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An interglacial period (or alternatively interglacial , interglaciation ) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age . The current Holocene interglacial began at the end of the Pleistocene , about 11,700 years ago.

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91-535: During the 2.5 million years of the Pleistocene , numerous glacials, or significant advances of continental ice sheets, in North America and Europe , occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter interglacials. During interglacials, such as the present one, the climate warms and the tundra recedes polewards following

182-473: A monotreme , appeared. The predatory South American phorusrhacids were rare in this time; among the last was Titanis , a large phorusrhacid that migrated to North America and rivaled mammals as top predator. Other birds probably evolved at this time, some modern (such as the genera Cygnus , Bubo , Struthio and Corvus ), some now extinct. Alligators and crocodiles died out in Europe as

273-611: A change from low-amplitude glacial cycles with a dominant periodicity of 41,000 years to asymmetric high-amplitude cycles dominated by a periodicity of 100,000 years. However, a 2020 study concluded that ice age terminations might have been influenced by obliquity since the Mid-Pleistocene Transition, which caused stronger summers in the Northern Hemisphere . Glaciation in the Pleistocene

364-574: A deviation from today's annual mean temperature, taken as zero. This sort of graph is based on another isotope ratio versus time. Ratios are converted to a percentage difference from the ratio found in standard mean ocean water (SMOW). The graph in either form appears as a waveform with overtones . One half of a period is a Marine isotopic stage (MIS). It indicates a glacial (below zero) or an interglacial (above zero). Overtones are stadials or interstadials. According to this evidence, Earth experienced 102 MIS stages beginning at about 2.588 Ma BP in

455-421: A few regions had been studied and the names were relatively few. Today the geologists of different nations are taking more of an interest in Pleistocene glaciology. As a consequence, the number of names is expanding rapidly and will continue to expand. Many of the advances and stadials remain unnamed. Also, the terrestrial evidence for some of them has been erased or obscured by larger ones, but evidence remains from

546-406: A large neocortex relative to body mass and stone tool manufacture. Improvements in dating methods and in the use of climate proxies have provided scientists with the means to test hypotheses of the evolution of human ancestors. Early hypotheses of the evolution of human traits emphasized the selective pressures produced by particular habitats. For example, many scientists have long favored

637-429: A particular habitat. The change to a cooler, drier, more seasonal climate had considerable impacts on Pliocene vegetation, reducing tropical species worldwide. Deciduous forests proliferated, coniferous forests and tundra covered much of the north, and grasslands spread on all continents (except Antarctica). Eastern Africa in particular saw a huge expansion of C 4 grasslands. Tropical forests were limited to

728-578: A team of Russian scientists in collaboration with Princeton University announced that they had brought two female nematodes frozen in permafrost , from around 42,000 years ago, back to life. The two nematodes, at the time, were the oldest confirmed living animals on the planet. The evolution of anatomically modern humans took place during the Pleistocene. At the beginning of the Pleistocene Paranthropus species were still present, as well as early human ancestors, but during

819-968: A tight band around the equator, and in addition to dry savannahs , deserts appeared in Asia and Africa. Both marine and continental faunas were essentially modern, although continental faunas were a bit more primitive than today. The land mass collisions meant great migration and mixing of previously isolated species, such as in the Great American Interchange . Herbivores got bigger, as did specialized predators. In North America, rodents , large mastodons and gomphotheres , and opossums continued successfully, while hoofed animals ( ungulates ) declined, with camel , deer , and horse all seeing populations recede. Three-toed horses ( Nannippus ), oreodonts , protoceratids , and chalicotheres became extinct. Borophagine dogs and Agriotherium became extinct, but other carnivores including

910-583: Is a change in Earth's orbit around the Sun, or eccentricity . The second is a shift in the tilt of Earth's axis, or obliquity . The third is the wobbling motion of Earth's axis, or precession . In the Southern Hemisphere , warmer summers occur when the lower-half of Earth is tilted toward the Sun and the planet is nearest the Sun in its elliptical orbit. Cooler summers occur when Earth is farthest from

1001-667: Is estimated that, at maximum glacial extent, 30% of the Earth's surface was covered by ice. In addition, a zone of permafrost stretched southward from the edge of the glacial sheet, a few hundred kilometres in North America , and several hundred in Eurasia . The mean annual temperature at the edge of the ice was −6 °C (21 °F); at the edge of the permafrost, 0 °C (32 °F). Each glacial advance tied up huge volumes of water in continental ice sheets 1,500 to 3,000 metres (4,900–9,800 ft) thick, resulting in temporary sea-level drops of 100 metres (300 ft) or more over

