184-607: Hops are the flowers (also called seed cones or strobiles ) of the hop plant Humulus lupulus , a member of the Cannabaceae family of flowering plants. They are used primarily as a bittering, flavouring, and stability agent in beer , to which, in addition to bitterness, they impart floral, fruity, or citrus flavours and aromas. Hops are also used for various purposes in other beverages and herbal medicine . The hops plants have separate female and male plants, and only female plants are used for commercial production. The hop plant
368-696: A Protestant convert, but he was distrusted by Scottish Calvinists who noticed the physical displays of affection between him and the king and alleged that Lennox "went about to draw the King to carnal lust". In August 1582, in what became known as the Ruthven Raid , the Protestant earls William Ruthven, Earl of Gowrie and Archibald Douglas, Earl of Angus lured James into Ruthven Castle , imprisoned him, and forced Lennox to leave Scotland. On 19 September 1582, during James's imprisonment, John Craig , whom
552-526: A diploid (two copies of each chromosome ) cell. Whereas in fertilization only plasmogamy, or the fusion of the whole sex cells, results, in Angiosperms (flowering plants) a process known as double fertilization, which involves both karyogamy and plasmogamy, occurs. In double fertilization the second sperm cell subsequently also enters the synergid and fuses with the two polar nuclei of the central cell. Since all three nuclei are haploid , they result in
736-411: A filament , or stalk. The anther contains microsporocytes which become pollen , the male gametophyte , after undergoing meiosis . Although they exhibit the widest variation among floral organs, the androecium is usually confined just to one whorl and to two whorls only in rare cases. Stamens range in number, size, shape, orientation, and in their point of connection to the flower. In general, there
920-684: A secret correspondence with James to prepare in advance for a smooth succession. With the queen clearly dying, Cecil sent James a draft proclamation of his accession to the English throne in March 1603. Elizabeth died in the early hours of 24 March, and James was proclaimed king in London later the same day. On 5 April, James left Edinburgh for London, promising to return every three years (a promise that he did not keep), and progressed slowly southwards. Local lords received him with lavish hospitality along
1104-528: A Catholic ceremony held at Stirling Castle . His godparents were Charles IX of France (represented by John, Count of Brienne ), Elizabeth I of England (represented by the Francis Russell, Earl of Bedford ), and Emmanuel Philibert, Duke of Savoy (represented by ambassador Philibert du Croc ). Mary refused to let the Archbishop of St Andrews , whom she referred to as "a pocky priest", spit in
1288-411: A broad base, stomata and chlorophyll and may have stipules . Sepals are often waxy and tough, and grow quickly to protect the flower as it develops. They may be deciduous , but will more commonly grow on to assist in fruit dispersal. If the calyx is fused it is called gamosepalous. The petals , or corolla, are almost or completely fiberless leaf-like structures that form the innermost whorl of
1472-691: A classic example of the Bohemian Pilsener style, which showcases noble hops. As with grapes, the location where hops are grown affects the hops' characteristics. Much as Dortmunder beer may within the EU be labelled "Dortmunder" only if it has been brewed in Dortmund , noble hops may officially be considered "noble" only if they were grown in the areas for which the hop varieties ( races ) were named. Noble hops are characterized through analysis as having an aroma quality resulting from numerous factors in
1656-456: A closer political union, the kingdoms of Scotland and England remained sovereign states , with their own parliaments, judiciaries, and laws, ruled by James in personal union . James was the son of Mary, Queen of Scots , and a great-great-grandson of Henry VII , King of England and Lord of Ireland, and thus a potential successor to all three thrones. He acceded to the Scottish throne at
1840-522: A description of the poetic tradition in his mother tongue of Scots , applying Renaissance principles. He also made statutory provision to reform and promote the teaching of music, seeing the two in connection. One act of his reign urges the Scottish burghs to reform and support the teaching of music in Sang Sculis . In furtherance of these aims, James was both patron and head of a loose circle of Scottish Jacobean court poets and musicians known as
2024-404: A different flower of the same plant, but others have mechanisms to prevent self-pollination and rely on cross-pollination , when pollen is transferred from the anther of one flower to the stigma of another flower on a different individual of the same species. Self-pollination happens in flowers where the stamen and carpel mature at the same time, and are positioned so that the pollen can land on
SECTION 10
#17327839431202208-418: A different plant of the same species. Because the genetic make-up of the sperm contained within the pollen from the other plant is different, their combination will result in a new, genetically distinct, plant, through the process of sexual reproduction . Since each new plant is genetically distinct, the different plants show variation in their physiological and structural adaptations and so the population as
2392-559: A flavor that improves as they age during periods of poor storage. In addition to beer, hops are used in herbal teas and in soft drinks. These soft drinks include Julmust (a carbonated beverage similar to soda that is popular in Sweden during December), Malta (a Latin American soft drink) and kvass . Hops can be eaten; the young shoots of the vine are edible and can be cooked like asparagus . Hops may be used in herbal medicine in
2576-462: A great deal of money, his account of the circumstances was not universally believed. In 1586, James signed the Treaty of Berwick with England. That and his mother's execution in 1587, which he denounced as a "preposterous and strange procedure", helped clear the way for his succession south of the border. Queen Elizabeth was unmarried and childless, and James was her most likely successor . Securing
2760-435: A head when James finally called a Parliament in 1621 to fund a military expedition in support of his son-in-law. The Commons on the one hand granted subsidies inadequate to finance serious military operations in aid of Frederick, and on the other—remembering the profits gained under Elizabeth by naval attacks on Spanish gold shipments—called for a war directly against Spain. In November 1621, roused by Edward Coke , they framed
2944-545: A hop garden in the will of Charlemagne 's father, Pepin the Short . Hops are also used in brewing for their antibacterial effect over less desirable microorganisms and for purported benefits including balancing the sweetness of the malt with bitterness and a variety of flavours and aromas. It is believed that traditional herb combinations for beers were abandoned after it was noticed that beers made with hops were less prone to spoilage. The first documented hop cultivation
3128-484: A king may impose new laws by royal prerogative but must also pay heed to tradition and to God, who would "stirre up such scourges as pleaseth him, for punishment of wicked kings". Basilikon Doron was written as a book of instruction for the four-year-old Prince Henry and provides a more practical guide to kingship. The work is considered to be well written and perhaps the best example of James's prose. James's advice concerning parliaments, which he understood as merely
3312-432: A large endosperm nucleus which is triploid . Following the formation of zygote it begins to grow through nuclear and cellular divisions, called mitosis , eventually becoming a small group of cells. One section of it becomes the embryo , while the other becomes the suspensor; a structure which forces the embryo into the endosperm and is later undetectable. Two small primordia also form at this time, that later become
3496-638: A large part of George Orwell 's A Clergyman's Daughter contain a vivid description of London families participating in this annual hops harvest. In England, many of those picking hops in Kent were from eastern areas of London. This provided a break from urban conditions that was spent in the countryside. People also came from Birmingham and other Midlands cities to pick hops in the Malvern area of Worcestershire. Some photographs have been preserved. The often-appalling living conditions endured by hop pickers during
3680-433: A lump sum of £600,000 to pay off the king's debts plus an annual grant of £200,000. The ensuing prickly negotiations became so protracted that James eventually lost patience and dismissed Parliament on 31 December 1610. "Your greatest error", he told Salisbury, "hath been that ye ever expected to draw honey out of gall". The same pattern was repeated with the so-called " Addled Parliament " of 1614, which James dissolved after
3864-1340: A mate). In pursuing this attractant from many flowers of the same species, the pollinator transfers pollen to the stigmas—arranged with equally pointed precision—of all of the flowers it visits. Many flowers rely on simple proximity between flower parts to ensure pollination, while others have elaborate designs to ensure pollination and prevent self-pollination . Flowers use animals including: insects ( entomophily ), birds ( ornithophily ), bats ( chiropterophily ), lizards, and even snails and slugs ( malacophilae ). Plants cannot move from one location to another, thus many flowers have evolved to attract animals to transfer pollen between individuals in dispersed populations. Most commonly, flowers are insect-pollinated, known as entomophilous ; literally "insect-loving" in Greek. To attract these insects flowers commonly have glands called nectaries on various parts that attract animals looking for nutritious nectar . Some flowers have glands called elaiophores , which produce oils rather than nectar. Birds and bees have color vision , enabling them to seek out colorful flowers. Some flowers have patterns, called nectar guides , that show pollinators where to look for nectar; they may be visible only under ultraviolet light, which
SECTION 20
#17327839431204048-514: A meeting with Tycho Brahe , James and Anne returned to Scotland on 1 May 1590. By all accounts, James was at first infatuated with Anne and, in the early years of their marriage, seems always to have shown her patience and affection. The royal couple produced three children who survived to adulthood: Henry Frederick, Prince of Wales , who died of typhoid fever in 1612, aged 18; Elizabeth , later queen of Bohemia ; and Charles , James's successor. Anne suffered from recurrent bouts of sickness and
