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Island ecology

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Island ecology is the study of island organisms and their interactions with each other and the environment. Islands account for nearly 1/6 of earth’s total land area, yet the ecology of island ecosystems is vastly different from that of mainland communities. Their isolation and high availability of empty niches lead to increased speciation . As a result, island ecosystems comprise 30% of the world’s biodiversity hotspots , 50% of marine tropical diversity, and some of the most unusual and rare species. Many species still remain unknown.

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135-672: The diversity of species on islands is highly impacted by human activities such as deforestation and introduction of the exotic species . In response, ecologists and managers are directing attention towards conservation and restoration of island species. Because they are simple systems, islands provide an opportunity to study processes of extinction that can be extrapolated to larger ecosystems. Islands are attractive sites for ecological research because they provide clear examples of evolution in action. They show interesting patterns of colonization, adaptation, and speciation. Islands are surrounded by water, and may or may not exist as part of

270-451: A genealogical branching of a single evolutionary tree , in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent lineages progressing to higher forms. While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his Journal , he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained

405-562: A 230-page "Essay", to be expanded with his research results if he died prematurely. In November, the anonymously published sensational best-seller Vestiges of the Natural History of Creation brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists. Darwin completed his third geological book in 1846. He now renewed

540-833: A 3.2% rise in global deforestation. Massive wildfires in Canada , exacerbated by climate change , contributed to a 24% increase in global tree cover loss, highlighting the ongoing threats to forests essential for carbon storage and biodiversity . Despite some progress, the overall trends in forest destruction and climate impacts remain off track. The IPCC Sixth Assessment Report stated in 2022: “Over 420 million ha of forest were lost to deforestation from 1990 to 2020; more than 90% of that loss took place in tropical areas (high confidence), threatening biodiversity, environmental services, livelihoods of forest communities and resilience to climate shocks (high confidence).” See also: Global deforestation sharply accelerated around 1852. As of 1947,

675-845: A Bachelor of Arts degree as the first step towards becoming an Anglican country parson . Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying. During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens ' Illustrations of British entomology (1829–1932). Through Fox, Darwin became

810-673: A Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle , a sum equivalent to about £115,000 in 2021. He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher. As the Victorian era began, Darwin pressed on with writing his Journal , and in August 1837 began correcting printer's proofs . As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable palpitation of

945-591: A beginning to each crop of species". Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection. To escape the pressures of London, the family moved to rural Down House in Kent in September. On 11 January 1844, Darwin mentioned his theorising to

1080-547: A book on geology. When they reached Brazil, Darwin was delighted by the tropical forest , but detested the sight of slavery there , and disputed this issue with FitzRoy. The survey continued to the south in Patagonia . They stopped at Bahía Blanca , and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like

1215-718: A break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a proglacial lake . Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry" . Advantages under "Marry" included "constant companion and

1350-471: A calm spell. On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell 's Principles of Geology , which set out uniformitarian concepts of land slowly rising or falling over immense periods, and Darwin saw things Lyell's way, theorising and thinking of writing

1485-562: A close friend and follower of botany professor John Stevens Henslow . He met other leading parson-naturalists who saw scientific work as religious natural theology , becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley 's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for

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1620-682: A continental land mass. Oceanic islands arise due to volcanic activity or reef growth, and usually subside over time due to erosion and changing sea levels. When islands emerge, they undergo the process of ecological succession as species colonize the island (see theory of island biogeography ). New species cannot immigrate via land, and instead must arrive via air, water, or wind. As a result, organisms with high dispersal capabilities, such as plants and birds, are much more common on islands than are poorly dispersing taxa like mammals. However, some mammals are present on islands, presumably from swimming or riding on natural “rafts” that are washed away from

1755-412: A copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during

1890-596: A fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation . In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully 's Malvern spa and

2025-529: A fortnight mapping strata in Wales. After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth . He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS  Beagle with captain Robert FitzRoy , a position for a gentleman rather than "a mere collector". The ship

2160-532: A freed black slave who had accompanied Charles Waterton in the South American rainforest . In Darwin's second year at the university, he joined the Plinian Society , a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant 's investigations of

2295-473: A friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time". Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation. He married Emma on 29 January 1839 and they were

2430-530: A giant capybara . The armour fragments were actually from Glyptodon , a huge armadillo-like creature, as Darwin had initially thought. These extinct creatures were related to living species in South America. In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into

2565-556: A giant version of the armour on local armadillos . From a jaw and tooth he identified the gigantic Megatherium , then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest. In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles. On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at

2700-506: A great name among the Naturalists of Europe". On 2 October 1836, Beagle anchored at Falmouth , Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to Cambridge to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be

2835-410: A majority of the educated public had accepted evolution as a fact . However, many initially favoured competing explanations that gave only a minor role to natural selection, and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution. Darwin's scientific discovery