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1092-459: Is no systematic correspondence between pluvials to glacials, however. Moreover, regional pluvials do not correspond to each other globally. For example, some have used the term "Riss pluvial" in Egyptian contexts. Any coincidence is an accident of regional factors. Only a few of the names for pluvials in restricted regions have been stratigraphically defined. The sum of transient factors acting at

1183-906: Is sometimes referred to as the Late Pliocene, whereas the Zanclean is referred to as the Early Pliocene. In the system of In the Paratethys area (central Europe and parts of western Asia) the Pliocene contains the Dacian (roughly equal to the Zanclean) and Romanian (roughly equal to the Piacenzian and Gelasian together) stages. As usual in stratigraphy, there are many other regional and local subdivisions in use. In Britain ,

1274-488: Is the period within an interglacial that experienced the most 'favourable' climate and often occurs during the middle of that interglacial. The climatic optimum of an interglacial both follows and is followed by phases within the same interglacial that experienced a less favourable climate (but still a 'better' climate than the one during the preceding or succeeding glacials ). During an interglacial optimum, sea levels rise to their highest values, but not necessarily exactly at

1365-592: The Ice Age ) is the geological epoch that lasted from c.  2.58 million to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations . Before a change was finally confirmed in 2009 by the International Union of Geological Sciences , the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of

1456-659: The Alpine ice sheet on the Alps . Scattered domes stretched across Siberia and the Arctic shelf. The northern seas were ice-covered. South of the ice sheets large lakes accumulated because outlets were blocked and the cooler air slowed evaporation. When the Laurentide Ice Sheet retreated, north-central North America was completely covered by Lake Agassiz . Over a hundred basins, now dry or nearly so, were overflowing in

1547-524: The Hardangervidda plateau to 1200 m in the Early Pliocene. In Southern Sweden similar movements elevated the South Swedish highlands leading to a deflection of the ancient Eridanos river from its original path across south-central Sweden into a course south of Sweden. The Pliocene is bookended by two significant events in the evolution of human ancestors. The first is the appearance of

1638-529: The ICS timescale, the Pleistocene is divided into four stages or ages , the Gelasian , Calabrian , Chibanian (previously the unofficial "Middle Pleistocene"), and Upper Pleistocene (unofficially the "Tarantian"). In addition to these international subdivisions, various regional subdivisions are often used. In 2009 the International Union of Geological Sciences (IUGS) confirmed a change in time period for

1729-481: The International Commission on Stratigraphy (ICS) stages is not established. During the Pliocene epoch (5.3 to 2.6 million years ago (Ma)), the Earth's climate became cooler and drier, as well as more seasonal, marking a transition between the relatively warm Miocene to the cooler Pleistocene . However, the beginning of the Pliocene was marked by an increase in global temperatures relative to

1820-686: The Isthmus of Panama during the Pliocene, making possible the Great American Interchange and bringing a nearly complete end to South America's distinctive native ungulate fauna, though other South American lineages like its predatory mammals were already extinct by this point and others like xenarthrans continued to do well afterwards. The formation of the Isthmus had major consequences on global temperatures, since warm equatorial ocean currents were cut off and an Atlantic cooling cycle began, with cold Arctic and Antarctic waters decreasing temperatures in

1911-603: The Isthmus of Panama , causing a faunal interchange between the two regions and changing ocean circulation patterns, with the onset of glaciation in the Northern Hemisphere occurring around 2.7 million years ago. During the Early Pleistocene (2.58–0.8 Ma), archaic humans of the genus Homo originated in Africa and spread throughout Afro-Eurasia . The end of the Early Pleistocene is marked by

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2002-677: The Laurentide Ice Sheet . Charles Lyell introduced the term "Pleistocene" in 1839 to describe strata in Sicily that had at least 70% of their molluscan fauna still living today. This distinguished it from the older Pliocene Epoch , which Lyell had originally thought to be the youngest fossil rock layer. He constructed the name "Pleistocene" ('most new' or 'newest') from the Greek πλεῖστος ( pleīstos ) 'most' and καινός ( kainós ( Latinized as cænus ) 'new'). This contrasts with

2093-615: The Mid-Pleistocene Transition , with the cyclicity of glacial cycles changing from 41,000-year cycles to asymmetric 100,000-year cycles, making the climate variation more extreme. The Late Pleistocene witnessed the spread of modern humans outside of Africa as well as the extinction of all other human species. Humans also spread to the Australian continent and the Americas for the first time, co-incident with