4232-454: A mere nine weeks when the Commons hesitated to grant him the money he required. James then ruled without parliament until 1621, employing officials such as the merchant Lionel Cranfield , who were astute at raising and saving money for the crown, and sold baronetcies and other dignities, many created for the purpose, as an alternative source of income. Another potential source of income was
4416-540: A new translation and compilation of approved books of the Bible to resolve discrepancies among different translations then being used. The King James Version , as it came to be known, was completed in 1611 and is considered a masterpiece of Jacobean prose. It is still in widespread use. In Scotland, James attempted to bring the Scottish Kirk "so neir as can be" to the English church and to reestablish episcopacy ,
4600-458: A noticeable scent. Because of this, plants typically have many thousands of tiny flowers which have comparatively large, feathery stigmas; to increase the chance of pollen being received. Whereas the pollen of entomophilous flowers is usually large, sticky, and rich in protein (to act as a "reward" for pollinators), anemophilous flower pollen is typically small-grained, very light, smooth, and of little nutritional value to insects . In order for
4784-632: A peace policy, and tried to avoid involvement in religious wars , especially the Thirty Years' War that devastated much of Central Europe. He tried but failed to prevent the rise of hawkish elements in the English Parliament who wanted war with Spain. He was succeeded by his second son, Charles I. James was the only son of Mary, Queen of Scots , and her second husband, Henry Stuart, Lord Darnley . Mary and Darnley were great-grandchildren of Henry VII of England through Margaret Tudor ,
4968-573: A period of decline and disrepute. Salisbury's passing gave James the notion of governing in person as his own chief Minister of State, with his young Scottish favourite Robert Carr carrying out many of Salisbury's former duties, but James's inability to attend closely to official business exposed the government to factionalism. The Howard party (consisting of Henry Howard, Earl of Northampton; Thomas Howard, Earl of Suffolk; Suffolk's son-in-law William Knollys, Lord Knollys ; Charles Howard, Earl of Nottingham ; and Thomas Lake ) soon took control of much of
5152-405: A petition asking not only for war with Spain but also for Prince Charles to marry a Protestant, and for enforcement of the anti-Catholic laws. James flatly told them not to interfere in matters of royal prerogative or they would risk punishment, which provoked them into issuing a statement protesting their rights, including freedom of speech. Urged on by George Villiers, Duke of Buckingham , and
5336-470: A plant can interpret important endogenous and environmental cues such as changes in levels of plant hormones and seasonable temperature and photoperiod changes. Many perennial and most biennial plants require vernalization to flower. The molecular interpretation of these signals is through the transmission of a complex signal known as florigen , which involves a variety of genes , including Constans, Flowering Locus C, and Flowering Locus T. Florigen
5520-706: A policy that met with strong opposition from presbyterians . James returned to Scotland in 1617 for the only time after his accession in England, in the hope of implementing Anglican ritual. James's bishops forced his Five Articles of Perth through a General Assembly the following year, but the rulings were widely resisted. James left the church in Scotland divided at his death, a source of future problems for his son. Throughout his life James had close relationships with male courtiers, which has caused debate among historians about their exact nature. In Scotland Anne Murray
5704-581: A principal cause of the Highlanders' shortcomings and sought to abolish it. It was against this background that James VI authorised the " Gentleman Adventurers of Fife " to civilise the "most barbarous Isle of Lewis " in 1598. James wrote that the colonists were to act "not by agreement" with the local inhabitants, but "by extirpation of thame". Their landing at Stornoway began well, but the colonists were driven out by local forces commanded by Murdoch and Neil MacLeod. The colonists tried again in 1605 with
Hops - Misplaced Pages Continue
5888-722: A process "specifically aimed at the extirpation of the Gaelic language, the destruction of its traditional culture and the suppression of its bearers." In the Northern Isles , James's cousin Patrick Stewart, Earl of Orkney , resisted the Statutes of Iona and was consequently imprisoned. His natural son Robert led an unsuccessful rebellion against James, and the Earl and his son were hanged. Their estates were forfeited, and
6072-498: A serious adverse effect on hops production, but remnants of this significant industry in the western states are still noticeable in the form of old hop kilns that survive throughout Sonoma County, California , among others. Florian Dauenhauer, of Santa Rosa in Sonoma County, became a manufacturer of hop-harvesting machines in 1940, in part because of the hop industry's importance to the county. This mechanization helped destroy
6256-657: A single parliament for England and Scotland. In his reign, the Plantation of Ulster and English colonisation of the Americas began. At 57 years and 246 days, James's reign in Scotland was the longest of any Scottish monarch . He achieved most of his aims in Scotland but faced great difficulties in England, including the Gunpowder Plot in 1605 and conflicts with the English Parliament . Under James,
6440-404: A sonar-reflecting petal above its flowers, which helps the bat find them, and one species, the cactus Espostoa frutescens , has flowers that are surrounded by an area of sound-absorbent and woolly hairs called the cephalium, which absorbs the bat's ultrasound instead. Flowers are also specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to
6624-550: A stock as to praise fools ... You see how many things you did not well ... I wish you would make use of your liberty with more modesty in time to come". As James's reign progressed, his government faced growing financial pressures, partly due to creeping inflation but also to the profligacy and financial incompetence of James's court. In February 1610, Salisbury proposed a scheme, known as the Great Contract , whereby Parliament, in return for ten royal concessions, would grant
6808-407: A third in 1944. Knuth named his two groups 'Hyphydrogamy' and the more common 'Ephydrogamy'. In hyphydrogamy pollination occurs below the surface of the water and so the pollen grains are typically negatively buoyant . For marine plants that exhibit this method, the stigmas are usually stiff, while freshwater species have small and feathery stigmas. In ephydrogamy pollination occurs on the surface of
6992-419: A way similar to valerian , as a treatment for anxiety, restlessness, and insomnia. A pillow filled with hops is a popular folk remedy for sleeplessness, and animal research has shown a sedative effect. The relaxing effect of hops may be due, in part, to the specific degradation product from alpha acids , 2-methyl-3-buten-2-ol, as demonstrated from nighttime consumption of non-alcoholic beer. 2-methyl-3-buten-2-ol
7176-435: A whole is better prepared for an adverse occurrence in the environment. Cross-pollination, therefore, increases the survival of the species and is usually preferred by flowers for this reason. The principal adaptive function of flowers is the promotion of cross-pollination or outcrossing, a process that allows the masking of deleterious mutations in the genome of progeny. The masking effect of outcrossing sexual reproduction
7360-416: Is a loss of B gene function, mutant flowers are produced with sepals in the first whorl as usual, but also in the second whorl instead of the normal petal formation. In the third whorl, the lack of the B function but the presence of the C function mimics the fourth whorl, leading to the formation of carpels also in the third whorl. The principal purpose of a flower is the reproduction of the individual and
7544-422: Is a much rarer method, occurring in only around 2% of abiotically pollinated flowers. Common examples of this include Calitriche autumnalis , Vallisneria spiralis and some sea-grasses . One characteristic which most species in this group share is a lack of an exine , or protective layer, around the pollen grain. Paul Knuth identified two types of hydrophilous pollination in 1906 and Ernst Schwarzenbach added
Hops - Misplaced Pages Continue
7728-532: Is a potent phytoestrogen . Hops are usually dried in an oast house before they are used in the brewing process. Undried or "wet" hops are sometimes (since c. 1990) used. The wort (sugar-rich liquid produced from malt ) is boiled with hops before it is cooled down and yeast is added, to start fermentation . The effect of hops on the finished beer varies by type and use, though there are two main hop types: bittering and aroma. Bittering hops have higher concentrations of alpha acids, and are responsible for
7912-514: Is a vigorous climbing herbaceous perennial , usually trained to grow up strings in a field called a hopfield, hop garden (in the South of England), or hop yard (in the West Country and United States) when grown commercially. Many different varieties of hops are grown by farmers around the world, with different types used for particular styles of beer. The first documented use of hops in beer
8096-417: Is a way to represent the structure of a flower using specific letters, numbers, and symbols, presenting substantial information about the flower in a compact form. It can represent a taxon , usually giving ranges of the numbers of different organs, or particular species. Floral formulae have been developed in the early 19th century and their use has declined since. Prenner et al. (2010) devised an extension of
8280-404: Is an example of coevolution , as the flower and pollinator have developed together over a long period to match each other's needs. This close relationship compounds the negative effects of extinction , however, since the extinction of either member in such a relationship would almost certainly mean the extinction of the other member as well. Flowers that use abiotic, or non-living, vectors use
8464-427: Is because it is directly connected to the method of seed dispersal; that being the purpose of fruit - to encourage or enable the seed's dispersal and protect the seed while doing so. Following the pollination of a flower, fertilization, and finally the development of a seed and fruit, a mechanism is typically used to disperse the fruit away from the plant. In Angiosperms (flowering plants) seeds are dispersed away from
