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Charles Darwin

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138-532: Charles Robert Darwin ( / ˈ d ɑːr w ɪ n / DAR -win ; 12 February 1809 – 19 April 1882) was an English naturalist , geologist , and biologist , widely known for his contributions to evolutionary biology . His proposition that all species of life have descended from a common ancestor is now generally accepted and considered a fundamental scientific concept. In a joint publication with Alfred Russel Wallace , he introduced his scientific theory that this branching pattern of evolution resulted from

276-450: A genealogical branching of a single evolutionary tree , in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent lineages progressing to higher forms. While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his Journal , he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained

414-559: A 230-page "Essay", to be expanded with his research results if he died prematurely. In November, the anonymously published sensational best-seller Vestiges of the Natural History of Creation brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists. Darwin completed his third geological book in 1846. He now renewed

552-839: A Bachelor of Arts degree as the first step towards becoming an Anglican country parson . Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying. During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens ' Illustrations of British entomology (1829–1932). Through Fox, Darwin became

690-688: A Fellow of the Royal Society (FRS). On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home. Darwin now had the framework of his theory of natural selection "by which to work", as his "prime hobby". His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory. For fifteen years this work

828-670: A Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle , a sum equivalent to about £115,000 in 2021. He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher. As the Victorian era began, Darwin pressed on with writing his Journal , and in August 1837 began correcting printer's proofs . As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable palpitation of

966-587: A beginning to each crop of species". Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection. To escape the pressures of London, the family moved to rural Down House in Kent in September. On 11 January 1844, Darwin mentioned his theorising to

1104-544: A book on geology. When they reached Brazil, Darwin was delighted by the tropical forest , but detested the sight of slavery there , and disputed this issue with FitzRoy. The survey continued to the south in Patagonia . They stopped at Bahía Blanca , and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like

1242-717: A break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a proglacial lake . Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry" . Advantages under "Marry" included "constant companion and

1380-468: A calm spell. On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell 's Principles of Geology , which set out uniformitarian concepts of land slowly rising or falling over immense periods, and Darwin saw things Lyell's way, theorising and thinking of writing

1518-560: A close friend and follower of botany professor John Stevens Henslow . He met other leading parson-naturalists who saw scientific work as religious natural theology , becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley 's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for

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1656-410: A copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during

1794-497: A cultural continuity possibly going back as far as the 8th century. The centre has a largely undisturbed medieval street plan and over 660 listed buildings , including several examples of timber framing from the 15th and 16th centuries. Shrewsbury Castle , a red sandstone fortification, and Shrewsbury Abbey , were founded in 1074 and 1083 respectively by the Norman Earl of Shrewsbury , Roger de Montgomery . The town

1932-486: A dozen Irish prisoners were selected to be killed after picking lots. This prompted Rupert to respond by executing Parliamentarian prisoners in Oswestry . By the 18th century Shrewsbury had become an important market town and stopping point for stagecoaches travelling between London and Holyhead with passengers on their way to Ireland; this led to the establishment of a number of coaching inns, many of which, such as

2070-594: A fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation . In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully 's Malvern spa and

2208-413: A few miles north of the town centre, between King Henry IV and Henry Percy (Hotspur) , with the king emerging victorious. Hotspur's body was taken by Thomas Neville , to Whitchurch , for burial. However, when rumours circulated that Percy was still alive, the king "had the corpse exhumed and displayed it, propped upright between two millstones, in the market place at Shrewsbury". That being done, Percy

2346-527: A fortnight mapping strata in Wales. After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth . He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS  Beagle with captain Robert FitzRoy , a position for a gentleman rather than "a mere collector". The ship

2484-530: A freed black slave who had accompanied Charles Waterton in the South American rainforest . In Darwin's second year at the university, he joined the Plinian Society , a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant 's investigations of

2622-471: A friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time". Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation. He married Emma on 29 January 1839 and they were

2760-527: A giant capybara . The armour fragments were actually from Glyptodon , a huge armadillo-like creature, as Darwin had initially thought. These extinct creatures were related to living species in South America. In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into

2898-553: A giant version of the armour on local armadillos . From a jaw and tooth he identified the gigantic Megatherium , then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest. In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles. On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at

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3036-504: A great name among the Naturalists of Europe". On 2 October 1836, Beagle anchored at Falmouth , Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to Cambridge to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be

3174-565: A hillfort near Baschurch . The Historia Divae Monacellae , composed in the 14th or 15th century, says that Brochwel Ysgithrog , the 6th-century king of Powis, had a palace at Shrewsbury that became the site of the foundation of St Chad. In the late 8th century, it is said that Offa took the town for the Mercians in 778, and he is associated by some sources with establishing the town's first church and dedicating it to St Chad . If so, then there may have been an ecclesiastical foundation in