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2970-632: A poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded. Both families were largely Unitarian , though the Wedgwoods were adopting Anglicanism . Robert Darwin, a freethinker , had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury , but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had

3105-609: A process he called natural selection , in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding . Darwin has been described as one of the most influential figures in human history and was honoured by burial in Westminster Abbey . Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh ; instead, he helped to investigate marine invertebrates . His studies at

3240-539: A programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and a close friend of the writer Harriet Martineau , who promoted the Malthusianism that underpinned the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the radical implications of transmutation of species , promoted by Grant and younger surgeons influenced by Geoffroy . Transmutation

3375-591: A sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls . On the geologically new Galápagos Islands , Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia,

3510-517: A self-funded gentleman scientist , and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage. Charles Lyell eagerly met Darwin for

3645-719: A similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected", thinking this comparison "a beautiful part of my theory". He later called his theory natural selection , an analogy with what he termed the "artificial selection" of selective breeding. On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in

3780-524: A species is released from its ancestral community interactions and then colonizes new niches. In response to these changing ecological pressures, island species can become much more docile than their mainland counterparts, and may grow larger (see island gigantism ) or smaller (see island dwarfism ). Some of these unique adaptations are reflected in charismatic island species such as the Malagasy hippopotamus , Komodo dragon , or pygmy mammoths . Although,

3915-537: A state of change. As resource depletion is a major issue on islands, the needs of human populations must also be taken into account. On many islands, scientists and managers are studying traditional practices of indigenous populations as potential conservation solutions. In some cases, limited-take systems that serve the community may provide a better alternative to fully closed protected areas, if there are not enough resources for proper enforcement. Public education plays an important role. A significant advancement in

4050-705: A taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending the nearby Anglican Shrewsbury School as a boarder . Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone ,

4185-468: A time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species. Three Fuegians on board, who had been seized during

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4320-440: A whole is closely linked to restoration . Active restoration on islands can be done for both animal species (translocations, induced breeding) and plant species (reforestation). Creating goals for restoration can be challenging because it is often impossible to restore the ecosystem to its “historic” or “normal” state, if that state can even be clearly defined. Restoration is never complete, as ecological communities are always in

4455-407: Is agreement that destruction of rainforests remains a significant environmental problem. The rate of net forest loss declined from 7.8 million ha per year in the decade 1990–2000 to 5.2 million ha per year in 2000–2010 and 4.7 million ha per year in 2010–2020. The rate of decline of net forest loss slowed in the most recent decade due to a reduction in the rate of forest expansion. In many parts of

4590-636: Is agriculture by far. More than 80% of deforestation was attributed to agriculture in 2018. Forests are being converted to plantations for coffee , palm oil , rubber and various other popular products. Livestock grazing also drives deforestation. Further drivers are the wood industry ( logging ), urbanization and mining . The effects of climate change are another cause via the increased risk of wildfires (see deforestation and climate change ). Deforestation results in habitat destruction which in turn leads to biodiversity loss . Deforestation also leads to extinction of animals and plants, changes to

4725-407: Is also important for the agricultural sector. The reason for this linkage is because the effects of climate change on agriculture pose new risks to global food systems . Since 1990, it is estimated that some 420 million hectares of forest have been lost through conversion to other land uses , although the rate of deforestation has decreased over the past three decades. Between 2015 and 2020,

4860-758: Is around 38%. Since 1960, roughly 15% of the Amazon has been removed with the intention of replacing the land with agricultural practices. It is no coincidence that Brazil has recently become the world's largest beef exporter at the same time that the Amazon rainforest is being clear cut. Another prevalent method of agricultural deforestation is slash-and-burn agriculture , which was primarily used by subsistence farmers in tropical regions but has now become increasingly less sustainable. The method does not leave land for continuous agricultural production but instead cuts and burns small plots of forest land which are then converted into agricultural zones. The farmers then exploit

4995-687: Is especially threatened by logging, hunting, fishing and plant gathering. However, islands’ vulnerability to introduced species is also due to agricultural, economic, and health differences as compared to continental land (Russel et al.). The smaller land area and population sizes of islands compared to continents create greater vulnerability to the impacts of introduced species (Russel et al.). Introduced species negatively impact ecosystems through altered predator-prey interactions that can cause harm or even local extinction to native species populations (Towns et al.). There are many examples of animal species such as birds, reptiles, and aquatic insects being harmed by

5130-471: Is known as a Malthusian catastrophe . Darwin was well-prepared to compare this to Augustin de Candolle 's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that

5265-472: Is known for its ability to survive under harsh conditions, and it is highly adaptable, as it will eat a wide variety of organisms (Platenberg). Native frog and anole lizard populations have declined, likely due to Cuban tree frog interference (Platenberg). In contrast to the accidental introduction of the Cuban treefrog, the cane toad was deliberately introduced to control agricultural pests (Platenberg). Similar to