8648-622: Is from the Middle English flour , which referred to both the ground grain and the reproductive structure in plants, before splitting off in the 17th century. It comes originally from the Latin name of the Italian goddess of flowers, Flora . The early word for flower in English was blossom , though it now refers to flowers only of fruit trees . The morphology of a flower, or its form and structure, can be considered in two parts:
8832-427: Is from the 9th century, though Hildegard of Bingen , 300 years later, is often cited as the earliest documented source. Before this period, brewers used a " gruit ", composed of a wide variety of bitter herbs and flowers, including dandelion , burdock root , marigold , horehound (the old German name for horehound, Berghopfen , means "mountain hops"), ground ivy , and heather . Early documents include mention of
9016-541: Is highly reduced or absent). The stem or stalk subtending a flower, or an inflorescence of flowers, is called a peduncle . If a peduncle supports more than one flower, the stems connecting each flower to the main axis are called pedicels . The apex of a flowering stem forms a terminal swelling which is called the torus or receptacle. In the majority of species, individual flowers have both carpels and stamens. These flowers are described by botanists as being perfect, bisexual, or hermaphrodite . In some species of plants,
9200-494: Is known as "genetic complementation". This beneficial effect of outcrossing on progeny is also recognized as hybrid vigour or heterosis. Once outcrossing is established due to the benefits of genetic complementation, subsequent switching to inbreeding becomes disadvantageous because it allows the expression of the previously masked deleterious recessive mutations, usually referred to as inbreeding depression. Charles Darwin in his 1889 book The Effects of Cross and Self-Fertilization in
9384-408: Is only one type of stamen, but there are plant species where the flowers have two types; a "normal" one and one with anthers that produce sterile pollen meant to attract pollinators. The gynoecium , or the carpels , is the female part of the flower found on the innermost whorl. Each carpel consists of a stigma , which receives pollen, a style , which acts as a stalk, and an ovary , which contains
SECTION 50
#17327839431209568-443: Is preferred because it allows for genetic variation , which contributes to the survival of the species. Many flowers depend on external factors for pollination, such as the wind, water, animals, and especially insects . Larger animals such as birds, bats, and even some pygmy possums , however, can also be employed. To accomplish this, flowers have specific designs which encourage the transfer of pollen from one plant to another of
9752-429: Is produced in the leaves in reproductively favorable conditions and acts in buds and growing tips to induce several different physiological and morphological changes. The first step of the transition is the transformation of the vegetative stem primordia into floral primordia. This occurs as biochemical changes take place to change the cellular differentiation of leaf, bud and stem tissues into tissue that will grow into
9936-517: Is reflected morphologically in the presence of more pulp , an aril , and sometimes an elaiosome (primarily for ants), which are other fleshy structures. James VI and I James VI and I (James Charles Stuart; 19 June 1566 – 27 March 1625) was King of Scotland as James VI from 24 July 1567 and King of England and Ireland as James I from the union of the Scottish and English crowns on 24 March 1603 until his death in 1625. Although he long tried to get both countries to adopt
10120-472: Is responsible for the pungent smell of fresh hops. Humulene and its oxidative reaction products may give beer its prominent hop aroma. Together, myrcene, humulene, and caryophyllene represent 80 to 90% of the total hops essential oil. Xanthohumol is the principal flavonoid in hops. The other well-studied prenylflavonoids are 8-Prenylnaringenin and isoxanthohumol . Xanthohumol is under basic research for its potential properties, while 8-prenylnaringenin
10304-459: Is structurally similar to tert-amyl alcohol which was historically used as an anesthetic. Hops tend to be unstable when exposed to light or air and lose their potency after a few months' storage. Flower A flower , also known as a bloom or blossom , is the reproductive structure found in flowering plants (plants of the division Angiospermae ). Flowers consist of a combination of vegetative organs – sepals that enclose and protect
10488-488: Is that male and female flowers develop on separate plants, although occasionally a fertile individual will develop which contains both male and female flowers. Because pollinated seeds are undesirable for brewing beer, only female plants are grown in hop fields, thus preventing pollination. Female plants are propagated vegetatively , and male plants are culled if plants are grown from seeds. Hop plants are planted in rows about 2 to 2.5 metres (7 to 8 ft) apart. Each spring,
10672-418: Is typically another part. In some families, such as the grasses , the petals are greatly reduced; in many species, the sepals are colorful and petal-like. Other flowers have modified petal-like stamens; the double flowers of peonies and roses are mostly petaloid stamens. Many flowers have symmetry. When the perianth is bisected through the central axis from any point and symmetrical halves are produced,
10856-574: Is visible to bees and some other insects. Flowers also attract pollinators by scent , though not all flower scents are appealing to humans; several flowers are pollinated by insects that are attracted to rotten flesh and have flowers that smell like dead animals. These are often called carrion flowers , including plants in the genus Rafflesia , and the titan arum . Flowers pollinated by night visitors, including bats and moths, are likely to concentrate on scent to attract pollinators and so most such flowers are white. Some plants pollinated by bats have
11040-482: Is well known that the king of England / fucks the Duke of Buckingham". Buckingham himself provides evidence that he slept in the same bed as the king, writing to James many years later that he had pondered "whether you loved me now ... better than at the time which I shall never forget at Farnham, where the bed's head could not be found between the master and his dog". Buckingham's words may be interpreted as non-sexual, in
11224-606: The Authorized King James Version ), and the 1604 revision of the Book of Common Prayer . Anthony Weldon claimed that James had been termed "the wisest fool in Christendom ", an epithet associated with his character ever since. Since the latter half of the 20th century, historians have tended to revise James's reputation and treat him as a serious and thoughtful monarch. He was strongly committed to
SECTION 60
#173278394312011408-454: The Battle of Langside , forcing her to flee to England, where she was subsequently kept in confinement by Elizabeth. On 23 January 1570, Moray was assassinated by James Hamilton of Bothwellhaugh . The next regent was James's paternal grandfather, Matthew Stewart, Earl of Lennox , who was carried fatally wounded into Stirling Castle a year later after a raid by Mary's supporters. His successor,
11592-564: The Castalian Band , which included William Fowler and Alexander Montgomerie among others, Montgomerie being a favourite of the king. James was himself a poet, and was happy to be seen as a practising member of the group. By the late 1590s, James's championing of native Scottish tradition was reduced to some extent by the increasing likelihood of his succession to the English throne. William Alexander and other courtier poets started to anglicise their written language, and followed
11776-433: The English Parliament , "compassed with one sea and of itself by nature indivisible?" In April 1604, however, the Commons refused his request to be titled "King of Great Britain" on legal grounds. In October 1604, he assumed the title "King of Great Britain" instead of "King of England" and "King of Scotland", though Francis Bacon told him that he could not use the style in "any legal proceeding, instrument or assurance" and
11960-752: The Hallertau in Germany, the Žatec ( Saaz ) in the Czech Republic , the Yakima ( Washington ) and Willamette (Oregon) valleys, and western Canyon County, Idaho (including the communities of Parma , Wilder , Greenleaf , and Notus ). The principal production centres in the UK are in Kent (which produces Kent Goldings hops), Herefordshire , and Worcestershire . Essentially all of the harvested hops are used in beer making. Although hops are grown in most of
12144-569: The Lordship of the Isles by James IV of Scotland in 1493 had led to troubled times for the western seaboard. James IV had subdued the organised military might of the Hebrides , but he and his immediate successors lacked the will or ability to provide an alternative form of governance. As a result, the 16th century became known as linn nan creach , the time of raids. Furthermore, the effects of
12328-588: The Neomexicanus hops of New Mexico . Others were named for the farmer who is recognized as first cultivating them, including Goldings or Fuggles from England, or by their growing habit like the Oregon Cluster. Around 1900, a number of institutions began to experiment with breeding specific hop varieties. The breeding program at Wye College in Wye, Kent , was started in 1904 and rose to prominence through
12512-638: The Orkney and Shetland islands were annexed to the Crown. In 1597–98, James wrote The True Law of Free Monarchies and Basilikon Doron ( Royal Gift ), in which he argues a theological basis for monarchy. In the True Law , he sets out the divine right of kings , explaining that kings are higher beings than other men for Biblical reasons, though "the highest bench is the sliddriest to sit upon". The document proposes an absolutist theory of monarchy, by which
12696-628: The Popish Recusants Act , which could require any subject to take an Oath of Allegiance denying the pope's authority over the king. James was conciliatory towards Catholics who took the Oath of Allegiance, and tolerated crypto-Catholicism even at court. Henry Howard, for example, was a crypto-Catholic, received back into the Catholic Church in his final months. On ascending the English throne, James suspected that he might need
