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Archaeogenetics is the study of ancient DNA using various molecular genetic methods and DNA resources. This form of genetic analysis can be applied to human, animal, and plant specimens. Ancient DNA can be extracted from various fossilized specimens including bones, eggshells, and artificially preserved tissues in human and animal specimens. In plants, ancient DNA can be extracted from seeds and tissue. Archaeogenetics provides us with genetic evidence of ancient population group migrations, domestication events, and plant and animal evolution. The ancient DNA cross referenced with the DNA of relative modern genetic populations allows researchers to run comparison studies that provide a more complete analysis when ancient DNA is compromised.

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128-790: [REDACTED] Look up hunnic in Wiktionary, the free dictionary. Hunnic may refer to: Huns , a former nomadic tribe of the Eurasian steppe Hunnic language , spoken by the Huns Hunnic grapes , a class of grapes grown in German-speaking countries during the Middle Ages See also [ edit ] Hun (disambiguation) Hunnic War (disambiguation) Hunno, Alamannic king in 385 Topics referred to by

256-748: A hemorrhage on his wedding night. After Attila's death in 453, the Hunnic Empire faced an internal power struggle between its vassalized Germanic peoples and the Hunnic ruling body. Led by Ellak , Attila's favored son and ruler of the Akatziri , the Huns engaged the Gepid king Ardaric at the Battle of Nedao , who led a Germanic coalition to overthrow Hunnic imperial authority. The Amali Goths would revolt

384-431: A basic laboratory setup and chemicals. It is also independent of sample size, as the process can be scaled to accommodate larger or smaller quantities. Another benefit is that the process can be executed at room temperature. However, this method does contain some drawbacks. Mainly, silica-based DNA extraction can only be applied to bone and teeth samples; they cannot be used on soft tissue . While they work well with

512-545: A closely related extant species can be used to estimate the divergence time of those two species from their last common ancestor . The phylogeny of some extinct species, such as Australian marsupial wolves and American ground sloths , has been constructed by this method. Mitochondrial DNA in animals and chloroplast DNA in plants are usually used for this purpose because they have hundreds of copies per cell and thus are more easily accessible in ancient fossils. Another method to investigate relationship between two species

640-531: A connection. The issue remains controversial, but recent archaeogenetic studies show some Hun-era individuals to have DNA similar to populations in ancient Mongolia. Their relationships with other entities, such as the Iranian Huns and the Huna people of South Asia , have also been disputed. Very little is known about Hunnic culture, and very few archaeological remains have been conclusively associated with

768-477: A date of over 300 kya. Examination of mitochondrial DNA (mtDNA), Y-chromosome DNA, and X-chromosome DNA indicate that the earliest population to leave Africa consisted of approximately 1500 males and females. It has been suggested by various studies that populations were geographically “structured” to some degree prior to the expansion out of Africa; this is suggested by the antiquity of shared mtDNA lineages. One study of 121 populations from various places throughout

896-437: A fixed rank with fixed duties. Kim affirms the importance of the logades for Hunnic administration, but notes that there were differences of rank between them, and suggests that it was more likely lower ranking officials who gathered taxes and tribute. He suggests that various Roman defectors to the Huns may have worked in a sort of imperial bureaucracy. Unlike the Iranian Huns , who quickly began to mint their own coinage,

1024-542: A genetic study of individuals from the around the Tian Shan mountains of central Asia dating from the late second century CE, Damgaard et al. 2018 found that these individuals represented a population of mixed East Asian and West Eurasian origin. They argued that this population descended from Xiongnu who expanded westward and mixed with Iranian Sakas . This population in the Tian Shan mountains may be connected to

1152-725: A group of Huns and Alans fighting against Radagaisus in defense of Italy. Uldin was also known for defeating Gothic rebels who troubled the East Romans around the Danube and for beheading the Goth Gainas around 400–401. The East Romans began to feel the pressure from Uldin's Huns again in 408. Uldin crossed the Danube and pillaged Thrace. The East Romans tried to buy off Uldin, but his sum was too high so they instead bought off Uldin's subordinates. This resulted in many desertions from Uldin's group of Huns. Uldin himself escaped back across

1280-729: A horse takes him on his back". They appear to have spent so much time riding that they walked clumsily, something observed in other nomadic groups. Roman sources characterize the Hunnic horses as ugly. It is not possible to determine the exact breed of horse the Huns used, despite a relatively good description by the Roman writer Vegetius . Sinor believes that it was likely a breed of Mongolian pony. However, horse remains are absent from all identified Hun burials. Based on anthropological descriptions and archaeological finds of other nomadic horses, Maenchen-Helfen believes that they rode mostly geldings . Apart from horses, ancient sources indicate that

1408-717: A legend developed based on medieval chronicles that the Hungarians , and the Székely ethnic group in particular, are descended from the Huns. However, mainstream scholarship dismisses a close connection between the Hungarians and Huns. Modern culture generally associates the Huns with extreme cruelty and barbarism. The origins of the Huns and their links to other steppe people remain uncertain: scholars generally agree that they originated in Central Asia but disagree on

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1536-708: A million years. Ludwik Hirszfeld was a Polish microbiologist and serologist who was the President of the Blood Group Section of the Second International Congress of Blood Transfusion. He founded blood group inheritance with Erich von Dungern in 1910, and contributed to it greatly throughout his life. He studied ABO blood groups . In one of his studies in 1919, Hirszfeld documented the ABO blood groups and hair color of people at

1664-537: A people who practiced artificial cranial deformation as evidence of Hunnic agriculture. Kim similarly argues that all steppe empires have possessed both pastoralist and sedentary populations, classifying the Huns as "agro-pastoralist". As a nomadic people, the Huns spent a great deal of time riding horses: Ammianus claimed that the Huns "are almost glued to their horses", Zosimus claimed that they "live and sleep on their horses", and Sidonius claimed that "[s]carce had an infant learnt to stand without his mother's aid when

1792-442: A plural suffix "supposedly meaning 'people'", qun (force), and hün (ferocious). Maenchen-Helfen dismisses all of these Turkic etymologies as "mere guesses" and proposes an Iranian etymology, from a word akin to Avestan hūnarā (skill), hūnaravant- (skillful). He suggests that it may originally have designated a rank rather than an ethnicity. Robert Werner has advanced an etymology from Tocharian ku (dog), suggesting—as