5400-605: Is not an issue because of longer fallow periods and lesser overall deforestation. The relatively small size of the plots allowed for no net input of CO 2 to be released. Consumption and production of beef is the primary driver of deforestation in the Amazon , with around 80% of all converted land being used to rear cattle. 91% of Amazon land deforested since 1970 has been converted to cattle ranching. Livestock ranching requires large portions of land to raise herds of animals and livestock crops for consumer needs. According to

5535-741: Is particularly troubling as yields of coral reef fish species are an important food source for island populations. Humans have contributed to globalization and decreased effective isolation of island communities, allowing for invasion of exotic species. This can have a profound effect on the native species. In Guam , the introduced brown tree snake ate nearly all of the native vertebrate species to extinction. Feral cats and dogs have also greatly diminished native vertebrate populations on islands, through both predation and disease. Introduced ungulates are another major threat, as they graze on native vegetation and can destroy entire forests. Exotic grasses can out-compete native understory species and increase

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5670-453: Is projected to occur by the mid-21st century. In the early 2000s, some scientists predicted that unless significant measures (such as seeking out and protecting old growth forests that have not been disturbed) are taken on a worldwide basis, by 2030 there will only be 10% remaining, with another 10% in a degraded condition . 80% will have been lost, and with them hundreds of thousands of irreplaceable species. Estimates vary widely as to

5805-727: Is responsible for 48% of deforestation; commercial agriculture is responsible for 32%; logging is responsible for 14%, and fuel wood removals make up 5%. More than 80% of deforestation was attributed to agriculture in 2018. Forests are being converted to plantations for coffee, tea, palm oil , rice, rubber , and various other popular products. The rising demand for certain products and global trade arrangements causes forest conversions , which ultimately leads to soil erosion . The top soil oftentimes erodes after forests are cleared which leads to sediment increase in rivers and streams. Most deforestation also occurs in tropical regions. The estimated amount of total land mass used by agriculture

5940-621: Is termed Island syndrome . After immigration, birds, and some reptiles or mammals, tend to become larger and predatory, showing relaxed intraspecific competition . For mammals, small species will increase in size and large species will decrease in size. This is referred to as the “island rule,” and is suggested to minimize energy expenditure. Other adaptations to life on islands include increased poikilothermy , relaxed anti-predator behavior, and reduced sexual selection in animals, and loss of herbivore defenses and reduced dispersal in plants. The formation of new islands and their isolation from

6075-399: Is the removal and destruction of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms , ranches , or urban use. About 31% of Earth's land surface is covered by forests at present. This is one-third less than the forest cover before the expansion of agriculture, with half of that loss occurring in

6210-571: Is the unifying theory of the life sciences , explaining the diversity of life . Darwin was born in Shrewsbury , Shropshire, on 12 February 1809, at his family's home, The Mount . He was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists . Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794),

6345-634: Is when a single species colonizes an area and rapidly diversifies to fill all of the available niches. A common example is the assemblage of finch species documented by Charles Darwin in the Galapagos Islands . Darwin’s finches exhibited adaptive radiation by evolving different beak sizes to exploit the diversity of seeds present on the different islands. Because the distributions of these populations are limited by their island habitats, they tend to have fewer individuals than their mainland counterparts and lower genetic variation. This, along with

6480-581: The California Channel Islands , where DDT poisoning reduced bald eagle populations. This, along with an abundance of introduced feral pigs for prey, allowed golden eagles to colonize the island and replace bald eagles. However, the golden eagles also ate native island foxes . Fox population levels decreased to near extinction, while skunk populations increased due to relaxed competition with foxes. Since island ecosystems are self-contained, it should be possible to mitigate many of

6615-560: The Congolese rainforest increased by 5%. The World Wildlife Fund 's ecoregion project catalogues habitat types throughout the world, including habitat loss such as deforestation, showing for example that even in the rich forests of parts of Canada such as the Mid-Continental Canadian forests of the prairie provinces half of the forest cover has been lost or altered. In 2011, Conservation International listed

6750-554: The Narrative were resolved at the end of the month when FitzRoy accepted Broderip's advice to make it a separate volume, and Darwin began work on his Journal and Remarks . Darwin's first paper showed that the South American landmass was slowly rising, and with Lyell's enthusiastic backing he read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to

6885-730: The Philippines , Indonesia , Thailand , Burma , Malaysia , Bangladesh , China, Sri Lanka , Laos , Nigeria , the Democratic Republic of the Congo , Liberia , Guinea , Ghana and the Ivory Coast , have lost large areas of their rainforest. Much of what remains of the world's rainforests is in the Amazon basin , where the Amazon Rainforest covers approximately 4 million square kilometres. Some 80% of

