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The Llanos ( Spanish Los Llanos , "The Plains"; Spanish pronunciation: [los ˈʝanos] ) is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela , in northwestern South America. It is an ecoregion of the tropical and subtropical grasslands, savannas, and shrublands biome .

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113-708: The Llanos occupy a lowland that extends mostly east and west. The Llanos are bounded on the west and northwest by the Andes , and on the north by the Venezuelan Coastal Range . The Guiana Highlands are to the southeast, and the Negro-Branco moist forests are to the southwest. To the east the Orinoco wetlands and Orinoco Delta swamp forests occupy the Orinoco Delta . The Llanos' main river

226-711: A sink or a source of carbon, depending on the specific wetland. If they function as a carbon sink, they can help with climate change mitigation . However, wetlands can also be a significant source of methane emissions due to anaerobic decomposition of soaked detritus , and some are also emitters of nitrous oxide . Humans are disturbing and damaging wetlands in many ways, including oil and gas extraction , building infrastructure, overgrazing of livestock , overfishing , alteration of wetlands including dredging and draining, nutrient pollution , and water pollution . Wetlands are more threatened by environmental degradation than any other ecosystem on Earth, according to

339-653: A canopy 8 to 20 meters high, and include Inga spp., Combretum frangulifolium , Gustavia augusta , Pterocarpus sp., Pterocarpus dubius , Spondias mombin , and Copaifera pubiflora . Semi-deciduous forests occur above flood level, and form a canopy 12 to 15 meters high. Common trees include Tabebuia billbergii , Godmania aesculifolia , Cassia moschata , Spondias mombin, Copaifera pubiflora, Bourreria cumanensis , Cordia spp., Bursera simaruba , Cochlospermum vitifolium , Hura crepitans , and Acacia glomerosa . "Matorrales" are deciduous and semi-deciduous shrublands 5 to 8 meters high which cover large areas in

452-402: A frequency and duration to support, and that under normal circumstances do support, a prevalence of vegetation typically adapted for life in saturated soil conditions. Wetlands generally included swamps, marshes, bogs, and similar areas.' For each of these definitions and others, regardless of the purpose, hydrology is emphasized (shallow waters, water-logged soils). The soil characteristics and

565-534: A hectare. During the rainy season from May to October, parts of the Llanos can flood up to a meter. This turns some savannass and grasslands into temporary wetlands , comparable to the Pantanal of central South America. This flooding also creates habitat for water birds and other wildlife. These seasonally flooded grasslands and savannas, known as llano bajo, typically have richer soils. They are characterized by

678-555: A higher potential for larger, high-magnitude earthquakes in the future. The mountain range is also a source of shallow intraplate earthquakes within the South American Plate. The largest such earthquake (as of 2024) struck Peru in 1947 and measured M s   7.5. In the Peruvian Andes, these earthquakes display normal ( 1946 ), strike-slip (1976), and reverse ( 1969 , 1983) mechanisms. The Amazonian Craton

791-566: A joint ice stream network. The Andes' dendritic glacier arms, components of valley glaciers, were up to 112.5 km (69.9 mi) long and over 1,250 m (4,101 ft) thick, and spanned a vertical distance of 5,150 m (16,896 ft). The climatic glacier snowline (ELA) was lowered from 4,600 m (15,092 ft) to 3,200 m (10,499 ft) at glacial times. The Andean region cuts across several natural and floristic regions, due to its extension, from Caribbean Venezuela to cold, windy, and wet Cape Horn passing through

904-594: A north–south axis of cultural influences. A long series of cultural development culminated in the expansion of the Inca civilization and Inca Empire in the central Andes during the 15th century. The Incas formed this civilization through imperialistic militarism as well as careful and meticulous governmental management. The government sponsored the construction of aqueducts and roads in addition to pre-existing installations. Some of these constructions still exist today. Devastated by European diseases and by civil war ,

1017-1259: A polar climate, wetland temperatures can be as low as −50 °C (−58 °F). Peatlands in arctic and subarctic regions insulate the permafrost , thus delaying or preventing its thawing during summer, as well as inducing its formation. The amount of precipitation a wetland receives varies widely according to its area. Wetlands in Wales , Scotland , and western Ireland typically receive about 1,500 mm (59 in) per year. In some places in Southeast Asia , where heavy rains occur, they can receive up to 10,000 mm (390 in). In some drier regions, wetlands exist where as little as 180 mm (7.1 in) precipitation occurs each year. Temporal variation: Surface flow may occur in some segments, with subsurface flow in other segments. Wetlands vary widely due to local and regional differences in topography , hydrology , vegetation , and other factors, including human involvement. Other important factors include fertility, natural disturbance, competition, herbivory , burial and salinity. When peat accumulates, bogs and fens arise. The most important factor producing wetlands

1130-540: A portion of the overall water cycle, which also includes atmospheric water (precipitation) and groundwater . Many wetlands are directly linked to groundwater and they can be a crucial regulator of both the quantity and quality of water found below the ground. Wetlands that have permeable substrates like limestone or occur in areas with highly variable and fluctuating water tables have especially important roles in groundwater replenishment or water recharge. Substrates that are porous allow water to filter down through

1243-711: A series of bends or oroclines . The Bolivian Orocline is a seaward-concave bending in the coast of South America and the Andes Mountains at about 18° S. At this point, the orientation of the Andes turns from northwest in Peru to south in Chile and Argentina . The Andean segments north and south of the Orocline have been rotated 15° counter-clockwise to 20° clockwise respectively. The Bolivian Orocline area overlaps with

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1356-422: A strong influence on wetland water chemistry, particularly in coastal wetlands and in arid and semiarid regions with large precipitation deficits. Natural salinity is regulated by interactions between ground and surface water, which may be influenced by human activity. Carbon is the major nutrient cycled within wetlands. Most nutrients, such as sulfur , phosphorus , carbon , and nitrogen are found within

