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122-708: The Oxford Clay (or Oxford Clay Formation ) is a Jurassic marine sedimentary rock formation underlying much of southeast England , from as far west as Dorset and as far north as Yorkshire . The Oxford Clay Formation dates to the Jurassic, specifically, the Callovian and Oxfordian ages, and comprises two main facies . The lower facies comprises the Peterborough Member , a fossiliferous organic-rich mudstone . This facies and its rocks are commonly known as lower Oxford Clay. The upper facies comprises

244-655: A 70 km diameter impact structure buried beneath the Kalahari desert in northern South Africa. The impact is dated to the Tithonian, approximately 146.06 ± 0.16 Mya. Another major structure is the Puchezh-Katunki crater , 40 kilometres in diameter, buried beneath Nizhny Novgorod Oblast in western Russia. The impact has been dated to the Sinemurian, 195.9 ± 1.0 Ma. At the beginning of the Jurassic, all of

366-499: A dominant component of Jurassic floras. The Late Triassic and Jurassic was a major time of diversification of conifers, with most modern conifer groups appearing in the fossil record by the end of the Jurassic, having evolved from voltzialean ancestors. Araucarian conifers have their first unambiguous records during the Early Jurassic, and members of the modern genus Araucaria were widespread across both hemispheres by

488-618: A global value of around -3% to -4%. In addition, numerous smaller scale carbon isotope excursions are globally recorded on the falling limb of the larger negative δ C excursion. Although the PTo-E is not associated with a decrease in δ C analogous to the TOAE's, volcanism is nonetheless believed to have been responsible for its onset as well, with the carbon injection most likely having an isotopically heavy, mantle-derived origin. The Karoo-Ferrar magmatism released so much carbon dioxide that it disrupted

610-419: A higher diversity ecological assemblage of lycophytes , conifers , seed ferns , and wet-adapted ferns is observed in the palaeobotanical and palynological record over the course of the TOAE. The coincidence of the zenith of Classopolis and the decline of seed ferns and spore producing plants with increased mercury loading implicates heavy metal poisoning as a key contributor to the floristic crisis during

732-647: A kink in one of the plate boundaries, resulting in the formation of the Pacific Plate at the centre of the junction. During the Middle to early Late Jurassic, the Sundance Seaway , a shallow epicontinental sea , covered much of northwest North America. The eustatic sea level is estimated to have been close to present levels during the Hettangian and Sinemurian, rising several tens of metres during

854-545: A mitigating factor that ameliorated to a degree the oppressively anoxic conditions that were widespread across much of the Tethys. The enhanced hydrological cycle during early Toarcian warming caused lakes to grow in size. During the anoxic event, the Sichuan Basin was transformed into a giant lake, which was believed to be approximately thrice as large as modern-day Lake Superior . Lacustrine sediments deposited as

976-453: A result of high ocean acidity and temperature inhibiting its mineralisation into apatite; the abundance of phosphorus in marine environments caused further eutrophication and consequent anoxia in a positive feedback loop. The end-Jurassic transition was originally considered one of eight mass extinctions, but is now considered to be a complex interval of faunal turnover, with the increase in diversity of some groups and decline in others, though

1098-664: A result of this lake's existence are represented by the Da’anzhai Member of the Ziliujing Formation . Roughly ~460 gigatons (Gt) of organic carbon and ~1,200 Gt of inorganic carbon were likely sequestered by this lake over the course of the TOAE. The TOAE and the Palaeocene-Eocene Thermal Maximum have been proposed as analogues to modern anthropogenic global warming based on the comparable quantity of greenhouse gases released into

1220-750: A skeleton." Eustreptospondylus E. oxoniensis Upper Disarticulated skull and skeleton, with some referred limb elements. Sauropoda Indeterminate The caudal vertebrae from Cambridgeshire were mistakenly considered part of the syntypic series of "Ornithopsis" leedsi by Upchurch and Martin (2003). Theropoda Indeterminate Middle Metriacanthosaurus Ophthalmosaurus O. icenicus A ophthalmosaurid ichthyosaur Cryptoclidus C. eurymerus A cryptoclidid C. richardsoni A cryptoclidid E. powelli A thalassophonean pliosaurid Liopleurodon L. ferox A thalassophonean pliosaurid L. pachydeirus A thalassophonean pliosaurid Jurassic The start of