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7020-610: The Royal Society 's Royal Medal, and it made his reputation as a biologist . Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before." In 1854, he became a Fellow of the Linnean Society of London , gaining postal access to its library. He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in

7155-410: The University of Cambridge 's Christ's College from 1828 to 1831 encouraged his passion for natural science . However, it was his five-year voyage on HMS  Beagle from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported Charles Lyell 's concept of gradual geological change . Publication of his journal of

7290-479: The World Wildlife Fund , "Extensive cattle ranching is the number one culprit of deforestation in virtually every Amazon country, and it accounts for 80% of current deforestation." The cattle industry is responsible for a significant amount of methane emissions since 60% of all mammals on earth are livestock cows. Replacing forest land with pastures creates a loss of forest stock , which leads to

7425-942: The Zoological Society . The ornithologist John Gould soon announced that the Galápagos birds that Darwin had thought a mixture of blackbirds , " gros-beaks " and finches , were, in fact, twelve separate species of finches . On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas. Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage , who described God as

7560-521: The first Beagle voyage then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there

7695-519: The marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aborigines "good-humoured & pleasant", their numbers depleted by European settlement. FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising. FitzRoy began writing

7830-598: The ordinary degree. Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature , explaining adaptation as God acting through laws of nature . He read John Herschel 's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described

7965-596: The "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones." This would result in the formation of new species. As he later wrote in his Autobiography : In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate

8100-424: The Cuban treefrog, the cane toad is also omnivorous. Cane toad tadpoles compete for limited freshwater resources with the white-lipped frog, a native species, thus limiting their resources (Platenberg). In general, when the introduction of a nonnative species results in extinction, the ecosystem experiences losses in some trophic levels (Platenberg). For example, vertebrate herbivores that are prone to extinction change

8235-622: The Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species . By the 1870s, the scientific community and

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8370-500: The Southern Ocean Islands, indirect effects such as invasive species and global warming can play a greater role in influencing populations than overexploitation , pollution and habitat loss . Human activities and the introduction of non-native species often cause trophic cascades , where direct effects on one species result in indirect effects on other species in the food web. An example is on Santa Cruz Island of

8505-459: The afterlife. While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street , then moved his "museum" in over Christmas. On 24 January 1839, Darwin

8640-506: The anatomy and life cycle of marine invertebrates in the Firth of Forth , and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech . One day, Grant praised Lamarck 's evolutionary ideas . Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin

8775-431: The behavioral and ecological factors mentioned above, makes island species more vulnerable to extinction. The continued survival of species on islands depends on factors such as natural selection , genetic variation, natural disturbances (hurricanes, volcanic eruptions) and human-caused disturbances (introduced species, habitat loss ). Human-caused disturbances tend to be the greatest cause of mortality, and understanding

8910-437: The botanist Joseph Dalton Hooker , writing with melodramatic humour "it is like confessing a murder". Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject." By July, Darwin had expanded his "sketch" into

9045-424: The causes of extinction facilitates conservation efforts. The movement of humans to islands has led to rapid extinction of native island species either from hunting, habitat destruction , or introduced species . Many large animals on islands have been hunted to extinction by humans. A well-known example is the dodo , once found on the island of Mauritius . It evolved to become large, flightless and docile, and

9180-642: The commodity is consumed. For example, consumption patterns in G7 countries are estimated to cause an average loss of 3.9 trees per person per year. In other words, deforestation can be directly related to imports—for example, coffee. In 2023, the Global Forest Watch reported a 9% decline in tropical primary forest loss compared to the previous year, with significant regional reductions in Brazil and Colombia overshadowed by increases elsewhere, leading to

9315-536: The community (Towns). Even if the number of species introduced to an island is roughly the same as the number of extinct ones, the ecological traits are not comparable enough to make up for losses (Towns). Global warming is emerging as a strong cause of species loss on islands. This can be due to sea level rise , the intrusion of salt water into freshwater habitats, or species inability to adapt to increasing temperatures and extreme weather events. Plant species are particularly susceptible. In more isolated areas, such as

9450-407: The conversion of forest to other land uses (regardless of whether it is human-induced). Deforestation and forest area net change are not the same: the latter is the sum of all forest losses (deforestation) and all forest gains (forest expansion) in a given period. Net change, therefore, can be positive or negative, depending on whether gains exceed losses, or vice versa. The FAO estimates that

9585-478: The country's western regions at a rate of 15,000 hectares at a randomly studied 2-month period in 2010. In 2009, Paraguay's parliament refused to pass a law that would have stopped cutting of natural forests altogether. As of 2007, less than 50% of Haiti's forests remained . From 2015 to 2019, the rate of deforestation in the Democratic Republic of the Congo doubled. In 2021, deforestation of