3312-605: A lock of the emperor's hair, presented to an officer of the 53rd, remains to this day in the collections of the Soldiers of Shropshire Museum at Shrewsbury Castle. HM Prison Shrewsbury , when new in the Georgian period, was considered a national example of improved conditions and more enlightened penal policy. Times change and a 2005 report on prison population found that it was the most overcrowded in England and Wales , despite

3450-408: A majority of the educated public had accepted evolution as a fact . However, many initially favoured competing explanations that gave only a minor role to natural selection, and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution. Darwin's scientific discovery

3588-568: A member of the Stanley family he was admitted. Thomas Mytton, the Bailiff of the town, a supporter of Richard III , had vowed that the only way he would get through was "over his dead body". Thomas then lay down and allowed Henry to step over him, to free himself from his oath. Henry was accommodated in the building now known as Henry Tudor House on Wyle Cop. In 1490, Henry VII, accompanied by his queen and his son, Prince Arthur , celebrated

3726-476: A naturalist, studies the world by observing plants and animals directly. Because organisms are functionally inseparable from the environment in which they live and because their structure and function cannot be adequately interpreted without knowing some of their evolutionary history, the study of natural history embraces the study of fossils as well as physiographic and other aspects of the physical environment". A common thread in many definitions of natural history

3864-556: A part of science proper. In Victorian Scotland, the study of natural history was believed to contribute to good mental health. Particularly in Britain and the United States, this grew into specialist hobbies such as the study of birds , butterflies, seashells ( malacology / conchology ), beetles, and wildflowers; meanwhile, scientists tried to define a unified discipline of biology (though with only partial success, at least until

4002-515: A plague in Viroconium around this time. It is claimed that Pengwern , sometime capital of the Kingdom of Powis (itself established by the 440s), was at Shrewsbury. The first attested association of Pengwern with Shrewsbury is mentioned by Giraldus Cambrensis in the 12th century. Alternative suggestions as to the location of Pengwern include Whittington Castle near Oswestry , and Berth,

4140-631: A poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded. Both families were largely Unitarian , though the Wedgwoods were adopting Anglicanism . Robert Darwin, a freethinker , had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury , but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had

4278-606: A process he called natural selection , in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding . Darwin has been described as one of the most influential figures in human history and was honoured by burial in Westminster Abbey . Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh ; instead, he helped to investigate marine invertebrates . His studies at

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4416-539: A programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and a close friend of the writer Harriet Martineau , who promoted the Malthusianism that underpinned the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the radical implications of transmutation of species , promoted by Grant and younger surgeons influenced by Geoffroy . Transmutation

4554-587: A sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls . On the geologically new Galápagos Islands , Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia,

4692-516: A self-funded gentleman scientist , and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage. Charles Lyell eagerly met Darwin for

4830-716: A similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected", thinking this comparison "a beautiful part of my theory". He later called his theory natural selection , an analogy with what he termed the "artificial selection" of selective breeding. On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in

4968-703: A taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending the nearby Anglican Shrewsbury School as a boarder . Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone ,

5106-464: A time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species. Three Fuegians on board, who had been seized during

5244-528: Is a market town , civil parish and the county town of Shropshire , England. It is sited on the River Severn , 33 miles (53 km) northwest of Wolverhampton , 15 miles (24 km) west of Telford , 31 miles (50 km) southeast of Wrexham and 53 miles (85 km) north of Hereford . At the 2021 census , the parish had a population of 76,782. Shrewsbury has Anglo-Saxon roots and institutions whose foundations dating from that time represent

5382-460: Is also echoed by H.W. Greene and J.B. Losos: "Natural history focuses on where organisms are and what they do in their environment, including interactions with other organisms. It encompasses changes in internal states insofar as they pertain to what organisms do". Some definitions go further, focusing on direct observation of organisms in their environments, both past and present, such as this one by G.A. Bartholomew: "A student of natural history, or

5520-471: Is known as a Malthusian catastrophe . Darwin was well-prepared to compare this to Augustin de Candolle 's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that

5658-400: Is most often defined as a type of observation and a subject of study, it can also be defined as a body of knowledge, and as a craft or a practice, in which the emphasis is placed more on the observer than on the observed. Definitions from biologists often focus on the scientific study of individual organisms in their environment, as seen in this definition by Marston Bates: "Natural history is

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5796-442: Is not limited to it. It involves the systematic study of any category of natural objects or organisms, so while it dates from studies in the ancient Greco-Roman world and the mediaeval Arabic world , through to European Renaissance naturalists working in near isolation, today's natural history is a cross-discipline umbrella of many specialty sciences; e.g., geobiology has a strong multidisciplinary nature. The meaning of

