residents of that continent. In Asia, Portuguese is co-official with other languages in East Timor and Macau , while most Portuguese-speakers in Asia—some 400,000 —are in Japan due to return immigration of Japanese Brazilians . In North America 1,000,000 people speak Portuguese as their home language, mainly immigrants from Brazil, Portugal, and other Portuguese-speaking countries and their descendants. In Oceania, Portuguese

1139-742: Is Spanish , followed by Portuguese , French , Italian and Romanian , which together cover a vast territory in Europe and beyond, and work as official and national languages in dozens of countries. In Europe, at least one Romance language is official in France , Portugal , Spain , Italy , Switzerland , Belgium , Romania , Moldova , Transnistria , Monaco , Andorra , San Marino and Vatican City . In these countries, French, Portuguese, Italian, Spanish, Romanian, Romansh and Catalan have constitutional official status. French, Italian, Portuguese, Spanish, and Romanian are also official languages of

1206-565: Is a synapomorphy of the earliest taxa to be included within Tetrapoda: did all the earliest members of the Tetrapoda inherit four limbs from a common ancestor, whereas all other vertebrates did not, or at least not homologously? By contrast, for a group within the tetrapods, such as birds, having four limbs is a plesiomorphy. Using these two terms allows a greater precision in the discussion of homology, in particular allowing clear expression of

1273-408: Is an approach to biological classification in which organisms are categorized in groups (" clades ") based on hypotheses of most recent common ancestry . The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies ) that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on

1340-764: Is an official language in Spain and in nine countries of South America , home to about half that continent's population; in six countries of Central America (all except Belize ); and in Mexico . In the Caribbean , it is official in Cuba , the Dominican Republic , and Puerto Rico . In all these countries, Latin American Spanish is the vernacular language of the majority of the population, giving Spanish

1407-440: Is correct, then the last common ancestor of turtles and birds, at the branch near the ▼ lived earlier than the last common ancestor of lizards and birds, near the ♦ . Most molecular evidence , however, produces cladograms more like this: lizards turtles crocodilians birds If this is accurate, then the last common ancestor of turtles and birds lived later than the last common ancestor of lizards and birds. Since

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1474-578: Is counter-productive, as they typically do not reflect actual mutual relationships precisely at all. E.g. Archaea, Asgard archaea, protists, slime molds, worms, invertebrata, fishes, reptilia, monkeys, Ardipithecus , Australopithecus , Homo erectus all contain Homo sapiens cladistically, in their sensu lato meaning. For originally extinct stem groups, sensu lato generally means generously keeping previously included groups, which then may come to include even living species. A pruned sensu stricto meaning

1541-422: Is limited, and the literature is often hard to interpret or generalize. Many of its speakers were soldiers, slaves, displaced peoples, and forced resettlers, and more likely to be natives of conquered lands than natives of Rome. In Western Europe, Latin gradually replaced Celtic and other Italic languages , which were related to it by a shared Indo-European origin. Commonalities in syntax and vocabulary facilitated

1608-528: Is no evidence that they recover more "true" or "correct" results from actual empirical data sets Every cladogram is based on a particular dataset analyzed with a particular method. Datasets are tables consisting of molecular , morphological, ethological and/or other characters and a list of operational taxonomic units (OTUs), which may be genes, individuals, populations, species, or larger taxa that are presumed to be monophyletic and therefore to form, all together, one large clade; phylogenetic analysis infers

1675-500: Is often adopted instead, but the group would need to be restricted to a single branch on the stem. Other branches then get their own name and level. This is commensurate to the fact that more senior stem branches are in fact closer related to the resulting group than the more basal stem branches; that those stem branches only may have lived for a short time does not affect that assessment in cladistics. The comparisons used to acquire data on which cladograms can be based are not limited to