1469-634: A typical location for calc-alkalic and subduction volcanism, the Andean Volcanic Belt has a large range of volcano-tectonic settings, such as rift systems, extensional zones, transpressional faults , subduction of mid-ocean ridges , and seamount chains apart from a large range of crustal thicknesses and magma ascent paths, and different amount of crustal assimilations. The Andes Mountains host large ore and salt deposits, and some of their eastern fold and thrust belts act as traps for commercially exploitable amounts of hydrocarbons . In

1582-410: A unique kind of wetland where lush plant growth and slow decay of dead plants (under anoxic conditions) results in organic peat accumulating; bogs, fens, and mires are different names for peatlands. Variations of names for wetland systems: Some wetlands have localized names unique to a region such as the prairie potholes of North America's northern plain, pocosins , Carolina bays and baygalls of

1695-515: A wetland is "an ecosystem that arises when inundation by water produces soils dominated by anaerobic and aerobic processes, which, in turn, forces the biota, particularly rooted plants, to adapt to flooding". Sometimes a precise legal definition of a wetland is required. The definition used for regulation by the United States government is: 'The term "wetlands" means those areas that are inundated or saturated by surface or ground water at

1808-422: A wetland system includes its plants ( flora ) and animals ( fauna ) and microbes (bacteria, fungi). The most important factor is the wetland's duration of flooding. Other important factors include fertility and salinity of the water or soils. The chemistry of water flowing into wetlands depends on the source of water, the geological material that it flows through and the nutrients discharged from organic matter in

1921-403: A wetland) affects hydro-periods (temporal fluctuations in water levels) by controlling the water balance and water storage within a wetland. Landscape characteristics control wetland hydrology and water chemistry. The O 2 and CO 2 concentrations of water depend upon temperature , atmospheric pressure and mixing with the air (from winds or water flows). Water chemistry within wetlands

2034-536: A whole glacier in the ablation area or in the accumulation area . In the high Andes of Central Chile and Mendoza Province , rock glaciers are larger and more common than glaciers; this is due to the high exposure to solar radiation . In these regions, glaciers occur typically at higher altitudes than rock glaciers. The lowest active rock glaciers occur at 900 m a.s.l. in Aconcagua . Though precipitation increases with height, there are semiarid conditions in

2147-558: A whole. To replace these wetland ecosystem services , enormous amounts of money would need to be spent on water purification plants, dams, levees, and other hard infrastructure, and many of the services are impossible to replace. Floodplains and closed-depression wetlands can provide the functions of storage reservoirs and flood protection. The wetland system of floodplains is formed from major rivers downstream from their headwaters . "The floodplains of major rivers act as natural storage reservoirs, enabling excess water to spread out over

2260-594: A wide area, which reduces its depth and speed. Wetlands close to the headwaters of streams and rivers can slow down rainwater runoff and spring snowmelt so that it does not run straight off the land into water courses. This can help prevent sudden, damaging floods downstream." Notable river systems that produce wide floodplains include the Nile River , the Niger river inland delta, the Zambezi River flood plain,

2373-780: A zone of volcanic activity that encompasses the Pacific rim of the Americas as well as the Asia-Pacific region. The Andes are the result of tectonic plate processes extending during the Mesozoic and Tertiary eras, caused by the subduction of oceanic crust beneath the South American Plate as the Nazca Plate and South American Plate converge. These processes were accelerated by the effects of climate. As

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2486-561: Is a result of the subduction of the Nazca Plate and Antarctic Plate underneath the South American Plate. The belt is subdivided into four main volcanic zones that are separated from each other by volcanic gaps. The volcanoes of the belt are diverse in terms of activity style, products, and morphology. While some differences can be explained by which volcanic zone a volcano belongs to, there are significant differences inside volcanic zones and even between neighboring volcanoes. Despite being

2599-551: Is actively underthrusted beneath the sub-Andes region of Peru, producing thrust faults. In Colombia, Ecuador, and Peru, thrust faulting occurs along the sub-Andes due in response to compression brought on by subduction, while in the high Andes, normal faulting occurs in response to gravitational forces. In the extreme south, a major transform fault separates Tierra del Fuego from the small Scotia Plate . The Andes range has many active volcanoes distributed in four volcanic zones separated by areas of inactivity. The Andean volcanism

2712-440: Is determined by the pH , salinity , nutrients, conductivity , soil composition, hardness , and the sources of water. Water chemistry varies across landscapes and climatic regions. Wetlands are generally minerotrophic (waters contain dissolved materials from soils) with the exception of ombrotrophic bogs that are fed only by water from precipitation. Because bogs receive most of their water from precipitation and humidity from

2825-633: Is either freshwater , brackish or saltwater . The main types of wetland are defined based on the dominant plants and the source of the water. For example, marshes are wetlands dominated by emergent herbaceous vegetation such as reeds , cattails and sedges . Swamps are dominated by woody vegetation such as trees and shrubs (although reed swamps in Europe are dominated by reeds, not trees). Mangrove forest are wetlands with mangroves , halophytic woody plants that have evolved to tolerate salty water . Examples of wetlands classified by

2938-412: Is hydrology, or flooding . The duration of flooding or prolonged soil saturation by groundwater determines whether the resulting wetland has aquatic, marsh or swamp vegetation . Other important factors include soil fertility, natural disturbance, competition, herbivory , burial, and salinity. When peat from dead plants accumulates, bogs and fens develop. Wetland hydrology is associated with

3051-471: Is natural, or the result of clearing which began during the Incan period. Regardless, in modern times , the clearance has accelerated, and the trees are now considered highly endangered , with some believing that as little as 10% of the original woodland remains. The Andes are rich in fauna: With almost 1,000 species, of which roughly 2/3 are endemic to the region, the Andes are the most important region in

3164-557: Is the Orinoco , which runs from west to east through the ecoregion and forms part of the border between Colombia and Venezuela. The Orinoco is the major river system of Venezuela. The ecoregion has a tropical savanna climate that grades into a tropical monsoon climate in the Colombian Llanos. Rainfall is highly seasonal, with a rainy season from April to November, and a dry season between December and March. The wettest months are typically June and July. Rainfall varies across