5934-501: Is owing to provision made by its head master, the excellent and worthy Thomas Ashton ", the school's first head master. During the English Civil War , Shrewsbury was a Royalist stronghold, under the command of Sir Francis Ottley . In the autumn of 1642 Charles I had a temporary base in the town. Prince Rupert established his headquarters in the town on 18 February 1644, being welcomed by Shrewsbury's aldermen. He

6072-497: Is the birthplace of Charles Darwin . It has had a role in nurturing aspects of English culture, including drama , ballet , dance and pantomime . Located 9 miles (14 km) east of the England–Wales border , Shrewsbury serves as the commercial centre for Shropshire and parts of mid-Wales, with a retail output of over £299 million per year and light industry and distribution centres, such as Battlefield Enterprise Park , on

6210-399: Is the inclusion of a descriptive component, as seen in a recent definition by H.W. Greene: "Descriptive ecology and ethology". Several authors have argued for a more expansive view of natural history, including S. Herman, who defines the field as "the scientific study of plants and animals in their natural environments. It is concerned with levels of organization from the individual organism to

6348-569: Is the unifying theory of the life sciences , explaining the diversity of life . Darwin was born in Shrewsbury , Shropshire, on 12 February 1809, at his family's home, The Mount . He was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists . Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794),

6486-550: The American Society of Naturalists and Polish Copernicus Society of Naturalists . Professional societies have recognized the importance of natural history and have initiated new sections in their journals specifically for natural history observations to support the discipline. These include "Natural History Field Notes" of Biotropica , "The Scientific Naturalist" of Ecology , "From the Field" of Waterbirds , and

6624-808: The Earls of Tankerville . St John the Baptist hospital passed to the Wood family and became almshouses. At this time the shrine and relics of St Winifred were destroyed. The Council of Wales and the Marches was established during the 1470s by Edward IV with a headquarters in the town. Its buildings partly survive near the castle and were later adapted to be an episcopal palace, the council also met at Ludlow Castle . Members of this council included John Dudley, Earl of Warwick , Sir Henry Sidney , William Herbert, 1st Earl of Pembroke , and Sir Rowland Hill , publisher of

6762-554: The Forest of Arden , a thickly wooded area, unpenetrated by Roman roads and somewhat dangerous in medieval times, so that travellers would pray at Coughton before entering. In 1102, Robert of Bellême, 3rd Earl of Shrewsbury was deposed and the title forfeited, as a consequence of him rebelling against Henry I and joining the Duke of Normandy 's invasion of England in 1101. William Pantulf , Lord of Wem , assisted Henry in putting down

6900-542: The Geneva Bible and potential inspiration for a hero in Shakespeare's As You Like It . Its functions were interpreted widely. It was to hear all suits, civil and criminal, which were brought by individuals too poor to sue at common law; it was to try all cases of murder, felony, piracy, wrecking and such crimes as were likely to disturb the peace; it was to investigate charges of misgovernment by officials and

7038-552: The Narrative were resolved at the end of the month when FitzRoy accepted Broderip's advice to make it a separate volume, and Darwin began work on his Journal and Remarks . Darwin's first paper showed that the South American landmass was slowly rising, and with Lyell's enthusiastic backing he read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to

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7176-486: The Natural History Society of Northumbria founded in 1829, London Natural History Society (1858), Birmingham Natural History Society (1859), British Entomological and Natural History Society founded in 1872, Glasgow Natural History Society, Manchester Microscopical and Natural History Society established in 1880, Whitby Naturalists' Club founded in 1913, Scarborough Field Naturalists' Society and

7314-531: The Renaissance , making it one of the longest-lasting of all natural history books. From the ancient Greeks until the work of Carl Linnaeus and other 18th-century naturalists, a major concept of natural history was the scala naturae or Great Chain of Being , an arrangement of minerals, vegetables, more primitive forms of animals, and more complex life forms on a linear scale of supposedly increasing perfection, culminating in our species. Natural history

7452-606: The Royal Society 's Royal Medal, and it made his reputation as a biologist . Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before." In 1854, he became a Fellow of the Linnean Society of London , gaining postal access to its library. He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in

7590-409: The University of Cambridge 's Christ's College from 1828 to 1831 encouraged his passion for natural science . However, it was his five-year voyage on HMS  Beagle from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported Charles Lyell 's concept of gradual geological change . Publication of his journal of

7728-940: The Zoological Society . The ornithologist John Gould soon announced that the Galápagos birds that Darwin had thought a mixture of blackbirds , " gros-beaks " and finches , were, in fact, twelve separate species of finches . On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas. Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage , who described God as

7866-519: The biological and geological sciences. The two were strongly associated. During the heyday of the gentleman scientists , many people contributed to both fields, and early papers in both were commonly read at professional science society meetings such as the Royal Society and the French Academy of Sciences —both founded during the 17th century. Natural history had been encouraged by practical motives, such as Linnaeus' aspiration to improve