12880-487: The Witchcraft Act 1563 . Several people were convicted of using witchcraft to send storms against James's ship , most notably Agnes Sampson . James became concerned with the threat posed by witches and wrote Daemonologie in 1597, a tract inspired by his personal involvement that opposed the practice of witchcraft and that provided background material for Shakespeare's Macbeth . James personally supervised
13064-414: The calyx , corolla , androecium , and gynoecium . Together the calyx and corolla make up the non-reproductive part of the flower called the perianth , and in some cases may not be differentiated. If this is the case, then they are described as tepals . The sepals , collectively called the calyx, are modified leaves that occur on the outermost whorl of the flower. They are leaf-like, in that they have
13248-522: The coronation was preached by John Knox . In accordance with the religious beliefs of most of the Scottish ruling class, James was brought up as a member of the Protestant Church of Scotland , the Kirk. The Privy Council selected George Buchanan , Peter Young , Adam Erskine (lay abbot of Cambuskenneth ), and David Erskine (lay abbot of Dryburgh ) as James's preceptors or tutors. As
13432-499: The cotyledon , which is used as an energy store. Plants which grow out one of these primordia are called monocotyledons , while those that grow out two are dicotyledons . The next stage is called the Torpedo stage and involves the growth of several key structures, including: the radicle (embryotic root), the epicotyl (embryotic stem), and the hypocotyl , (the root/shoot junction). In the final step vascular tissue develops around
13616-439: The gizzard of animals or even to germinate better after passing through them. They can be eaten by birds ( ornithochory) , bats ( chiropterochory) , rodents , primates, ants ( myrmecochory ), non-bird sauropsids ( saurochory) , mammals in general (mammaliochory) , and even fish . Typically their fruit are fleshy, have a high nutritional value, and may have chemical attractants as an additional "reward" for dispersers. This
13800-518: The wind or, much less commonly, water , to move pollen from one flower to the next. In wind-dispersed ( anemophilous ) species, the tiny pollen grains are carried, sometimes many thousands of kilometers, by the wind to other flowers. Common examples include the grasses , birch trees , along with many other species in the order Fagales , ragweeds , and many sedges . They do not need to attract pollinators and therefore tend not to grow large, showy, or colorful flowers, and do not have nectaries, nor
13984-442: The "Golden Age" of Elizabethan literature and drama continued, with writers such as William Shakespeare , John Donne , Ben Jonson , and Francis Bacon contributing to a flourishing literary culture. James was a prolific writer, authoring works such as Daemonologie (1597), The True Law of Free Monarchies (1598), and Basilikon Doron (1599). He sponsored the translation of the Bible into English (later named after him,
14168-564: The Czech Republic may have noticeable noble hop aroma. Certain ales (particularly the highly hopped style known as India Pale Ale , or IPA) can have high levels of hop bitterness. Brewers may use software tools to control the bittering levels in the boil and adjust recipes to account for a change in the hop bill or seasonal variations in the crop that may lead to the need to compensate for a difference in alpha acid contribution. Data may be shared with other brewers via BeerXML allowing
14352-645: The Duke of Buckingham" whom the king would, recalled Sir Edward Peyton , "tumble and kiss as a mistress". Restoration of Apethorpe Palace , Northamptonshire, undertaken in 2004–08 revealed a previously unknown passage linking the bedchambers of James and Villiers. Some biographers of James argue that the relationships were not sexual. James's Basilikon Doron lists sodomy among crimes "ye are bound in conscience never to forgive", and James's wife Anne gave birth to seven live children, as well as suffering two stillbirths and at least three other miscarriages. Contemporary Huguenot poet Théophile de Viau observed that "it
14536-541: The Earl of Mar, "took a vehement sickness" and died on 28 October 1572 at Stirling. Mar's illness, wrote James Melville , followed a banquet at Dalkeith Palace given by James Douglas, Earl of Morton . Morton was elected to Mar's office and proved in many ways the most effective of James's regents, but he made enemies by his rapacity. He fell from favour when Frenchman Esmé Stewart, Sieur d'Aubigny , first cousin of James's father Lord Darnley and future Earl of Lennox , arrived in Scotland and quickly established himself as
14720-472: The English succession became a cornerstone of his policy. During the Spanish Armada crisis of 1588, he assured Elizabeth of his support as "your natural son and compatriot of your country". Elizabeth sent James an annual subsidy from 1586 which gave her some leverage over affairs in Scotland. Throughout his youth, James was praised for his chastity, since he showed little interest in women. After
14904-607: The European cultivars or races Hallertau, Tettnanger, Spalt, and Saaz . Some proponents assert that the English varieties Fuggle, East Kent Goldings and Goldings might qualify as "noble hops" due to the similar composition, but such terms are not applied to English varieties. Their low relative bitterness, but strong aroma, are often distinguishing characteristics of European-style lagers , such as Pilsener , Dunkel , and Oktoberfest/Märzen . In beer, they are considered aroma hops (as opposed to bittering hops); see Pilsner Urquell as
15088-544: The Gunpowder Plot was atypical. Instead, it was the previous session of 1604 that shaped the attitudes of both sides for the rest of the reign, though the initial difficulties owed more to mutual incomprehension than conscious enmity. On 7 July 1604, James had angrily prorogued Parliament after failing to win its support either for full union or financial subsidies. "I will not thank where I feel no thanks due", he had remarked in his closing speech. "... I am not of such
15272-480: The Privy Council, as well as five Scottish nobles. In the early years of James's reign, the day-to-day running of the government was tightly managed by the shrewd Cecil, later Earl of Salisbury , ably assisted by the experienced Thomas Egerton , whom James made Baron Ellesmere and Lord Chancellor , and by Thomas Sackville , soon Earl of Dorset , who continued as Lord Treasurer . As a consequence, James
15456-694: The Reformation were slow to affect the Gàidhealtachd , driving a religious wedge between this area and centres of political control in the Central Belt . In 1540, James V had toured the Hebrides, forcing the clan chiefs to accompany him. There followed a period of peace, but the clans were soon at loggerheads with one another again. During James VI's reign, the citizens of the Hebrides were portrayed as lawless barbarians rather than being
15640-557: The Spanish ambassador Diego Sarmiento de Acuña, Count of Gondomar , James ripped the protest out of the record book and dissolved Parliament. In early 1623, Prince Charles, now 22, and Buckingham decided to seize the initiative and travel to Spain incognito, to win Infanta Maria Anna directly, but the mission proved an ineffectual mistake. Maria Anna detested Charles, and the Spanish confronted them with terms that included
15824-642: The Tower of London, where he had been placed at the king's request. Among those convicted of the murder were the Earl and Countess of Somerset; the Earl had been replaced as the king's favourite in the meantime by Villiers. James pardoned the Countess and commuted the Earl's sentence of death, eventually pardoning him in 1624. The implication of the king in such a scandal provoked much public and literary conjecture and irreparably tarnished James's court with an image of corruption and depravity. The subsequent downfall of
16008-485: The US have lately been the more significant breeders of new hop varieties, including dwarf hop varieties. Hops from New Zealand, such as Pacific Gem, Motueka and Nelson Sauvin, are used in a "Pacific Pale Ale" style of beer with increasing production in 2014. The term "noble hops" is a marketing term that traditionally refers to certain varieties of hops that became known for being low in bitterness and high in aroma . They are
16192-470: The Vegetable Kingdom at the beginning of chapter XII noted, "The first and most important of the conclusions which may be drawn from the observations given in this volume, is that generally cross-fertilisation is beneficial and self-fertilisation often injurious, at least with the plants on which I experimented." Self-pollination is the pollination of the carpel of a flower by pollen from either
16376-470: The age of thirteen months, after his mother was forced to abdicate in his favour. Four regents governed during his minority , which ended officially in 1578, though he did not gain full control of his government until 1583. In 1589, he married Anne of Denmark . Three of their children survived to adulthood: Henry Frederick , Elizabeth , and Charles . In 1603, James succeeded his cousin Elizabeth I ,
16560-425: The alpha acids or humulones . During wort boiling, the humulones are thermally isomerized into iso-alpha acids or isohumulones , which are responsible for the bitter taste of beer. Hops contain beta acids or lupulones . These are desirable for their aroma contributions to beer. The main components of hops essential oils are terpene hydrocarbons consisting of myrcene , humulene and caryophyllene . Myrcene
16744-490: The apical meristem, which becomes a whorl of sepals. In the second whorl, both A and B genes are expressed, leading to the formation of petals. In the third whorl, B and C genes interact to form stamens and in the center of the flower C genes alone give rise to carpels. The model is based upon studies of aberrant flowers and mutations in Arabidopsis thaliana and the snapdragon, Antirrhinum majus . For example, when there
16928-485: The aromatic compounds evaporate during the boil. The degree of bitterness imparted by hops depends on the degree to which alpha acids are isomerized during the boil, and the impact of a given amount of hops is specified in International Bitterness Units . On the other hand, unboiled hops are only mildly bitter . Aroma hops are typically added to the wort later to prevent the evaporation of
17112-582: The bodies of the pollinator when it lands in search of its attractant. Other flowers use mimicry or pseudocopulation to attract pollinators. Many orchids, for example, produce flowers resembling female bees or wasps in color, shape, and scent. Males move from one flower to the next in search of a mate, pollinating the flowers. Many flowers have close relationships with one or a few specific pollinating organisms. Many flowers, for example, attract only one specific species of insect and therefore rely on that insect for successful reproduction. This close relationship