1920-537: A ranking hierarchy, much like Germanic societies. Denis Sinor similarly notes that, with the exception of the historically uncertain Balamber , no Hun leaders are named in the sources until Uldin , indicating their relative unimportance. Thompson argues that permanent kingship only developed with the Huns' invasion of Europe and the near-constant warfare that followed. Regarding the organization of Hunnic rule under Attila, Peter Golden comments "it can hardly be called

2048-562: A sample, even when it is highly fragmented and of low concentration. It involves attaching a generic sequence to every single strand that generic primers can bond to, and thus all of the DNA present is amplified. This is generally more costly and time intensive than PCR but due to the difficulties involved in ancient DNA amplification it is cheaper and more efficient. One method of massive parallel sequencing , developed by Margulies et al., employs bead-based emulsion PCR and pyrosequencing , and

2176-608: A similarity in 2.18 and 1.62 bases per 10,000 respectively, suggesting Vi-80 sample was from a male individual. Other similar studies include finding of a mutation associated with dwarfism in Arabidopsis in ancient Nubian cotton , and investigation on the bitter taste perception locus in Neanderthals. Modern humans are thought to have evolved in Africa at least 200 kya (thousand years ago), with some evidence suggesting

2304-607: A single migratory event between 60 and 70 kya. Genetic evidence shows that occupation of the Near East and Europe happened no earlier than 50 kya. Studying haplogroup U has shown separate dispersals from the Near East both into Europe and into North Africa. Much of the work done in archaeogenetics focuses on the Neolithic transition in Europe. Cavalli-Svorza's analysis of genetic-geographic patterns led him to conclude that there

2432-402: A specimen is collected from an archaeological site, DNA can be extracted through a series of processes. One of the more common methods utilizes silica and takes advantage of polymerase chain reactions in order to collect ancient DNA from bone samples. There are several challenges that add to the difficulty when attempting to extract ancient DNA from fossils and prepare it for analysis. DNA

2560-495: A state, much less an empire". Golden speaks instead of a "Hunnic confederacy". Kim, however, argues that the Huns were far more organized and centralized, with some basis in organization of the Xiongnu state. Walter Pohl notes the correspondences of Hunnic government to those of other steppe empires, but nevertheless argues that the Huns do not appear to have been a unified group when they arrived in Europe. Ammianus wrote that

2688-493: A trading network for the production and consumption of readily available resources. Archaeogenetics has been used to study the domestication of animals. By analyzing genetic diversity in domesticated animal populations researchers can search for genetic markers in DNA to give valuable insight about possible traits of progenitor species. These traits are then used to help distinguish archaeological remains between wild and domesticated specimens. The genetic studies can also lead to

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2816-492: A variety of different fossils, they may be less effective in fossils that are not fresh (e.g. treated fossils for museums ). Also, contamination poses a risk for all DNA replication in general, and this method may result in misleading results if applied to contaminated material. Polymerase chain reaction is a process that can amplify segments of DNA and is often used on extracted ancient DNA. It has three main steps: denaturation , annealing , and extension. Denaturation splits

2944-524: A variety of genetic signatures. Maróti et al. 2022 showed that the genomes of nine Hun-era individuals who lived in the basin varied from European to Northeast Asian connections, with those individuals showing associations with Northeast Asia being most similar to groups found in Mongolia such as the Xiongnu and the Xianbei . An analysis of Hun-era genomes by Gnecchi-Ruscone et al. 2022 likewise found

3072-529: A wide range of genetic variability, with two individuals showing a connection to ancient Northeast Asians and others showing European ancestry. The history of the Huns in the fourth century is not very clear, and the Huns left no sources themselves. The Romans became aware of the Huns when the latter's invasion of the Pontic steppes forced thousands of Goths to move to the Lower Danube to seek refuge in

3200-399: A “pioneer colonization” model of European occupation, with incorporation of foraging populations into arriving Neolithic populations. Furthermore, analysis of ancient DNA, not just extant DNA, is shedding light on some issues. For instance, comparison of neolithic and mesolithic DNA has indicated that the development of dairying preceded widespread lactose tolerance. South Asia has served as

3328-471: Is a recent admixture of some Negrito groups with their local populations. Archaeogenetics has been used to better understand the populating of the Americas from Asia. Native American mtDNA haplogroups have been estimated to be between 15 and 20 kya, although there is some variation in these estimates. Genetic data has been used to propose various theories regarding how the Americas were colonized. Although

3456-403: Is also difficult due to the lack of repeatability caused by the uniqueness of specimens. Silica-based DNA extraction is a method used as a purification step to extract DNA from archaeological bone artifacts and yield DNA that can be amplified using polymerase chain reaction (PCR) techniques. This process works by using silica as a means to bind DNA and separate it from other components of

3584-411: Is continuously being split up. While the organism is alive these splits are repaired; however, once an organism has died, the DNA will begin to deteriorate without repair. This results in samples having strands of DNA measuring around 100 base pairs in length. Contamination is another significant challenge at multiple steps throughout the process. Often other DNA, such as bacterial DNA, will be present in

3712-583: Is different from Wikidata All article disambiguation pages All disambiguation pages Huns The Huns were a nomadic people who lived in Central Asia , the Caucasus , and Eastern Europe between the 4th and 6th centuries AD. According to European tradition, they were first reported living east of the Volga River, in an area that was part of Scythia at the time. By 370 AD,

3840-421: Is difficult because the bone fossilisation degrades and DNA is chemically modified, usually by bacteria and fungi in the soil. The best time to extract DNA from a fossil is when it is freshly out of the ground as it contains six times the DNA when compared to stored bones. The temperature of extraction site also affects the amount of obtainable DNA, evident by a decrease in success rate for DNA amplification if

3968-555: Is especially helpful when the morphology of the fossil is ambiguous. Apart from that, species identification can also be done by finding specific genetic markers in an aDNA sequence. For example, the American indigenous population is characterized by specific mitochondrial RFLPs and deletions defined by Wallace et al. aDNA comparison study can also reveal the evolutionary relationship between two species. The number of base differences between DNA of an ancient species and that of