3277-1647: Is the largest city of all Andean countries. It is the seat of the Andean Community of Nations . La Paz , Bolivia 's seat of government, is the highest capital city in the world, at an elevation of approximately 3,650 m (11,975 ft). Parts of the La Paz conurbation, including the city of El Alto , extend up to 4,200 m (13,780 ft). Other cities in or near the Andes include Bariloche , Catamarca , Jujuy , Mendoza , Salta , San Juan , Tucumán , and Ushuaia in Argentina; Calama and Rancagua in Chile; Cochabamba , Oruro , Potosí , Sucre , Tarija , and Yacuiba in Bolivia; Arequipa , Cajamarca , Cusco , Huancayo , Huánuco , Huaraz , Juliaca , and Puno in Peru; Ambato , Cuenca , Ibarra , Latacunga , Loja , Riobamba , and Tulcán in Ecuador; Armenia , Cúcuta , Bucaramanga , Duitama , Ibagué , Ipiales , Manizales , Palmira , Pasto , Pereira , Popayán , Sogamoso , Tunja , and Villavicencio in Colombia; and Barquisimeto , La Grita , Mérida , San Cristóbal , Tovar, Trujillo , and Valera in Venezuela. The cities of Caracas , Valencia , and Maracay are in

3390-451: Is the only place in the world where both crocodiles and alligators coexist. The saltwater crocodile inhabits estuaries and mangroves. Snapping turtles also inhabit wetlands. Birds , particularly waterfowl and waders use wetlands extensively. Mammals of wetlands include numerous small and medium-sized species such as voles , bats , muskrats and platypus in addition to large herbivorous and apex predator species such as

3503-545: Is usually saturated with water". More precisely, wetlands are areas where "water covers the soil , or is present either at or near the surface of the soil all year or for varying periods of time during the year, including during the growing season". A patch of land that develops pools of water after a rain storm would not necessarily be considered a "wetland", even though the land is wet. Wetlands have unique characteristics: they are generally distinguished from other water bodies or landforms based on their water level and on

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3616-404: The 2010 Maule earthquake (M8.8), the 2015 Illapel earthquake (M8.2), and the 1960 Valdivia earthquake (M9.5), which as of 2024 was the strongest ever recorded on seismometers. The amount, magnitude, and type of seismic activity varies greatly along the subduction zone. These differences are due to a wide range of factors, including friction between the plates, angle of subduction, buoyancy of

3729-682: The Americas and Antarctica . The etymology of the word Andes has been debated. The majority consensus is that it derives from the Quechua word anti "east" as in Antisuyu (Quechua for "east region"), one of the four regions of the Inca Empire . The term cordillera comes from the Spanish word cordel "rope" and is used as a descriptive name for several contiguous sections of

3842-456: The Andean cock-of-the-rock , while mixed-species flocks dominated by tanagers and furnariids are commonly seen—in contrast to several vocal but typically- cryptic species of wrens , tapaculos , and antpittas . A number of species such as the royal cinclodes and white-browed tit-spinetail are associated with Polylepis , and consequently also threatened . The Andes Mountains form

3955-592: The Caribbean Sea off the coast of Venezuela, were formerly thought to represent the submerged peaks of the extreme northern edge of the Andes range, but ongoing geological studies indicate that such a simplification does not do justice to the complex tectonic boundary between the South American and Caribbean plates . The Andes are an orogenic belt of mountains along the Pacific Ring of Fire ,

4068-535: The Cenozoic . Tectonic forces above the subduction zone along the entire west coast of South America where the Nazca Plate and a part of the Antarctic Plate are sliding beneath the South American Plate continue to produce an ongoing orogenic event resulting in minor to major earthquakes and volcanic eruptions to this day. Many high-magnitude earthquakes have been recorded in the region, such as

4181-505: The Millennium Ecosystem Assessment from 2005. Methods exist for assessing wetland ecological health . These methods have contributed to wetland conservation by raising public awareness of the functions that wetlands can provide. Since 1971, work under an international treaty seeks to identify and protect " wetlands of international importance ." A simplified definition of wetland is "an area of land that

4294-633: The Pantanal in South America, and the Sundarbans in the Ganges - Brahmaputra delta. Wetlands contribute many ecosystem services that benefit people. These include for example water purification , stabilization of shorelines, storm protection and flood control . In addition, wetlands also process and condense carbon (in processes called carbon fixation and sequestration ), and other nutrients and water pollutants . Wetlands can act as

4407-694: The Tropical Andes , the Dry Andes , and the Wet Andes . The Andes are the highest mountain range which is outside of Asia . The range's highest peak, Argentina's Aconcagua , rises to an elevation of about 6,961 m (22,838 ft) above sea level. The peak of Chimborazo in the Ecuadorian Andes is farther from the Earth's center than any other location on the Earth's surface, due to

4520-506: The Venezuelan Coastal Range , which is a debatable extension of the Andes at the northern extremity of South America. Cities and large towns are connected with asphalt -paved roads, while smaller towns are often connected by dirt roads, which may require a four-wheel-drive vehicle. The rough terrain has historically put the costs of building highways and railroads that cross the Andes out of reach of most neighboring countries, even with modern civil engineering practices. For example,

4633-674: The War of the Pacific between Chile and the allied Bolivia and Peru, in a diplomatic deal to keep Peru out of the war. The Chilean Army and Chilean Navy defeated the combined forces of Bolivia and Peru, and Chile took over Bolivia's only province on the Pacific Coast, some land from Peru that was returned to Peru decades later. Bolivia has been completely landlocked ever since. It mostly uses seaports in eastern Argentina and Uruguay for international trade because its diplomatic relations with Chile have been suspended since 1978. Because of

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4746-444: The atmosphere , their water usually has low mineral ionic composition. In contrast, wetlands fed by groundwater or tides have a higher concentration of dissolved nutrients and minerals. Fen peatlands receive water both from precipitation and ground water in varying amounts so their water chemistry ranges from acidic with low levels of dissolved minerals to alkaline with high accumulation of calcium and magnesium . Salinity has