17296-467: The brewing process the papery cones are discarded. The need for massed labor at harvest time meant hop-growing had a big social impact. Around the world, the labor-intensive harvesting work involved large numbers of migrant workers who would travel for the annual hop harvest. Whole families would participate and live in hoppers' huts , with even the smallest children helping in the fields. The final chapters of W. Somerset Maugham 's Of Human Bondage and
17480-454: The child's mouth, as was then the custom. The subsequent entertainment , devised by Frenchman Bastian Pagez , featured men dressed as satyrs and sporting tails, to which the English guests took offence, thinking the satyrs "done against them". Lord Darnley was murdered on 10 February 1567 at Kirk o' Field, Edinburgh, perhaps in revenge for the killing of Rizzio. James inherited his father's titles of Duke of Albany and Earl of Ross . Mary
17664-517: The collective cluster of flowers is called an inflorescence . Some inflorescences are composed of many small flowers arranged in a formation that resembles a single flower. A common example of this is most members of the very large composite ( Asteraceae ) group. A single daisy or sunflower , for example, is not a flower but a flower head —an inflorescence composed of numerous flowers (or florets). An inflorescence may include specialized stems and modified leaves known as bracts . A floral formula
17848-503: The context of 17th-century court life, and remain ambiguous despite their fondness. It is also possible that James was bisexual. When the Earl of Salisbury died in 1612, he was little mourned by those who jostled to fill the power vacuum. Until Salisbury's death, the Elizabethan administrative system over which he had presided continued to function with relative efficiency; from this time forward, however, James's government entered
18032-464: The continental United States and Canada, cultivation of hops for commercial production requires a particular environment. As hops are a climbing plant, they are trained to grow up trellises made from strings or wires that support the plants and allow them significantly greater growth with the same sunlight profile. In this way, energy that would have been required to build structural cells is also freed for crop growth. The hop plant's reproduction method
18216-491: The cradle of Scottish Christianity and nationhood. Official documents describe the peoples of the Highlands as "void of the knawledge and feir of God" who were prone to "all kynd of barbarous and bestile cruelteis". The Gaelic language, spoken fluently by James IV and probably by James V, became known in the time of James VI as "Erse" or Irish, implying that it was foreign in nature. Parliament decided that Gaelic had become
18400-566: The crossing had been abandoned, James sailed from Leith with a 300-strong retinue to fetch Anne personally in what historian David Harris Willson called "the one romantic episode of his life". The couple were married formally at the Bishop's Palace in Oslo on 23 November. James received a dowry of 75,000 Danish dalers and a gift of 10,000 dalers from his mother-in-law, Sophie of Mecklenburg-Güstrow . After stays at Elsinore and Copenhagen and
18584-619: The developing flower. These petals attract pollinators, and reproductive organs that produce gametophytes , which in flowering plants produce gametes . The male gametophytes, which produce sperm, are enclosed within pollen grains produced in the anthers . The female gametophytes are contained within the ovules produced in the ovary . Most flowering plants depend on animals, such as bees, moths, and butterflies, to transfer their pollen between different flowers, and have evolved to attract these pollinators by various strategies, including brightly colored, conspicuous petals, attractive scents, and
18768-440: The eldest son and heir apparent of the monarch automatically became Duke of Rothesay and Prince and Great Steward of Scotland . Five days later, the English diplomat Henry Killigrew saw the queen, who had not fully recovered and could only speak faintly. The baby was "sucking at his nurse" and was "well proportioned and like to prove a goodly prince". He was baptised "Charles James" or "James Charles" on 17 December 1566 in
18952-407: The essential oil, such as an alpha:beta ratio of 1:1, low alpha-acid levels (2–5%) with a low cohumulone content, low myrcene in the hop oil, high humulene in the oil, a ratio of humulene:caryophyllene above three, and poor storability resulting in them being more prone to oxidation. In reality, this means they have a relatively consistent bittering potential as they age, due to beta-acid oxidation, and
19136-439: The essential oils, to impart "hop taste" (if during the final 30 minutes of boil) or "hop aroma" (if during the final 10 minutes, or less, of boil). Aroma hops are often added after the wort has cooled and while the beer ferments, a technique known as "dry hopping", which contributes to the hop aroma. Farnesene is a major component in some hops. The composition of hop essential oils can differ between varieties and between years in
19320-484: The existing model to broaden the descriptive capability of the formula. The format of floral formulae differs in different parts of the world, yet they convey the same information. The structure of a flower can also be expressed by the means of floral diagrams . The use of schematic diagrams can replace long descriptions or complicated drawings as a tool for understanding both floral structure and evolution. Such diagrams may show important features of flowers, including
19504-977: The first dwarf hop varieties, which are easier to pick by machine and far more economical to grow. Wye College have also been responsible for breeding hop varieties that will grow with only 12 hours of daily light for the South African hop farmers. Wye College was closed in 2009 but the legacy of their hop breeding programs, particularly that of the dwarf varieties, is continuing as already the US private and public breeding programs are using their stock material. Particular hop varieties are associated with beer regions and styles, for example pale lagers are usually brewed with European (often German, Polish or Czech) noble hop varieties such as Saaz , Hallertau and Strissel Spalt. British ales use hop varieties such as Fuggles, Goldings and W.G.V. North American beers often use Cascade hops , Columbus hops, Centennial hops, Willamette, Amarillo hops and about forty more varieties as
19688-455: The first of James's powerful favourites. James was proclaimed an adult ruler in a ceremony of Entry to Edinburgh on 19 October 1579. Morton was executed on 2 June 1581, belatedly charged with complicity in Darnley's murder. On 8 August, James made Lennox the only duke in Scotland. The king, then fifteen years old, remained under the influence of Lennox for about one more year. Lennox was
19872-587: The first year of his reign, despite the smoothness of the succession and the warmth of his welcome: the Bye Plot and Main Plot , which led to the arrest of Henry Brooke, Lord Cobham and Walter Raleigh , among others. Those hoping for a change in government from James were disappointed at first when he kept Elizabeth's Privy Councillors in office, as secretly planned with Cecil, but James soon added long-time supporter Henry Howard and his nephew Thomas Howard to
20056-413: The flower is said to be actinomorphic or regular. This is an example of radial symmetry . When flowers are bisected and produce only one line that produces symmetrical halves, the flower is said to be irregular or zygomorphic . If, in rare cases, they have no symmetry at all they are called asymmetric. Flowers may be directly attached to the plant at their base ( sessile —the supporting stalk or stem
20240-497: The flower's stigma. This pollination does not require an investment from the plant to provide nectar and pollen as food for pollinators. Some flowers produce diaspores without fertilization ( parthenocarpy ). After fertilization, the ovary of the flower develops into fruit containing seeds . Flowers have long been appreciated for their beauty and pleasant scents, and also hold cultural significance as religious, ritual, or symbolic objects, or sources of medicine and food. Flower
20424-511: The flowers are imperfect or unisexual: having only either male (stamen) or female (carpel) parts. If unisexual male and female flowers appear on the same plant, the species is called monoecious . However, if an individual plant is either female or male, the species is called dioecious . Many flowers have nectaries , which are glands that produce a sugary fluid used to attract pollinators. They are not considered as an organ on their own. In those species that have more than one flower on an axis,
20608-424: The following two broad groups of pollination methods: Flowers that use biotic vectors attract and use insects , bats , birds , or other animals to transfer pollen from one flower to the next. Often they are specialized in shape and have an arrangement of the stamens that ensures that pollen grains are transferred to the bodies of the pollinator when it lands in search of its attractant (such as nectar, pollen, or
20792-451: The gametophytes also develop inside the spores, i.e., they are endosporic. Since the flowers are the reproductive organs of the plant, they mediate the joining of the sperm, contained within pollen, to the ovules — contained in the ovary. Pollination is the movement of pollen from the anthers to the stigma. Normally pollen is moved from one plant to another, known as cross-pollination , but many plants can self-pollinate. Cross-pollination
20976-623: The government and its patronage. Even the powerful Carr fell into the Howard camp, hardly experienced for the responsibilities thrust upon him and often dependent on his intimate friend Thomas Overbury for assistance with government papers. Carr had an adulterous affair with Frances Howard, Countess of Essex , daughter of the Earl of Suffolk. James assisted Frances by securing an annulment of her marriage to free her to marry Carr, now Earl of Somerset. In summer 1615, however, it emerged that Overbury had been poisoned. He had died on 15 September 1613 in
21160-649: The harvest became a matter of scandal across Kent and other hop-growing counties. Eventually, the Rev. John Young Stratton , Rector of Ditton, Kent, began to gather support for reform, resulting in 1866 in the formation of the Society for the Employment and Improved Lodging of Hop Pickers. The hop-pickers were given very basic accommodation, with very poor sanitation. This led to the spread of infectious diseases and led to contaminated water. The 1897 Maidstone typhoid epidemic
