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63-813: The Huronian Supergroup is a Proterozoic assemblage of geologic groups of the Superior Craton of the Canadian Shield in Ontario and Quebec . It extends from west of the city of Sault Ste. Marie in the west to the Ontario-Quebec border to the east and is part of the Southern Geologic Province. The Huronian Supergroup consists of four groups. All of the rocks are low grade metamorphic sedimentary rocks or igneous rocks . The four geologic groups that make up

126-472: A positive feedback known as a Fisherian runaway . Thus sexual reproduction, as a form of natural selection , has an effect on evolution . Sexual dimorphism is where the basic phenotypic traits vary between males and females of the same species . Dimorphism is found in both sex organs and in secondary sex characteristics , body size, physical strength and morphology, biological ornamentation , behavior and other bodily traits. However, sexual selection

189-472: A bacterium to bind, take up, and recombine exogenous DNA into its chromosome, it must enter a special physiological state referred to as competence (see Natural competence ). Sexual reproduction in early single-celled eukaryotes may have evolved from bacterial transformation, or from a similar process in archaea (see below). On the other hand, bacterial conjugation is a type of direct transfer of DNA between two bacteria mediated by an external appendage called

252-589: A few billion years in age. It is believed that 43% of modern continental crust was formed in the Proterozoic, 39% formed in the Archean, and only 18% in the Phanerozoic . Studies by Condie (2000) and Rino et al. (2004) harvp error: no target: CITEREFRinoKomiyaWindleyet_al2004 ( help ) suggest that crust production happened episodically. By isotopically calculating the ages of Proterozoic granitoids it

315-481: A few plausible models that explain tectonics of the early Earth prior to the formation of Columbia, but the current most plausible hypothesis is that prior to Columbia, there were only a few independent cratons scattered around the Earth (not necessarily a supercontinent, like Rodinia or Columbia). The Proterozoic can be roughly divided into seven biostratigraphic zones which correspond to informal time periods. The first

378-403: A haploid spore that grows into the dominant gametophyte form, which is a multicellular haploid body with leaf-like structures that photosynthesize . Haploid gametes are produced in antheridia (male) and archegonia (female) by mitosis. The sperm released from the antheridia respond to chemicals released by ripe archegonia and swim to them in a film of water and fertilize the egg cells thus producing

441-440: A multicellular gametophyte phase that produces gametes at maturity. The gametophytes of different groups of plants vary in size. Mosses and other pteridophytic plants may have gametophytes consisting of several million cells, while angiosperms have as few as three cells in each pollen grain. Flowering plants are the dominant plant form on land and they reproduce either sexually or asexually. Often their most distinctive feature

504-526: A period of increasing crustal recycling, suggesting subduction . Evidence for this increased subduction activity comes from the abundance of old granites originating mostly after 2.6  Ga . The occurrence of eclogite (a type of metamorphic rock created by high pressure, > 1 GPa), is explained using a model that incorporates subduction. The lack of eclogites that date to the Archean Eon suggests that conditions at that time did not favor

567-531: A population because they are better at securing mates for sexual reproduction. It has been described as "a powerful evolutionary force that does not exist in asexual populations". The first fossilized evidence of sexual reproduction in eukaryotes is from the Stenian period, about 1.05 billion years old. Biologists studying evolution propose several explanations for the development of sexual reproduction and its maintenance. These reasons include reducing

630-407: A zygote. The zygote divides by mitotic division and grows into a multicellular, diploid sporophyte. The sporophyte produces spore capsules ( sporangia ), which are connected by stalks ( setae ) to the archegonia. The spore capsules produce spores by meiosis and when ripe the capsules burst open to release the spores. Bryophytes show considerable variation in their reproductive structures and the above

693-444: Is a fitness disadvantage of sexual reproduction. The two-fold cost of sex includes this cost and the fact that any organism can only pass on 50% of its own genes to its offspring. However, one definite advantage of sexual reproduction is that it increases genetic diversity and impedes the accumulation of harmful genetic mutations . Sexual selection is a mode of natural selection in which some individuals out-reproduce others of