4859-576: The beavers , coypu , swamp rabbit , Florida panther , jaguar , and moose . Wetlands attract many mammals due to abundant seeds, berries, and other vegetation as food for herbivores, as well as abundant populations of invertebrates, small reptiles and amphibians as prey for predators. Invertebrates of wetlands include aquatic insects such as dragonflies , aquatic bugs and beetles , midges, mosquitos , crustaceans such as crabs, crayfish, shrimps, microcrustaceans, mollusks like clams, mussels, snails and worms. Invertebrates comprise more than half of

4972-602: The equatorial bulge resulting from the Earth's rotation . The world's highest volcanoes are in the Andes, including Ojos del Salado on the Chile-Argentina border, which rises to 6,893 m (22,615 ft). The Andes are also part of the American Cordillera , a chain of mountain ranges ( cordillera ) that consists of an almost continuous sequence of mountain ranges that form the western "backbone" of

5085-517: The longest continental mountain range in the world, forming a continuous highland along the western edge of South America . The range is 8,900 km (5,530 mi) long and 200 to 700 km (124 to 435 mi) wide (widest between 18°S and 20°S latitude ) and has an average height of about 4,000 m (13,123 ft). The Andes extend from South to North through seven South American countries: Argentina , Chile , Bolivia , Peru , Ecuador , Colombia , and Venezuela . Along their length,

5198-592: The lowland paca ( Cuniculus paca ), agoutis ( Dasyprocta spp.), and Brazilian porcupine ( Coendou prehensilis ), and large cats like the puma ( Puma concolor ), jaguar ( Panthera onca ), and ocelot ( Leopardus pardalis ). The endangered giant otter ( Pteronura brasiliensis ) lives along the Orinoco and its tributaries. Some of the largest jaguars in the world are found in the Llanos, with average weights of over 100 kg (220 lb) for males. The Llanos long-nosed armadillo ( Dasypus sabanicola ) and

5311-483: The rodent order, inhabit the Andes' alpine regions. The Andean condor , the largest bird of its kind in the Western Hemisphere , occurs throughout much of the Andes but generally in very low densities. Other animals found in the relatively open habitats of the high Andes include the huemul , cougar , foxes in the genus Pseudalopex , and, for birds, certain species of tinamous (notably members of

5424-607: The soils . Wetlands form a transitional zone between waterbodies and dry lands , and are different from other terrestrial or aquatic ecosystems due to their vegetation 's roots having adapted to oxygen-poor waterlogged soils . They are considered among the most biologically diverse of all ecosystems, serving as habitats to a wide range of aquatic and semi-aquatic plants and animals , with often improved water quality due to plant removal of excess nutrients such as nitrates and phosphorus . Wetlands exist on every continent , except Antarctica . The water in wetlands

5537-703: The 1,000-kilometer-wide (620 mi) Drake Passage lie the mountains of the Antarctic Peninsula south of the Scotia Plate , which appear to be a continuation of the Andes chain. The far east regions of the Andes experience a series of changes resulting from the Andean orogeny. Parts of the Sunsás Orogen in Amazonian craton disappeared from the surface of the earth, being overridden by

5650-651: The African grass Melinis minutiflora , and now cover large areas. Agriculture, particularly rice and maize, now cover extensive areas, including rice fields in former seasonal wetlands. In Los Llanos, the governments of Venezuela and Colombia had developed a strong oil and gas industry in the zones of Arauca, Casanare, Guárico, Anzoátegui, Apure and Monagas. The Orinoco Belt , entirely in Venezuelan territory, consists of large deposits of extra heavy crude ( oil sands ). The Orinoco belt oil sands are known to be one of

5763-469: The Andean peoples are those of the Quechua and Aymara language families. Woodbine Parish and Joseph Barclay Pentland surveyed a large part of the Bolivian Andes from 1826 to 1827. In modern times, the largest cities in the Andes are Bogotá , with a metropolitan population of over ten million, and Santiago , Medellín , Cali , and Quito . Lima is a coastal city adjacent to the Andes and

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5876-405: The Andes are tobacco and potatoes . The high-altitude Polylepis forests and woodlands are found in the Andean areas of Colombia , Ecuador , Peru , Bolivia , and Chile . These trees, by locals referred to as Queñua, Yagual, and other names, can be found at altitudes of 4,500 m (14,760 ft) above sea level. It remains unclear if the patchy distribution of these forests and woodlands

5989-510: The Andes are split into several ranges, separated by intermediate depressions . The Andes are the location of several high plateaus —some of which host major cities such as Quito , Bogotá , Cali , Arequipa , Medellín , Bucaramanga , Sucre , Mérida , El Alto , and La Paz . The Altiplano Plateau is the world's second highest after the Tibetan Plateau . These ranges are in turn grouped into three major divisions based on climate:

6102-521: The Andes end at the Pacific Ocean , although the Peru-Chile trench can be considered their ultimate western limit. From a geographical approach, the Andes are considered to have their western boundaries marked by the appearance of coastal lowlands and less-rugged topography. The Andes also contain large quantities of iron ore located in many mountains within the range. The Andean orogen has

6215-462: The Andes such as the Incas have practiced irrigation techniques for over 6,000 years. Because of the mountain slopes, terracing has been a common practice. Terracing, however, was only extensively employed after Incan imperial expansions to fuel their expanding realm. The potato holds a very important role as an internally-consumed staple crop. Maize was also an important crop for these people, and

6328-526: The Andes varies greatly depending on latitude, altitude, and proximity to the sea. Temperature, atmospheric pressure, and humidity decrease in higher elevations. The southern section is rainy and cool, while the central section is dry. The northern Andes are typically rainy and warm, with an average temperature of 18 °C (64 °F) in Colombia . The climate is known to change drastically in rather short distances. Rainforests exist just kilometers away from