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4096-556: Is genetic evidence that Chad-speaking descendants of Nilo-Saharan speakers migrated from Sudan to Lake Chad about 8 kya. Genetic evidence has also indicated that non-African populations made significant contributions to the African gene pool. For example, the Saharan African Beja people have high levels of Middle-Eastern as well as East African Cushitic DNA. Analysis of mtDNA shows that modern humans occupied Eurasia in

4224-574: Is likewise disputed, but probably in 406/407 and 431/433 respectively. Otherwise, the Huns made no attempt to conquer or settle on Roman territory. Following Attila's death, the Huns were driven out of Pannonia and some appear to have returned to the Pontic Steppe, while one group settled in Dobruja . One of the only written sources for the size of Attila's domain is given by the Roman historian Priscus. Priscus refers to Attila ruling as far as

4352-592: Is native to India. Analysis of mtDNA and NRY (non-recombining region of Y chromosome) sequences have indicated that the first major dispersal out of Africa went through Saudi Arabia and the Indian coast 50–100 kya, and a second major dispersal occurred 15–50 kya north of the Himalayas. Much work has been done to discover the extent of north-to-south and south-to-north migrations within Eastern Asia. Comparing

4480-468: Is that it requires overlapping primer pairs for ancient DNA due to the short sequences. There can also be “jumping PCR” which causes recombination during the PCR process which can make analyzing the DNA more difficult in inhomogeneous samples. DNA extracted from fossil remains is primarily sequenced using Massive parallel sequencing , which allows simultaneous amplification and sequencing of all DNA segments in

4608-415: Is therefore futile to speculate about identity or blood relationships between H(s)iung-nu, Hephthalites, and Attila's Huns, for instance. All we can safely say is that the name Huns , in late antiquity, described prestigious ruling groups of steppe warriors. Today, there is "no general consensus" and "scholarship is divided" on the issue of a Hun-Xiongnu connection. Recent supporters of a connection between

4736-728: Is through DNA hybridization . Single-stranded DNA segments of both species are allowed to form complementary pair bonding with each other. More closely related species have a more similar genetic makeup, and thus a stronger hybridization signal. Scholz et al. conducted southern blot hybridization on Neanderthal aDNA (extracted from fossil remain W-NW and Krapina). The results showed weak ancient human-Neanderthal hybridization and strong ancient human-modern human hybridization. The human-chimpanzee and neanderthal-chimpanzee hybridization are of similarly weak strength. This suggests that humans and neanderthals are not as closely related as two individuals of

4864-577: Is used over mitochondrial and chloroplast DNA because of its faster mutation rate as well as its intraspecific variation due to a higher consistency of polymorphism genetic markers . Findings in crop 'domestication genes' (traits that were specifically selected for or against) include Through the study of archaeogenetics in plant domestication, signs of the first global economy can also be uncovered. The geographical distribution of new crops highly selected in one region found in another where it would have not originally been introduced serve as evidence of

4992-678: The Eastern Roman Empire . In 451, they invaded the Western Roman province of Gaul , where they fought a combined army of Romans and Visigoths at the Battle of the Catalaunian Fields , and in 452, they invaded Italy. After the death of Attila in 453, the Huns ceased to be a major threat to Rome and lost much of their empire following the Battle of Nedao (c. 454). Descendants of the Huns, or successors with similar names, are recorded by neighboring populations to

5120-626: The Kerch Strait into Crimea . Discovering the land fertile, they then attacked the Goths. Jordanes ' Getica relates that the Goths held the Huns to be offspring of "unclean spirits" and Gothic witches ( Getica 24:121). Since Joseph de Guignes in the 18th century , modern historians have associated the Huns who appeared on the borders of Europe in the 4th century AD with the Xiongnu who had invaded numerous Central Plain polities from

5248-582: The Mongolian Plateau between the 3rd century BC and the 2nd century AD . After the devastating defeat by the Han dynasty , the northern branch of the Xiongnu retreated north-westward; their descendants may have migrated through the Eurasian Steppe and consequently they may have some degree of cultural and genetic continuity with the Huns. Scholars also discussed the relationship between

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5376-606: The North Caucasian Huns , were genuine Huns. The rulers of various post-Hunnic steppe peoples are known to have claimed descent from Attila in order to legitimize their right to the power, and various steppe peoples were also called "Huns" by Western and Byzantine sources from the fourth century onward. The Huns have traditionally been described as pastoral nomads , living off of herding and moving from pasture to pasture to graze their animals. Hyun Jin Kim, however, holds

5504-617: The Peace of Anatolius with the two Hun kings. Bleda died in 445, and Attila became the sole ruler of the Huns. In 447, Attila invaded the Balkans and Thrace. The war came to an end in 449 with an agreement in which the East Romans agreed to pay Attila an annual tribute of 2100 pounds of gold. Throughout their raids on the Eastern Roman Empire , the Huns had maintained good relations with the Western Empire. However, Honoria , sister of

5632-736: The Sabirs . In 463, the Saragurs defeated the Akatziri, or Akatir Huns, and asserted dominance in the Pontic region. The western Huns under Dengizich experienced difficulties in 461 when they were defeated by Valamir in a war against the Sadages , a people allied with the Huns. His campaigning was also met with dissatisfaction from Ernak , ruler of the Akatziri Huns, who wanted to focus on

5760-671: The Sogdian merchants under their rule, who were involved in the trade along the Silk Road to China. Atwood notes that Jordanes describes how the Crimean city of Cherson , "where the avaricious traders bring in the goods of Asia", was under the control of the Akatziri Huns in the sixth century. Hunnic governmental structure has long been debated. Peter Heather argues that the Huns were a disorganized confederation in which leaders acted completely independently and that eventually established

5888-507: The Yakut or Tungus . He notes that archaeological finds of presumed Huns suggest that they were a racially mixed group containing only some individuals with East Asian features. Kim similarly cautions against seeing the Huns as a homogenous racial group, while still arguing that they were "partially or predominantly of Mongoloid extraction (at least initially)." Some archaeologists have argued that archaeological finds have failed to prove that

6016-430: The 1950s. During the 1940s, Boyd and Karl O. Renkonen independently discovered that lectins react differently to various blood types, after finding that the crude extracts of the lima bean and tufted vetch agglutinated the red blood cells from blood type A but not blood types B or O. This ultimately led to the disclosure of thousands of plants that contained these proteins. In order to examine racial differences and