2184-748: The Neogene Period in the Cenozoic Era . The Pliocene follows the Miocene Epoch and is followed by the Pleistocene Epoch. Prior to the 2009 revision of the geologic time scale, which placed the four most recent major glaciations entirely within the Pleistocene, the Pliocene also included the Gelasian Stage, which lasted from 2.59 to 1.81 Ma, and is now included in the Pleistocene. As with other older geologic periods,

2275-662: The Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of Ethiopia and to the west in the Atlas Mountains . In the northern hemisphere, many glaciers fused into one. The Cordilleran Ice Sheet covered the North American northwest; the east was covered by the Laurentide . The Fenno-Scandian ice sheet rested on northern Europe , including much of Great Britain;

2366-455: The geological strata that define the start and end are well-identified but the exact dates of the start and end of the epoch are slightly uncertain. The boundaries defining the Pliocene are not set at an easily identified worldwide event but rather at regional boundaries between the warmer Miocene and the relatively cooler Pleistocene. The upper boundary was set at the start of the Pleistocene glaciations. Charles Lyell (later Sir Charles) gave

2457-497: The hominin Australopithecus anamensis in the early Pliocene, around 4.2 million years ago. The second is the appearance of Homo , the genus that includes modern humans and their closest extinct relatives, near the end of the Pliocene at 2.6 million years ago. Key traits that evolved among hominins during the Pliocene include terrestrial bipedality and, by the end of the Pliocene, encephalized brains (brains with

2548-477: The last glacial period ended about 10,000 years ago. Over 11 major glacial events have been identified, as well as many minor glacial events. A major glacial event is a general glacial excursion, termed a "glacial." Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials". These events are defined differently in different regions of

2639-634: The savannah hypothesis . This proposes that the evolution of terrestrial bipedality and other traits was an adaptive response to Pliocene climate change that transformed forests into more open savannah . This was championed by Grafton Elliot Smith in his 1924 book, The Evolution of Man , as "the unknown world beyond the trees", and was further elaborated by Raymond Dart as the killer ape theory . Other scientists, such as Sherwood L. Washburn , emphasized an intrinsic model of hominin evolution. According to this model, early evolutionary developments triggered later developments. The model placed little emphasis on

2730-493: The weasel family diversified, and dogs and short-faced bears did well. Ground sloths , huge glyptodonts , and armadillos came north with the formation of the Isthmus of Panama. The latitudinal diversity gradient among terrestrial North American mammals became established during this epoch some time after 4 Ma. In Eurasia rodents did well, while primate distribution declined. Elephants , gomphotheres and stegodonts were successful in Asia (the largest land mammals of

2821-599: The woolly rhinoceros , various giraffids , such as the Sivatherium ; ground sloths , Irish elk , cave lions , cave bears , Gomphotheres , American lions , dire wolves , and short-faced bears , began late in the Pleistocene and continued into the Holocene. Neanderthals also became extinct during this period. At the end of the last ice age, cold-blooded animals, smaller mammals like wood mice , migratory birds, and swifter animals like whitetail deer had replaced

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2912-569: The Early Pleistocene Gelasian . Early Pleistocene stages were shallow and frequent. The latest were the most intense and most widely spaced. By convention, stages are numbered from the Holocene, which is MIS1. Glacials receive an even number and interglacials receive an odd number. The first major glacial was MIS2-4 at about 85–11 ka BP. The largest glacials were 2, 6, 12, and 16. The warmest interglacials were 1, 5, 9 and 11. For matching of MIS numbers to named stages, see under

3003-409: The Earth's surface is cyclical: climate, ocean currents and other movements, wind currents, temperature, etc. The waveform response comes from the underlying cyclical motions of the planet, which eventually drag all the transients into harmony with them. The repeated glaciations of the Pleistocene were caused by the same factors. The Mid-Pleistocene Transition , approximately one million years ago, saw

3094-516: The Late Miocene and Pliocene, indicating increased aridification. Further aridification of Central Asia was caused by the development of Northern Hemisphere glaciation during the Late Pliocene. A sediment core from the northern South China Sea shows an increase in dust storm activity during the middle Pliocene. The South Asian Summer Monsoon (SASM) increased in intensity after 2.95 Ma, likely because of enhanced cross-equatorial pressure caused by