8004-520: The first Beagle voyage then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there

8142-497: The humanities (primarily what is now known as classics ) and divinity , with science studied largely through texts rather than observation or experiment. The study of nature revived in the Renaissance , and quickly became a third branch of academic knowledge, itself divided into descriptive natural history and natural philosophy , the analytical study of nature. In modern terms, natural philosophy roughly corresponded to modern physics and chemistry , while natural history included

8280-516: The marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aborigines "good-humoured & pleasant", their numbers depleted by European settlement. FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising. FitzRoy began writing

8418-642: The modern evolutionary synthesis ). Still, the traditions of natural history continue to play a part in the study of biology, especially ecology (the study of natural systems involving living organisms and the inorganic components of the Earth's biosphere that support them), ethology (the scientific study of animal behavior), and evolutionary biology (the study of the relationships between life forms over very long periods of time), and re-emerges today as integrative organismal biology. Amateur collectors and natural history entrepreneurs played an important role in building

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8556-596: The ordinary degree. Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature , explaining adaptation as God acting through laws of nature . He read John Herschel 's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described

8694-802: The "Natural History Miscellany section" of the American Naturalist . Natural history observations have contributed to scientific questioning and theory formation. In recent times such observations contribute to how conservation priorities are determined. Mental health benefits can ensue, as well, from regular and active observation of chosen components of nature, and these reach beyond the benefits derived from passively walking through natural areas. Shrewsbury 52°42′29″N 2°45′14″W  /  52.708°N 2.754°W  / 52.708; -2.754 Shrewsbury ( / ˈ ʃ r oʊ z b ər i / SHROHZ -bər-ee , also / ˈ ʃ r uː z -/ SHROOZ - )

8832-592: The "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones." This would result in the formation of new species. As he later wrote in his Autobiography : In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate

8970-520: The 1220s, there was also a general hospital dedicated to St John the Baptist. In January 1234 Prince Llywelyn ab Iorwerth of Wales and Richard Marshal, 3rd Earl of Pembroke burned down the town and laid siege to its castle. In 1283, Edward I summoned a parliament in Shrewsbury, later adjourned to Acton Burnell , to try and condemn Dafydd ap Gruffydd , last of the native Princes of Wales, to execution by hanging, drawing and quartering within

9108-402: The 14th and 15th centuries and a new shrine for her relics was built in the late 1300s. Around this time the abbey illegally acquired the relics of St Beuno , uncle of St Winifred, by stealing them. As a result the abbey was fined but allowed to keep the relics. From 1155, during the reign of Henry II , there was a leper hospital dedicated to St Giles and associated with Shrewsbury Abbey. From

9246-684: The Bailiffs in a most royal and hospitable manner. Shrewsbury's monastic institutions were disbanded with the Dissolution of the Monasteries and the Abbey was closed in 1540. Henry VIII intended to make Shrewsbury one of his 13 new bishoprics (serving Shropshire and Staffordshire ) and hence a cathedral city, after the formation of the Church of England but the citizens of the town declined

9384-607: The Council House to St Chad’s Church, the choir of which was fitted up in imitation of St George's Chapel, Windsor Castle with the stalls decorated with the arms of the Knights of the Garter ; on the conclusion of divine service Sir Henry devoted the afternoon to feasting the burgesses. Shrewsbury thrived throughout the 16th and 17th centuries, a period when the town was at the height of its commercial importance. This success

9522-528: The Council: represented a remarkable experiment in regional government. It administered the law cheaply and rapidly; it dealt with up to twenty cases a day and George Owen stated that the 'oppressed poor' flocked to it. In 1551 there was a notable outbreak of sweating sickness in the town, which Dr John Caius was in the town to attend to at the command of the council. The following year, after his return to London, Caius published A Boke or Counseill Against

9660-551: The Disease Commonly Called the Sweate, or Sweatyng Sicknesse . The president of the council was the dedicatee of the book and the dedicatory epistle explains his appointment. This text became the main source of knowledge of this disease, now understood to be influenza . In 1581, Sir Henry Sidney, celebrated the feast of St George, on 23 April, in this town, with great splendour: a solemn procession went from

9798-620: The Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species . By the 1870s, the scientific community and

9936-819: The English term "natural history" (a calque of the Latin historia naturalis ) has narrowed progressively with time, while, by contrast, the meaning of the related term "nature" has widened (see also History below). In antiquity , "natural history" covered essentially anything connected with nature , or used materials drawn from nature, such as Pliny the Elder 's encyclopedia of this title , published c.  77 to 79 AD , which covers astronomy , geography , humans and their technology , medicine , and superstition , as well as animals and plants. Medieval European academics considered knowledge to have two main divisions:

10074-548: The Lion Hotel, are extant to this day. A town hall was built in the Market Place on the site of an ancient guildhall in 1730; it was demolished and a new combined guildhall and shirehall was erected on the site in 1837. Local soldier and statesman Robert Clive served as the town's mayor in 1762 and was Shrewsbury's MP from 1762 until his death in 1774. St Chad's Church collapsed in 1788 after attempts to expand

10212-670: The Natural History Institute (Prescott, Arizona): Natural history – a practice of intentional focused attentiveness and receptivity to the more-than-human world, guided by honesty and accuracy – is the oldest continuous human endeavor. In the evolutionary past of our species, the practice of natural history was essential for our survival, imparting critical information on habits and chronologies of plants and animals that we could eat or that could eat us. Natural history continues to be critical to human survival and thriving. It contributes to our fundamental understanding of how

10350-750: The Quarry. In the Industrial Revolution the Shrewsbury Canal opened in 1797, initially connecting the town to Trench . By 1835 it had been linked up to the Shropshire Canal and thence to the rest of the canal network . In the period directly after Napoleon 's surrender after the Battle of Waterloo , the town's own 53rd (Shropshire) Regiment of Foot was sent to guard him in his exile on St Helena . A locket containing

10488-556: The Sorby Natural History Society, Sheffield , founded in 1918. The growth of natural history societies was also spurred due to the growth of British colonies in tropical regions with numerous new species to be discovered. Many civil servants took an interest in their new surroundings, sending specimens back to museums in the Britain . (See also: Indian natural history ) Societies in other countries include

10626-471: The afterlife. While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street , then moved his "museum" in over Christmas. On 24 January 1839, Darwin was elected

10764-505: The anatomy and life cycle of marine invertebrates in the Firth of Forth , and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech . One day, Grant praised Lamarck 's evolutionary ideas . Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin

10902-434: The botanist Joseph Dalton Hooker , writing with melodramatic humour "it is like confessing a murder". Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject." By July, Darwin had expanded his "sketch" into

11040-410: The crypt compromised the structural integrity of the tower above. Now known as Old St Chad's, the remains of the church building and its churchyard are on the corner of Princess Street, College Hill and Belmont. A new St Chad's Church was built just four years after the collapse, but as a large neo-classical round church and in a different and more elevated location, at the top of Claremont Hill close to

11178-403: The diversity of the natural world. Natural history was understood by Pliny the Elder to cover anything that could be found in the world, including living things, geology, astronomy, technology, art, and humanity. De Materia Medica was written between 50 and 70 AD by Pedanius Dioscorides , a Roman physician of Greek origin. It was widely read for more than 1,500 years until supplanted in

11316-539: The duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in selective breeding such as farmers and pigeon fanciers . Over time, his research drew on information from his relatives and children,

11454-426: The economic condition of Sweden. Similarly, the Industrial Revolution prompted the development of geology to help find useful mineral deposits. Modern definitions of natural history come from a variety of fields and sources, and many of the modern definitions emphasize a particular aspect of the field, creating a plurality of definitions with a number of common themes among them. For example, while natural history

11592-422: The economy of nature". Natural history#Before 1900 Natural history is a domain of inquiry involving organisms , including animals , fungi , and plants , in their natural environment , leaning more towards observational than experimental methods of study. A person who studies natural history is called a naturalist or natural historian . Natural history encompasses scientific research but

11730-410: The ecosystem, and stresses identification, life history, distribution, abundance, and inter-relationships. It often and appropriately includes an esthetic component", and T. Fleischner, who defines the field even more broadly, as "A practice of intentional, focused attentiveness and receptivity to the more-than-human world, guided by honesty and accuracy". These definitions explicitly include the arts in

11868-710: The emergence of professional biological disciplines and research programs. Particularly in the 19th century, scientists began to use their natural history collections as teaching tools for advanced students and the basis for their own morphological research. The term "natural history" alone, or sometimes together with archaeology, forms the name of many national, regional, and local natural history societies that maintain records for animals (including birds (ornithology), insects ( entomology ) and mammals (mammalogy)), fungi ( mycology ), plants (botany), and other organisms. They may also have geological and microscopical sections. Examples of these societies in Britain include

12006-419: The false verdicts of juries; it was to enforce the laws against livery and maintenance, to punish rumour mongers and adulterers, and to deal with disputes concerning enclosures, villein service and manorial questions; it heard appeals from the common law courts; and it was responsible for administering the legislation dealing with religion. According to historian John Davies , at its peak in the mid-16th century,

12144-710: The family butler, neighbours, colonists and former shipmates. He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour. The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils , palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of Darwin's illness remained unknown, and attempts at treatment had only ephemeral success. On 23 June, he took