6144-528: The Altai Mountains of Siberia between 17.2 and 10.1 kya, after the LGM. Analysis of both mtDNA and Y-chromosome DNA reveals evidence of “small, founding populations.” Studying haplogroups has led some scientists to conclude that a southern migration into the Americas from one small population was impossible, although separate analysis has found that such a model is feasible if such a migration happened along

6272-491: The Battle of Chalons (451), "the vast majority" of Attila's entourage and troops appears to have been of European origin, while Attila himself seems to have had East Asian features. Genetic data is difficult to apply to steppe nomad societies, because they frequently migrated, intermixed, and were assimilated into each other. Nevertheless, genetics can supply information on migrations from East Asia to Europe and vice versa. In

6400-646: The Carpathian Mountains, have been attributed to the time of Attila and associated with the nomadic milieu of the Huns. While scholars have speculated about direct Hunnic control and settlement here, it is entirely unclear what kind of relationship the population of these regions had to the Huns. The Huns ruled over numerous other groups, including Goths , Gepids , Sarmatians , Heruli , Alans , Rugii , Suevi , and Sciri , alongside other groups where they occasionally asserted control. Peter Heather suggests that some of these groups were resettled along

6528-597: The Chinese called the Xiongnu dogs—that the dog was the totem animal of the Hunnic tribe. He also compares the name Massagetae , noting that the element saka in that name means dog. Others such as Harold Bailey, S. Parlato, and Jamsheed Choksy have argued that the name derives from an Iranian word akin to Avestan Ẋyaona , and was a generalized term meaning "hostiles, opponents". Christopher Atwood dismisses this possibility on phonological and chronological grounds. While not arriving at an etymology per se , Atwood derives

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6656-465: The DNA into two single strands at high temperatures. Annealing involves attaching primer strands of DNA to the single strands that allow Taq polymerase to attach to the DNA. Extension occurs when Taq polymerase is added to the sample and matches base pairs to turn the two single strands into two complete double strands. This process is repeated many times, and is usually repeated a higher number of times when used with ancient DNA . Some issues with PCR

6784-418: The Danube by the Huns. Subject peoples of the Huns were led by their own kings. Those recognized as ethnic Huns appear to have had more rights and status, as evidenced by the account of Priscus. One of the principal sources of information on Hunnic warfare is Ammianus Marcellinus , who includes an extended description of the Huns' methods of war: They also sometimes fight when provoked, and then they enter

6912-523: The Danube, after which he is not mentioned again in history. Hunnish mercenaries are mentioned on several occasions being employed by the East and West Romans, as well as the Goths, during the late 4th and 5th century. In 433 some parts of Pannonia were ceded to them by Flavius Aetius , the magister militum of the Western Roman Empire . From 434 the brothers Attila and Bleda ruled

7040-533: The European Huns by individual burials that contains objects stylistically related to those used by the European Huns, although this could be a sign of the exchange of goods and the connections between elites rather than a sign of migration. As of 2023, there is little genetic data from the Carpathian basin in the Hunnic period (5th century), and the population living there during the Hunnic period shows

7168-620: The European Huns did not strike their own coins. The extent of Hunnish control in Barbarian Europe is poorly understood, as it is not much covered in Roman sources. It is generally assumed that they established an empire that stretched as far West as the Rhine and perhaps as far north as the Baltic, though it is difficult to establish its boundaries with certainty. Some scholars, such as Otto Maenchen-Helfen and Peter Golden, believe that

7296-541: The Huns and Romans, and the Huns overcame a weak Roman army to raze the cities of Margus, Singidunum and Viminacium . Although a truce was concluded in 441, two years later Constantinople again failed to deliver the tribute and war resumed. In the following campaign, Hun armies approached Constantinople and sacked several cities before defeating the Romans at the Battle of Chersonesus . The Eastern Roman Emperor Theodosius II gave in to Hun demands and in autumn 443 signed

7424-434: The Huns and Xiongnu include Hyun Jin Kim and Etienne de la Vaissière . De la Vaissière argues that ancient Chinese and Indian sources used Xiongnu and Hun to translate each other, and that the various "Iranian Huns" were similarly identified with the Xiongnu. Kim believes that the term Hun was "not primarily an ethnic group, but a political category" and argues for a fundamental political and cultural continuity between

7552-515: The Huns are known to have practiced a form of nomadic pastoralism . As their contact with the Roman world grew, their economy became increasingly tied with Rome through tribute, raiding, and trade. They do not seem to have had a unified government when they entered Europe but rather to have developed a unified tribal leadership in the course of their wars with the Romans. The Huns ruled over a variety of peoples who spoke numerous languages, and some maintained their own rulers. Their main military technique

7680-526: The Huns had any "Mongoloid" features at all, and some scholars have argued that the Huns were predominantly " Caucasian " in appearance. Other archaeologists have argued that "Mongoloid" features are found primarily among members of the Hunnic aristocracy, which, however, also included Germanic leaders who were integrated into the Hun polity. Kim argues that the composition of the Huns became progressively more "Caucasian" during their time in Europe; he notes that by

7808-642: The Huns had arrived on the Volga, causing the westwards movement of Goths and Alans . By 430, they had established a vast, but short-lived, empire on the Danubian frontier of the Roman empire in Europe. Either under Hunnic hegemony , or fleeing from it, several central and eastern European peoples established kingdoms in the region, including not only Goths and Alans, but also Vandals , Gepids , Heruli , Suebians and Rugians . The Huns, especially under their King Attila , made frequent and devastating raids into

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7936-565: The Huns had small eyes and flat noses. The Roman writer Priscus gives the following eyewitness description of Attila: "Short of stature, with a broad chest and a large head; his eyes were small, his beard thin and sprinkled with grey; and he had a flat nose and tanned skin, showing evidence of his origin." Many scholars take these to be unflattering depictions of East Asian (obsolete " Mongoloid ") racial characteristics. Maenchen-Helfen argues that, while many Huns had East Asian racial characteristics, they were unlikely to have looked as Asiatic as

8064-423: The Huns having several kings, with one being the "first of the kings". Ammianus also mentions that the Huns made their decisions in a general council ( omnes in commune ) while seated on horseback. He makes no mention of the Huns being organized into tribes, but Priscus and other writers do, naming some of them. The first Hunnic ruler known by name is Uldin . Thompson takes Uldin's sudden disappearance after he