3185-479: The North American west. Lake Bonneville , for example, stood where Great Salt Lake now does. In Eurasia, large lakes developed as a result of the runoff from the glaciers. Rivers were larger, had a more copious flow, and were braided . African lakes were fuller, apparently from decreased evaporation. Deserts, on the other hand, were drier and more extensive. Rainfall was lower because of the decreases in oceanic and other evaporation. It has been estimated that during

3276-528: The Pleistocene glacial-interglacial cycles. During the late Pliocene and early Pleistocene, 3.6 to 2.6 Ma, the Arctic was much warmer than it is at the present day (with summer temperatures some 8 °C warmer than today). That is a key finding of research into a lake-sediment core obtained in Eastern Siberia, which is of exceptional importance because it has provided the longest continuous late Cenozoic land-based sedimentary record thus far. During

3367-434: The Pleistocene to 2.58 Ma, results in the inclusion of all the recent repeated glaciations within the Pleistocene. Radiocarbon dating is considered to be inaccurate beyond around 50,000 years ago. Marine isotope stages (MIS) derived from Oxygen isotopes are often used for giving approximate dates. Pleistocene non-marine sediments are found primarily in fluvial deposits , lakebeds, slope and loess deposits as well as in

3458-496: The Pleistocene's overall climate could be characterised as a continuous El Niño with trade winds in the south Pacific weakening or heading east, warm air rising near Peru , warm water spreading from the west Pacific and the Indian Ocean to the east Pacific, and other El Niño markers. Pleistocene climate was marked by repeated glacial cycles in which continental glaciers pushed to the 40th parallel in some places. It

3549-503: The Pleistocene, changing the start date from 1.806 to 2.588 million years BP, and accepted the base of the Gelasian as the base of the Pleistocene, namely the base of the Monte San Nicola GSSP . The start date has now been rounded down to 2.580 million years BP. The IUGS has yet to approve a type section , Global Boundary Stratotype Section and Point (GSSP), for the upper Pleistocene/Holocene boundary ( i.e.

3640-752: The Pleistocene, the East Antarctic Ice Sheet thinned by at least 500 meters, and that thinning since the Last Glacial Maximum is less than 50 meters and probably started after ca 14 ka. During the 2.5 million years of the Pleistocene, numerous cold phases called glacials ( Quaternary ice age ), or significant advances of continental ice sheets, in Europe and North America, occurred at intervals of approximately 40,000 to 100,000 years. The long glacial periods were separated by more temperate and shorter interglacials which lasted about 10,000–15,000 years. The last cold episode of

3731-411: The Pleistocene, the plates upon which they sit probably having moved no more than 100 km (62 mi) relative to each other since the beginning of the period. In glacial periods, the sea level would drop by up to 120 m (390 ft) lower than today during peak glaciation, exposing large areas of the present continental shelf as dry land. According to Mark Lynas (through collected data),

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3822-565: The Pliocene is divided into the following stages (old to young): Gedgravian, Waltonian , Pre-Ludhamian, Ludhamian, Thurnian, Bramertonian or Antian, Pre-Pastonian or Baventian, Pastonian and Beestonian . In the Netherlands the Pliocene is divided into these stages (old to young): Brunssumian C, Reuverian A, Reuverian B, Reuverian C, Praetiglian , Tiglian A, Tiglian B, Tiglian C1-4b, Tiglian C4c, Tiglian C5, Tiglian C6 and Eburonian . The exact correlations between these local stages and

3913-553: The Pliocene its name in Principles of Geology (volume 3, 1833). The word pliocene comes from the Greek words πλεῖον ( pleion , "more") and καινός ( kainos , "new" or "recent") and means roughly "continuation of the recent", referring to the essentially modern marine mollusc fauna. In the official timescale of the ICS , the Pliocene is subdivided into two stages . From youngest to oldest they are: The Piacenzian

4004-429: The Pliocene may have accelerated on the disappearance of forests and the spread of grasslands and savannas. During the Pliocene the earth climate system response shifted from a period of high frequency-low amplitude oscillation dominated by the 41,000-year period of Earth's obliquity to one of low-frequency, high-amplitude oscillation dominated by the 100,000-year period of the orbital eccentricity characteristic of

4095-503: The Pliocene were such proboscideans as Deinotherium , Anancus , and Mammut borsoni , ) though proboscidean diversity declined significantly during the Late Pliocene. Hyraxes migrated north from Africa. Horse diversity declined, while tapirs and rhinos did fairly well. Bovines and antelopes were successful; some camel species crossed into Asia from North America. Hyenas and early saber-toothed cats appeared, joining other predators including dogs, bears, and weasels. Africa