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756-400: Is a basic outline. Also in some species each plant is one sex ( dioicous ) while other species produce both sexes on the same plant ( monoicous ). Fungi are classified by the methods of sexual reproduction they employ. The outcome of sexual reproduction most often is the production of resting spores that are used to survive inclement times and to spread. There are typically three phases in

819-508: Is a transfer of plasmid DNA between bacteria, but the plasmids are rarely incorporated into the bacterial chromosome; and gene transfer and genetic exchange in archaea . Bacterial transformation involves the recombination of genetic material and its function is mainly associated with DNA repair . Bacterial transformation is a complex process encoded by numerous bacterial genes, and is a bacterial adaptation for DNA transfer. This process occurs naturally in at least 40 bacterial species. For

882-574: Is a unisex species that uses a form of parthenogenesis called gynogenesis , where unfertilized eggs develop into embryos that produce female offspring. Poecilia formosa mate with males of other fish species that use internal fertilization, the sperm does not fertilize the eggs but stimulates the growth of the eggs which develops into embryos. Animals have life cycles with a single diploid multicellular phase that produces haploid gametes directly by meiosis. Male gametes are called sperm, and female gametes are called eggs or ova. In animals, fertilization of

945-568: Is believed to have developed in an ancient eukaryotic ancestor. In eukaryotes, diploid precursor cells divide to produce haploid cells in a process called meiosis . In meiosis, DNA is replicated to produce a total of four copies of each chromosome. This is followed by two cell divisions to generate haploid gametes. After the DNA is replicated in meiosis, the homologous chromosomes pair up so that their DNA sequences are aligned with each other. During this period before cell divisions, genetic information

1008-399: Is entirely beneficial. Larger populations appear to respond more quickly to some of the benefits obtained through sexual reproduction than do smaller population sizes. However, newer models presented in recent years suggest a basic advantage for sexual reproduction in slowly reproducing complex organisms . Sexual reproduction allows these species to exhibit characteristics that depend on

1071-431: Is exchanged between homologous chromosomes in genetic recombination . Homologous chromosomes contain highly similar but not identical information, and by exchanging similar but not identical regions, genetic recombination increases genetic diversity among future generations. During sexual reproduction, two haploid gametes combine into one diploid cell known as a zygote in a process called fertilization . The nuclei from

1134-595: Is only implied over an extended period of time leading to sexual dimorphism. A few arthropods, such as barnacles , are hermaphroditic , that is, each can have the organs of both sexes . However, individuals of most species remain of one sex their entire lives. A few species of insects and crustaceans can reproduce by parthenogenesis , especially if conditions favor a "population explosion". However, most arthropods rely on sexual reproduction, and parthenogenetic species often revert to sexual reproduction when conditions become less favorable. The ability to undergo meiosis

1197-645: Is the third of the four geologic eons of Earth's history , spanning the time interval from 2500 to 538.8   Mya , the longest eon of the Earth's geologic time scale . It is preceded by the Archean and followed by the Phanerozoic , and is the most recent part of the Precambrian "supereon". The Proterozoic is subdivided into three geologic eras (from oldest to youngest): the Paleoproterozoic , Mesoproterozoic and Neoproterozoic . It covers

1260-412: Is their reproductive organs, commonly called flowers. The anther produces pollen grains which contain the male gametophytes that produce sperm nuclei. For pollination to occur, pollen grains must attach to the stigma of the female reproductive structure ( carpel ), where the female gametophytes are located within ovules enclose within the ovary . After the pollen tube grows through the carpel's style,

1323-459: Is widespread among arthropods including both those that reproduce sexually and those that reproduce parthenogenetically . Although meiosis is a major characteristic of arthropods, understanding of its fundamental adaptive benefit has long been regarded as an unresolved problem, that appears to have remained unsettled. Aquatic arthropods may breed by external fertilization, as for example horseshoe crabs do, or by internal fertilization , where

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1386-560: The gametophyte , which produces gametes directly by mitosis. This type of life cycle, involving alternation between two multicellular phases, the sexual haploid gametophyte and asexual diploid sporophyte, is known as alternation of generations . The evolution of sexual reproduction is considered paradoxical, because asexual reproduction should be able to outperform it as every young organism created can bear its own young. This implies that an asexual population has an intrinsic capacity to grow more rapidly with each generation. This 50% cost