21344-507: The hop industry derive from the Dutch language . Hops were then grown as far north as Aberdeen , near breweries for convenience of infrastructure. According to Thomas Tusser 's 1557 Five Hundred Points of Good Husbandry : The hop for his profit I thus do exalt, It strengtheneth drink and it flavored malt; And being well-brewed long kept it will last, And drawing abide, if ye draw not too fast. In England there were many complaints over
21528-486: The hops are spread out on the upper floor of the oast house and heated by heating units on the lower floor. The dried hops are then compressed into bales by a baler . Hop cones contain different oils, such as lupulin, a yellowish, waxy substance, an oleoresin , that imparts flavour and aroma to beer. Lupulin contains lupulone and humulone , which possess antibiotic properties, suppressing bacterial growth favoring brewer's yeast to grow. After lupulin has been extracted in
21712-561: The king had personally appointed royal chaplain in 1579, rebuked him so sharply from the pulpit for having issued a proclamation so offensive to the clergy "that the king wept". After James escaped from Falkland on 27 June 1583, he assumed increasing control of his kingdom. He pushed through the Black Acts to assert royal authority over the Kirk, and denounced the writings of his former tutor Buchanan. Between 1584 and 1603, he established effective royal government and relative peace among
21896-504: The king to London after 1603. James's role as active literary participant and patron made him a defining figure in many respects for English Renaissance poetry and drama, which reached a pinnacle of achievement in his reign, but his patronage of the high style in the Scottish tradition, which included his ancestor James I of Scotland , became largely sidelined. From 1601, in the last years of Elizabeth's life, certain English politicians—notably her chief minister Robert Cecil —maintained
22080-473: The king's "head court", foreshadows his difficulties with the English House of Commons : "Hold no Parliaments," he tells Henry, "but for the necesitie of new Lawes, which would be but seldome". In the True Law , James maintains that the king owns his realm as a feudal lord owns his fief, because kings arose "before any estates or ranks of men, before any parliaments were holden, or laws made, and by them
22264-530: The king's person occurred in August 1600, when James was apparently assaulted by Alexander Ruthven , the younger brother of John Ruthven, Earl of Gowrie at Gowrie House , the seat of the Ruthvens. Ruthven was run through by James's page John Ramsay , and the Earl of Gowrie was killed in the ensuing fracas; there were few surviving witnesses. Given James's history with the Ruthvens and the fact that he owed them
22448-478: The large majority of the bitter flavour of a beer. European (so-called "noble") hops typically average 5–9% alpha acids by weight (AABW), and the newer American cultivars typically range from 8–19% AABW. Aroma hops usually have a lower concentration of alpha acids (~5%) and are the primary contributors of hop aroma and (nonbitter) flavour. Bittering hops are boiled for a longer period of time, typically 60–90 minutes, and often have inferior aromatic properties, as
22632-577: The last Tudor monarch of England and Ireland, who died childless. He continued to reign in all three kingdoms for 22 years, a period known as the Jacobean era , until his death in 1625. After the Union of the Crowns, he based himself in England (the largest of the three realms) from 1603, returning to Scotland only once, in 1617, and styled himself " King of Great Britain and Ireland ". He was an advocate of
22816-641: The leading potato-growing states in the United States are also major hops-producing areas; however, not all potato-growing areas can produce good hops naturally: soils in the Maritime Provinces of Canada, for example, lack the boron that hops prefer. Historically, hops were not grown in Ireland, but were imported from England. In 1752 more than 500 tons of English hops were imported through Dublin alone. Important production centres today are
23000-416: The local industry by enabling large-scale mechanized production, which moved to larger farms in other areas. Dauenhauer Manufacturing Company remains a current producer of hop harvesting machines. In addition to water, cellulose, and various proteins, the chemical composition of hops consists of compounds important for imparting character to beer. Probably the most important chemical compound within hops are
23184-565: The lords, ably assisted by John Maitland of Thirlestane , who led the government until 1592. An eight-man commission known as the Octavians brought some control over the ruinous state of James's finances in 1596, but it drew opposition from vested interests. It was disbanded within a year after a riot in Edinburgh, which was stoked by anti-Catholicism and led the court to withdraw to Linlithgow temporarily. One last Scottish attempt against
23368-528: The loss of Lennox, he continued to prefer male company. A suitable marriage, however, was necessary to reinforce his rule, and the choice fell on fourteen-year-old Anne of Denmark , younger daughter of the Protestant Danish king Frederick II . Shortly after a proxy marriage in Copenhagen in August 1589, Anne sailed for Scotland but was forced by storms to the coast of Norway. On hearing that
23552-478: The necessary finance, they prevailed upon James to call another Parliament, which met in February 1624. For once, the outpouring of anti-Catholic sentiment in the Commons was echoed in court, where control of policy was shifting from James to Charles and Buckingham, who pressured the king to declare war and engineered the impeachment of Lord Treasurer Lionel Cranfield, by now made Earl of Middlesex , when he opposed
23736-653: The negotiations for almost a decade. The policy was supported by the Howards and other Catholic-leaning ministers and diplomats—together known as the Spanish Party—but deeply distrusted in Protestant England. When Walter Raleigh was released from imprisonment in 1616, he embarked on a hunt for gold in South America with strict instructions from James not to engage the Spanish. Raleigh's expedition
23920-446: The older sister of Henry VIII . Mary's rule over Scotland was insecure, and she and her husband, being Roman Catholics , faced a rebellion by Protestant noblemen. During Mary's and Darnley's difficult marriage, Darnley secretly allied himself with the rebels and conspired in the murder of the queen's private secretary, David Rizzio , just three months before James's birth. James was born on 19 June 1566 at Edinburgh Castle , and as
24104-400: The ovary. After penetrating the center-most part of the ovary it enters the egg apparatus and into one synergid . At this point the end of the pollen tube bursts and releases the two sperm cells, one of which makes its way to an egg, while also losing its cell membrane and much of its protoplasm . The sperm's nucleus then fuses with the egg's nucleus, resulting in the formation of a zygote ,
24288-504: The ovule is fertilized by pollen from the same flower or plant, occurs in flowers that always self-fertilize, such as many dandelions . Some flowers are self-pollinated and have flowers that never open or are self-pollinated before the flowers open; these flowers are called cleistogamous ; many species in the genus Viola exhibit this, for example. Conversely, many species of plants have ways of preventing self-pollination and hence, self-fertilization. Unisexual male and female flowers on
24472-524: The ovules. Carpels may occur in one to several whorls, and when fused are often described as a pistil . Inside the ovary, the ovules are attached to the placenta by structures called funiculi . Although this arrangement is considered "typical", plant species show a wide variation in floral structure. The four main parts of a flower are generally defined by their positions on the receptacle and not by their function. Many flowers lack some parts or parts may be modified into other functions or look like what
24656-534: The perianth. They are often delicate and thin and are usually colored, shaped, or scented to encourage pollination. Although similar to leaves in shape, they are more comparable to stamens in that they form almost simultaneously with one another, but their subsequent growth is delayed. If the corolla is fused together it is called sympetalous. The androecium , or stamens, is the whorl of pollen-producing male parts. Stamens consist typically of an anther , made up of four pollen sacs arranged in two thecae , connected to
24840-481: The plan on grounds of cost. The outcome of the Parliament of 1624 was ambiguous: James still refused to declare or fund a war, but Charles believed the Commons had committed themselves to finance a war against Spain, a stance that was to contribute to his problems with Parliament in his own reign. After the Gunpowder Plot, James sanctioned harsh measures to control English Catholics. In May 1606, Parliament passed
25024-451: The plant so as to not force competition between the mother and the daughter plants, as well as to enable the colonization of new areas. They are often divided into two categories, though many plants fall in between or in one or more of these: In allochory, plants use an external vector , or carrier, to transport their seeds away from them. These can be either biotic (living), such as by birds and ants, or abiotic (non-living), such as by
25208-400: The process of pollination. Fertilization , also called Synagmy, occurs following pollination, which is the movement of pollen from the stamen to the carpel. It encompasses both plasmogamy , the fusion of the protoplasts , and karyogamy , the fusion of the nuclei . When pollen lands on the stigma of the flower it begins creating a pollen tube which runs down through the style and into
25392-453: The production of nectar , a food source for pollinators. In this way, many flowering plants have co-evolved with pollinators to be mutually dependent on services they provide to one another—in the plant's case, a means of reproduction; in the pollinator's case, a source of food. When pollen from the anther of a flower is deposited on the stigma , this is called pollination. Some flowers may self-pollinate , producing seed using pollen from
25576-476: The prospect of a Spanish dowry from a marriage between Charles, Prince of Wales, and Infanta Maria Anna of Spain . The policy of the Spanish match , as it was called, was also attractive to James as a way to maintain peace with Spain and avoid the additional costs of a war. Peace could be maintained as effectively by keeping the negotiations alive as by consummating the match—which may explain why James protracted