8192-443: The Huns of his day had no kings, but rather that each group of Huns instead had a group of leading men ( primates ) for times of war . E.A. Thompson supposes that, even in war, the leading men had little actual power. He further argues that they most likely did not acquire their position purely hereditarily. Heather, however, argues that Ammianus merely meant that the Huns didn't have a single ruler; he notes that Olympiodorus mentions

8320-575: The Huns together. Attila and Bleda were as ambitious as their uncle Rugila . In 435 they forced the Eastern Roman Empire to sign the Treaty of Margus , giving the Huns trade rights and an annual tribute from the Romans. When the Romans breached the treaty in 440, Attila and Bleda attacked Castra Constantias, a Roman fortress and marketplace on the banks of the Danube . War broke out between

8448-420: The Huns traded their horses for what he considered to have been "a very considerable source of income in gold", he is otherwise skeptical of Thompson's argument. He notes that the Romans strictly regulated trade with the barbarians and that, according to Priscus, trade only occurred at a fair once a year. While he notes that smuggling also likely occurred, he argues that "the volume of both legal and illegal trade

8576-417: The Huns used wagons for transportation. Maenchen-Helfen suggests that these wagons were mainly utilized to carry their tents, loot, as well as the elderly, women, and children. The Huns received a large amount of gold from the Romans, either in exchange for fighting for them as mercenaries or as tribute. Raiding and looting also furnished the Huns with gold and other valuables. Denis Sinor has argued that at

8704-409: The Huns. They are believed to have used bronze cauldrons and to have performed artificial cranial deformation . No description exists of the Hunnic religion of the time of Attila, but practices such as divination are attested, and the existence of shamans is likely. It is also known that the Huns had a language of their own ; however, only three words and personal names attest to it. Economically,

8832-514: The Macedonian front, leading to his discovery that the hair color and blood type had no correlation. In addition to that he observed that there was a decrease of blood group A from western Europe to India and the opposite for blood group B. He hypothesized that the east-to-west blood group ratio stemmed from two blood groups consisting of mainly A or B mutating from blood group O, and mixing through migration or intermingling. A majority of his work

8960-539: The Neolithic. Most mtDNA's were “already established” among existing Mesolithic and Paleolithic groups. Most “control-region lineages” of modern European mtDNA are traced to a founder event of reoccupying northern Europe towards the end of the Last Glacial Maximum (LGM). One study of extant European mtDNA's suggest this reoccupation occurred after the end of the LGM, although another suggests it occurred before. Analysis of haplogroups V, H, and U5 support

9088-584: The Pontic Steppe north of the Black Sea. They had conquered the Hungarian Plain in stages. The precise date that they conquered the north bank of the Danube is unclear. Maenchen-Helfen argued that they may have already taken control of it in the 370s. The dates when they gained control of the Roman territory south of the Middle Danube, Pannonia Valeria and the other provinces of Pannonia ,

9216-762: The Roman Empire in 376. The Huns conquered the Alans , most of the Greuthungi or Eastern Goths, and then most of the Thervingi or Western Goths, with many fleeing into the Roman Empire . In 395 the Huns began their first large-scale attack on the Eastern Roman Empire . Huns attacked in Thrace, overran Armenia , and pillaged Cappadocia . They entered parts of Syria , threatened Antioch , and passed through

9344-698: The Western Roman Emperor Valentinian III , sent Attila a ring and requested his help to escape her betrothal to a senator. Attila claimed her as his bride and half the Western Roman Empire as dowry. Additionally, a dispute arose about the rightful heir to a king of the Salian Franks . In 451, Attila's forces entered Gaul . Once in Gaul, the Huns first attacked Metz , then their armies continued westward, passing both Paris and Troyes to lay siege to Orléans . Flavius Aetius

9472-481: The Xiongnu and the European Huns, as well as between the Xiongnu and the "Iranian Huns". The name Hun is attested in classical European sources as Greek Οὖννοι ( Ounnoi ) and Latin Hunni or Chuni . John Malalas records their name as Οὖννα ( Ounna ). Another possible Greek variant may be Χοὖνοι ( Khounoi ), although this group's identification with the Huns is disputed. Classical sources also frequently use

9600-575: The Xiongnu, the Huns, and a number of people in central Asia who were also known as or came to be identified with the name "Hun" or " Iranian Huns ". The most prominent of these were Chionites , the Kidarites , and the Hephthalites . Otto J. Maenchen-Helfen was the first to challenge the traditional approach, based primarily on the study of written sources, and to emphasize the importance of archaeological research. Since Maenchen-Helfen's work,

9728-463: The Y-chromosome lineages indicate that primarily males partook in these migrations. The discovery of two subbranches U2i and U2e of the U mtDNA lineage, which arose in Central Asia has “modulated” views of a large migration from Central Asia into India, as the two branches diverged 50 kya. Furthermore, U2e is found in large percentages in Europe but not India, and vice versa for U2i, implying U2i

9856-532: The assistance of the settled agricultural population at the edge of the steppe they could not have survived". He argues that the Huns were forced to supplement their diet by hunting and gathering. Maenchen-Helfen, however, notes that archaeological finds indicate that various steppe nomad populations did grow grain; in particular, he identifies a find at Kunya Uaz in Khwarezm on the Ob River of agriculture among

9984-485: The battle drawn up in wedge-shaped masses, while their medley of voices makes a savage noise. And as they are lightly equipped for swift motion, and unexpected in action, they purposely divide suddenly into scattered bands and attack, rushing about in disorder here and there, dealing terrific slaughter; and because of their extraordinary rapidity of movement they are never seen to attack a rampart or pillage an enemy's camp. And on this account you would not hesitate to call them

10112-414: The chemical composition of bone and soil, and hydrology . There are three perseveration diagenetic phases. The first phase is bacterial putrefaction , which is estimated to cause a 15-fold degradation of DNA. Phase 2 is when bone chemically degrades, mostly by depurination . The third diagenetic phase occurs after the fossil is excavated and stored, in which bone DNA degradation occurs most rapidly. Once