9720-780: The decades 1990–2000 and 2000–2010. Some claim that rainforests are being destroyed at an ever-quickening pace. The London-based Rainforest Foundation notes that "the UN figure is based on a definition of forest as being an area with as little as 10% actual tree cover, which would therefore include areas that are actually savanna-like ecosystems and badly damaged forests". Other critics of the FAO data point out that they do not distinguish between forest types, and that they are based largely on reporting from forestry departments of individual countries, which do not take into account unofficial activities like illegal logging. Despite these uncertainties, there

9855-498: The deforestation of the Amazon can be attributed to cattle ranching, as Brazil is the largest exporter of beef in the world. The Amazon region has become one of the largest cattle ranching territories in the world. The regions with the highest tropical deforestation rate between 2000 and 2005 were Central America —which lost 1.3% of its forests each year—and tropical Asia. In Central America , two-thirds of lowland tropical forests have been turned into pasture since 1950 and 40% of all

9990-542: The duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in selective breeding such as farmers and pigeon fanciers . Over time, his research drew on information from his relatives and children,

10125-623: The ecosystem depend on such as seabirds, can also have significant effects (Towns). Seabirds are essential to the structure of the ecosystem because they transport large amounts of nutrients into ecosystems and burrow into soils, helping vegetation (Towns). However, since human settlement, seabird communities have been severely impacted on islands across the globe (Towns). While some exotic species may perform ecological roles similar to those of extinct species, there are many roles that cannot be fulfilled by other species. Therefore, similar exotic species do not offer complete replacement for extinct species in

10260-548: The ecosystem function in plant communities. This phenomenon is seen in New Zealand, where the loss of bird species may have changed dynamics in avian-induced vegetation communities and impacted abundances of forest plants (Platenberg). Also in New Zealand, small mammalian predators such as rodents have little direct effect on vegetation but have greater effect on island faunas through extinction and displacement (Wood et al.). Pacific rats, for example, were thought to be causing

10395-463: The extent of deforestation in the tropics. In 2019, the world lost nearly 12 million hectares of tree cover. Nearly a third of that loss, 3.8 million hectares, occurred within humid tropical primary forests, areas of mature rainforest that are especially important for biodiversity and carbon storage. This is equivalent to losing an area of primary forest the size of a football pitch every six seconds. A 2002 analysis of satellite imagery suggested that

10530-713: The family butler, neighbours, colonists and former shipmates. He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour. The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils , palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of Darwin's illness remained unknown, and attempts at treatment had only ephemeral success. On 23 June, he took

10665-528: The field is the shift from 2-D to 3-D science in ecology. This approach offers a comprehensive understanding of ecological dynamics and human impacts on both terrestrial and marine environments. The adoption of 3-D models supports better natural resource management and informs conservation and management plans. This transition also affects how policymakers evaluate regulations, providing a more dynamic perspective on ecological and environmental challenges. Deforestation Deforestation or forest clearance

10800-597: The first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen , who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium Darwin had identified, a near complete skeleton of the unknown Scelidotherium and a hippopotamus -sized rodent -like skull named Toxodon resembling

10935-432: The form of charcoal or timber ), while cleared land is used as pasture for livestock and agricultural crops. The vast majority of agricultural activity resulting in deforestation is subsidized by government tax revenue . Disregard of ascribed value, lax forest management , and deficient environmental laws are some of the factors that lead to large-scale deforestation. The types of drivers vary greatly depending on

11070-717: The giant tortoises of the Galápagos Islands and the Seychelles (the Galápagos tortoise and Aldabrachelys respectively) are sometimes given as examples of insular gigantism, they are now thought to represent the last remaining populations of historically widespread giant tortoises i.e. gigantism is an ancestral trait that occurred in the absence of insular selection pressures. The collection of differences in morphology , ecology , physiology and behaviour of insular species compared to their continental counterparts

11205-568: The global forest carbon stock has decreased 0.9%, and tree cover 4.2% between 1990 and 2020. As of 2019 there is still disagreement about whether the global forest is shrinking or not: "While above-ground biomass carbon stocks are estimated to be declining in the tropics, they are increasing globally due to increasing stocks in temperate and boreal forest. Deforestation in many countries —both naturally occurring and human-induced —is an ongoing issue. Between 2000 and 2012, 2.3 million square kilometres (890,000 square miles) of forests around

11340-535: The growth of shearwater chicks, such as loss of tissue in their feet (Towns et al.). Other examples of high-impact species to island ecosystems are the Cuban treefrog and cane toad, which were introduced to the US Virgin Islands under various circumstances (Platenberg). The Cuban treefrog has been present in the USVI since arriving unintentionally on a cargo ship in the 1970s (Platenberg). The Cuban tree frog

11475-582: The heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at Maer Hall , Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood , nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been