25760-651: The quality of imported hops, the sacks of which were often contaminated by stalks, sand or straw to increase their weight. As a result, in 1603, King James I approved an Act of Parliament banning the practice by which "the Subjects of this Realm have been of late years abused &c. to the Value of £20,000 yearly, besides the Danger of their Healths". Hop cultivation was begun in the present-day United States in 1629 by English and Dutch farmers. Before prohibition , cultivation
25944-600: The relative positions of the various organs, including the presence of fusion and symmetry, as well as structural details. A flower develops on a modified shoot or axis from a determinate apical meristem ( determinate meaning the axis grows to a set size). It has compressed internodes, bearing structures that in classical plant morphology are interpreted as highly modified leaves . Detailed developmental studies, however, have shown that stamens are often initiated more or less like modified stems (caulomes) that in some cases may even resemble branchlets . Taking into account
26128-592: The repeal of anti-Catholic legislation by Parliament. Though a treaty was signed, Charles and Buckingham returned to England in October without the infanta and immediately renounced the treaty, much to the delight of the British people. Disillusioned by the visit to Spain, Charles and Buckingham now turned James's Spanish policy upon its head and called for a French match and a war against the Habsburg empire. To raise
26312-479: The reproduction of a recipe allowing for differences in hop availability. Lately the dried pucks, extracts and pellets replace whole hops in brewing processes because of efficiency and cost. There are many different varieties of hops used in brewing today. Historically, hops varieties were identified by geography, i.e., from the towns of Hallertau , Spalt , and Tettnang in Germany , or the region writ large like
26496-420: The reproductive organs. Growth of the central part of the stem tip stops or flattens out and the sides develop protuberances in a whorled or spiral fashion around the outside of the stem end. These protuberances develop into the sepals, petals, stamens, and carpels . Once this process begins, in most plants, it cannot be reversed and the stems develop flowers, even if the initial start of the flower formation event
26680-477: The roots send forth new bines that are started up strings from the ground to an overhead trellis. The cones grow high on the bine, and in the past, these cones were picked by hand. Harvesting of hops became much more efficient with the invention of the mechanical hops separator, patented by Emil Clemens Horst in 1909. Hops are harvested at the end of summer. The bines are cut down, separated, and then dried in an oast house to reduce moisture content. To be dried,
26864-639: The route and James was amazed by the wealth of his new land and subjects, claiming that he was "swapping a stony couch for a deep feather bed". James arrived in the capital on 7 May, nine days after Elizabeth's funeral. His new subjects flocked to see him, relieved that the succession had triggered neither unrest nor invasion. On arrival at London, he was mobbed by a crowd of spectators. James's English coronation took place on 25 July at Westminster Abbey . An outbreak of plague restricted festivities. The Royal Entry to London with elaborate allegories provided by dramatic poets such as Thomas Dekker and Ben Jonson
27048-490: The same flower or another flower on the same plant, leading to the creation of a genetic clone through asexual reproduction . This increases the reliability of producing seeds, the rate at which they can be produced, and lowers the amount energy needed. But, most importantly, it limits genetic variation . In addition, self-pollination causes inbreeding depression , due largely to the expression of recessive deleterious mutations . The extreme case of self-fertilization, when
27232-412: The same plant may not appear or mature at the same time, or pollen from the same plant may be incapable of fertilizing its ovules. The latter flower types, which have chemical barriers to their own pollen, are referred to as self-incompatible. In Clianthus puniceus , self-pollination is used strategically as an "insurance policy". When a pollinator, in this case a bird, visits C. puniceus , it rubs off
27416-481: The same result, although a third attempt in 1607 was more successful. The Statutes of Iona were enacted in 1609, which required clan chiefs to provide support for Protestant ministers to Highland parishes; to outlaw bards; to report regularly to Edinburgh to answer for their actions; and to send their heirs to Lowland Scotland , to be educated in English-speaking Protestant schools. So began
27600-769: The same species. The period during which this process can take place (when the flower is fully expanded and functional) is called anthesis , hence the study of pollination biology is called anthecology . Flowering plants usually face evolutionary pressure to optimize the transfer of their pollen , and this is typically reflected in the morphology of the flowers and the behavior of the plants. Pollen may be transferred between plants via several 'vectors,' or methods. Around 80% of flowering plants make use of biotic or living vectors. Others use abiotic, or non-living, vectors and some plants make use of multiple vectors, but most are highly specialized. Though some fit between or outside of these groups, most flowers can be divided between
27784-624: The same variety, having a significant influence on flavour and aroma. Today, a substantial amount of "dual-use" hops are used, as well. These have high concentrations of alpha acids and good aromatic properties. These can be added to the boil at any time, depending on the desired effect. Hop acids also contribute to and stabilize the foam qualities of beer. Flavours and aromas are described appreciatively using terms which include "grassy", "floral", "citrus", "spicy", "piney", "lemony", "grapefruit", and "earthy". Many pale lagers have fairly low hop influence, while lagers marketed as Pilsener or brewed in
27968-482: The seed. The ovary, inside which the seed is forming from the ovule, grows into a fruit . All the other main floral parts die during this development, including: the style, stigma, sepals, stamens, and petals. The fruit contains three structures: the exocarp , or outer layer, the mesocarp , or the fleshy part, and the endocarp , or innermost layer, while the fruit wall is called the pericarp . The size, shape, toughness, and thickness varies among different fruit. This
28152-492: The skilled diplomacy of the delegation, in particular Robert Cecil and Henry Howard, now Earl of Northampton . James celebrated the treaty by hosting a great banquet. Freedom of worship for Catholics in England, however, continued to be a major objective of Spanish policy, causing constant dilemmas for James, distrusted abroad for repression of Catholics while at home being encouraged by the Privy Council to show even less tolerance towards them. A dissident Catholic, Guy Fawkes ,
28336-439: The species. All flowering plants are heterosporous , that is, every individual plant produces two types of spores . Microspores are produced by meiosis inside anthers and megaspores are produced inside ovules that are within an ovary. Anthers typically consist of four microsporangia and an ovule is an integumented megasporangium. Both types of spores develop into gametophytes inside sporangia. As with all heterosporous plants,
28520-428: The stigmatic covering and allows for pollen from the bird to enter the stigma. If no pollinators visit, however, then the stigmatic covering falls off naturally to allow for the flower's own anthers to pollinate the flower through self-pollination. Pollen is a large contributor to asthma and other respiratory allergies which combined affect between 10 and 50% of people worldwide. This number appears to be growing, as
28704-525: The support of Catholics in England, so he assured Henry Percy, Earl of Northumberland , a prominent sympathiser of the old religion, that he would not persecute "any that will be quiet and give but an outward obedience to the law". In the Millenary Petition of 1603, the Puritan clergy demanded the abolition of confirmation , wedding rings, and the term "priest", among other things, and that
28888-463: The temperature increases due to climate change mean that plants are producing more pollen , which is also more allergenic. Pollen is difficult to avoid, however, because of its small size and prevalence in the natural environment. Most of the pollen which causes allergies is that produced by wind-dispersed pollinators such as the grasses , birch trees , oak trees, and ragweeds ; the allergens in pollen are proteins which are thought to be necessary in
29072-627: The third consecutive Scottish monarch to ascend to the throne as an infant. The care of James was entrusted to the Earl and Countess of Mar , "to be conserved, nursed, and upbrought" in the security of Stirling Castle. James was anointed King of Scotland at the age of thirteen months at the Church of the Holy Rude in Stirling, by Adam Bothwell , Bishop of Orkney , on 29 July 1567. The sermon at
29256-490: The title was not used on English statutes. James forced the Scottish Parliament to use it, and it was used on proclamations, coinage, letters, and treaties in both realms. James achieved more success in foreign policy. Never having been at war with Spain, he devoted his efforts to bringing the long Anglo–Spanish War to an end, and a peace treaty was signed between the two countries in August 1604, thanks to
29440-435: The torture of women accused of being witches. After 1599, his views became more sceptical. In a later letter written in England to his son Henry, James congratulates the prince on "the discovery of yon little counterfeit wench. I pray God ye may be my heir in such discoveries ... most miracles now-a-days prove but illusions, and ye may see by this how wary judges should be in trusting accusations". The forcible dissolution of
29624-428: The vegetative part, consisting of non-reproductive structures such as petals ; and the reproductive or sexual parts. A stereotypical flower is made up of four kinds of structures attached to the tip of a short stalk or axis, called a receptacle . Each of these parts or floral organs is arranged in a spiral called a whorl . The four main whorls (starting from the base of the flower or lowest node and working upwards) are
