Entomology (from Ancient Greek ἔντομον (entomon) 'insect' and - λογία ( -logia ) 'study') is the scientific study of insects , a branch of zoology . In the past the term insect was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arachnids , myriapods , and crustaceans . This wider meaning may still be encountered in informal use. The field is also referred to as insectology in American English, while in British English insectology implies the study of the relationships between insects and humans.
84-492: This glossary of entomology describes terms used in the formal study of insect species by entomologists . [REDACTED] The dictionary definition of thesaurus:insect#See also at Wiktionary Entomology Like several of the other fields that are categorized within zoology , entomology is a taxon -based category; any form of scientific study in which there is a focus on insect-related inquiries is, by definition, entomology. Entomology, therefore, overlaps with
168-442: A common ancestor is now generally accepted and considered a fundamental scientific concept. In a joint publication with Alfred Russel Wallace , he introduced his scientific theory that this branching pattern of evolution resulted from a process he called natural selection , in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding . Darwin has been described as one of
252-451: A genealogical branching of a single evolutionary tree , in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent lineages progressing to higher forms. While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his Journal , he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained
336-562: A 230-page "Essay", to be expanded with his research results if he died prematurely. In November, the anonymously published sensational best-seller Vestiges of the Natural History of Creation brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists. Darwin completed his third geological book in 1846. He now renewed
420-845: A Bachelor of Arts degree as the first step towards becoming an Anglican country parson . Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying. During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens ' Illustrations of British entomology (1829–1932). Through Fox, Darwin became
504-673: A Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle , a sum equivalent to about £115,000 in 2021. He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher. As the Victorian era began, Darwin pressed on with writing his Journal , and in August 1837 began correcting printer's proofs . As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable palpitation of
588-591: A beginning to each crop of species". Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection. To escape the pressures of London, the family moved to rural Down House in Kent in September. On 11 January 1844, Darwin mentioned his theorising to
672-547: A book on geology. When they reached Brazil, Darwin was delighted by the tropical forest , but detested the sight of slavery there , and disputed this issue with FitzRoy. The survey continued to the south in Patagonia . They stopped at Bahía Blanca , and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like
756-718: A break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a proglacial lake . Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry" . Advantages under "Marry" included "constant companion and
840-471: A calm spell. On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell 's Principles of Geology , which set out uniformitarian concepts of land slowly rising or falling over immense periods, and Darwin saw things Lyell's way, theorising and thinking of writing
924-562: A close friend and follower of botany professor John Stevens Henslow . He met other leading parson-naturalists who saw scientific work as religious natural theology , becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley 's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for
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#17327866976821008-538: A code of ethical behavior" (ESA Certification Corporation). Individuals who are planning to become Certified IPM Technicians (CITs), need to obtain at around 1-4 years of experience in pest management and successfully pass an exam, that is based on the information, that they are acquainted with (ESA Certification Corporation). Like in Public Health Entomology (PHE), those who want to become Certified IPM Technicians (CITs) also have to "agree to ascribe to
1092-499: A code of ethical behavior" (ESA Certification Corporation). These individuals must also be approved to use pesticides (ESA Certification Corporation). For those who plan on becoming Board Certified Entomologists (BCEs), individuals have to pass two exams and "agree to ascribe to a code of ethical behavior" (ESA Certification Corporation). As with this, they also have to fulfill a certain amount of educational requirements every 12 months (ESA Certification Corporation). Forensic entomology
1176-412: A copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during
1260-635: A cross-section of topics as diverse as molecular genetics , environmental archaeology , behavior , neuroscience , biomechanics , biochemistry , systematics , physiology , developmental biology , ecology , morphology , and paleontology . Over 1.3 million insect species have been described, more than two-thirds of all known species. Some insect species date back to around 400 million years ago. They have many kinds of interactions with humans and other forms of life on Earth. For example, species such as P. pyralis conduct bioluminescent reactions in their light-emitting organs, which have been
1344-596: A fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation . In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully 's Malvern spa and
1428-529: A fortnight mapping strata in Wales. After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth . He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS Beagle with captain Robert FitzRoy , a position for a gentleman rather than "a mere collector". The ship
1512-532: A freed black slave who had accompanied Charles Waterton in the South American rainforest . In Darwin's second year at the university, he joined the Plinian Society , a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant 's investigations of
1596-473: A friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time". Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation. He married Emma on 29 January 1839 and they were