1742-504: Is similar to the traditional comparative method of historical linguistics, but is more explicit in its use of parsimony and allows much faster analysis of large datasets ( computational phylogenetics ). Textual criticism or stemmatics : Cladistic methods have been used to reconstruct the phylogeny of manuscripts of the same work (and reconstruct the lost original) using distinctive copying errors as apomorphies. This differs from traditional historical-comparative linguistics in enabling

1809-1088: Is spoken by many residents of Alghero , on the island of Sardinia , and it is co-official in that city. Galician , with more than three million speakers, is official together with Spanish in Galicia , and has legal recognition in neighbouring territories in Castilla y León . A few other languages have official recognition on a regional or otherwise limited level; for instance, Asturian and Aragonese in Spain; Mirandese in Portugal; Friulian , Sardinian and Franco-Provençal in Italy; and Romansh in Switzerland. The remaining Romance languages survive mostly as spoken languages for informal contact. National governments have historically viewed linguistic diversity as an economic, administrative or military liability, as well as

1876-496: Is the second most spoken Romance language, after French, due mainly to the number of speakers in East Timor . Its closest relative, Galician, has official status in the autonomous community of Galicia in Spain , together with Spanish. Outside Europe, French is spoken natively most in the Canadian province of Quebec , and in parts of New Brunswick and Ontario . Canada is officially bilingual , with French and English being

1943-628: The European Union . Spanish, Portuguese, French, Italian, Romanian, and Catalan were the official languages of the defunct Latin Union ; and French and Spanish are two of the six official languages of the United Nations . Outside Europe, French , Portuguese and Spanish are spoken and enjoy official status in various countries that emerged from the respective colonial empires . With almost 500 million speakers worldwide, Spanish

2010-568: The Latin or Neo-Latin languages , are the languages that are directly descended from Vulgar Latin . They are the only extant subgroup of the Italic branch of the Indo-European language family . The five most widely spoken Romance languages by number of native speakers are: The Romance languages spread throughout the world owing to the period of European colonialism beginning in

2077-593: The Occitan varieties). Since the late twentieth and early twenty-first centuries, increased sensitivity to the rights of minorities has allowed some of these languages to start recovering their prestige and lost rights. Yet it is unclear whether these political changes will be enough to reverse the decline of minority Romance languages. Between 350 BC and 150 AD, the expansion of the Roman Empire , together with its administrative and educational policies, made Latin

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2144-659: The Republic of Moldova , where it is the dominant language and spoken by a majority of the population, but neighboring areas in Serbia ( Vojvodina and the Bor District ), Bulgaria, Hungary, and Ukraine ( Bukovina , Budjak ) and in some villages between the Dniester and Bug rivers. As with Italian, Romanian is spoken outside of its ethnic range by immigrant communities. In Europe, Romanian-speakers form about two percent of

2211-417: The declension system of Latin and, as a result, have SVO sentence structure and make extensive use of prepositions . By most measures, Sardinian and Italian are the least divergent languages from Latin, while French has changed the most. However, all Romance languages are closer to each other than to classical Latin . Documentary evidence about Vulgar Latin for the purposes of comprehensive research

2278-482: The 15th century; there are more than 900 million native speakers of Romance languages found worldwide, mainly in the Americas , Europe , and parts of Africa . Portuguese, French and Spanish also have many non-native speakers and are in widespread use as lingua francas . There are also numerous regional Romance languages and dialects. All of the five most widely spoken Romance languages are also official languages of

2345-523: The 1990s, the development of effective polymerase chain reaction techniques allowed the application of cladistic methods to biochemical and molecular genetic traits of organisms, vastly expanding the amount of data available for phylogenetics. At the same time, cladistics rapidly became popular in evolutionary biology, because computers made it possible to process large quantities of data about organisms and their characteristics. The cladistic method interprets each shared character state transformation as

2412-694: The European Union (with France, Italy, Portugal, Romania and Spain being part of it). The term Romance derives from the Vulgar Latin adverb romanice , "in Roman ", derived from romanicus : for instance, in the expression romanice loqui , "to speak in Roman" (that is, the Latin vernacular ), contrasted with latine loqui , "to speak in Latin" (Medieval Latin, the conservative version of