10240-455: The coasts. Finally, archaeogenetics has been used to study the occupation of Australia and New Guinea. The Indigenous people of Australia and New Guinea are phenotypically very similar, but mtDNA has shown that this is due to convergence from living in similar conditions. Non-coding regions of mt-DNA have shown “no similarities” between the aboriginal populations of Australia and New Guinea. Furthermore, no major NRY lineages are shared between

10368-601: The continent found 14 genetic and linguistic “clusters,” suggesting an ancient geographic structure to African populations. In general, genotypic and phenotypic analysis have shown “large and subdivided throughout much of their evolutionary history.” Genetic analysis has supported archaeological hypotheses of a large-scale migrations of Bantu speakers into Southern Africa approximately 5 kya. Microsatellite DNA, single nucleotide polymorphisms (SNPs), and insertion/deletion polymorphisms (INDELS) have shown that Nilo-Saharan speaking populations originate from Sudan. Furthermore, there

10496-415: The development of domestication of dogs. Genetic studies have shown that all dogs are descendants from the gray wolf, however, it is currently unknown when, where, and how many times dogs were domesticated. Some genetic studies have indicated multiple domestications while others have not. Archaeological findings help better understand this complicated past by providing solid evidence about the progression of

10624-542: The distribution and migration patterns of various racial groups, Boyd systematically collected and classified blood samples from around the world, leading to his discovery that blood groups are not influenced by the environment, and are inherited. In his book Genetics and the Races of Man (1950), Boyd categorized the world population into 13 distinct races, based on their different blood type profiles and his idea that human races are populations with differing alleles . One of

10752-490: The existing data on blood group gene frequencies, and largely contributing to the genetic map of the world through his investigation of blood groups in many populations. Mourant discovered the new blood group antigens of the Lewis , Henshaw , Kell , and Rhesus systems, and analyzed the association of blood groups and various other diseases. He also focused on the biological significance of polymorphisms . His work provided

10880-513: The extant of Attila's empire has been exaggerated and he probably only controlled Pannonia and some adjacent areas. In the 390s, the majority of the Huns were probably based around the Volga and Don on the Pontic Steppe. But by the 420s, the Huns were based on Great Hungarian Plain , the only large grassland near the Roman empire capable of supporting large numbers of horses. However, Aleksander Paroń believes that they likely continued to control

11008-416: The fossil is found in warmer regions. A drastic change of a fossil's environment also affects DNA preservation. Since excavation causes an abrupt change in the fossil's environment, it may lead to physiochemical change in the DNA molecule. Moreover, DNA preservation is also affected by other factors such as the treatment of the unearthed fossil like (e.g. washing, brushing and sun drying), pH , irradiation ,

11136-433: The fossil process that inhibit PCR amplification. However, silica itself is also a strong PCR inhibitor , so careful measures must be taken to ensure that silica is removed from the DNA after extraction. The general process for extracting DNA using the silica-based method is outlined by the following: One of the main advantages of silica-based DNA extraction is that it is relatively quick and efficient, requiring only

11264-422: The foundation for archaeogenetics because it facilitated the separation of genetic evidence for biological relationships between people. This genetic evidence was previously used for that purpose. It also provided material that could be used to appraise the theories of population genetics . William Boyd was an American immunochemist and biochemist who became famous for his research on the genetics of race in

11392-517: The genetic diversity of northeastern groups with southeastern groups has allowed archaeologists to conclude many of the northeast Asian groups came from the southeast. The Pan-Asian SNP (single nucleotide polymorphism) study found “a strong and highly significant correlation between haplotype diversity and latitude,” which, when coupled with demographic analysis, supports the case for a primarily south-to-north occupation of East Asia. Archaeogenetics has also been used to study hunter-gatherer populations in

11520-614: The help of his bucellarii , then attacked the quarreling Goths and Huns, defeating them. In 469, Dengizich was defeated and killed in Thrace. After Dengizich's death, the Huns seem to have been absorbed by other ethnic groups such as the Bulgars . Kim, however, argues that the Huns continued under Ernak, becoming the Kutrigur and Utigur Hunno- Bulgars . This conclusion is still subject to some controversy. Some scholars also argue that another group identified in ancient sources as Huns,

11648-455: The high civilian officers Gennadius Avienus and Trigetius, as well as Pope Leo I , who met Attila at Mincio in the vicinity of Mantua , and obtained from him the promise that he would withdraw from Italy and negotiate peace with the emperor. The new Eastern Roman Emperor Marcian then halted tribute payments, resulting in Attila planning to attack Constantinople. However, in 453 Attila died of

11776-427: The hinge upon which the entire administration of the Hun empire turned": he argues for their existence in the government of Uldin, and that each had command over detachments of the Hunnic army and ruled over specific portions of the Hunnic empire, where they were responsible also for collecting tribute and provisions. Maenchen-Helfen, however, argues that the word logades denotes simply prominent individuals and not

11904-497: The identification of ancestors for domesticated animals. The information gained from genetics studies on current populations helps guide the Archaeologist's search for documenting these ancestors. Archaeogenetics has been used to trace the domestication of pigs throughout the old world. These studies also reveal evidence about the details of early farmers. Methods of Archaeogenetics have also been used to further understand

12032-450: The identification of the Xiongnu as the Huns' ancestors has become controversial. Additionally, several scholars have questioned the identification of the "Iranian Huns" with the European Huns. Walter Pohl cautions that none of the great confederations of steppe warriors was ethnically homogenous, and the same name was used by different groups for reasons of prestige, or by outsiders to describe their lifestyle or geographic origin. [...] It

12160-462: The incoming Oghur speaking peoples. Dengizich attacked the Romans in 467, without the assistance of Ernak. He was surrounded by the Romans and besieged, and came to an agreement that he would surrender if his people were given land for their herds and his starving forces given food. During the negotiations, a Hun in service of the Romans named Chelchel persuaded the enemy Goths to attack their Hun overlords. The Romans, under their General Aspar and with

12288-619: The islands in the "ocean" ( Ὠκεανός ), but it is unclear if this meant the Baltic Sea or the world-encircling Ocean that the Greeks and Romans believed in. In either case, the description of Attila ruling as far as the islands in the "ocean" may be hyperbole. Archaeology is often used to argue for an area having been under Hunnic control; however, nomadic peoples often control territories beyond their immediate settlement. A large number of major finds from Silesia and Lesser Poland , north of