1342-411: A sparse Pliensbachian marine vertebrate fossil record. The TOAE is suggested to have caused the extinction of various clades of dinosaurs, including coelophysids , dilophosaurids , and many basal sauropodomorph clades, as a consequence of the remodelling of terrestrial ecosystems caused by global climate change. Some heterodontosaurids and thyreophorans also perished in the extinction event. In

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1464-468: A transition from icehouse to greenhouse conditions further retarded ocean circulation, aiding the establishment of anoxic conditions. Geochemical evidence from what was then the northwestern European epicontinental sea suggests that a shift from cooler, more saline water conditions to warmer, fresher conditions prompted the development of significant density stratification of the water column and induced anoxia. Extensive organic carbon burial induced by anoxia

1586-492: Is Austrohamia minuta from the Early Jurassic (Pliensbachian) of Patagonia, known from many parts of the plant. The reproductive structures of Austrohamia have strong similarities to those of the primitive living cypress genera Taiwania and Cunninghamia . By the Middle to Late Jurassic Cupressaceae were abundant in warm temperate–tropical regions of the Northern Hemisphere, most abundantly represented by

1708-494: Is divided into three epochs : Early, Middle, and Late. Similarly, in stratigraphy , the Jurassic is divided into the Lower Jurassic , Middle Jurassic , and Upper Jurassic series . Geologists divide the rocks of the Jurassic into a stratigraphic set of units called stages , each formed during corresponding time intervals called ages. Stages can be defined globally or regionally. For global stratigraphic correlation,

1830-565: Is followed by the Kimmeridgian Warm Interval (KWI) between 164 and 150 million years ago. Based on fossil wood distribution, this was one of the wettest intervals of the Jurassic. The Pangaean interior had less severe seasonal swings than in previous warm periods as the expansion of the Central Atlantic and Western Indian Ocean provided new sources of moisture. A prominent drop in temperatures occurred during

1952-470: Is fourth order, with a periodicity of approximately 410,000 years. During the Early Jurassic the world's oceans transitioned from an aragonite sea to a calcite sea chemistry, favouring the dissolution of aragonite and precipitation of calcite . The rise of calcareous plankton during the Middle Jurassic profoundly altered ocean chemistry, with the deposition of biomineralized plankton on

2074-618: Is linked to the Jura Mountains , a forested mountain range that mainly follows the France–Switzerland border . The name "Jura" is derived from the Celtic root * jor via Gaulish *iuris "wooded mountain", which was borrowed into Latin as a name of a place and evolved into Juria and finally Jura . During a tour of the region in 1795, German naturalist Alexander von Humboldt recognized carbonate deposits within

2196-475: Is marked by a globally documented high amplitude negative carbon isotope excursion, as well as the deposition of black shales and the extinction and collapse of carbonate-producing marine organisms, associated with a major rise in global temperatures. The TOAE is often attributed to the eruption of the Karoo-Ferrar large igneous provinces and the associated increase of carbon dioxide concentration in

2318-577: Is named after the city of Oxford in England and was named by Alcide d'Orbigny in 1844 in reference to the Oxford Clay . The base of the Oxfordian lacks a defined GSSP. W. J. Arkell in studies in 1939 and 1946 placed the lower boundary of the Oxfordian as the first appearance of the ammonite Quenstedtoceras mariae (then placed in the genus Vertumniceras ). Subsequent proposals have suggested

2440-707: Is the setting of the world's largest oil reserves, including the Ghawar Field , the world's largest oil field. The Jurassic-aged Sargelu and Naokelekan formations are major source rocks for oil in Iraq . Over 1500 gigatons of Jurassic coal reserves are found in north-west China, primarily in the Turpan-Hami Basin and the Ordos Basin . Major impact structures include the Morokweng impact structure ,