4186-650: The Sun during the Southern Hemisphere summer. Such effects are more pronounced when the eccentricity of the orbit is large. When the obliquity is large, seasonal changes are more extreme. Interglacials are a useful tool for geological mapping and for anthropologists, as they can be used as a dating method for hominid fossils. Brief periods of milder climate that occurred during the last glacial are called interstadials . Most, but not all, interstadials are shorter than interglacials. Interstadial climates may have been relatively warm, but not necessarily. Because

4277-546: The Western Atlantic, assemblages of bivalves exhibited remarkable stasis with regards to their basal metabolic rates throughout the various climatic changes of the Pliocene. The Pliocene was a high water mark for species diversity among Caribbean corals. From 5 to 2 Ma, coral species origination rates were relatively high in the Caribbean, although a noticeable extinction event and drop in diversity occurred at

4368-582: The Zanclean until around 3.5 Ma. During the Piacenzian, from about 3.5 to 2.6 Ma, the region was forested at irregular intervals and contained a significant Saharan palaeoriver until 3.35 Ma, when trade winds began to dominate over fluvial transport of pollen. Around 3.26 Ma, a strong aridification event that was followed by a return to more humid conditions, which was itself followed by another aridification around 2.7 Ma. From 2.6 to 2.4 Ma, vegetation zones began repeatedly shifting latitudinally in response to glacial-interglacial cycles. The climate of eastern Africa

4459-706: The appearance of Homo sapiens about 300,000 years ago. Artifacts associated with modern human behavior are unambiguously attested starting 40,000–50,000 years ago. According to mitochondrial timing techniques, modern humans migrated from Africa after the Riss glaciation in the Middle Palaeolithic during the Eemian Stage , spreading all over the ice-free world during the late Pleistocene. A 2005 study posits that humans in this migration interbred with archaic human forms already outside of Africa by

4550-527: The articles for those names. Both marine and continental faunas were essentially modern but with many more large land mammals such as Mammoths , Mastodons , Diprotodons , Smilodons , tigers , lions , Aurochs , short-faced bears , giant sloths , species within Gigantopithecus and others. Isolated landmasses such as Australia , Madagascar , New Zealand and islands in the Pacific saw

4641-735: The climate cooled. Venomous snake genera continued to increase as more rodents and birds evolved. Rattlesnakes first appeared in the Pliocene. The modern species Alligator mississippiensis , having evolved in the Miocene, continued into the Pliocene, except with a more northern range; specimens have been found in very late Miocene deposits of Tennessee . Giant tortoises still thrived in North America, with genera like Hesperotestudo . Madtsoid snakes were still present in Australia. The amphibian order Allocaudata became extinct. In

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4732-426: The colder periods (stadials) have often been very dry, wetter (not necessarily warmer) periods have been registered in the sedimentary record as interstadials as well. The oxygen isotope ratio obtained from seabed sediment core samples , a proxy for the average global temperature, is an important source of information for changes in Earth's climate. An interglacial optimum, or climatic optimum of an interglacial,

4823-501: The cooler Messinian . This increase was related to the 1.2 million year obliquity amplitude modulation cycle . By 3.3-3.0 Ma, during the Mid-Piacenzian Warm Period (mPWP), global average temperature was 2–3 °C higher than today, while carbon dioxide levels were the same as today (400 ppm). Global sea level was about 25 m higher, though its exact value is uncertain. The northern hemisphere ice sheet

4914-475: The early-mid Pliocene (5 Ma – 3 Ma), after three-million-year intervals with modern or glacial ice volume became longer and collapse occurs only at times when warmer global temperature coincide with strong austral summer insolation anomalies. Continents continued to drift , moving from positions possibly as far as 250 km from their present locations to positions only 70 km from their current locations. South America became linked to North America through

5005-570: The end of this interval. Oceans continued to be relatively warm during the Pliocene, though they continued cooling. The Arctic ice cap formed , drying the climate and increasing cool shallow currents in the North Atlantic. Deep cold currents flowed from the Antarctic. The formation of the Isthmus of Panama about 3.5 million years ago cut off the final remnant of what was once essentially a circum-equatorial current that had existed since

5096-618: The entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions. The effects of glaciation were global. Antarctica was ice-bound throughout the Pleistocene as well as the preceding Pliocene. The Andes were covered in the south by the Patagonian ice cap. There were glaciers in New Zealand and Tasmania . The current decaying glaciers of Mount Kenya , Mount Kilimanjaro , and