2479-715: The Romanian and Italian languages, as well as their common Latin origin. The total of 880 million native speakers of Romance languages (ca. 2020) are divided as follows: Catalan is the official language of Andorra . In Spain, it is co-official with Spanish in Catalonia , the Valencian Community (under the name Valencian ), and the Balearic Islands , and it is recognized, but not official, in an area of Aragon known as La Franja . In addition, it

2546-546: The adoption of Latin. To some scholars, this suggests the form of Vulgar Latin that evolved into the Romance languages was around during the time of the Roman Empire (from the end of the first century BC), and was spoken alongside the written Classical Latin which was reserved for official and formal occasions. Other scholars argue that the distinctions are more rightly viewed as indicative of sociolinguistic and register differences normally found within any language. With

2613-429: The basis of morphological characters and originally calculated by hand, genetic sequencing data and computational phylogenetics are now commonly used in phylogenetic analyses, and the parsimony criterion has been abandoned by many phylogeneticists in favor of more "sophisticated" but less parsimonious evolutionary models of character state transformation. Cladists contend that these models are unjustified because there

2680-531: The branching pattern within that clade. Different datasets and different methods, not to mention violations of the mentioned assumptions, often result in different cladograms. Only scientific investigation can show which is more likely to be correct. Until recently, for example, cladograms like the following have generally been accepted as accurate representations of the ancestral relations among turtles, lizards, crocodilians, and birds: turtles lizards crocodilians birds If this phylogenetic hypothesis

2747-525: The clade, but in principle each level stands on its own, to be assigned a unique name. For a fully bifurcated tree, adding a group to a tree also adds an additional (named) clade, and a new level on that branch. Specifically, also extinct groups are always put on a side-branch, not distinguishing whether an actual ancestor of other groupings was found. The techniques and nomenclature of cladistics have been applied to disciplines other than biology. (See phylogenetic nomenclature .) Cladistics findings are posing

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2814-443: The cladograms show two mutually exclusive hypotheses to describe the evolutionary history, at most one of them is correct. The cladogram to the right represents the current universally accepted hypothesis that all primates , including strepsirrhines like the lemurs and lorises , had a common ancestor all of whose descendants are or were primates, and so form a clade; the name Primates is therefore recognized for this clade. Within

2881-462: The criterion of mutual intelligibility can become ambiguous when it comes to determining whether two language varieties belong to the same language or not. The following is a list of groupings of Romance languages, with some languages chosen to exemplify each grouping. Not all languages are listed, and the groupings should not be interpreted as well-separated genetic clades in a tree model . The Romance language most widely spoken natively today

2948-402: The development of cultures or artifacts using groups of cultural traits or artifact features. Comparative mythology and folktale use cladistic methods to reconstruct the protoversion of many myths. Mythological phylogenies constructed with mythemes clearly support low horizontal transmissions (borrowings), historical (sometimes Palaeolithic) diffusions and punctuated evolution. They also are

3015-468: The dominant native language in continental Western Europe. Latin also exerted a strong influence in southeastern Britain , the Roman province of Africa , western Germany , Pannonia and the whole Balkans . During the Empire's decline, and after its fragmentation and the collapse of its Western half in the fifth and sixth centuries, the spoken varieties of Latin became more isolated from each other, with

3082-755: The eighth and ninth centuries Latin gave way to Romance. During the political decline of the Western Roman Empire in the fifth century, there were large-scale migrations into the empire, and the Latin-speaking world was fragmented into several independent states. Central Europe and the Balkans were occupied by Germanic and Slavic tribes, as well as by Huns . Cladistics#In disciplines other than biology Cladistics ( / k l ə ˈ d ɪ s t ɪ k s / klə- DIST -iks ; from Ancient Greek κλάδος kládos 'branch')