2562-622: Is used to mark the beginnings of stages, as well as smaller timespans within stages, referred to as "ammonite zones"; these, in turn, are also sometimes subdivided further into subzones. Global stratigraphy is based on standard European ammonite zones, with other regions being calibrated to the European successions. The oldest part of the Jurassic Period has historically been referred to as the Lias or Liassic, roughly equivalent in extent to

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2684-593: The Cleveland , West Netherlands, and South German Basins. Valdorbia, a site in the Umbria-Marche Apennines, also exhibited euxinia during the anoxic event. There is less evidence of euxinia outside the northwestern Tethys, and it likely only occurred transiently in basins in Panthalassa and the southwestern Tethys. Due to the clockwise circulation of the oceanic gyre in the western Tethys and

2806-814: The Early Toarcian palaeoenvironmental crisis , or the Jenkyns Event , was an extinction event that occurred during the early part of the Toarcian age, approximately 183 million years ago, during the Early Jurassic . The extinction event had two main pulses, the first being the Pliensbachian-Toarcian boundary event ( PTo-E ). The second, larger pulse, the Toarcian Oceanic Anoxic Event ( TOAE ),

2928-513: The International Commission on Stratigraphy (ICS) ratify global stages based on a Global Boundary Stratotype Section and Point (GSSP) from a single formation (a stratotype ) identifying the lower boundary of the stage. The ages of the Jurassic from youngest to oldest are as follows: Jurassic stratigraphy is primarily based on the use of ammonites as index fossils . The first appearance datum of specific ammonite taxa

3050-517: The Karoo-Ferrar Large Igneous Province is generally attributed to have caused the surge in atmospheric carbon dioxide levels. Argon-argon dating of Karoo-Ferrar rhyolites points to a link between Karoo-Ferrar volcanism and the extinction event, a conclusion reinforced by uranium-lead dating and palaeomagnetism. Occurring during a broader, gradual positive carbon isotope excursion as measured by δ C values,

3172-728: The Redcar Mudstone Formation , and was ratified in 2005. The beginning of the Pliensbachian is defined by the first appearance of the ammonite Bifericeras donovani . The village Thouars (Latin: Toarcium ), just south of Saumur in the Loire Valley of France , lends its name to the Toarcian Stage. The Toarcian was named by Alcide d'Orbigny in 1842, with the original locality being Vrines quarry around 2 km northwest of Thouars. The GSSP for

3294-435: The elegantulum subzone of the serpentinum ammonite zone, during a marked, pronounced warming interval. The TOAE lasted for approximately 500,000 years, though a range of estimates from 200,000 to 1,000,000 years have also been given. The PTo-E primarily affected shallow water biota, while the TOAE was the more severe event for organisms living in deep water. Geological, isotopic, and palaeobotanical evidence suggests

3416-475: The mirabile subzone of the tenuicostatum ammonite zone, coinciding with a slight drop in oxygen concentrations and the beginning of warming following a late Pliensbachian cool period. This first pulse, occurring near the Pliensbachian-Toarcian boundary, is referred to as the PTo-E. The TOAE itself occurred near the tenuicostatum – serpentinum ammonite biozonal boundary, specifically in

3538-636: The Aalenian, precessionally forced climatic changes dictated peatland wildfire magnitude and frequency. The European climate appears to have become noticeably more humid at the Aalenian-Bajocian boundary but then became more arid during the middle Bajocian. A transient ice age possibly occurred in the late Bajocian. The Callovian-Oxfordian boundary at the end of the MJCI witnessed particularly notable global cooling, potentially even an ice age. This

3660-748: The Bajocian Stage after the town of Bayeux (Latin: Bajoce ) in Normandy, France. The GSSP for the base of the Bajocian is located in the Murtinheira section at Cabo Mondego , Portugal; it was ratified in 1997. The base of the Bajocian is defined by the first appearance of the ammonite Hyperlioceras mundum . The Bathonian is named after the city of Bath , England, introduced by Belgian geologist d'Omalius d'Halloy in 1843, after an incomplete section of oolitic limestones in several quarries in

3782-593: The Black Jurassic and the overlying clayey sandstone and ferruginous oolite of the Brown Jurassic sequences of southwestern Germany. The GSSP for the base of the Aalenian is located at Fuentelsaz in the Iberian range near Guadalajara, Spain , and was ratified in 2000. The base of the Aalenian is defined by the first appearance of the ammonite Leioceras opalinum . Alcide d'Orbigny in 1842 named