1680-530: A giant capybara . The armour fragments were actually from Glyptodon , a huge armadillo-like creature, as Darwin had initially thought. These extinct creatures were related to living species in South America. In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into
1764-556: A giant version of the armour on local armadillos . From a jaw and tooth he identified the gigantic Megatherium , then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest. In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles. On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at
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#17327866976821848-506: A great name among the Naturalists of Europe". On 2 October 1836, Beagle anchored at Falmouth , Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to Cambridge to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be
1932-515: A minor role to natural selection, and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution. Darwin's scientific discovery is the unifying theory of the life sciences , explaining the diversity of life . Darwin was born in Shrewsbury , Shropshire, on 12 February 1809, at his family's home, The Mount . He
2016-539: A programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and a close friend of the writer Harriet Martineau , who promoted the Malthusianism that underpinned the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the radical implications of transmutation of species , promoted by Grant and younger surgeons influenced by Geoffroy . Transmutation
2100-591: A sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls . On the geologically new Galápagos Islands , Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia,
2184-517: A self-funded gentleman scientist , and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage. Charles Lyell eagerly met Darwin for
2268-579: A series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species . By the 1870s, the scientific community and a majority of the educated public had accepted evolution as a fact . However, many initially favoured competing explanations that gave only
2352-719: A similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected", thinking this comparison "a beautiful part of my theory". He later called his theory natural selection , an analogy with what he termed the "artificial selection" of selective breeding. On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in
2436-468: A time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species. Three Fuegians on board, who had been seized during
2520-411: Is a branch of forensic science that studies insects found on corpses or elsewhere around crime scenes. This includes studying the types of insects commonly found on cadavers , their life cycles, their presence in different environments, and how insect assemblages change with decomposition . Medical entomology is focused upon insects and arthropods that impact human health. Veterinary entomology
2604-434: Is included in this category, because many animal diseases can "jump species" and become a human health threat, for example, bovine encephalitis. Medical entomology also includes scientific research on the behavior, ecology, and epidemiology of arthropod disease vectors , and involves a tremendous outreach to the public, including local and state officials and other stake holders in the interest of public safety. Anthecology
Glossary of entomology terms - Misplaced Pages Continue
2688-471: Is known as a Malthusian catastrophe . Darwin was well-prepared to compare this to Augustin de Candolle 's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that
2772-428: Is the study of pollination and the relationships between flowers and their pollinators . It has received increasing attention in the interest of agriculture science amid the impacts of pollinator decline from human actions. Many entomologists specialize in a single order or even a family of insects, and a number of these subspecialties are given their own informal names, typically (but not always) derived from
2856-686: The Linnean Society of London . Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex , followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in
2940-554: The Narrative were resolved at the end of the month when FitzRoy accepted Broderip's advice to make it a separate volume, and Darwin began work on his Journal and Remarks . Darwin's first paper showed that the South American landmass was slowly rising, and with Lyell's enthusiastic backing he read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to
3024-586: The Royal Entomological Society in London in 1833, one of the earliest such societies in the world; earlier antecedents, such as the Aurelian society date back to the 1740s. In the late 19th century, the growth of agriculture, and colonial trade spawned the "era of economic entomology" which created the professional entomologist associated with the rise of the university and training in the field of biology. Entomology developed rapidly in
3108-602: The Royal Society 's Royal Medal, and it made his reputation as a biologist . Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before." In 1854, he became a Fellow of the Linnean Society of London , gaining postal access to its library. He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in
3192-942: The Zoological Society . The ornithologist John Gould soon announced that the Galápagos birds that Darwin had thought a mixture of blackbirds , " gros-beaks " and finches , were, in fact, twelve separate species of finches . On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas. Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage , who described God as
3276-521: The first Beagle voyage then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there
3360-519: The marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aborigines "good-humoured & pleasant", their numbers depleted by European settlement. FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising. FitzRoy began writing
3444-598: The ordinary degree. Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature , explaining adaptation as God acting through laws of nature . He read John Herschel 's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described
Glossary of entomology terms - Misplaced Pages Continue