6441-542: The Andes, as well as the entire Andean range, and the combined mountain chain along the western part of the North and South American continents. The Andes can be divided into three sections: At the northern end of the Andes, the separate Sierra Nevada de Santa Marta range is often, but not always, treated as part of the Northern Andes. The Leeward Antilles islands Aruba , Bonaire , and Curaçao , which lie in

6554-524: The Andes. The Sierras de Córdoba , where the effects of the ancient Pampean orogeny can be observed, owe their modern uplift and relief to the Andean orogeny in the Tertiary . Further south in southern Patagonia , the onset of the Andean orogeny caused the Magallanes Basin to evolve from being an extensional back-arc basin in the Mesozoic to being a compressional foreland basin in

6667-491: The Incas were defeated in 1532 by an alliance composed of tens of thousands of allies from nations they had subjugated (e.g. Huancas , Chachapoyas , Cañaris ) and a small army of 180 Spaniards led by Francisco Pizarro . One of the few Inca sites the Spanish never found in their conquest was Machu Picchu , which lay hidden on a peak on the eastern edge of the Andes where they descend to the Amazon. The main surviving languages of

6780-639: The Okavango River inland delta, the Kafue River flood plain, the Lake Bangweulu flood plain (Africa), Mississippi River (US), Amazon River (South America), Yangtze River (China), Danube River (Central Europe) and Murray-Darling River (Australia). Groundwater replenishment can be achieved for example by marsh , swamp , and subterranean karst and cave hydrological systems. The surface water visibly seen in wetlands only represents

6893-478: The Patagonian Orocline. The western rim of the South American Plate has been the place of several pre-Andean orogenies since at least the late Proterozoic and early Paleozoic , when several terranes and microcontinents collided and amalgamated with the ancient cratons of eastern South America, by then the South American part of Gondwana . The formation of the modern Andes began with

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7006-470: The Southeastern US, mallines of Argentina, Mediterranean seasonal ponds of Europe and California, turloughs of Ireland, billabongs of Australia, among many others. Wetlands are found throughout the world in different climates. Temperatures vary greatly depending on the location of the wetland. Many of the world's wetlands are in the temperate zones , midway between the North or South Poles and

7119-502: The US, the best known classifications are the Cowardin classification system and the hydrogeomorphic (HGM) classification system. The Cowardin system includes five main types of wetlands: marine (ocean-associated), estuarine (mixed ocean- and river-associated), riverine (within river channels), lacustrine (lake-associated) and palustrine (inland nontidal habitats). Peatlands are

7232-610: The United States' commercial fish and shellfish stocks depend solely on estuaries to survive. Amphibians such as frogs and salamanders need both terrestrial and aquatic habitats in which to reproduce and feed. Because amphibians often inhabit depressional wetlands like prairie potholes and Carolina bays, the connectivity among these isolated wetlands is an important control of regional populations. While tadpoles feed on algae, adult frogs forage on insects. Frogs are sometimes used as an indicator of ecosystem health because their thin skin permits absorption of nutrients and toxins from

7345-520: The Venezuelan llanos, and is also widespread in the Colombian llanos. Grasses of genus Trachypogon are predominant, and species include Trachypogon plumosus , T. vestitus , Axonopus canescens , A. anceps , Andropogon selloanus , Aristida spp., Leptocoryphium lanatum , Paspalum carinatum , Sporobolus indicus , and S. cubensis , and sedges in the genera Rhynchospora and Bulbostylis . Shrubs and herbs are most commonly legumes in

7458-592: The area of the maximum width of the Altiplano Plateau , and according to Isacks (1988) the Orocline is related to crustal shortening . The specific point at 18° S where the coastline bends is known as the Arica Elbow. Further south lies the Maipo Orocline, a more subtle orocline between 30° S and 38°S with a seaward-concave break in the trend at 33° S. Near the southern tip of the Andes lies

7571-1106: The central Venezuelan llanos, and may be a form of secondary vegetation in areas that were formerly dry deciduous forest. Typical shrubs are Bourreria cumanensis , Randia aculeata , Godmania aesculifolia , Pereskia guamacho , Prosopis spp., Xylosma benthamii , Erytroxylum sp., and Cereus hexagonus . Mammals of the grassland and savanna include white-tailed deer ( Odocoileus virginianus ), giant anteater ( Myrmecophaga tridactyla ), capybara ( Hydrochoerus hydrochaeris ), giant armadillo ( Priodontes maximus ), Eastern cottontail ( Sylvilagus floridanus ), Alston's cotton rat ( Sigmodon alstoni ), Hispid cotton rat ( S. hispidus ), Zygodontomys brevicauda , and Oecomys bicolor . The gallery forests are home to more diverse large and medium-sized mammals, including collared peccary ( Tayassu tajacu ), white-lipped peccary ( T. pecari ), South American tapir ( Tapirus terrestris ), white-tailed deer, red brocket ( Mazama americana ), wedge-capped capuchin ( Cebus olivaceus ), Venezuelan red howler ( Alouatta seniculus ), large rodents like

7684-480: The cowhands that took care of the herds, and had some cultural similarities with the gauchos of the Pampas or the vaqueros of Spanish and Mexican Texas . Decades of extensive cattle raising has altered the ecology of the Llanos. Grasslands and savannas are frequently burned to make them more suitable for grazing and eliminate trees and shrubs. Non-native grasses have been introduced for cattle fodder, including

7797-399: The dry climate that reduced the disturbing actions of meteoric water . The dry climate in the central western Andes has also led to the creation of extensive saltpeter deposits which were extensively mined until the invention of synthetic nitrates . Yet another result of the dry climate are the salars of Atacama and Uyuni , the former being the largest source of lithium and the latter

7910-532: The dry season (winter) possible and allows the cultivation of frost-resistant vegetable crops like onion and carrot . The Andes rose to fame for their mineral wealth during the Spanish conquest of South America . Although Andean Amerindian peoples crafted ceremonial jewelry of gold and other metals, the mineralizations of the Andes were first mined on a large scale after the Spanish arrival. Potosí in present-day Bolivia and Cerro de Pasco in Peru were among