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3904-560: The Black Jurassic in England by William Conybeare and William Phillips in 1822. William Phillips, the geologist, worked with William Conybeare to find out more about the Black Jurassic in England. The French palaeontologist Alcide d'Orbigny in papers between 1842 and 1852 divided the Jurassic into ten stages based on ammonite and other fossil assemblages in England and France, of which seven are still used, but none has retained its original definition. The German geologist and palaeontologist Friedrich August von Quenstedt in 1858 divided

4026-655: The Callovian–Oxfordian boundary, peaking possibly as high as 140 metres above present sea level at the Kimmeridgian–Tithonian boundary. The sea levels falls in the late Tithonian, perhaps to around 100 metres, before rebounding to around 110 metres at the Tithonian–Berriasian boundary. The sea level within the long-term trends across the Jurassic was cyclical, with 64 fluctuations, 15 of which were over 75 metres. The most noted cyclicity in Jurassic rocks

4148-464: The Early Jurassic, but also including part of the preceding Rhaetian . The Hettangian Stage was named by Swiss palaeontologist Eugène Renevier in 1864 after Hettange-Grande in north-eastern France. The GSSP for the base of the Hettangian is located at the Kuhjoch Pass, Karwendel Mountains , Northern Calcareous Alps , Austria; it was ratified in 2010. The beginning of the Hettangian, and thus

4270-465: The Early Toarcian diversity collapse. Belemnite richness in the northwestern Tethys dropped during the PTo-E but slightly increased across the TOAE. Belemnites underwent a major change in habitat preference from cold, deep waters to warm, shallow waters. Their average rostrum size also increased, though this trend heavily varied depending on the lineage of belemnites. The Toarcian extinction

4392-519: The Greek goddess of dawn . His name was chosen by Albert Oppel for this stratigraphical stage because the Tithonian finds itself hand in hand with the dawn of the Cretaceous. The base of the Tithonian currently lacks a GSSP. The working definition for the base of the Tithonian is the first appearance of the ammonite genus Gravesia . The upper boundary of the Jurassic is currently undefined, and

4514-563: The Hispanic Corridor into European seas after the extinction event, aided in their dispersal by higher sea levels. The TOAE had minor effects on marine reptiles, in stark contrast to the major impact it had on many clades of marine invertebrates. In fact, in the Southwest German Basin, ichthyosaur diversity was higher after the extinction interval, although this may be in part a sampling artefact resulting from

4636-792: The Jura Mountains as geologically distinct from the Triassic aged Muschelkalk of southern Germany , but he erroneously concluded that they were older. He then named them Jura-Kalkstein ('Jura limestone') in 1799. In 1829, the French naturalist Alexandre Brongniart published a book entitled Description of the Terrains that Constitute the Crust of the Earth or Essay on the Structure of

4758-483: The Jurassic as a whole, is marked by the first appearance of the ammonite Psiloceras spelae tirolicum in the Kendlbach Formation exposed at Kuhjoch. The base of the Jurassic was previously defined as the first appearance of Psiloceras planorbis by Albert Oppel in 1856–58, but this was changed as the appearance was seen as too localised an event for an international boundary. The Sinemurian Stage

4880-407: The Jurassic of Asia has strap-shaped ginkgo-like leaves with highly distinct reproductive structures with similarities to those of peltasperm and corystosperm seed ferns, has been suggested to be a member of Ginkgoales sensu lato. Toarcian Oceanic Anoxic Event The Toarcian extinction event , also called the Pliensbachian-Toarcian extinction event , the Early Toarcian mass extinction ,

5002-572: The Jurassic was marked by the major Triassic–Jurassic extinction event , associated with the eruption of the Central Atlantic Magmatic Province (CAMP). The beginning of the Toarcian Age started around 183 million years ago and is marked by the Toarcian Oceanic Anoxic Event , a global episode of oceanic anoxia , ocean acidification , and elevated global temperatures associated with extinctions, likely caused by