29808-619: The water and so the pollen has a low density to enable floating, though many also use rafts, and are hydrophobic . Marine flowers have floating thread-like stigmas and may have adaptations for the tide, while freshwater species create indentations in the water. The third category, set out by Schwarzenbach, is those flowers which transport pollen above the water through conveyance. This ranges from floating plants, ( Lemnoideae ), to staminate flowers ( Vallisneria ). Most species in this group have dry, spherical pollen which sometimes forms into larger masses, and female flowers which form depressions in
29992-401: The water; the method of transport varies. Flowers can be pollinated by two mechanisms; cross-pollination and self-pollination. No mechanism is indisputably better than the other as they each have their advantages and disadvantages. Plants use one or both of these mechanisms depending on their habitat and ecological niche . Cross-pollination is the pollination of the carpel by pollen from
30176-514: The wearing of cap and surplice become optional. James was strict in enforcing conformity at first, inducing a sense of persecution amongst many Puritans; but ejections and suspensions from livings became rarer as the reign continued. As a result of the Hampton Court Conference of 1604, some Puritan demands were acceded to in the 1604 Book of Common Prayer , though many remained displeased. The conference also commissioned
30360-517: The whole body of the State in general". The sensational discovery of the "Gunpowder Plot", as it quickly became known, aroused a mood of national relief at the delivery of the king and his sons. The Earl of Salisbury exploited this to extract higher subsidies from the ensuing Parliament than any but one granted to Elizabeth. Fawkes and others implicated in the unsuccessful conspiracy were executed. The co-operation between monarch and Parliament following
30544-480: The whole diversity in the development of the androecium of flowering plants, we find a continuum between modified leaves (phyllomes), modified stems (caulomes), and modified branchlets (shoots). The transition to flowering is one of the major phase changes that a plant makes during its life cycle. The transition must take place at a time that is favorable for fertilization and the formation of seeds , hence ensuring maximal reproductive success. To meet these needs
30728-460: The wind or water. Many plants use biotic vectors to disperse their seeds away from them. This method falls under the umbrella term zoochory , while endozoochory , also known as fruigivory, refers specifically to plants adapted to grow fruit in order to attract animals to eat them. Once eaten they go through typically go through animal's digestive system and are dispersed away from the plant. Some seeds are specially adapted either to last in
30912-419: The wind to effectively pick up and transport the pollen, the flowers typically have anthers loosely attached to the end of long thin filaments, or pollen forms around a catkin which moves in the wind. Rarer forms of this involve individual flowers being moveable by the wind ( pendulous ), or even less commonly; the anthers exploding to release the pollen into the wind. Pollination through water ( hydrophily )
31096-606: The work of Prof. E. S. Salmon . Salmon released Brewer's Gold and Brewer's Favorite for commercial cultivation in 1934, and went on to release more than two dozen new cultivars before his death in 1959. Brewer's Gold has become the ancestor of the bulk of new hop releases around the world since its release. Wye College continued its breeding program and again received attention in the 1970s, when Dr. Ray A. Neve released Wye Target , Wye Challenger , Wye Northdown , Wye Saxon and Wye Yeoman . More recently, Wye College and its successor institution Wye Hops Ltd., have focused on breeding
31280-456: The young king's senior tutor, Buchanan subjected James to regular beatings but also instilled in him a lifelong passion for literature and learning. Buchanan sought to turn James into a God-fearing, Protestant king who accepted the limitations of monarchy, as outlined in his treatise De Jure Regni apud Scotos . In 1568, Mary escaped from Lochleven Castle, leading to several years of sporadic violence. The Earl of Moray defeated Mary's troops at
31464-522: Was partly as a result of hop-pickers camping near the Farleigh Springs which supplied Maidstone with water. Particularly in Kent, because of a shortage of small-denomination coin of the realm, many growers issued their own currency to those doing the labor. In some cases, the coins issued were adorned with fanciful hops images, making them quite beautiful. In the United States, Prohibition had
31648-622: Was a disastrous failure, and his son Walter was killed fighting the Spanish. On Raleigh's return to England, James had him executed to the indignation of the public, who opposed the appeasement of Spain. James's policy was further jeopardised by the outbreak of the Thirty Years' War , especially after his Protestant son-in-law, Frederick V, Elector Palatine , was ousted from Bohemia by the Catholic Emperor Ferdinand II in 1620, and Spanish troops simultaneously invaded Frederick's Rhineland home territory. Matters came to
31832-552: Was already unpopular, and her marriage on 15 May 1567 to James Hepburn, Earl of Bothwell , who was widely suspected of murdering Darnley, heightened widespread bad feeling towards her. In June 1567, Protestant rebels arrested Mary and imprisoned her in Lochleven Castle ; she never saw her son again. She was forced to abdicate on 24 July 1567 in favour of the infant James and to appoint her illegitimate half-brother James Stewart, Earl of Moray , as regent . This made James
32016-611: Was also a religious and political choice in the early 16th century. There was no tax on hops to be paid to the Catholic church, unlike on gruit. For this reason the Protestants preferred hopped beer. Hops used in England were imported from France, Holland and Germany and were subject to import duty ; it was not until 1524 that hops were first grown in the southeast of England ( Kent ), when they were introduced as an agricultural crop by Dutch farmers. Consequently, many words used in
32200-472: Was deferred to 15 March 1604. Dekker wrote that "the streets seemed to be paved with men; stalls instead of rich wares were set out with children; open casements filled up with women". The kingdom to which James succeeded, however, had its problems. Monopolies and taxation had engendered a widespread sense of grievance, and the costs of war in Ireland had become a heavy burden on the government, which had debts of £400,000. James survived two conspiracies in
32384-416: Was dependent on some environmental cue. The ABC model is a simple model that describes the genes responsible for the development of flowers. Three gene activities interact in a combinatorial manner to determine the developmental identities of the primordia organ within the floral apical meristem . These gene functions are called A, B, and C. Genes are expressed in only the outer and lower most section of
32568-437: Was discovered in the cellars of the parliament buildings on the night of 4–5 November 1605, the eve of the state opening of the second session of James's first English Parliament. Fawkes was guarding a pile of wood not far from 36 barrels of gunpowder with which he intended to blow up Parliament House the following day and cause the destruction, as James put it, "not only ... of my person, nor of my wife and posterity also, but of
32752-455: Was free to concentrate on bigger issues, such as a scheme for a closer union between England and Scotland and matters of foreign policy, as well as to enjoy his leisure pursuits, particularly hunting. James was ambitious to build on the personal union of Scotland and England to establish a single country under one monarch, one parliament, and one law, a plan that met opposition in both realms. "Hath He not made us all in one island," James told
32936-496: Was in 736, in the Hallertau region of present-day Germany, although the first mention of the use of hops in brewing in that country was 1079. However, in a will of Pepin the Short , the father of Charlemagne , hop gardens were left to the Cloister of Saint-Denis in 768. Not until the 13th century did hops begin to start threatening the use of gruit for flavouring. Gruit was used when the nobility levied taxes on hops. Whichever
33120-667: Was known as the king's mistress. After his accession in England, his peaceful and scholarly attitude contrasted strikingly with the bellicose and flirtatious behaviour of Elizabeth, as indicated by the contemporary epigram Rex fuit Elizabeth, nunc est regina Iacobus (Elizabeth was King, now James is Queen). Some of James's biographers conclude that Esmé Stewart, Duke of Lennox; Robert Carr, Earl of Somerset ; and George Villiers, Duke of Buckingham, were his lovers. John Oglander observed that he "never yet saw any fond husband make so much or so great dalliance over his beautiful spouse as I have seen King James over his favourites, especially
33304-418: Was mainly centred around New York, California, Oregon, and Washington state . Problems with powdery mildew and downy mildew devastated New York's production by the 1920s, and California only produces hops on a small scale. Hops production is concentrated in moist temperate climates, with much of the world's production occurring near the 48th parallel north . Hop plants prefer the same soils as potatoes and
33488-464: Was seriously ill from 1617. James visited Anne only three times during her last illness. She died before her husband , in March 1619. James's visit to Denmark, a country familiar with witch-hunts , sparked an interest in the study of witchcraft , which he considered a branch of theology. He attended the North Berwick witch trials , the first major persecution of witches in Scotland under
33672-435: Was taxed made the brewer then quickly switch to the other. In Britain, hopped beer was first imported from Holland around 1400, yet hops were condemned as late as 1519 as a "wicked and pernicious weed". In 1471, Norwich , England, banned use of the plant in the brewing of ale ("beer" was the name for fermented malt liquors bittered with hops; only in recent times are the words often used as synonyms). In Germany, using hops
33856-481: Was the land distributed, which at first was wholly theirs. And so it follows of necessity that kings were the authors and makers of the laws, and not the laws of the kings." In the 1580s and 1590s, James promoted the literature of his native country. He published his treatise Some Rules and Cautions to be Observed and Eschewed in Scottish Prosody in 1584 at the age of 18. It was both a poetic manual and
#119880