8023-476: The ecoregion's wetlands. Indigenous peoples of the Llanos include the Guahibo in the western Llanos of Colombia and Venezuela, and the Yaruro in the eastern Llanos in Venezuela. The primary economic activity in the Llanos since the Spanish colonial era is the herding of millions of cattle. An 1856 watercolor by Manuel María Paz depicts sparsely populated open grazing lands with cattle and palm trees. The term llanero ("plainsman") became synonymous with

8136-553: The ecoregion, from up to 3,000 millimetres (120 in) per year in the southwest, 1,200 to 1,600 millimetres (47 to 63 in) in Apure State, and 800 to 1,200 millimetres (31 to 47 in) per year in the Llanos of Monagas State in the northeast. Mean annual temperature is 27 °C or 80.6 °F, and the average monthly temperature varies little throughout the year; the lowest-temperature months (June, July, December, and January) are only 2 °C or 3.6 °F cooler than

8249-634: The equator. In these zones, summers are warm and winters are cold, but temperatures are not extreme. In subtropical zone wetlands, such as along the Gulf of Mexico , average temperatures might be 11 °C (52 °F). Wetlands in the tropics are subjected to much higher temperatures for a large portion of the year. Temperatures for wetlands on the Arabian Peninsula can exceed 50 °C (122 °F) and these habitats would therefore be subject to rapid evaporation. In northeastern Siberia , which has

8362-501: The establishment of the century-long Inca Empire . This all changed in the 16th century, when the Spanish conquistadors colonized the mountains in advance of the mining economy. In the tide of anti-imperialist nationalism, the Andes became the scene of a series of independence wars in the 19th century, when rebel forces swept through the region to overthrow Spanish colonial rule. Since then, many former Spanish territories have become five independent Andean states. The climate in

8475-705: The events of the Triassic , when Pangaea began the breakup that resulted in developing several rifts . The development continued through the Jurassic Period. It was during the Cretaceous Period that the Andes began to take their present form, by the uplifting , faulting , and folding of sedimentary and metamorphic rocks of the ancient cratons to the east. The rise of the Andes has not been constant, as different regions have had different degrees of tectonic stress, uplift, and erosion . Across

8588-423: The extreme in the slopes near the virtually-lifeless Atacama Desert. About 30,000 species of vascular plants live in the Andes, with roughly half being endemic to the region, surpassing the diversity of any other hotspot . The small tree Cinchona pubescens , a source of quinine which is used to treat malaria , is found widely in the Andes as far south as Bolivia. Other important crops that originated from

8701-504: The following areas: According to the Ramsar Convention: The economic worth of the ecosystem services provided to society by intact, naturally functioning wetlands is frequently much greater than the perceived benefits of converting them to 'more valuable' intensive land use – particularly as the profits from unsustainable use often go to relatively few individuals or corporations, rather than being shared by society as

8814-464: The forelands of the Atacama Desert , some of the largest porphyry copper mineralizations occur, making Chile and Peru the first- and second-largest exporters of copper in the world. Porphyry copper in the western slopes of the Andes has been generated by hydrothermal fluids (mostly water) during the cooling of plutons or volcanic systems. The porphyry mineralization further benefited from

8927-417: The genera Mimosa , Cassia , Desmodium , Eriosema , Galactia , Indigofera , Phaseolus , Stylosanthes , Tephrosia , and Zornia . The trees manteco ( Byrsonima crassifolia ), chaparro ( Curatella americana ), and alcornoque ( Bowdichia virgilioides ) are the most common, growing either as scattered trees or in woodland patches known as matas which range in area from 12 meters in diameter up to

9040-555: The genus Nothoprocta ), Andean goose , giant coot , flamingos (mainly associated with hypersaline lakes), lesser rhea , Andean flicker , diademed sandpiper-plover , miners , sierra-finches and diuca-finches . Lake Titicaca hosts several endemics, among them the highly endangered Titicaca flightless grebe and Titicaca water frog . A few species of hummingbirds , notably some hillstars , can be seen at altitudes above 4,000 m (13,100 ft), but far higher diversities can be found at lower altitudes, especially in

9153-512: The glaciers were over ten times longer. On the east side of this section of the Mendozina Andes, they flowed down to 2,060 m (6,759 ft) and on the west side to about 1,220 m (4,003 ft) above sea level. The massifs of Aconcagua (6,961 m (22,838 ft)), Tupungato (6,550 m (21,490 ft)), and Nevado Juncal (6,110 m (20,046 ft)) are tens of kilometres away from each other and were connected by

9266-424: The grass Paspalum fasciculatum . Trees include the palm Copernicia tectorum and gallery forest species. Gallery forests include evergreen seasonally flooded forests, and semi-deciduous forests on higher ground. Morichales are seasonally flooded forests characterized by the moriche palm ( Mauritia flexuosa ). Vegas are seasonally flooded evergreen forests found along the Orinoco and its tributaries. Trees form

9379-597: The hottest months. The plant communities in the Llanos include open grasslands, savannas with scattered trees or clumps of trees, and small areas of forest, typically gallery forests along rivers and streams. There are seasonally flooded grasslands and savannas ( llano bajo ) and grasslands and savannas that remain dry throughout the year ( llano alto ). The llano alto grasslands and savannas are characterized by grasses and shrubs 30–100 cm high, forming tussocks 10 to 30 cm apart. Soils are typically sandy and nutrient-poor. Llano alto covers approximately two-thirds of

9492-556: The humid Andean forests (" cloud forests ") growing on slopes in Colombia, Ecuador, Peru, Bolivia, and far northwestern Argentina. These forest-types, which includes the Yungas and parts of the Chocó, are very rich in flora and fauna, although few large mammals exist, exceptions being the threatened mountain tapir , spectacled bear , and yellow-tailed woolly monkey . Birds of humid Andean forests include mountain toucans , quetzals , and