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5124-513: The Jurassic was warmer than the present, and there were no ice caps . Forests grew close to the poles, with large arid expanses in the lower latitudes. On land, the fauna transitioned from the Triassic fauna, dominated jointly by dinosauromorph and pseudosuchian archosaurs , to one dominated by dinosaurs alone. The first stem-group birds appeared during the Jurassic, evolving from a branch of theropod dinosaurs. Other major events include

5246-797: The Jurassic, both the North and South Pole were covered by oceans. Beginning in the Early Jurassic, the Boreal Ocean was connected to the proto-Atlantic by the "Viking corridor" or Transcontinental Laurasian Seaway, a passage between the Baltic Shield and Greenland several hundred kilometers wide. During the Callovian, the Turgai Epicontinental Sea formed, creating a marine barrier between Europe and Asia. Madagascar and Antarctica began to rift away from Africa during

5368-686: The Jurassic, the North Atlantic Ocean remained relatively narrow, while the South Atlantic did not open until the Cretaceous. The continents were surrounded by Panthalassa , with the Tethys Ocean between Gondwana and Asia. At the end of the Triassic, there was a marine transgression in Europe, flooding most parts of central and western Europe transforming it into an archipelago of islands surrounded by shallow seas. During

5490-572: The Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined GSSP. Placing a GSSP for this boundary has been difficult because of the strong regionality of most biostratigraphic markers, and lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes ), that could be used to define or correlate a boundary. Calpionellids , an enigmatic group of planktonic protists with urn-shaped calcitic tests briefly abundant during

5612-713: The Known Lands of the Earth. In this book, Brongniart used the phrase terrains jurassiques when correlating the "Jura-Kalkstein" of Humboldt with similarly aged oolitic limestones in Britain, thus coining and publishing the term "Jurassic". The German geologist Leopold von Buch in 1839 established the three-fold division of the Jurassic, originally named from oldest to the youngest: the Black Jurassic , Brown Jurassic , and White Jurassic . The term " Lias " had previously been used for strata of equivalent age to

5734-526: The Late Jurassic (Kimmeridgian) of Scotland, which remains the only known unequivocal fossil of the group before the Cretaceous. Despite being the earliest known member of the Pinaceae, Eathiestrobus appears to be a member of the pinoid clade of the family, suggesting that the initial diversification of Pinaceae occurred earlier than has been found in the fossil record. The earliest record of

5856-513: The Lower Oxford Clay. Geographically, they are located in Cambridgeshire, England. Indeterminate euronithopod remains stratigraphically present in the Lower Oxford Clay and geographically located in Cambridgeshire, England. Callovosaurus C. leedsi Lower "Femur." Lexovisaurus L. durobivensis Lower Indeterminate Loricatosaurus L. priscus Lower Sarcolestes S. leedsi Lower "Partial mandible." Cetiosauriscus C. stewarti Lower "Rear half of

5978-416: The Middle Jurassic. Also abundant during the Jurassic is the extinct family Cheirolepidiaceae , often recognised through their highly distinctive Classopolis pollen. Jurassic representatives include the pollen cone Classostrobus and the seed cone Pararaucaria . Araucarian and Cheirolepidiaceae conifers often occur in association. The oldest definitive record of the cypress family ( Cupressaceae )

6100-445: The Middle to Late Jurassic, corresponding to the increasing aridity of the region. Ginkgoales , of which the sole living species is Ginkgo biloba , were more diverse during the Jurassic: they were among the most important components of Eurasian Jurassic floras and were adapted to a wide variety of climatic conditions. The earliest representatives of the genus Ginkgo , represented by ovulate and pollen organs similar to those of

6222-405: The North Atlantic Ocean with eastern Panthalassa. Palaeontological data suggest that the seaway had been open since the Early Jurassic. As part of the Nevadan orogeny , which began during the Triassic, the Cache Creek Ocean closed, and various terranes including the large Wrangellia Terrane accreted onto the western margin of North America. By the Middle Jurassic the Siberian plate and