12416-443: The major early corridor for geographical dispersal of modern humans from out-of-Africa. Based on studies of mtDNA line M, some have suggested that the first occupants of India were Austro-Asiatic speakers who entered about 45–60 kya. The Indian gene pool has contributions from earliest settlers, as well as West Asian and Central Asian populations from migrations no earlier than 8 kya. The lack of variation in mtDNA lineages compared to

12544-507: The meat of these animals, with Maenchen-Helfen arguing, on the basis of what is known of other steppe nomads, that they likely mostly ate mutton, along with sheep's cheese and milk. They also "certainly" ate horse meat, drank mare's milk, and likely made cheese and kumis . In times of starvation, they may have boiled their horses' blood for food. Ancient sources uniformly deny that the Huns practiced any sort of agriculture. Thompson, taking these accounts at their word, argues that "[w]ithout

12672-545: The most abundant information sources regarding inheritable traits linked to race remains the study of blood groups. Fossil retrieval starts with selecting an excavation site . Potential excavation sites are usually identified with the mineralogy of the location and visual detection of bones in the area. However, there are more ways to discover excavation zones using technology such as field portable x-ray fluorescence and Dense Stereo Reconstruction. Tools used include knives , brushes , and pointed trowels which assist in

12800-497: The most terrible of all warriors, because they fight from a distance with missiles having sharp bone, instead of their usual points, joined to the shafts with wonderful skill; then they gallop over the intervening spaces and fight hand to hand with swords, regardless of their own lives; and while the enemy are guarding against wounds from the sabre-thrusts, they throw strips of cloth plaited into nooses over their opponents and so entangle them that they fetter their limbs and take from them

12928-463: The most widely held theory suggests “three waves” of migration after the LGM through the Bering Strait, genetic data have given rise to alternative hypotheses. For example, one hypothesis proposes a migration from Siberia to South America 20–15 kya and a second migration that occurred after glacial recession. Y-chromosome data has led some to hold that there was a single migration starting from

13056-551: The name from the Ongi River in Mongolia, which was pronounced the same as, or similarly to, the name Xiongnu, and suggests that it was originally a dynastic name rather than an ethnic name. Most of the ancient descriptions of the Huns stress their strange appearance from a Roman perspective. These descriptions typically caricature the Huns as monsters. Jordanes stresses that the Huns were short of stature, had tanned skin and round and shapeless heads. Various writers mention that

13184-469: The names of older and unrelated steppe nomads instead of the name Hun , calling them Massagetae , Scythians , and Cimmerians , among other names. The etymology of Hun is unclear. Various proposed etymologies generally assume at least that the names of the various Eurasian groups known as Huns are related. There have been a number of proposed Turkic etymologies, deriving the name variously from Turkic ön , öna (to grow), qun (glutton), kün , gün ,

13312-428: The original sample. To avoid contamination it is necessary to take many precautions such as separate ventilation systems and workspaces for ancient DNA extraction work. The best samples to use are fresh fossils as uncareful washing can lead to mold growth. DNA coming from fossils also occasionally contains a compound that inhibits DNA replication. Coming to a consensus on which methods are best at mitigating challenges

13440-636: The pastures may vary, the winter quarters always remained the same. This is, in fact, what Jordanes writes of the Hunnic Altziagiri tribe: they pastured near Cherson on the Crimea and then wintered further north, with Maenchen-Helfen holding the Syvash as a likely location. Ancient sources mention that the Huns' herds consisted of various animals, including cattle, horses, and goats; sheep, though unmentioned in ancient sources, "are more essential to

13568-518: The power of riding or walking. Archaeogenetics Archaeogenetics receives its name from the Greek word arkhaios , meaning "ancient", and the term genetics , meaning "the study of heredity". The term archaeogenetics was conceived by archaeologist Colin Renfrew . In February 2021, scientists reported the oldest DNA ever sequenced was successfully retrieved from a mammoth dating back over

13696-565: The province of Euphratesia . At the same time, the Huns invaded the Sasanian Empire . This invasion was initially successful, coming close to the capital of the empire at Ctesiphon ; however, they were defeated badly during the Persian counterattack. During their brief diversion from the Eastern Roman Empire, the Huns may have threatened tribes further west. Uldin , the first Hun identified by name in contemporary sources, headed

13824-790: The region, such as the Ainu from Japan and Negrito groups in the Philippines. For example, the Pan-Asian SNP study found that Negrito populations in Malaysia and the Negrito populations in the Philippines were more closely related to non-Negrito local populations than to each other, suggesting Negrito and non-Negrito populations are linked by one entry event into East Asia; although other Negrito groups do share affinities, including with Indigenous Australians . A possible explanation of this

13952-608: The removal of fossils from the earth. To avoid contaminating the ancient DNA , specimens are handled with gloves and stored in -20 °C immediately after being unearthed. Ensuring that the fossil sample is analyzed in a lab that has not been used for other DNA analysis could prevent contamination as well. Bones are milled to a powder and treated with a solution before the polymerase chain reaction (PCR) process. Samples for DNA amplification may not necessarily be fossil bones. Preserved skin, salt-preserved or air-dried, can also be used in certain situations. DNA preservation

14080-505: The same species are, but they are more related to each other than to chimpanzees. There have also been some attempts to decipher aDNA to provide valuable phenotypic information of ancient species. This is always done by mapping aDNA sequence onto the karyotype of a well-studied closely related species, which share a lot of similar phenotypic traits. For example, Green et al. compared the aDNA sequence from Neanderthal Vi-80 fossil with modern human X and Y chromosome sequence, and they found

14208-409: The same term [REDACTED] This disambiguation page lists articles associated with the title Hunnic . If an internal link led you here, you may wish to change the link to point directly to the intended article. Retrieved from " https://en.wikipedia.org/w/index.php?title=Hunnic&oldid=1225961223 " Category : Disambiguation pages Hidden categories: Short description

14336-466: The same year under Valamir , allegedly defeating the Huns in a separate engagement. However, this did not result in the complete collapse of Hunnic power in the Carpathian region, but did result in the loss of many of their Germanic vassals. At the same time, the Huns were also dealing with the arrival of more Oghur Turkic-speaking peoples from the East, including the Oghurs , Saragurs , Onogurs , and