9605-517: The hyperarid Atacama Desert . Rainforests and tropical dry forests used to encircle much of the northern Andes but are now greatly diminished , especially in the Chocó and inter-Andean valleys of Colombia. Opposite the humid Andean slopes are the relatively dry Andean slopes in most of western Peru, Chile, and Argentina. Along with several Interandean Valles , they are typically dominated by deciduous woodland, shrub and xeric vegetation, reaching

9718-468: The interface between truly terrestrial ecosystems and aquatic systems, making them inherently different from each other, yet highly dependent on both." In environmental decision-making, there are subsets of definitions that are agreed upon to make regulatory and policy decisions. Under the Ramsar international wetland conservation treaty , wetlands are defined as follows: An ecological definition of

9831-462: The known animal species in wetlands, and are considered the primary food web link between plants and higher animals (such as fish and birds). Depending on a wetland's geographic and topographic location, the functions it performs can support multiple ecosystem services , values, or benefits. United Nations Millennium Ecosystem Assessment and Ramsar Convention described wetlands as a whole to be of biosphere significance and societal importance in

9944-555: The larger glaciers descend to sea level. The Andes of Chile and Argentina can be divided into two climatic and glaciological zones: the Dry Andes and the Wet Andes . Since the Dry Andes extend from the latitudes of the Atacama Desert to the area of the Maule River , precipitation is more sporadic, and there are strong temperature oscillations. The line of equilibrium may shift drastically over short periods of time, leaving

10057-1187: The largest, behind that of the Athabasca Oil Sands in Alberta , Canada. Venezuela's non-conventional oil deposits of about 1,200 billion barrels (1.9 × 10 m), found primarily in the Orinoco oil sands, are estimated to approximately equal the world's reserves of conventional oil. A 2017 assessment found that 105,323 km², or 28%, of the ecoregion, is in protected areas. Protected areas include Aguaro-Guariquito National Park (5,857.5 km²), Cinaruco-Capanaparo National Park (5,843.68 km²), Tortuga Arrau Reserve (98.56 km²), and Caño Guaritico Wildlife Refuge (93.0 km²) in Venezuela, and El Tuparro National Natural Park (5,549.08 km²) in Colombia. 6°49′23.2″N 67°38′57.4″W  /  6.823111°N 67.649278°W  / 6.823111; -67.649278 Andes The Andes ( / ˈ æ n d iː z / AN -deez ), Andes Mountains or Andean Mountain Range ( Spanish : Cordillera de los Andes ; Quechua : Anti ) are

10170-523: The main crossover of the Andes between Argentina and Chile is still accomplished through the Paso Internacional Los Libertadores . Only recently have the ends of some highways that came rather close to one another from the east and the west been connected. Much of the transportation of passengers is done via aircraft. However, there is one railroad that connects Chile with Peru via the Andes, and there are others that make

10283-576: The nearly-7,000-metre (22,966 ft) highest mountains of the Andes. This dry steppe climate is considered to be typical of the subtropical position at 32–34° S. The valley bottoms have no woods, just dwarf scrub. The largest glaciers, for example the Plomo Glacier and the Horcones Glaciers, do not even reach 10 km (6.2 mi) in length and have only insignificant ice thickness. At glacial times, however, c. 20,000 years ago,

10396-837: The plants and animals controlled by the wetland hydrology are often additional components of the definitions. Wetlands can be tidal (inundated by tides) or non-tidal. The water in wetlands is either freshwater , brackish , saline , or alkaline . There are four main kinds of wetlands – marsh , swamp , bog , and fen (bogs and fens being types of peatlands or mires ). Some experts also recognize wet meadows and aquatic ecosystems as additional wetland types. Sub-types include mangrove forests , carrs , pocosins , floodplains , peatlands, vernal pools , sinks , and many others. The following three groups are used within Australia to classify wetland by type: Marine and coastal zone wetlands, inland wetlands and human-made wetlands. In

10509-541: The principal mines of the Spanish Empire in the New World. Río de la Plata and Argentina derive their names from the silver of Potosí. Wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water , either permanently, for years or decades, or only seasonally. Flooding results in oxygen -poor ( anoxic ) processes taking place, especially in

10622-402: The rainy season (summer), part of the rangeland is used for cropping (mainly potatoes, barley, broad beans, and wheat). Irrigation is helpful in advancing the sowing data of the summer crops, which guarantees an early yield in periods of food shortage. Also, by early sowing, maize can be cultivated higher up in the mountains (up to 3,800 m (12,500 ft)). In addition, it makes cropping in

10735-460: The same connection via southern Bolivia. There are multiple highways in Bolivia that cross the Andes. Some of these were built during a period of war between Bolivia and Paraguay , in order to transport Bolivian troops and their supplies to the war front in the lowlands of southeastern Bolivia and western Paraguay. For decades, Chile claimed ownership of land on the eastern side of the Andes. However, these claims were given up in about 1870 during

10848-548: The short-tailed opossum Monodelphis orinoci are endemic to the Llanos. The Llanos' wetlands supports around 70 species of water birds , including the scarlet ibis . A large portion of the distribution of the sharp-tailed ibis ( Cercibis oxycerca ) and white-bearded flycatcher ( Phelpsia inornata ) is in the Llanos. Native reptiles include the Orinoco crocodile ( Crocodylus intermedius ), spectacled caiman ( Caiman crocodilus ), Green anaconda ( Eunectes murinus ), and Arrau turtle ( Podocnemis expansa ), which live in

10961-721: The snow-covered peak of Cotopaxi . The mountains have a large effect on the temperatures of nearby areas. The snow line depends on the location. It is between 4,500 and 4,800 m (14,764 and 15,748 ft) in the tropical Ecuadorian, Colombian, Venezuelan, and northern Peruvian Andes, rising to 4,800–5,200 m (15,748–17,060 ft) in the drier mountains of southern Peru and northern Chile south to about 30°S before descending to 4,500 m (14,760 ft) on Aconcagua at 32°S , 2,000 m (6,600 ft) at 40°S , 500 m (1,640 ft) at 50°S , and only 300 m (980 ft) in Tierra del Fuego at 55°S ; from 50°S, several of