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6344-462: The North China-Amuria block had collided, resulting in the closure of the Mongol-Okhotsk Ocean . During the Early Jurassic, around 190 million years ago, the Pacific Plate originated at the triple junction of the Farallon , Phoenix , and Izanagi tectonic plates , the three main oceanic plates of Panthalassa. The previously stable triple junction had converted to an unstable arrangement surrounded on all sides by transform faults because of

6466-410: The PTo-E and TOAE. In northeastern Panthalassa, in what is now British Columbia , euxinia dominated anoxic bottom waters. The early stages of the TOAE were accompanied by a decrease in the acidity of seawater following a substantial decrease prior to the TOAE. Seawater pH then dropped close to the middle of the event, strongly acidifying the oceans. The sudden decline of carbonate production during

6588-463: The South Midlands. The Oxford Clay is well known for its rich fossil record of fish and invertebrates . Many of the fossils are well preserved, occasionally some are found exceptionally well preserved. Animals which lived in the Oxford Clay Sea include plesiosaurs , marine crocodiles , ichthyosaurs , cephalopods (such as belemnites ), bivalves (such as Gryphaea ), and a variety of gastropods . Dinosaur eggs are stratigraphically present in

6710-402: The TOAE is preceded by a global negative δ C excursion recognised in fossil wood, organic carbon, and carbonate carbon in the tenuicostatum ammonite zone of northwestern Europe, with this negative δ C shift being the result of volcanic discharge of light carbon. The global ubiquity of this negative δ C excursion has been called into question, however, due to its absence in certain deposits from

6832-402: The TOAE is widely believed to be the result of this abrupt episode of ocean acidification . Additionally, the enhanced recycling of phosphorus back into seawater as a result of high temperatures and low seawater pH inhibited its mineralisation into apatite, helping contribute to oceanic anoxia. The abundance of phosphorus in marine environments created a positive feedback loop whose consequence

6954-502: The TOAE represented one of the most severe extinctions in their evolutionary history. While the event had significant impact on marine invertebrates, it had little effect on marine reptiles. During the TOAE, the Sichuan Basin was transformed into a giant lake , probably three times the size of modern-day Lake Superior , represented by the Da'anzhai Member of the Ziliujing Formation . The lake likely sequestered ~460 gigatons (Gt) of organic carbon and ~1,200 Gt of inorganic carbon during

7076-434: The TOAE, as shown by a positive δ S excursion in carbonate-associated sulphate occurs synchronously with the positive δ C excursion in carbonate carbon during the falciferum ammonite zone. This positive δ S excursion has been attributed to the depletion of isotopically light sulphur in the marine sulphate reservoir that resulted from microbial sulphur reduction in anoxic waters. Similar positive δ S excursions corresponding to

7198-466: The TOAE. Carbonate platforms collapsed during both the PTo-E and the TOAE. Enhanced continental weathering and nutrient runoff was the dominant driver of carbonate platform decline in the PTo-E, while the biggest culprits during the TOAE were heightened storm activity and a decrease in the pH of seawater. The recovery from the mass extinction among benthos commenced with the recolonisation of barren locales by opportunistic pioneer taxa. Benthic recovery

7320-445: The TOAE. Concentrations of phosphorus, magnesium, and manganese rose in the oceans. A -0.5% excursion in δ Ca provides further evidence of increased continental weathering. Osmium isotope ratios confirm further still a major increase in weathering. The enhanced continental weathering in turn led to increased eutrophication that helped drive the anoxic event in the oceans. Continual transport of continentally weathered nutrients into

7442-586: The TOAE; the increase in clastic sedimentation was synchronous with excursions in Os/ Os, Sr/ Sr, and δ Ca. Additionally, the Toarcian was punctuated by intervals of extensive kaolinite enrichment. These kaolinites correspond to negative oxygen isotope excursions and high Mg/Ca ratios and are thus reflective of climatic warming events that characterised much of the Toarcian. Likewise, illitic/smectitic clays were also common during this hyperthermal perturbation. The Intertropical Convergence Zone (ITCZ) migrated southwards across southern Gondwana, turning much of