3528-596: The "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones." This would result in the formation of new species. As he later wrote in his Autobiography : In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate
3612-500: The 19th and 20th centuries and was studied by large numbers of people, including such notable figures as Charles Darwin , Jean-Henri Fabre , Vladimir Nabokov , Karl von Frisch (winner of the 1973 Nobel Prize in Physiology or Medicine ), and twice Pulitzer Prize winner E. O. Wilson . There has also been a history of people becoming entomologists through museum curation and research assistance, such as Sophie Lutterlough at
3696-569: The Entomological Society of America have varying credential requirements. These different programs are known as Public Health Entomology (PHE), Certified IPM Technicians (CITs), and Board Certified Entomologists (BCEs) (ESA Certification Corporation). To be qualified in public health entomology (PHE), one must pass an exam on the types of arthropods that can spread diseases and lead to medical complications (ESA Certification Corporation). These individuals also have to "agree to ascribe to
3780-653: The Entomological Society of America launched a new professional certification program for the pest control industry called the Associate Certified Entomologist (ACE). To qualify as a "true entomologist" an individual would normally require an advanced degree, with most entomologists pursuing a PhD. While not true entomologists in the traditional sense, individuals who attain the ACE certification may be referred to as ACEs or Associate Certified Entomologists. As such, other credential programs managed by
3864-530: The Smithsonian National Museum of Natural History . Insect identification is an increasingly common hobby, with butterflies and (to a lesser extent) dragonflies being the most popular. Most insects can easily be allocated to order , such as Hymenoptera (bees, wasps, and ants) or Coleoptera (beetles). However, identifying to genus or species is usually only possible through the use of identification keys and monographs . Because
3948-618: The Wedgwoods were adopting Anglicanism . Robert Darwin, a freethinker , had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury , but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had a taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending
4032-459: The afterlife. While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street , then moved his "museum" in over Christmas. On 24 January 1839, Darwin
4116-506: The anatomy and life cycle of marine invertebrates in the Firth of Forth , and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech . One day, Grant praised Lamarck 's evolutionary ideas . Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin
4200-437: The botanist Joseph Dalton Hooker , writing with melodramatic humour "it is like confessing a murder". Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject." By July, Darwin had expanded his "sketch" into
4284-424: The class Insecta contains a very large number of species (over 330,000 species of beetles alone) and the characteristics distinguishing them are unfamiliar, and often subtle (or invisible without a microscope), this is often very difficult even for a specialist. This has led to the development of automated species identification systems targeted on insects, for example, Daisy , ABIS, SPIDA and Draw-wing. In 1994,
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#17327866976824368-448: The collectors tended to be from the aristocracy, and there developed a trade involving collectors around the world and traders. This has been called the "era of heroic entomology". William Kirby is widely considered as the father of entomology in England. In collaboration with William Spence , he published a definitive entomological encyclopedia, Introduction to Entomology , regarded as the subject's foundational text. He also helped found
4452-542: The duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in selective breeding such as farmers and pigeon fanciers . Over time, his research drew on information from his relatives and children,
4536-713: The family butler, neighbours, colonists and former shipmates. He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour. The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils , palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of Darwin's illness remained unknown, and attempts at treatment had only ephemeral success. On 23 June, he took
4620-597: The first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen , who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium Darwin had identified, a near complete skeleton of the unknown Scelidotherium and a hippopotamus -sized rodent -like skull named Toxodon resembling
4704-582: The heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at Maer Hall , Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood , nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been
4788-472: The highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt 's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick 's geology course, then on 4 August travelled with him to spend
4872-681: The individual entitled to that name. No man can be truly called an entomologist, sir; the subject is too vast for any single human intelligence to grasp." — Oliver Wendell Holmes Sr. , The Poet at the Breakfast Table . Charles Darwin Charles Robert Darwin ( / ˈ d ɑːr w ɪ n / DAR -win ; 12 February 1809 – 19 April 1882) was an English naturalist , geologist , and biologist , widely known for his contributions to evolutionary biology . His proposition that all species of life have descended from
4956-412: The modern sense began only relatively recently, in the 16th century. Ulisse Aldrovandi 's De Animalibus Insectis (Concerning Insect Animals) was published in 1602. Microscopist Jan Swammerdam published History of Insects , correctly describing the reproductive organs of insects and metamorphosis . In 1705, Maria Sibylla Merian published the book Metamorphosis Insectorum Surinamensium about
5040-538: The most influential figures in human history and was honoured by burial in Westminster Abbey . Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh ; instead, he helped to investigate marine invertebrates . His studies at the University of Cambridge 's Christ's College from 1828 to 1831 encouraged his passion for natural science . However, it
5124-503: The nearby Anglican Shrewsbury School as a boarder . Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone ,