12282-467: The feast of St George in the town. The town is recorded as having entertained Henry again in 1496, with attendants lodged in the Sextry of Old St Chads; more generally it is said of Henry VII's relations with the town that: The intercourse which had begun thus favourably was kept up in after years by Henry, who, with his queen and son, frequently visited this town, upon which occasions they were feasted by

12420-411: The field of botany, be it as authors, collectors, or illustrators. In modern Europe, professional disciplines such as botany, geology, mycology , palaeontology , physiology , and zoology were formed. Natural history , formerly the main subject taught by college science professors, was increasingly scorned by scientists of a more specialized manner and relegated to an "amateur" activity, rather than

12558-638: The field of natural history, and are aligned with the broad definition outlined by B. Lopez, who defines the field as the "Patient interrogation of a landscape" while referring to the natural history knowledge of the Eskimo ( Inuit ). A slightly different framework for natural history, covering a similar range of themes, is also implied in the scope of work encompassed by many leading natural history museums , which often include elements of anthropology, geology, paleontology, and astronomy along with botany and zoology, or include both cultural and natural components of

12696-597: The first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen , who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium Darwin had identified, a near complete skeleton of the unknown Scelidotherium and a hippopotamus -sized rodent -like skull named Toxodon resembling

12834-632: The guilds of the Ironmongers and Goldsmiths . This fraternity were patrons of the Altar of St Michael in St Chad's Church and they kept a Mercers Hall on the site of the Sextry of Old St Chads . In 1485, ahead of the Battle of Bosworth Field , Henry Tudor , while not yet king, marched his forces on a route that lay through Shrewsbury. He was initially denied access to the town, but on intervention by

12972-582: The heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at Maer Hall , Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood , nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been

13110-471: The highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt 's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick 's geology course, then on 4 August travelled with him to spend

13248-471: The offer, which is the point of origin of the term "Proud Salopians": the town leadership preferring to be the most senior town in the country and not the most junior city. As a consequence of the dissolution, the monastic hospitals were closed and the incomes from their endowments were transferred to secular owners. St Giles's leper hospital passed to the Prince family, who were succeeded by their descendants

13386-430: The official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history. In Cape Town , South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries,

13524-492: The origin of species". Without telling Darwin, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society , and reported in magazines, including The Athenaeum . Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have

13662-631: The outskirts. The A5 and A49 trunk roads come together as the town's by-pass and five railway lines meet at Shrewsbury railway station . In Old English the settlement was known as Scrobbesburh ( dative Scrobbesbyrig ), which may mean either "Scrobb's fort" or "the fortified place in the bushes" (or "shrubs", the modern derivate). This name gradually evolved in three directions, into Sciropscire , which became Shropshire ; into Sloppesberie , which became Salop / Salopia (an alternative name for both town and county), and into Schrosberie , which eventually became

13800-522: The parents of ten children, seven of whom survived to adulthood. Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus 's An Essay on the Principle of Population . On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what

13938-573: The possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal Macrauchenia , which resembled a giant guanaco , a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations – explaining the variations he had observed in Galápagos tortoises , mockingbirds, and rheas. He sketched branching descent, and then

14076-443: The rebellion. To deal with the thickly wooded local forests, ideal for the concealment of archers, Pantulf brought in 6,000 foot soldiers to cut down trees and open up the roads. Henry subsequently took the government of the town into his own hands and in 1116 the nobility of England did homage to William Ætheling , Henry’s son, at Shrewsbury, and swore allegiance to his father. The early death of William Ætheling without issue led to

14214-518: The replacement of extinct species by others" as "a natural in contradistinction to a miraculous process". When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on

14352-517: The same idea, prompting the immediate joint submission of both their theories to the Linnean Society of London . Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex , followed by The Expression of

14490-569: The strategic importance of the Roman Road link via the Via Devana ). Viking raiders from the north were reaching as far south as Bridgnorth by 910. In the early 10th century, the relics of St Alkmund were translated to the town from Derby , this was probably the work of Æthelflæd. (Later, after St Alkmund's Church became the property of Lilleshall Abbey in about 1145, the relics were retranslated back to Derby.) Roger de Montgomery

14628-407: The struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work... By mid-December, Darwin saw

14766-546: The study of animals and Plants—of organisms. ... I like to think, then, of natural history as the study of life at the level of the individual—of what plants and animals do, how they react to each other and their environment, how they are organized into larger groupings like populations and communities" and this more recent definition by D.S. Wilcove and T. Eisner: "The close observation of organisms—their origins, their evolution, their behavior, and their relationships with other species". This focus on organisms in their environment

14904-621: The succession crisis, known in history as the Anarchy , and during this period, in 1138, King Stephen successfully besieged the town's castle held by William FitzAlan for the Empress Maud . In 1138 the relics of St Winifred were brought to Shrewsbury from Gwytheryn , following their purchase by the Abbot of Shrewsbury , the abbey being ready for consecration but having no relics prior to that time. The popularity of St Winifred grew in