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7564-414: The Tithonian, known as the Early Tithonian Cooling Event (ETCE). The end of the Jurassic was marked by the Tithonian–early Barremian Cool Interval (TBCI), beginning 150 million years ago and continuing into the Early Cretaceous . The Toarcian Oceanic Anoxic Event (TOAE), also known as the Jenkyns Event, was an episode of widespread oceanic anoxia during the early part of the Toarcian Age, c. 183 Mya. It

7686-529: The Toarcian around 174 million years ago. During the Toarcian Warm Interval, ocean surface temperatures likely exceeded 30 °C (86 °F), and equatorial and subtropical (30°N–30°S) regions are likely to have been extremely arid, with temperatures in the interior of Pangea likely in excess of 40 °C (104 °F).The Toarcian Warm Interval is followed by the Middle Jurassic Cool Interval (MJCI) between 174 and 164 million years ago, which may have been punctuated by brief, ephemeral icehouse intervals. During

7808-431: The Toarcian mass extinction. Poisoning by mercury, along with chromium, copper, cadmium, arsenic, and lead is speculated to be responsible for heightened rates of spore malformation and dwarfism concomitant with enrichments in all these toxic metals. The TOAE was associated with widespread phosphatisation of marine fossils believed to result from the warming-induced increase in weathering that increased phosphate flux into

7930-466: The abrupt warming interval associated with the TOAE. This global warming, driven by rising atmospheric carbon dioxide, was the mainspring of the early Toarcian environmental crisis. Carbon dioxide levels rose from about 500 ppm to about 1,000 ppm. Seawater warmed by anywhere between 3 °C and 7 °C, depending on latitude. At the height of this supergreenhouse interval, global sea surface temperatures (SSTs) averaged about 21 °C. The eruption of

8052-489: The appearance of the earliest crabs and modern frogs , salamanders and lizards . Mammaliaformes , one of the few cynodont lineages to survive the end of the Triassic, continued to diversify throughout the period, with the Jurassic seeing the emergence of the first crown group mammals . Crocodylomorphs made the transition from a terrestrial to an aquatic life. The oceans were inhabited by marine reptiles such as ichthyosaurs and plesiosaurs , while pterosaurs were

8174-587: The approximate time intervals corresponding to the PTo-E and TOAE have likewise been invoked as tell-tale evidence of the ecological calamity's cause being a large igneous province, although some researchers attribute these elevated mercury levels to increased terrigenous flux. There is evidence that the motion of the African Plate suddenly changed in velocity, shifting from mostly northward movement to southward movement. Such shifts in plate motion are associated with similar large igneous provinces emplaced in other time intervals. A 2019 geochronological study found that

8296-428: The atmosphere, as well as the possible associated release of methane clathrates . This likely accelerated the hydrological cycle and increased silicate weathering , as evidenced by an increased amount of organic matter of terrestrial origin found in marine deposits during the TOAE. Groups affected include ammonites, ostracods , foraminifera , bivalves , cnidarians , and especially brachiopods , for which

8418-700: The atmosphere. Carbon release via metamorphic heating of coal has been criticised as a major driver of the environmental perturbation, however, on the basis that coal transects themselves do not show the δ C excursions that would be expected if significant quantities of thermogenic methane were released, suggesting that much of the degassed emissions were either condensed as pyrolytic carbon or trapped as coalbed methane. In addition, possible associated release of deep sea methane clathrates has been potentially implicated as yet another cause of global warming. Episodic melting of methane clathrates dictated by Milankovitch cycles has been put forward as an explanation fitting

8540-407: The base of the Toarcian is located at Peniche, Portugal , and was ratified in 2014. The boundary is defined by the first appearance of ammonites belonging to the subgenus Dactylioceras ( Eodactylites ) . The Aalenian is named after the city of Aalen in Germany. The Aalenian was defined by Swiss geologist Karl Mayer-Eymar in 1864. The lower boundary was originally between the dark clays of

8662-417: The carbon cycle disruption. It has also been hypothesised that the release of cryospheric methane trapped in permafrost amplified the warming and its detrimental effects on marine life. Obliquity-paced carbon isotope excursions have been interpreted as some researchers as reflective of permafrost decline and consequent greenhouse gas release. The TOAE is believed to be the second largest anoxic event of