3149-561: The end of the colonial domination. As a result, Italian outside Italy and Switzerland is now spoken only as a minority language by immigrant communities in North and South America and Australia . In some former Italian colonies in Africa—namely Libya , Eritrea and Somalia —it is spoken by a few educated people in commerce and government. Romania did not establish a colonial empire. The native range of Romanian includes not only

3216-410: The exact historic relationships between the groups. The following terms, coined by Hennig, are used to identify shared or distinct character states among groups: The terms plesiomorphy and apomorphy are relative; their application depends on the position of a group within a tree. For example, when trying to decide whether the tetrapods form a clade, an important question is whether having four limbs

3283-512: The field of biology. Any group of individuals or classes that are hypothesized to have a common ancestor, and to which a set of common characteristics may or may not apply, can be compared pairwise. Cladograms can be used to depict the hypothetical descent relationships within groups of items in many different academic realms. The only requirement is that the items have characteristics that can be identified and measured. Anthropology and archaeology : Cladistic methods have been used to reconstruct

3350-769: The foothills of the Alps , and several other foothills in other parts of the world are called "piedmont" , and include: Ecosystems of piedmonts (foothills) are often known as submontane zones, relating to the higher montane ecosystems . Romance languages Pontic Steppe Caucasus East Asia Eastern Europe Northern Europe Pontic Steppe Northern/Eastern Steppe Europe South Asia Steppe Europe Caucasus India Indo-Aryans Iranians East Asia Europe East Asia Europe Indo-Aryan Iranian Indo-Aryan Iranian Others European The Romance languages , also known as

3417-481: The fossil taxon is related to other fossil and extant taxa, as implied by the pattern of shared apomorphic features. An otherwise extinct group with any extant descendants, is not considered (literally) extinct, and for instance does not have a date of extinction. Anything having to do with biology and sex is complicated and messy, and cladistics is no exception. Many species reproduce sexually, and are capable of interbreeding for millions of years. Worse, during such

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3484-407: The hierarchical relationships among different homologous features. It can be difficult to decide whether a character state is in fact the same and thus can be classified as a synapomorphy, which may identify a monophyletic group, or whether it only appears to be the same and is thus a homoplasy, which cannot identify such a group. There is a danger of circular reasoning: assumptions about the shape of

3551-597: The language used in writing and formal contexts or as a lingua franca), and with barbarice loqui , "to speak in Barbarian " (the non-Latin languages of the peoples living outside the Roman Empire). From this adverb the noun romance originated, which applied initially to anything written romanice , or "in the Roman vernacular". Most of the Romance-speaking area in Europe has traditionally been

3618-460: The last common ancestor of the group, and thus emerged within the group. ("Evolved from" is misleading, because in cladistics all descendants stay in the ancestral group). To keep only valid clades, upon finding that the group is paraphyletic this way, either such excluded groups should be granted to the clade, or the group should be abolished. Branches down to the divergence to the next significant (e.g. extant) sister are considered stem-groupings of

3685-487: The latter contains Tarsiiformes and Anthropoidea. Lemurs and tarsiers may have looked closely related to humans, in the sense of being close on the evolutionary tree to humans. However, from the perspective of a tarsier, humans and lemurs would have looked close, in the exact same sense. Cladistics forces a neutral perspective, treating all branches (extant or extinct) in the same manner. It also forces one to try to make statements, and honestly take into account findings, about

3752-547: The most native speakers of any Romance language. In Africa it is one of the official languages of Equatorial Guinea . Spanish was one of the official languages in the Philippines in Southeast Asia until 1973. In the 1987 constitution, Spanish was removed as an official language (replaced by English), and was listed as an optional/voluntary language along with Arabic. It is currently spoken by a minority and taught in

3819-624: The official languages and government services in French theoretically mandated to be provided nationwide. In parts of the Caribbean, such as Haiti , French has official status, but most people speak creoles such as Haitian Creole as their native language. French also has official status in much of Africa, with relatively few native speakers but larger numbers of second language speakers. Although Italy also had some colonial possessions before World War II , its language did not remain official after