11074-429: The soil and underlying rock into aquifers which are the source of much of the world's drinking water . Wetlands can also act as recharge areas when the surrounding water table is low and as a discharge zone when it is high. Mangroves , coral reefs , salt marsh can help with shoreline stabilization and storm protection. Tidal and inter-tidal wetland systems protect and stabilize coastal zones. Coral reefs provide

11187-537: The soil of wetlands. Anaerobic and aerobic respiration in the soil influences the nutrient cycling of carbon, hydrogen, oxygen, and nitrogen, and the solubility of phosphorus thus contributing to the chemical variations in its water. Wetlands with low pH and saline conductivity may reflect the presence of acid sulfates and wetlands with average salinity levels can be heavily influenced by calcium or magnesium. Biogeochemical processes in wetlands are determined by soils with low redox potential. The life forms of

11300-412: The soils and plants at higher elevations. Plants and animals may vary within a wetland seasonally or in response to flood regimes. There are four main groups of hydrophytes that are found in wetland systems throughout the world. Submerged wetland vegetation can grow in saline and fresh-water conditions. Some species have underwater flowers, while others have long stems to allow the flowers to reach

11413-530: The sources of water include tidal wetlands , where the water source is ocean tides ); estuaries , water source is mixed tidal and river waters; floodplains , water source is excess water from overflowed rivers or lakes; and bogs and vernal ponds , water source is rainfall or meltwater . The world's largest wetlands include the Amazon River basin , the West Siberian Plain ,

11526-406: The spatial and temporal dispersion, flow, and physio-chemical attributes of surface and ground waters. Sources of hydrological flows into wetlands are predominantly precipitation , surface water (saltwater or freshwater), and groundwater. Water flows out of wetlands by evapotranspiration , surface flows and tides , and subsurface water outflow. Hydrodynamics (the movement of water through and from

11639-526: The subducting plate, rate of subduction, and hydration value of the mantle material. The highest rate of seismic activity is observed in the central portion of the boundary, between 33°S and 35°S. In this area, the angle of subduction is very low, meaning the subducting plate is nearly horizontal. Studies of mantle hydration across the subduction zone have shown a correlation between increased material hydration and lower-magnitude, more-frequent seismic activity. Zones exhibiting dehydration instead are thought to have

11752-637: The subduction of the Nazca Plate and the Antarctic Plate . To the east, the Andes range is bounded by several sedimentary basins , such as the Orinoco Basin , the Amazon Basin , the Madre de Dios Basin, and the Gran Chaco , that separate the Andes from the ancient cratons in eastern South America. In the south, the Andes share a long boundary with the former Patagonia Terrane . To the west,

11865-719: The surface of the water. When trees and shrubs comprise much of the plant cover in saturated soils, those areas in most cases are called swamps . The upland boundary of swamps is determined partly by water levels. This can be affected by dams Some swamps can be dominated by a single species, such as silver maple swamps around the Great Lakes . Others, like those of the Amazon basin , have large numbers of different tree species. Other examples include cypress ( Taxodium ) and mangrove swamps. Many species of fish are highly dependent on wetland ecosystems. Seventy-five percent of

11978-496: The surface. Submerged species provide a food source for native fauna, habitat for invertebrates, and also possess filtration capabilities. Examples include seagrasses and eelgrass . Floating water plants or floating vegetation are usually small, like those in the Lemnoideae subfamily (duckweeds). Emergent vegetation like the cattails ( Typha spp.), sedges ( Carex spp.) and arrow arum ( Peltandra virginica ) rise above

12091-464: The surrounding environment resulting in increased extinction rates in unfavorable and polluted environmental conditions. Reptiles such as snakes , lizards , turtles , alligators and crocodiles are common in wetlands of some regions. In freshwater wetlands of the Southeastern US, alligators are common and a freshwater species of crocodile occurs in South Florida. The Florida Everglades

12204-452: The tortuous terrain in places, villages and towns in the mountains—to which travel via motorized vehicles is of little use—are still located in the high Andes of Chile, Bolivia, Peru, and Ecuador . Locally, the relatives of the camel , the llama , and the alpaca continue to carry out important uses as pack animals, but this use has generally diminished in modern times. Donkeys , mules , and horses are also useful. The ancient peoples of

12317-440: The types of plants that live within them. Specifically, wetlands are characterized as having a water table that stands at or near the land surface for a long enough period each year to support aquatic plants . A more concise definition is a community composed of hydric soil and hydrophytes . Wetlands have also been described as ecotones , providing a transition between dry land and water bodies. Wetlands exist "...at

12430-477: The uplift of the Andes created a rain shadow on the western fringes of Chile, ocean currents and prevailing winds carried moisture away from the Chilean coast. This caused some areas of the subduction zone to be sediment-starved, causing excess friction and an increased rate of compressed coastal uplift. The main cause of the rise of the Andes is the compression of the western rim of the South American Plate due to

12543-642: The world for amphibians . The diversity of animals in the Andes is high, with almost 600 species of mammals (13% endemic), more than 1,700 species of birds (about 1/3 endemic), more than 600 species of reptiles (about 45% endemic), and almost 400 species of fish (about 1/3 endemic). The vicuña and guanaco can be found living in the Altiplano , while the closely related domesticated llama and alpaca are widely kept by locals as pack animals and for their meat and wool . The crepuscular (active during dawn and dusk) chinchillas , two threatened members of

12656-470: The world's largest reserve of the element. Early Mesozoic and Neogene plutonism in Bolivia's Cordillera Central created the Bolivian tin belt as well as the famous, now-mostly-depleted, deposits of Cerro Rico de Potosí . The Andes Mountains, initially inhabited by hunter-gatherers , experienced the development of agriculture and the rise of politically centralized civilizations , which culminated in

12769-408: Was used for the production of chicha , important to Andean native people. Currently, tobacco , cotton , and coffee are the main export crops. Coca , despite eradication programs in some countries, remains an important crop for legal local use in a mildly stimulating herbal tea , and illegally for the production of cocaine . In unirrigated land, pasture is the most common type of land use. In

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