15042-524: The system of the Swedish naturalist Carl Linnaeus . The British historian of Chinese science Joseph Needham calls Li Shizhen "the 'uncrowned king' of Chinese naturalists", and his Bencao gangmu "undoubtedly the greatest scientific achievement of the Ming". His works translated to many languages direct or influence many scholars and researchers. A significant contribution to English natural history

15180-408: The systematics of the taxa. He wrote his first examination of the species in 1846, but did not formally describe it until 1854. FitzRoy's long delayed Narrative was published in May 1839. Darwin's Journal and Remarks got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing

15318-557: The town after Dafydd was captured, ending his rebellion against the king. It is thought this parliament met in the Abbey. Shrewsbury was devastated by the Black Death , which, records suggest, arrived in the spring of 1349. Examining the number of local church benefices falling vacant due to death, 1349 alone saw twice as many vacancies as the previous ten years combined, suggesting a high death toll in Shrewsbury. "The Great Fire of Shrewsbury" took place in 1394: St Chad's church

15456-682: The town within a century of the death of Chad of Mercia . By the beginning of the 10th century, Shrewsbury was home to three moneyers who had permission to operate a mint in the town, using dies supplied by the royal government. In 914, Æthelflæd , daughter of Alfred the Great and known as the Lady of the Mercians, fortified Shrewsbury, along with Hereford and two other fortresses, at Scergeat (a currently unknown location) and Weardbyrig (thought to be Whitchurch , which would make sense given

15594-625: The town's name, Shrewsbury. Its later Welsh name Amwythig means "fortified place". The town's name can be pronounced as either 'Shrowsbury' or 'Shroosbury', the correct pronunciation being a matter of longstanding debate. Evidence of Neolithic occupation of a religious form dating back before 2,000 BC, was discovered in 2017 in the grounds of the medieval Church of the Holy Fathers in Sutton Farm , making it Britain's oldest place of worship. An Early Bronze Age urned burial

15732-470: The voyage made Darwin famous as a popular author. Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described

15870-418: The voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations, and at intervals during the voyage his specimens were sent to Cambridge together with letters including

16008-401: The work of earthworms , inspiring "a new & important theory" on their role in soil formation , which Darwin presented at the Geological Society on 1 November 1837. His Journal was printed and ready for publication by the end of February 1838, as was the first volume of the Narrative , but FitzRoy was still working hard to finish his own volume. William Whewell pushed Darwin to take on

16146-475: The world works by providing the empirical foundation of natural sciences, and it contributes directly and indirectly to human emotional and physical health, thereby fostering healthier human communities. It also serves as the basis for all conservation efforts, with natural history both informing the science and inspiring the values that drive these. As a precursor to Western science , natural history began with Aristotle and other ancient philosophers who analyzed

16284-687: The world's large natural history collections, such as the Natural History Museum, London , and the National Museum of Natural History in Washington, DC. Three of the greatest English naturalists of the 19th century, Henry Walter Bates , Charles Darwin , and Alfred Russel Wallace —who knew each other—each made natural history travels that took years, collected thousands of specimens, many of them new to science, and by their writings both advanced knowledge of "remote" parts of

16422-564: The world. The plurality of definitions for this field has been recognized as both a weakness and a strength, and a range of definitions has recently been offered by practitioners in a recent collection of views on natural history. Prior to the advent of Western science humans were engaged and highly competent in indigenous ways of understanding the more-than-human world that are now referred to as traditional ecological knowledge . 21st century definitions of natural history are inclusive of this understanding, such as this by Thomas Fleischner of

16560-799: The world—the Amazon basin , the Galápagos Islands , and the Indonesian Archipelago , among others—and in so doing helped to transform biology from a descriptive to a theory-based science. The understanding of "Nature" as "an organism and not as a mechanism" can be traced to the writings of Alexander von Humboldt (Prussia, 1769–1859). Humboldt's copious writings and research were seminal influences for Charles Darwin, Simón Bolívar , Henry David Thoreau , Ernst Haeckel , and John Muir . Natural history museums , which evolved from cabinets of curiosities , played an important role in

16698-416: Was a " wren " was in the finch group . Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands. The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards. By mid-March 1837, barely six months after his return to England, Darwin was speculating in his Red Notebook on

16836-470: Was anathema to Anglicans defending social order, but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species. Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought

16974-801: Was basically static through the Middle Ages in Europe—although in the Arabic and Oriental world, it proceeded at a much brisker pace. From the 13th century, the work of Aristotle was adapted rather rigidly into Christian philosophy , particularly by Thomas Aquinas , forming the basis for natural theology . During the Renaissance, scholars (herbalists and humanists, particularly) returned to direct observation of plants and animals for natural history, and many began to accumulate large collections of exotic specimens and unusual monsters . Leonhart Fuchs