11610-472: The highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt 's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick 's geology course, then on 4 August travelled with him to spend

11745-402: The implication of increased greenhouse gas emissions by burning agriculture methodologies and land-use change . A large contributing factor to deforestation is the lumber industry . A total of almost 4 million hectares (9.9 million acres) of timber, or about 1.3% of all forest land, is harvested each year. In addition, the increasing demand for low-cost timber products only supports

11880-478: The introduction of predators such as rats, cats, and ants. For example, seabirds on islands in Hawaii are impacted by non-native predators like barn owls and ants. At a colony in Hawaii, at least 20% of wedge-tailed shearwater eggs were taken by mynas, dark-plumaged birds of the starling family (Towns et al.). Also in Hawaii, more than forty four introduced ant species, especially tropical fire ants, caused detriments to

12015-639: The largest annual rate of net forest loss in 2010–2020, at 3.9 million ha, followed by South America, at 2.6 million ha. The rate of net forest loss has increased in Africa in each of the three decades since 1990. It has declined substantially in South America, however, to about half the rate in 2010–2020 compared with 2000–2010. Asia had the highest net gain of forest area in 2010–2020, followed by Oceania and Europe. Nevertheless, both Europe and Asia recorded substantially lower rates of net gain in 2010–2020 than in 2000–2010. Oceania experienced net losses of forest area in

12150-543: The last century. Between 15 million to 18 million hectares of forest, an area the size of Bangladesh , are destroyed every year. On average 2,400 trees are cut down each minute. Estimates vary widely as to the extent of deforestation in the tropics . In 2019, nearly a third of the overall tree cover loss, or 3.8 million hectares, occurred within humid tropical primary forests . These are areas of mature rainforest that are especially important for biodiversity and carbon storage . The direct cause of most deforestation

12285-483: The local climate, and displacement of indigenous people who live in forests. Deforested regions often also suffer from other environmental problems such as desertification and soil erosion . Another problem is that deforestation reduces the uptake of carbon dioxide ( carbon sequestration ) from the atmosphere. This reduces the potential of forests to assist with climate change mitigation . The role of forests in capturing and storing carbon and mitigating climate change

12420-429: The local extinction of large, nocturnal ground-dwelling lizards, and they were assumed to have minimal effect on diurnal species, those that sleep at night, such as shore skinks (Wood et al.). In places where Pacific rats were removed, however, shore skink populations rapidly changed, indicating that the rats’ impact has been underestimated (Wood et al.). The loss of a keystone species, a species that all other species in

12555-548: The lumber company to continue logging. Experts do not agree on whether industrial logging is an important contributor to global deforestation. Some argue that poor people are more likely to clear forest because they have no alternatives, others that the poor lack the ability to pay for the materials and labour needed to clear forest. Charles Darwin Charles Robert Darwin ( / ˈ d ɑːr w ɪ n / DAR -win ; 12 February 1809 – 19 April 1882)

12690-430: The main driver of deforestation and forest fragmentation and the associated loss of forest biodiversity. Large-scale commercial agriculture (primarily cattle ranching and cultivation of soya bean and oil palm) accounted for 40 percent of tropical deforestation between 2000 and 2010, and local subsistence agriculture for another 33 percent. Trees are cut down for use as building material, timber or sold as fuel (sometimes in

12825-420: The mainland provides many unoccupied niches for species to adapt to. Since immigration of predators and competitors is limited, many organisms are able to persist in these new niches. This results in a high occurrence of endemism , where species are unique to a localized area. For example, 50% of endemic bird areas are found on islands. Endemism is often the result of adaptive radiation . Adaptive radiation

12960-408: The mainland. Of the species that arrive, only some will be able to survive and establish populations. As a result, islands have fewer species than mainland habitats. Island populations are small and exhibit low genetic variability (see founder effect ), but are isolated from the predators and competitors that they initially evolved with. This can lead to a process called ecological release , where

13095-943: The many necessary interactions between people and agencies. Once a reserve is in place, managers can then practice adaptive management and do continued community education. On land, island conservation focuses on the protection of species and their habitat. In some cases, conservation can be integrated with agricultural production. For example, the Acacia koa plantations and wooded pastures in Hawaii are anthropogenically altered ecosystems, yet allow connectivity between forest fragments and thus maintain higher diversity than would open pasture. Other directions include habitat restoration, and eradication of introduced predators, ungulates, and exotic plants (via hunting, removal or biological control ). In marine ecosystems, there has been an increasing establishment of “no-take” reserves. This allows for

13230-460: The nutrients in the ashes of the burned plants. As well as, intentionally set fires can possibly lead to devastating measures when unintentionally spreading fire to more land, which can result in the destruction of the protective canopy. The repeated cycle of low yields and shortened fallow periods eventually results in less vegetation being able to grow on once burned lands and a decrease in average soil biomass. In small local plots sustainability