5187-782: The evolution of large birds and even reptiles such as the Elephant bird , moa , Haast's eagle , Quinkana , Megalania and Meiolania . The severe climatic changes during the Ice Age had major impacts on the fauna and flora. With each advance of the ice, large areas of the continents became depopulated, and plants and animals retreating southwards in front of the advancing glacier faced tremendous stress. The most severe stress resulted from drastic climatic changes, reduced living space, and curtailed food supply. A major extinction event of large mammals ( megafauna ), which included mammoths , mastodons , saber-toothed cats , glyptodons ,

5278-444: The extinction of most large-bodied animals in these regions. The aridification and cooling trends of the preceding Neogene were continued in the Pleistocene. The climate was strongly variable depending on the glacial cycle, with the sea levels being up to 120 metres (390 ft) lower than present at peak glaciation, allowing the connection of Asia and North America via Beringia and the covering of most of northern North America by

5369-549: The first time since the Cretaceous , with North American rodents and primates mixing with southern forms. Litopterns and the notoungulates , South American natives, were mostly wiped out, except for the macrauchenids and toxodonts , which managed to survive. Small weasel-like carnivorous mustelids , coatis and short-faced bears migrated from the north. Grazing glyptodonts , browsing giant ground sloths and smaller caviomorph rodents , pampatheres , and armadillos did

5460-414: The glacial range, which have their own glacial history depending on latitude, terrain and climate. There is a general correspondence between glacials in different regions. Investigators often interchange the names if the glacial geology of a region is in the process of being defined. However, it is generally incorrect to apply the name of a glacial in one region to another. For most of the 20th century, only

5551-450: The historical terminology was established. Corresponding to the terms glacial and interglacial, the terms pluvial and interpluvial are in use (Latin: pluvia , rain). A pluvial is a warmer period of increased rainfall; an interpluvial is of decreased rainfall. Formerly a pluvial was thought to correspond to a glacial in regions not iced, and in some cases it does. Rainfall is cyclical also. Pluvials and interpluvials are widespread. There

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5642-855: The ice sheets. Forests return to areas that once supported tundra vegetation. Interglacials are identified on land or in shallow epicontinental seas by their paleontology. Floral and faunal remains of species pointing to temperate climate and indicating a specific age are used to identify particular interglacials. Commonly used are mammalian and molluscan species, pollen and plant macro-remains (seeds and fruits). However, many other fossil remains may be helpful: insects, ostracods, foraminifera, diatoms, etc. Recently, ice cores and ocean sediment cores provide more quantitative and accurately-dated evidence for temperatures and total ice volumes. Interglacials and glacials coincide with cyclic changes in Earth's orbit . Three orbital variations contribute to interglacials. The first

5733-445: The immediately preceding Pliocene ("newer", from πλείων ( pleíōn , "more") and kainós ) and the immediately subsequent Holocene ("wholly new" or "entirely new", from ὅλος ( hólos , "whole") and kainós ) epoch , which extends to the present time. The Pleistocene has been dated from 2.580 million (±0.005) to 11,700 years BP with the end date expressed in radiocarbon years as 10,000 carbon-14 years BP. It covers most of

5824-413: The influence of seasonal variation. More recent research has emphasized the variability selection hypothesis, which proposes that variability in climate fostered development of hominin traits. Improved climate proxies show that the Pliocene climate of east Africa was highly variable, suggesting that adaptability to varying conditions was more important in driving hominin evolution than the steady pressure of

5915-622: The large amounts of material moved about by glaciers. Less common are cave deposits, travertines and volcanic deposits (lavas, ashes). Pleistocene marine deposits are found primarily in shallow marine basins mostly (but with important exceptions) in areas within a few tens of kilometres of the modern shoreline. In a few geologically active areas such as the Southern California coast, Pleistocene marine deposits may be found at elevations of several hundred metres. The modern continents were essentially at their present positions during

6006-408: The late Pleistocene, incorporating archaic human genetic material into the modern human gene pool. Pliocene The Pliocene ( / ˈ p l aɪ . ə s iː n , ˈ p l aɪ . oʊ -/ PLY -ə-seen, PLY -oh- ; also Pleiocene ) is the epoch in the geologic time scale that extends from 5.33 to 2.58 million years ago (Ma). It is the second and most recent epoch of

6097-467: The late Zanclean, Italy remained relatively warm and humid. Central Asia became more seasonal during the Pliocene, with colder, drier winters and wetter summers, which contributed to an increase in the abundance of C 4 plants across the region. In the Loess Plateau , δ13C values of occluded organic matter increased by 2.5% while those of pedogenic carbonate increased by 5% over the course of