1449-410: The ova remain in the female's body and the sperm must somehow be inserted. All known terrestrial arthropods use internal fertilization. Opiliones (harvestmen), millipedes , and some crustaceans use modified appendages such as gonopods or penises to transfer the sperm directly to the female. However, most male terrestrial arthropods produce spermatophores , waterproof packets of sperm , which

1512-420: The sex cell nuclei from the pollen grain migrate into the ovule to fertilize the egg cell and endosperm nuclei within the female gametophyte in a process termed double fertilization . The resulting zygote develops into an embryo, while the triploid endosperm (one sperm cell plus two female cells) and female tissues of the ovule give rise to the surrounding tissues in the developing seed. The ovary, which produced

1575-507: The sex organs present although not reproductively functional. After several months or years, depending on the species, the sex organs develop further to maturity and the animal becomes sexually mature . Most female mammals are only fertile during certain periods during their estrous cycle, at which point they are ready to mate. For most mammals, males and females exchange sexual partners throughout their adult lives . The vast majority of fish species lay eggs that are then fertilized by

1638-662: The Ediacaran, proving that multicellular life had already become widespread tens of millions of years before the Cambrian Explosion in what is known as the Avalon Explosion . Nonetheless, the upper boundary of the Proterozoic has remained fixed at the base of the Cambrian , which is currently placed at 538.8 Ma. Sexual reproduction In placental mammals , sperm cells exit the penis through

1701-779: The Huronian are the Elliot Lake Group, Hough Lake Group, Quirke Lake Group, and the Cobalt Group (from oldest to youngest). Major glacial periods, Huronian glaciation , occurred at the beginning of the Hough Lake, Quirke Lake and Cobalt. The bulk of the Huronian Group consists of mostly metamorphosed sandstones and mudstones . Since there are no index fossils, the sedimentary rocks cannot be dated using fossils as they can in Phanerozoic rocks. However,

1764-737: The Neoproterozoic Era at the end of the Proterozoic Eon, possibly climaxing with the hypothesized Snowball Earth of the Sturtian and Marinoan glaciations. One of the most important events of the Proterozoic was the accumulation of oxygen in the Earth's atmosphere. Though oxygen is believed to have been released by photosynthesis as far back as the Archean Eon, it could not build up to any significant degree until mineral sinks of unoxidized sulfur and iron had been exhausted. Until roughly 2.3 billion years ago, oxygen

1827-592: The Palaeoproterozoic or Mesoproterozoic, according to molecular data. Classically, the boundary between the Proterozoic and the Phanerozoic eons was set at the base of the Cambrian Period when the first fossils of animals, including trilobites and archeocyathids , as well as the animal-like Caveasphaera , appeared. In the second half of the 20th century, a number of fossil forms have been found in Proterozoic rocks, particularly in ones from

1890-602: The base contains volcanic rocks that have been dated at about 2.480 billion years ago (Gya). The top of the Huronian and cross cutting all of the units within are the Nipissing sills which date to 2.2185 Gya. The age at which the Huronian sediments were deposited is between 2.480 Gya and 2.219 Gya. Proterozoic The Proterozoic ( IPA : / ˌ p r oʊ t ər ə ˈ z oʊ ɪ k , ˌ p r ɒ t -, - ər oʊ -, - t r ə -, - t r oʊ -/ PROH -tər-ə- ZOH -ik, PROT-, -⁠ər-oh-, -⁠trə-, -⁠troh- )

1953-486: The breakup of the supercontinent Columbia and prior to the assemblage of the supercontinent Gondwana (~500 Ma). The defining orogenic event associated with the formation of Gondwana was the collision of Africa, South America, Antarctica and Australia forming the Pan-African orogeny . Columbia was dominant in the early-mid Proterozoic and not much is known about continental assemblages before then. There are