3886-584: The other continents to such an extent that about two-thirds of all Romance language speakers today live outside Europe. Despite other influences (e.g. substratum from pre-Roman languages, especially Continental Celtic languages ; and superstratum from later Germanic or Slavic invasions), the phonology , morphology , and lexicon of all Romance languages consist mainly of evolved forms of Vulgar Latin. However, some notable differences exist between today's Romance languages and their Roman ancestor. With only one or two exceptions, Romance languages have lost

3953-597: The population in Italy , Spain , and Portugal . Romanian is also spoken in Israel by Romanian Jews, where it is the native language of five percent of the population, and is spoken by many more as a secondary language. The Aromanian language is spoken today by Aromanians in Bulgaria, North Macedonia, Albania, Kosovo, and Greece. Flavio Biondo was the first scholar to have observed (in 1435) linguistic affinities between

4020-444: The primates, all anthropoids (monkeys, apes, and humans) are hypothesized to have had a common ancestor all of whose descendants are or were anthropoids, so they form the clade called Anthropoidea. The "prosimians", on the other hand, form a paraphyletic taxon. The name Prosimii is not used in phylogenetic nomenclature , which names only clades; the "prosimians" are instead divided between the clades Strepsirhini and Haplorhini , where

4087-432: The reciprocal host. There are several processes in nature which can cause horizontal gene transfer . This does typically not directly interfere with ancestry of the organism, but can complicate the determination of that ancestry. On another level, one can map the horizontal gene transfer processes, by determining the phylogeny of the individual genes using cladistics. If there is unclarity in mutual relationships, there are

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4154-554: The rise of the Roman Empire, spoken Latin spread first throughout Italy and then through southern , western , central , and southeastern Europe , and northern Africa along parts of western Asia . Latin reached a stage when innovations became generalised around the sixth and seventh centuries. After that time and within two hundred years, it became a dead language since "the Romanized people of Europe could no longer understand texts that were read aloud or recited to them." By

4221-434: The school curriculum. Portuguese, in its original homeland, Portugal , is spoken by almost the entire population of 10 million. As the official language of Brazil , it is spoken by more than 200 million people, as well as in neighboring parts of eastern Paraguay and northern Uruguay . This accounts for slightly more than half the population of South America, making Portuguese the most spoken official Romance language in

4288-492: The terms "cladistics" and "clade" were popularized by other researchers. Cladistics in the original sense refers to a particular set of methods used in phylogenetic analysis, although it is now sometimes used to refer to the whole field. What is now called the cladistic method appeared as early as 1901 with a work by Peter Chalmers Mitchell for birds and subsequently by Robert John Tillyard (for insects) in 1921, and W. Zimmermann (for plants) in 1943. The term " clade "

4355-469: The western dialects coming under heavy Germanic influence (the Goths and Franks in particular) and the eastern dialects coming under Slavic influence. The dialects diverged from Latin at an accelerated rate and eventually evolved into a continuum of recognizably different typologies. The colonial empires established by Portugal , Spain , and France from the fifteenth century onward spread their languages to

4422-419: Was championed at this time by the numerical taxonomists Peter Sneath and Robert Sokal , and evolutionary taxonomy by Ernst Mayr . Originally conceived, if only in essence, by Willi Hennig in a book published in 1950, cladistics did not flourish until its translation into English in 1966 (Lewin 1997). Today, cladistics is the most popular method for inferring phylogenetic trees from morphological data. In

4489-554: Was introduced in 1958 by Julian Huxley after having been coined by Lucien Cuénot in 1940, "cladogenesis" in 1958, "cladistic" by Arthur Cain and Harrison in 1960, "cladist" (for an adherent of Hennig's school) by Ernst Mayr in 1965, and "cladistics" in 1966. Hennig referred to his own approach as "phylogenetic systematics". From the time of his original formulation until the end of the 1970s, cladistics competed as an analytical and philosophical approach to systematics with phenetics and so-called evolutionary taxonomy . Phenetics

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