6188-540: The latest period of repeated glaciation , up to and including the Younger Dryas cold spell. The end of the Younger Dryas has been dated to about 9700 BCE (11,700 calendar years BP). The end of the Younger Dryas is the official start of the current Holocene Epoch . Although it is considered an epoch, the Holocene is not significantly different from previous interglacial intervals within the Pleistocene. In

6279-470: The lower Palaeolithic they disappeared, and the only hominin species found in fossilic records is Homo erectus for much of the Pleistocene. Acheulean lithics appear along with Homo erectus , some 1.8 million years ago, replacing the more primitive Oldowan industry used by A. garhi and by the earliest species of Homo . The Middle Paleolithic saw more varied speciation within Homo , including

6370-478: The megafauna and migrated north. Late Pleistocene bighorn sheep were more slender and had longer legs than their descendants today. Scientists believe that the change in predator fauna after the late Pleistocene extinctions resulted in a change of body shape as the species adapted for increased power rather than speed. The extinctions hardly affected Africa but were especially severe in North America where native horses and camels were wiped out. In July 2018,

6461-754: The now-separated Atlantic Ocean. Africa 's collision with Europe formed the Mediterranean Sea , cutting off the remnants of the Tethys Ocean . The border between the Miocene and the Pliocene is also the time of the Messinian salinity crisis . During the Late Pliocene, the Himalayas became less active in their uplift, as evidenced by sedimentation changes in the Bengal Fan . The land bridge between Alaska and Siberia ( Beringia )

6552-512: The onset of continental glaciation in the Arctic, suggesting that vegetation changes in Australia during the Pliocene behaved similarly to during the Late Pleistocene and were likely characterised by comparable cycles of aridity and humidity. The equatorial Pacific Ocean sea surface temperature gradient was considerably lower than it is today. Mean sea surface temperatures in the east were substantially warmer than today but similar in

6643-400: The opposite, migrating to the north and thriving there. The marsupials remained the dominant Australian mammals, with herbivore forms including wombats and kangaroos , and the huge Diprotodon . Carnivorous marsupials continued hunting in the Pliocene, including dasyurids , the dog-like thylacine and cat-like Thylacoleo . The first rodents arrived in Australia. The modern platypus ,

6734-478: The period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology . The name is a combination of Ancient Greek πλεῖστος ( pleîstos ) 'most' and καινός ( kainós ; Latinized as cænus ) 'new'. At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by

6825-417: The ratio of O to O (two isotopes of oxygen ) by mass (measured by a mass spectrometer ) present in the calcite of oceanic core samples is used as a diagnostic of ancient ocean temperature change and therefore of climate change. Cold oceans are richer in O , which is included in the tests of the microorganisms ( foraminifera ) contributing the calcite. A more recent version of

6916-461: The recent period of repeated glaciations. The name Plio-Pleistocene has, in the past, been used to mean the last ice age. Formerly, the boundary between the two epochs was drawn at the time when the foraminiferal species Hyalinea baltica first appeared in the marine section at La Castella, Calabria, Italy. However, the revised definition of the Quaternary , by pushing back the start date of

7007-737: The reorganisation of the Indonesian Throughflow. In the south-central Andes , an arid period occurred from 6.1 to 5.2 Ma, with another occurring from 3.6 to 3.3 Ma. These arid periods are coincident with global cold periods, during which the position of the Southern Hemisphere westerlies shifted northward and disrupted the South American Low Level Jet, which brings moisture to southeastern South America. From around 3.8 Ma to about 3.3 Ma, North Africa experienced an extended humid period. In northwestern Africa, tropical forests extended up to Cape Blanc during

7098-414: The same time as the climatic optimum. The last six interglacials are: Hypothetical runaway greenhouse state Tropical temperatures may reach poles Global climate during an ice age Earth's surface entirely or nearly frozen over Pleistocene The Pleistocene ( / ˈ p l aɪ s t ə ˌ s iː n , - s t oʊ -/ PLY -stə-seen, -⁠stoh- ; referred to colloquially as

7189-407: The sampling process makes use of modern glacial ice cores. Although less rich in O than seawater, the snow that fell on the glacier year by year nevertheless contained O and O in a ratio that depended on the mean annual temperature. Temperature and climate change are cyclical when plotted on a graph of temperature versus time. Temperature coordinates are given in the form of