17112-586: Was billeted in a building then the home of the family of Thomas Jones in the precincts of what is now the Prince Rupert Hotel. Shrewsbury only fell to Parliament forces after they were let in by a parliamentarian sympathiser at the St Mary's Water Gate (now also known as Traitor's Gate). After Thomas Mytton captured Shrewsbury in February 1645; in following with the ordnance of no quarter ;

17250-424: Was consumed by an accidental fire, which spread to a great portion of the town, then chiefly consisting of timber houses with thatched roofs. The damage was so considerable that Richard II remitted the town's taxes for three years towards the repairs. In 1398, Richard summoned a Great Parliament in the town, which is believed to have met in the Abbey. In 1403 the Battle of Shrewsbury was fought at Battlefield ,

17388-471: Was excavated at Crowmeole in 2015. An Iron Age double ring ditch has been excavated at Meole Brace . Amongst other finds, parts of an iron age sword and scabbard were recovered. At Meole Brace, an extensive roadside settlement along the line of the Roman military road connecting Viroconium Cornoviorum and Caersws was uncovered, with evidence of trading of amphorae and mortaria . A major discovery

17526-573: Was given the town as a gift from William the Conqueror and took the title of Earl of Shrewsbury. He built at Shrewsbury Castle in 1074, though archaeological excavations at the site of Shrewsbury castle in 2019 have indicated that the location may have been a fortified site in the time of the Anglo-Saxons. He also founded Shrewsbury Abbey as a Benedictine monastery in 1083. The town's position just off Watling Street placed it within

17664-424: Was in the background to his main occupation of writing on geology and publishing expert reports on the Beagle collections, in particular, the barnacles . The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed Mr. Arthrobalanus . His confusion over the relationship of this species ( Cryptophialus minutus ) to other barnacles caused him to fixate on

17802-447: Was in this period that Edward VI gave permission for the foundation of a free school, which was later to become Shrewsbury School . Later, William Camden , in his Britannia (begun 1577), remarked of the town that "Shrewsbury is inhabited both by Welsh and English, who speak each other's language; and among other things greatly to their praise is the grammar school founded by them, the best filled in all England, whose flourishing state

17940-688: Was largely due to the town's location, which allowed it to control the Welsh wool trade , a major industry at the time, with the rest of Britain and Europe, with the River Severn and Watling Street acting as trading routes. This trade was dominated by the Shrewsbury Drapers Company for many years. As a result, a number of grand edifices, including the Ireland's Mansion (built 1575) and Draper's Hall (1658), were constructed. It

18078-402: Was made by parson-naturalists such as Gilbert White , William Kirby , John George Wood , and John Ray , who wrote about plants, animals, and other aspects of nature. Many of these men wrote about nature to make the natural theology argument for the existence or goodness of God. Since early modern times, however, a great number of women made contributions to natural history, particularly in

18216-502: Was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England. Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel -beds stranded above high tide. High in the Andes he saw seashells, and several fossil trees that had grown on

18354-405: Was one of the three founding fathers of botany, along with Otto Brunfels and Hieronymus Bock . Other important contributors to the field were Valerius Cordus , Konrad Gesner ( Historiae animalium ), Frederik Ruysch , and Gaspard Bauhin . The rapid increase in the number of known organisms prompted many attempts at classifying and organizing species into taxonomic groups , culminating in

18492-576: Was rather bored by Robert Jameson 's natural-history course, which covered geology – including the debate between neptunism and plutonism . He learned the classification of plants and assisted with work on the collections of the University Museum , one of the largest museums in Europe at the time. Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge , in January 1828, to study for

18630-660: Was subjected to posthumous execution . One of the Princes in the Tower , Richard of Shrewsbury , was born in the town around 17 August 1473, the second son of King Edward IV of England and Elizabeth Woodville . In 1480, Edward V , then Prince of Wales (and the other prince of the Princes in the Tower), was resident in Shrewsbury. On 11 May Edward V confirmed the composition of the town's Mercer's Company , which had merged with

18768-571: Was surprised to find some benefit from hydrotherapy . Then, in 1851, his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises. In eight years of work on barnacles, Darwin's theory helped him to find " homologies " showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites , showing an intermediate stage in evolution of distinct sexes . In 1853, it earned him

18906-547: Was the finding of the Shrewsbury Hoard of more than 9000 Roman coins in a field near the town in 2009. Prior to the late 8th century, there is little in the way of reliable records. There is a tradition that the town was "founded in the 5th century, on occasion of the decay of the Roman Uriconium ." Historian John Wacher suggests that Shrewsbury may have been refortified by refugees fleeing an outbreak of

19044-442: Was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II , to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution. After delays,

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