14464-550: The slaves would have been used to manage the Huns' herds of cattle, sheep, and goats. Priscus attests that slaves were used as domestic servants, but also that educated slaves were used by the Huns in positions of administration or even architects. Some slaves were even used as warriors. The Huns also traded with the Romans. E. A. Thompson argued that this trade was very large scale, with the Huns trading horses, furs, meat, and slaves for Roman weapons, linen, and grain, and various other luxury goods. While Maenchen-Helfen concedes that

14592-569: The south, east, and west as having occupied parts of Eastern Europe and Central Asia from about the 4th to 6th centuries. Variants of the Hun name are recorded in the Caucasus until the early 8th century. In the 18th century, French scholar Joseph de Guignes became the first to propose a link between the Huns and the Xiongnu people, who lived in northern China from the 3rd century BC to the late 1st century AD. Since Guignes's time, considerable scholarly effort has been devoted to investigating such

14720-405: The specifics of their origins. Classical sources assert that they appeared in Europe suddenly around 370. Most typically, Roman writers' attempts to elucidate the origins of the Huns simply equated them with earlier steppe peoples. Roman writers repeated a tale that the Huns had entered the domain of the Goths while they were pursuing a wild stag, or else one of their cows that had escaped, across

14848-606: The splitting event between the two groups was over 50 kya, casting doubt on recent common ancestry between the two. Archaeogenetics has been used to understand the development of domestication of plants and animals. The combination of genetics and archeological findings have been used to trace the earliest signs of plant domestication around the world. However, since the nuclear, mitochondrial, and chloroplast genomes used to trace domestication's moment of origin have evolved at different rates, its use to trace genealogy have been somewhat problematic. Nuclear DNA in specific

14976-475: The steppe nomad even than horses" and must have been a large part of their herds. Sheep bones are frequently found in Hun period graves. Additionally, Maenchen-Helfen argues that the Huns may have kept small herds of Bactrian camels in the part of their territory in modern Romania and Ukraine, something attested for the Sarmatians. Ammianus Marcellinus says that the majority of the Huns' diet came from

15104-524: The term "nomad" to be misleading: [T]he term 'nomad', if it denotes a wandering group of people with no clear sense of territory, cannot be applied wholesale to the Huns. All the so-called 'nomads' of Eurasian steppe history were peoples whose territory/territories were usually clearly defined, who as pastoralists moved about in search of pasture, but within a fixed territorial space. Maenchen-Helfen notes that pastoral nomads (or "seminomads") typically alternate between summer pastures and winter quarters: while

15232-505: The time of Attila, the Hunnic economy became almost entirely dependent on plunder and tribute from the Roman provinces. Civilians and soldiers captured by the Huns might also be ransomed back, or else sold to Roman slave dealers as slaves. The Huns themselves, Maenchen-Helfen argued, had little use for slaves due to their nomadic pastoralist lifestyle. More recent scholarship, however, has demonstrated that pastoral nomadists are actually more likely to use slave labor than sedentary societies:

15360-420: The time of the report of Olympiodorus, the Huns at some point developed a system of ranked kings, including a senior king by the time of Charaton . Priscus also speaks of "picked men" or logades ( λογάδες ) forming part of Attila's government, naming five of them. Some of the "picked men" seem to have been chosen because of birth, others for reasons of merit. Thompson argued that these "picked men" "were

15488-483: The two populations. The high frequency of a single NRY lineage unique to Australia coupled with “low diversity of lineage-associated Y-chromosomal short tandem repeat (Y-STR) haplotypes” provide evidence for a “recent founder or bottleneck” event in Australia. But there is relatively large variation in mtDNA, which would imply that the bottleneck effect impacted males primarily. Together, NRY and mtDNA studies show that

15616-577: Was mounted archery . The Huns may have stimulated the Great Migration , a contributing factor in the collapse of the Western Roman Empire . The memory of the Huns also lived on in various Christian saints' lives , where the Huns play the roles of antagonists, as well as in Germanic heroic legend , where the Huns are variously antagonists or allies to the Germanic main figures. In Hungary ,

15744-401: Was a massive influx of Near Eastern populations into Europe at the start of the Neolithic. This view led him “to strongly emphasize the expanding early farmers at the expense of the indigenous Mesolithic foraging populations.” mtDNA analysis in the 1990s, however, contradicted this view. M.B. Richards estimated that 10–22% of extant European mtDNA's had come from Near Eastern populations during

15872-456: Was apparently modest". He does note that wine and silk appear to have been imported into the Hunnic Empire in large quantities, however. Roman gold coins appear to have been in circulation as currency within the whole of the Hunnic Empire. Christopher Atwood has suggested that the purpose of the original Hunnic incursion into Europe may have been to establish an outlet to the Black Sea for

16000-510: Was found to be powerful in analyses of aDNA because it avoids potential loss of sample, substrate competition for templates, and error propagation in replication. The most common way to analyze an aDNA sequence is to compare it with a known sequence from other sources, and this could be done in different ways for different purposes. The identity of the fossil remain can be uncovered by comparing its DNA sequence with those of known species using software such as BLASTN. This archaeogenetic approach

16128-573: Was given the duty of relieving Orléans by Emperor Valentinian III. A combined army of Roman and Visigoths then fought the Huns at the Battle of the Catalaunian Plains . The following year, Attila renewed his claims to Honoria and territory in the Western Roman Empire . Leading his army across the Alps and into Northern Italy, he sacked and razed a number of cities. Hoping to avoid the sack of Rome, Emperor Valentinian III sent three envoys,

16256-480: Was researching the links of blood types to sex, disease, climate, age, social class, and race. His work led him to discover that peptic ulcer was more dominant in blood group O, and that AB blood type mothers had a high male-to-female birth ratio. Arthur Mourant was a British hematologist and chemist . He received many awards, most notably Fellowship of the Royal Society . His work included organizing

16384-492: Was unsuccessful at war as a sign that the Hunnic kingship was "democratic" at this time rather than a permanent institution. Kim, however, argues that Uldin is actually a title and that he was likely merely a subking. Priscus calls Attila "king" or "emperor" ( βασιλέυς ), but it is unknown what native title he was translating. With the exception of the sole rule of Attila, the Huns often had two rulers; Attila himself later appointed his son Ellac as co-king. Heather argues that by

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