7280-474: The scene. Bears, dogs and weasels (originally from North America) joined cats, hyenas and civets as the African predators, forcing hyenas to adapt as specialized scavengers. Most mustelids in Africa declined as a result of increased competition from the new predators, although Enhydriodon omoensis remained an unusually successful terrestrial predator. South America was invaded by North American species for

7371-649: The sole factor responsible for the variations in climate since they explain neither the long-term cooling trend over the Plio-Pleistocene nor the millennial variations in the Greenland Ice Cores known as Dansgaard-Oeschger events and Heinrich events. Milankovitch pacing seems to best explain glaciation events with periodicity of 100,000, 40,000, and 20,000 years. Such a pattern seems to fit the information on climate change found in oxygen isotope cores. In oxygen isotope ratio analysis, variations in

7462-421: The study of cyclical climate changes. The glacials in the following tables show historical usages, are a simplification of a much more complex cycle of variation in climate and terrain, and are generally no longer used. These names have been abandoned in favour of numeric data because many of the correlations were found to be either inexact or incorrect and more than four major glacials have been recognised since

7553-400: The surrounding environment. Anthropologists tended to focus on intrinsic models while geologists and vertebrate paleontologists tended to put greater emphasis on habitats. Alternatives to the savanna hypothesis include the woodland/forest hypothesis, which emphasizes the evolution of hominins in closed habitats, or hypotheses emphasizing the influence of colder habitats at higher latitudes or

7644-648: The upper boundary). The proposed section is the North Greenland Ice Core Project ice core 75° 06' N 42° 18' W. The lower boundary of the Pleistocene Series is formally defined magnetostratigraphically as the base of the Matuyama (C2r) chronozone , isotopic stage 103. Above this point there are notable extinctions of the calcareous nannofossils : Discoaster pentaradiatus and Discoaster surculus . The Pleistocene covers

7735-505: The west. This condition has been described as a permanent El Niño state, or “El Padre.” Several mechanisms have been proposed for this pattern, including increased tropical cyclone activity . The extent of the West Antarctic Ice Sheet oscillated at the 40 kyr period of Earth's obliquity. Ice sheet collapse occurred when the global average temperature was 3 °C warmer than today and carbon dioxide concentration

7826-543: Was a series of glacials and interglacials, stadials and interstadials, mirroring periodic climate changes. The main factor at work in climate cycling is now believed to be Milankovitch cycles . These are periodic variations in regional and planetary solar radiation reaching the Earth caused by several repeating changes in the Earth's motion. The effects of Milankovitch cycles were enhanced by various positive feedbacks related to increases in atmospheric carbon dioxide concentrations and Earth's albedo. Milankovitch cycles cannot be

7917-511: Was at 400 ppmv. This resulted in open waters in the Ross Sea . Global sea-level fluctuation associated with ice-sheet collapse was probably up to 7 meters for the west Antarctic and 3 meters for the east Antarctic. Model simulations are consistent with reconstructed ice-sheet oscillations and suggest a progression from a smaller to a larger West Antarctic ice sheet in the last 5 million years. Intervals of ice sheet collapse were much more common in

8008-486: Was dominated by hoofed animals, and primates continued their evolution, with australopithecines (some of the first hominins ) and baboon-like monkeys such as the Dinopithecus appearing in the late Pliocene. Rodents were successful, and elephant populations increased. Cows and antelopes continued diversification and overtook pigs in numbers of species. Early giraffes appeared. Horses and modern rhinos came onto

8099-484: Was ephemeral before the onset of extensive glaciation over Greenland that occurred in the late Pliocene around 3 Ma. The formation of an Arctic ice cap is signaled by an abrupt shift in oxygen isotope ratios and ice-rafted cobbles in the North Atlantic and North Pacific Ocean beds. Mid-latitude glaciation was probably underway before the end of the epoch. The global cooling that occurred during

8190-452: Was first flooded near the start of the Pliocene, allowing marine organisms to spread between the Arctic and Pacific Oceans. The bridge would continue to be periodically flooded and restored thereafter. Pliocene marine formations are exposed in northeast Spain , southern California , New Zealand , and Italy . During the Pliocene parts of southern Norway and southern Sweden that had been near sea level rose. In Norway this rise elevated

8281-545: Was very similar to what it is today. Unexpectedly, the expansion of grasslands in eastern Africa during this epoch appears to have been decoupled from aridification and not caused by it, as evidenced by their asynchrony. Southwestern Australia hosted heathlands , shrublands , and woodlands with a greater species diversity compared to today during the Middle and Late Pliocene. Three different aridification events occurred around 2.90, 2.59, and 2.56 Ma, and may have been linked to

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