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2016-783: The conjugation pilus. Bacterial conjugation is controlled by plasmid genes that are adapted for spreading copies of the plasmid between bacteria. The infrequent integration of a plasmid into a host bacterial chromosome, and the subsequent transfer of a part of the host chromosome to another cell do not appear to be bacterial adaptations. Exposure of hyperthermophilic archaeal Sulfolobus species to DNA damaging conditions induces cellular aggregation accompanied by high frequency genetic marker exchange Ajon et al. hypothesized that this cellular aggregation enhances species-specific DNA repair by homologous recombination. DNA transfer in Sulfolobus may be an early form of sexual interaction similar to

2079-522: The deciphering of Precambrian Supereon tectonics. It is known that tectonic processes of the Proterozoic Eon resemble greatly the evidence of tectonic activity, such as orogenic belts or ophiolite complexes, we see today. Hence, most geologists would conclude that the Earth was active at that time. It is also commonly accepted that during the Precambrian, the Earth went through several supercontinent breakup and rebuilding cycles ( Wilson cycle ). In

2142-464: The eggs are fertilized within the female and the eggs simply hatch within the female body, or in seahorses , the male carries the developing young within a pouch, and gives birth to live young. Fishes can also be viviparous , where the female supplies nourishment to the internally growing offspring. Some fish are hermaphrodites , where a single fish is both male and female and can produce eggs and sperm. In hermaphroditic fish, some are male and female at

2205-426: The eon continued the massive continental accretion that had begun late in the Archean Eon. The Proterozoic Eon also featured the first definitive supercontinent cycles and wholly modern mountain building activity ( orogeny ). There is evidence that the first known glaciations occurred during the Proterozoic. The first began shortly after the beginning of the Proterozoic Eon, and evidence of at least four during

2268-757: The evolution of eukaryotes via symbiogenesis ; several global glaciations , which produced the 300 million years-long Huronian glaciation (during the Siderian and Rhyacian periods of the Paleoproterozoic) and the hypothesized Snowball Earth (during the Cryogenian period in the late Neoproterozoic); and the Ediacaran period (635–538.8  Ma ), which is characterized by the evolution of abundant soft-bodied multicellular organisms such as sponges , algae , cnidarians , bilaterians and

2331-481: The female gametophyte(s), then grows into a fruit , which surrounds the seed(s). Plants may either self-pollinate or cross-pollinate . In 2013, flowers dating from the Cretaceous (100 million years before present) were found encased in amber, the oldest evidence of sexual reproduction in a flowering plant. Microscopic images showed tubes growing out of pollen and penetrating the flower's stigma. The pollen

2394-494: The females take into their bodies. A few such species rely on females to find spermatophores that have already been deposited on the ground, but in most cases males only deposit spermatophores when complex courtship rituals look likely to be successful. Insect species make up more than two-thirds of all extant animal species. Most insect species reproduce sexually, though some species are facultatively parthenogenetic . Many insect species have sexual dimorphism , while in others

2457-678: The first half of the Ediacaran from 0.63–0.55 Ga, and the Belomorian, spanning from 0.55–0.542 Ga. The emergence of advanced single-celled eukaryotes began after the Oxygen Catastrophe . This may have been due to an increase in the oxidized nitrates that eukaryotes use, as opposed to cyanobacteria . It was also during the Proterozoic that the first symbiotic relationships between mitochondria (found in nearly all eukaryotes) and chloroplasts (found in plants and some protists only) and their hosts evolved. By

2520-405: The formation of a zygote, and varying degrees of development, in many species the eggs are deposited outside the female; while in others, they develop further within the female and the young are born live. There are three extant kinds of mammals: monotremes , placentals and marsupials , all with internal fertilization. In placental mammals, offspring are born as juveniles: complete animals with

2583-419: The formation of high grade metamorphism and therefore did not achieve the same levels of subduction as was occurring in the Proterozoic Eon. As a result of remelting of basaltic oceanic crust due to subduction, the cores of the first continents grew large enough to withstand the crustal recycling processes. The long-term tectonic stability of those cratons is why we find continental crust ranging up to

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2646-409: The gametes fuse, and each gamete contributes half of the genetic material of the zygote. Multiple cell divisions by mitosis (without change in the number of chromosomes) then develop into a multicellular diploid phase or generation. In plants, the diploid phase, known as the sporophyte , produces spores by meiosis. These spores then germinate and divide by mitosis to form a haploid multicellular phase,