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#17327866976825208-433: The official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history. In Cape Town , South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries,
5292-496: The origin of species". Without telling Darwin, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society , and reported in magazines, including The Athenaeum . Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have
5376-522: The parents of ten children, seven of whom survived to adulthood. Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus 's An Essay on the Principle of Population . On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what
5460-573: The possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal Macrauchenia , which resembled a giant guanaco , a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations – explaining the variations he had observed in Galápagos tortoises , mockingbirds, and rheas. He sketched branching descent, and then
5544-520: The replacement of extinct species by others" as "a natural in contradistinction to a miraculous process". When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on
5628-448: The scientific name of the group: Like other scientific specialties, entomologists have a number of local, national, and international organizations. There are also many organizations specializing in specific subareas. Here is a list of selected very large insect collections, housed in museums, universities, or research institutes. "I suppose you are an entomologist?" "Not quite so ambitious as that, sir. I should like to put my eyes on
5712-408: The struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work... By mid-December, Darwin saw
5796-530: The subject of much research, especially in recent years. Entomology is rooted in nearly all human cultures from prehistoric times, primarily in the context of agriculture (especially biological control and beekeeping ). The natural Roman philosopher Pliny the Elder (23–79 CE) wrote a book on the kinds of insects, while the scientist of Kufa , Ibn al-A'rābī (760–845 CE) wrote a book on flies, Kitāb al-Dabāb ( كتاب الذباب ). However scientific study in
5880-409: The systematics of the taxa. He wrote his first examination of the species in 1846, but did not formally describe it until 1854. FitzRoy's long delayed Narrative was published in May 1839. Darwin's Journal and Remarks got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing
5964-465: The tropical insects of Dutch Surinam . Early entomological works associated with the naming and classification of species followed the practice of maintaining cabinets of curiosity , predominantly in Europe. This collecting fashion led to the formation of natural history societies, exhibitions of private collections, and journals for recording communications and the documentation of new species. Many of
6048-420: The voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations, and at intervals during the voyage his specimens were sent to Cambridge together with letters including
6132-421: The voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting the immediate joint submission of both their theories to
6216-403: The work of earthworms , inspiring "a new & important theory" on their role in soil formation , which Darwin presented at the Geological Society on 1 November 1837. His Journal was printed and ready for publication by the end of February 1838, as was the first volume of the Narrative , but FitzRoy was still working hard to finish his own volume. William Whewell pushed Darwin to take on
6300-707: Was elected a Fellow of the Royal Society (FRS). On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home. Darwin now had the framework of his theory of natural selection "by which to work", as his "prime hobby". His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory. For fifteen years this work
6384-418: Was a " wren " was in the finch group . Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands. The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards. By mid-March 1837, barely six months after his return to England, Darwin was speculating in his Red Notebook on
6468-472: Was anathema to Anglicans defending social order, but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species. Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought
6552-418: Was his five-year voyage on HMS Beagle from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported Charles Lyell 's concept of gradual geological change . Publication of his journal of the voyage made Darwin famous as a popular author. Puzzled by the geographical distribution of wildlife and fossils he collected on
6636-425: Was in the background to his main occupation of writing on geology and publishing expert reports on the Beagle collections, in particular, the barnacles . The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed Mr. Arthrobalanus . His confusion over the relationship of this species ( Cryptophialus minutus ) to other barnacles caused him to fixate on
6720-504: Was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England. Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel -beds stranded above high tide. High in the Andes he saw seashells, and several fossil trees that had grown on
6804-524: Was rather bored by Robert Jameson 's natural-history course, which covered geology – including the debate between neptunism and plutonism . He learned the classification of plants and assisted with work on the collections of the University Museum , one of the largest museums in Europe at the time. Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge , in January 1828, to study for
6888-574: Was surprised to find some benefit from hydrotherapy . Then, in 1851, his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises. In eight years of work on barnacles, Darwin's theory helped him to find " homologies " showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites , showing an intermediate stage in evolution of distinct sexes . In 1853, it earned him
6972-482: Was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists . Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794), a poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded. Both families were largely Unitarian , though
7056-446: Was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II , to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution. After delays,
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