13365-433: The official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history. In Cape Town , South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries,

13500-496: The origin of species". Without telling Darwin, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society , and reported in magazines, including The Athenaeum . Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have

13635-566: The original 16 million square kilometres (6 million square miles) of tropical rainforest that formerly covered the Earth. More than 3.6 million hectares of virgin tropical forest was lost in 2018. The global annual net loss of trees is estimated to be approximately 10 billion. According to the Global Forest Resources Assessment 2020 the global average annual deforested land in the 2015–2020 demi-decade

13770-418: The other hand, a 2005 analysis of satellite images reveals that deforestation of the Amazon rainforest is twice as fast as scientists previously estimated. From 2010 to 2015, worldwide forest area decreased by 3.3 million ha per year, according to FAO . During this five-year period, the biggest forest area loss occurred in the tropics, particularly in South America and Africa. Per capita forest area decline

13905-522: The parents of ten children, seven of whom survived to adulthood. Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus 's An Essay on the Principle of Population . On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what

14040-410: The planet had 15 to 16 million km (5.8 to 6.2 million sq mi) of mature tropical forests , but by 2015, it was estimated that about half of these had been destroyed. Total land coverage by tropical rainforests decreased from 14% to 6%. Much of this loss happened between 1960 and 1990, when 20% of all tropical rainforests were destroyed. At this rate, extinction of such forests

14175-517: The possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal Macrauchenia , which resembled a giant guanaco , a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations – explaining the variations he had observed in Galápagos tortoises , mockingbirds, and rheas. He sketched branching descent, and then

14310-492: The rainforests have been lost in the last 40 years. Brazil has lost 90–95% of its Mata Atlântica forest. Deforestation in Brazil increased by 88% for the month of June 2019, as compared with the previous year. However, Brazil still destroyed 1.3 million hectares in 2019. Brazil is one of several countries that have declared their deforestation a national emergency. Paraguay was losing its natural semi-humid forests in

14445-472: The rate of deforestation in the humid tropics (approximately 5.8 million hectares per year) was roughly 23% lower than the most commonly quoted rates. A 2005 report by the United Nations Food and Agriculture Organization (FAO) estimated that although the Earth's total forest area continued to decrease at about 13 million hectares per year, the global rate of deforestation had been slowing. On

14580-404: The rate of deforestation was estimated at 10 million hectares per year, down from 16 million hectares per year in the 1990s. The area of primary forest worldwide has decreased by over 80 million hectares since 1990. More than 100 million hectares of forests are adversely affected by forest fires, pests, diseases, invasive species , drought and adverse weather events. Deforestation is defined as

14715-710: The reestablishment of native species, and also the augmentation of commercially harvested species. However, in both terrestrial and marine systems, these actions are expensive and do not always result in the desired outcomes. For example, some non-natives become keystone species and their removal can cause more harm than good to the ecosystem. To be more effective, managers of island ecosystems should share information and learn from each other’s mistakes. Island conservation tends to focus on preservation of individual species and their habitats. However, many ecologists caution that ecological and evolutionary processes should be conserved as well. The conservation of island communities as

14850-476: The region in which they take place. The regions with the greatest amount of deforestation for livestock and row crop agriculture are Central and South America, while commodity crop deforestation was found mainly in Southeast Asia. The region with the greatest forest loss due to shifting agriculture was sub-Saharan Africa. The overwhelming direct cause of deforestation is agriculture. Subsistence farming

14985-520: The replacement of extinct species by others" as "a natural in contradistinction to a miraculous process". When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on

15120-415: The risk of fire. Lastly, social insects such as ants also cause major problems. Nonnative species introduced onto islands can have profound effects on an island's ecosystem, more so than nonnative species introduced to continental land (Platenberg). The higher impact of introduced and nonnative species on islands is largely a result of lower biodiversity levels (Platenburg et al.). Biodiversity on islands

15255-519: The same idea, prompting the immediate joint submission of both their theories to the Linnean Society of London . Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex , followed by The Expression of

15390-522: The size of India—by 2050. 36% of globally planted forest area is in East Asia – around 950,000 square kilometers. From those 87% are in China. Rates of deforestation vary around the world. Up to 90% of West Africa 's coastal rainforests have disappeared since 1900. Madagascar has lost 90% of its eastern rainforests. In South Asia , about 88% of the rainforests have been lost. Mexico , India ,

15525-408: The struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work... By mid-December, Darwin saw

15660-409: The systematics of the taxa. He wrote his first examination of the species in 1846, but did not formally describe it until 1854. FitzRoy's long delayed Narrative was published in May 1839. Darwin's Journal and Remarks got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing

15795-505: The threats to species. Ecologists and managers are working together to prioritize areas for conservation and to quickly design and implement action plans. Not everything can be put into a reserve, so it is important to first compile pertinent information and prioritize areas of concern. Once an area has been chosen, managers must then acquire ownership and gain support. Local experts and indigenous populations should also be involved in this process. Having clearly defined goals will facilitate

15930-467: The top 10 most endangered forests, characterized by having all lost 90% or more of their original habitat , and each harboring at least 1500 endemic plant species (species found nowhere else in the world). As of 2015 , it is estimated that 70% of the world's forests are within one kilometer of a forest edge, where they are most prone to human interference and destruction. Deforestation in particular countries: Agricultural expansion continues to be

16065-473: The voyage made Darwin famous as a popular author. Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described

16200-420: The voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations, and at intervals during the voyage his specimens were sent to Cambridge together with letters including

16335-403: The work of earthworms , inspiring "a new & important theory" on their role in soil formation , which Darwin presented at the Geological Society on 1 November 1837. His Journal was printed and ready for publication by the end of February 1838, as was the first volume of the Narrative , but FitzRoy was still working hard to finish his own volume. William Whewell pushed Darwin to take on

16470-473: The world were cut down. Deforestation and forest degradation continue to take place at alarming rates, which contributes significantly to the ongoing loss of biodiversity . Deforestation is more extreme in tropical and subtropical forests in emerging economies. More than half of all plant and land animal species in the world live in tropical forests . As a result of deforestation, only 6.2 million square kilometres (2.4 million square miles) remain of

16605-576: The world, especially in East Asian countries, reforestation and afforestation are increasing the area of forested lands. The amount of forest has increased in 22 of the world's 50 most forested nations. Asia as a whole gained 1 million hectares of forest between 2000 and 2005. Tropical forest in El Salvador expanded more than 20% between 1992 and 2001. Based on these trends, one study projects that global forestation will increase by 10%—an area

16740-707: Was elected a Fellow of the Royal Society (FRS). On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home. Darwin now had the framework of his theory of natural selection "by which to work", as his "prime hobby". His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory. For fifteen years this work

16875-456: Was 10 million hectares and the average annual forest area net loss in the 2000–2010 decade was 4.7 million hectares. The world has lost 178 million ha of forest since 1990, which is an area about the size of Libya. An analysis of global deforestation patterns in 2021 showed that patterns of trade, production, and consumption drive deforestation rates in complex ways. While the location of deforestation can be mapped, it does not always match where

17010-418: Was a " wren " was in the finch group . Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands. The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards. By mid-March 1837, barely six months after his return to England, Darwin was speculating in his Red Notebook on

17145-461: Was also greatest in the tropics and subtropics but is occurring in every climatic domain (except in the temperate) as populations increase. An estimated 420 million ha of forest has been lost worldwide through deforestation since 1990, but the rate of forest loss has declined substantially. In the most recent five-year period (2015–2020), the annual rate of deforestation was estimated at 10 million ha, down from 12 million ha in 2010–2015. Africa had

17280-426: Was an English naturalist , geologist , and biologist , widely known for his contributions to evolutionary biology . His proposition that all species of life have descended from a common ancestor is now generally accepted and considered a fundamental scientific concept. In a joint publication with Alfred Russel Wallace , he introduced his scientific theory that this branching pattern of evolution resulted from

17415-472: Was anathema to Anglicans defending social order, but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species. Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought

17550-425: Was in the background to his main occupation of writing on geology and publishing expert reports on the Beagle collections, in particular, the barnacles . The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed Mr. Arthrobalanus . His confusion over the relationship of this species ( Cryptophialus minutus ) to other barnacles caused him to fixate on

17685-504: Was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England. Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel -beds stranded above high tide. High in the Andes he saw seashells, and several fossil trees that had grown on

17820-577: Was rather bored by Robert Jameson 's natural-history course, which covered geology – including the debate between neptunism and plutonism . He learned the classification of plants and assisted with work on the collections of the University Museum , one of the largest museums in Europe at the time. Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge , in January 1828, to study for

17955-610: Was subsequently driven to extinction by humans and introduced predators. The depletion of natural resources can have dramatic effects on island ecology. On Easter Island , the depletion of the forest by humans not only resulted in widespread loss of species, but also the collapse of the island civilization. Today there are over 500 million people on islands, all dependent on local resources either directly (traditional use) or indirectly (ecotourism revenue). Population growth and development result in heavy deforestation , pollution , and over-exploitation . Overharvesting of ocean fauna

18090-574: Was surprised to find some benefit from hydrotherapy . Then, in 1851, his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises. In eight years of work on barnacles, Darwin's theory helped him to find " homologies " showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites , showing an intermediate stage in evolution of distinct sexes . In 1853, it earned him

18225-446: Was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II , to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution. After delays,

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