2709-665: The late Palaeoproterozoic, eukaryotic organisms had become moderately biodiverse. The blossoming of eukaryotes such as acritarchs did not preclude the expansion of cyanobacteria – in fact, stromatolites reached their greatest abundance and diversity during the Proterozoic, peaking roughly 1.2 billion years ago. The earliest fossils possessing features typical of fungi date to the Paleoproterozoic Era, some 2.4 billion years ago; these multicellular benthic organisms had filamentous structures capable of anastomosis . The Viridiplantae evolved sometime in

2772-625: The late Proterozoic (most recent), the dominant supercontinent was Rodinia (~1000–750 Ma). It consisted of a series of continents attached to a central craton that forms the core of the North American Continent called Laurentia . An example of an orogeny (mountain building processes) associated with the construction of Rodinia is the Grenville orogeny located in Eastern North America. Rodinia formed after

2835-421: The likelihood of the accumulation of deleterious mutations, increasing rate of adaptation to changing environments , dealing with competition , DNA repair , masking deleterious mutations, and reducing genetic variation on the genomic level. All of these ideas about why sexual reproduction has been maintained are generally supported, but ultimately the size of the population determines if sexual reproduction

2898-876: The male urethra and enter the vagina during copulation , while egg cells enter the uterus through the oviduct . Other vertebrates of both sexes possess a cloaca for the release of sperm or egg cells. Sexual reproduction is the most common life cycle in multicellular eukaryotes, such as animals , fungi and plants . Sexual reproduction also occurs in some unicellular eukaryotes. Sexual reproduction does not occur in prokaryotes , unicellular organisms without cell nuclei , such as bacteria and archaea . However, some processes in bacteria, including bacterial conjugation , transformation and transduction , may be considered analogous to sexual reproduction in that they incorporate new genetic information. Some proteins and other features that are key for sexual reproduction may have arisen in bacteria, but sexual reproduction

2961-454: The male. Some species lay their eggs on a substrate like a rock or on plants, while others scatter their eggs and the eggs are fertilized as they drift or sink in the water column. Some fish species use internal fertilization and then disperse the developing eggs or give birth to live offspring. Fish that have live-bearing offspring include the guppy and mollies or Poecilia . Fishes that give birth to live young can be ovoviviparous , where

3024-414: The most plausible reason for maintaining this capability is the benefit of repairing DNA damage , caused by a variety of stresses, through recombination that occurs during meiosis . Three distinct processes in prokaryotes are regarded as similar to eukaryotic sex : bacterial transformation , which involves the incorporation of foreign DNA into the bacterial chromosome; bacterial conjugation , which

3087-453: The ovum by a sperm results in the formation of a diploid zygote that develops by repeated mitotic divisions into a diploid adult. Plants have two multicellular life-cycle phases, resulting in an alternation of generations . Plant zygotes germinate and divide repeatedly by mitosis to produce a diploid multicellular organism known as the sporophyte. The mature sporophyte produces haploid spores by meiosis that germinate and divide by mitosis to form

3150-415: The same or different plants. After rains or when dew deposits a film of water, the motile sperm are splashed away from the antheridia, which are normally produced on the top side of the thallus, and swim in the film of water to the archegonia where they fertilize the egg. To promote out crossing or cross fertilization the sperm are released before the eggs are receptive of the sperm, making it more likely that

3213-402: The same time while in other fish they are serially hermaphroditic; starting as one sex and changing to the other. In at least one hermaphroditic species, self-fertilization occurs when the eggs and sperm are released together. Internal self-fertilization may occur in some other species. One fish species does not reproduce by sexual reproduction but uses sex to produce offspring; Poecilia formosa

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3276-554: The sessile Ediacaran biota (some of which had evolved sexual reproduction ) and provides the first obvious fossil evidence of life on Earth . The geologic record of the Proterozoic Eon is more complete than that for the preceding Archean Eon. In contrast to the deep-water deposits of the Archean, the Proterozoic features many strata that were laid down in extensive shallow epicontinental seas ; furthermore, many of those rocks are less metamorphosed than Archean rocks, and many are unaltered. Studies of these rocks have shown that

3339-450: The sexes look nearly identical. Typically they have two sexes with males producing spermatozoa and females ova. The ova develop into eggs that have a covering called the chorion , which forms before internal fertilization. Insects have very diverse mating and reproductive strategies most often resulting in the male depositing a spermatophore within the female, which she stores until she is ready for egg fertilization. After fertilization, and

3402-477: The sexual reproduction of fungi: plasmogamy , karyogamy and meiosis . The cytoplasm of two parent cells fuse during plasmogamy and the nuclei fuse during karyogamy. New haploid gametes are formed during meiosis and develop into spores. The adaptive basis for the maintenance of sexual reproduction in the Ascomycota and Basidiomycota ( dikaryon ) fungi was reviewed by Wallen and Perlin. They concluded that

3465-405: The specific environment that they inhabit, and the particular survival strategies that they employ. In order to reproduce sexually, both males and females need to find a mate . Generally in animals mate choice is made by females while males compete to be chosen. This can lead organisms to extreme efforts in order to reproduce, such as combat and display, or produce extreme features caused by

3528-541: The sperm will fertilize the eggs of different thallus. After fertilization, a zygote is formed which grows into a new sporophytic plant. The condition of having separate sporophyte and gametophyte plants is called alternation of generations . The bryophytes , which include liverworts , hornworts and mosses , reproduce both sexually and vegetatively . They are small plants found growing in moist locations and like ferns, have motile sperm with flagella and need water to facilitate sexual reproduction. These plants start as

3591-538: The time from the appearance of free oxygen in Earth's atmosphere to just before the proliferation of complex life on the Earth during the Cambrian Explosion . The name Proterozoic combines two words of Greek origin: protero- meaning "former, earlier", and -zoic , meaning "of life". Well-identified events of this eon were the transition to an oxygenated atmosphere during the Paleoproterozoic;

3654-567: The time was virtually all obligate anaerobic . A second, later surge in oxygen concentrations is called the Neoproterozoic Oxygenation Event , occurred during the Middle and Late Neoproterozoic and drove the rapid evolution of multicellular life towards the end of the era. The Proterozoic Eon was a very tectonically active period in the Earth's history. The late Archean Eon to Early Proterozoic Eon corresponds to

3717-444: Was determined that there were several episodes of rapid increase in continental crust production. The reason for these pulses is unknown, but they seemed to have decreased in magnitude after every period. Evidence of collision and rifting between continents raises the question as to what exactly were the movements of the Archean cratons composing Proterozoic continents. Paleomagnetic and geochronological dating mechanisms have allowed

3780-511: Was probably due to two factors: Exhaustion of the chemical sinks, and an increase in carbon sequestration , which sequestered organic compounds that would have otherwise been oxidized by the atmosphere. The first surge in atmospheric oxygen at the beginning of the Proterozoic is called the Great Oxygenation Event , or alternately the Oxygen Catastrophe – to reflect the mass extinction of almost all life on Earth, which at

3843-487: Was probably only 1% to 2% of its current level. The banded iron formations , which provide most of the world's iron ore , are one mark of that mineral sink process. Their accumulation ceased after 1.9 billion years ago, after the iron in the oceans had all been oxidized . Red beds , which are colored by hematite , indicate an increase in atmospheric oxygen 2 billion years ago. Such massive iron oxide formations are not found in older rocks. The oxygen buildup

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3906-424: Was sticky, suggesting it was carried by insects. Ferns produce large diploid sporophytes with rhizomes , roots and leaves. Fertile leaves produce sporangia that contain haploid spores . The spores are released and germinate to produce small, thin gametophytes that are typically heart shaped and green in color. The gametophyte prothalli , produce motile sperm in the antheridia and egg cells in archegonia on

3969-761: Was the Labradorian, lasting from 2.0–1.65  Ga . It was followed by the Anabarian, which lasted from 1.65–1.2 Ga and was itself followed by the Turukhanian from 1.2–1.03 Ga. The Turukhanian was succeeded by the Uchuromayan, lasting from 1.03–0.85 Ga, which was in turn succeeded by the Yuzhnouralian, lasting from 0.85–0.63 Ga. The final two zones were the Amadeusian, spanning

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