8784-447: The dominant flying vertebrates . Modern sharks and rays first appeared and diversified during the period, while the first known crown-group teleost fish appeared near the end of the period. The flora was dominated by ferns and gymnosperms , including conifers , of which many modern groups made their first appearance during the period, as well as other groups like the extinct Bennettitales . The chronostratigraphic term "Jurassic"

8906-623: The emplacement of the Karoo-Ferrar large igneous province and the TOAE were not causally linked, and simply happened to occur rather close in time, contradicting mainstream interpretations of the TOAE. The authors of the study conclude that the timeline of the TOAE does not match up with the course of activity of the Karoo-Ferrar magmatic event. The large igneous province also intruded into coal seams, releasing even more carbon dioxide and methane than it otherwise would have. Magmatic sills are also known to have intruded into shales rich in organic carbon, causing additional venting of carbon dioxide into

9028-688: The end of the Jurassic. The oldest unambiguous members of Podocarpaceae are known from the Jurassic, found across both hemispheres, including Scarburgia and Harrisiocarpus from the Middle Jurassic of England, as well as unnamed species from the Middle-Late Jurassic of Patagonia. During the Early Jurassic, the flora of the mid-latitudes of Eastern Asia were dominated by the extinct deciduous broad leafed conifer Podozamites , which appears to not be closely related to any living family of conifer. Its range extended northwards into polar latitudes of Siberia and then contracted northward in

9150-653: The end of the Triassic, the Peltaspermaceae became extinct in most parts of the world, with Lepidopteris persisting into the Early Jurassic in Patagonia. Dicroidium , a corystosperm seed fern that was a dominant part of Gondwanan floral communities during the Triassic, also declined at the Triassic–Jurassic boundary, surviving as a relict in Antarctica into the Early Jurassic. Conifers formed

9272-465: The eruption of the Karoo-Ferrar large igneous provinces . The end of the Jurassic, however, has no clear, definitive boundary with the Cretaceous and is the only boundary between geological periods to remain formally undefined. By the beginning of the Jurassic, the supercontinent Pangaea had begun rifting into two landmasses: Laurasia to the north and Gondwana to the south. The climate of

9394-417: The event. Seawater pH , which had already substantially decreased prior to the event, increased slightly during the early stages of the TOAE, before dropping to its lowest point around the middle of the event. This ocean acidification is the probable cause of the collapse of carbonate production. Additionally, anoxic conditions were exacerbated by enhanced recycling of phosphorus back into ocean water as

9516-506: The evidence for this is primarily European, probably controlled by changes in eustatic sea level. There is no evidence of a mass extinction of plants at the Triassic–Jurassic boundary. At the Triassic–Jurassic boundary in Greenland, the sporomorph (pollen and spores) record suggests a complete floral turnover. An analysis of macrofossil floral communities in Europe suggests that changes were mainly due to local ecological succession . At

9638-686: The first appearance of Cardioceras redcliffense as the lower boundary. The village of Kimmeridge on the coast of Dorset , England, is the origin of the name of the Kimmeridgian. The stage was named by Alcide d'Orbigny in 1842 in reference to the Kimmeridge Clay . The GSSP for the base of the Kimmeridgian is the Flodigarry section at Staffin Bay on the Isle of Skye , Scotland , which

9760-465: The first appearance of the ammonite Strambergella jacobi , formerly placed in the genus Berriasella , but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of C. alpina . The Kimmeridge Clay and equivalents are the major source rock for the North Sea oil . The Arabian Intrashelf Basin, deposited during the Middle and Late Jurassic,

9882-402: The genus Elatides . The Jurassic also saw the first appearances of some modern genera of cypresses, such as Sequoia . Members of the extinct genus Schizolepidopsis which likely represent a stem-group to the pine family ( Pinaceae ), were widely distributed across Eurasia during the Jurassic. The oldest unambiguous record of Pinaceae is the pine cone Eathiestrobus , known from

10004-403: The imprint of the 9 Myr long-term carbon cycle that was otherwise steady and stable during the Jurassic and Early Cretaceous. The values of Os/ Os rose from ~0.40 to ~0.53 during the PTo-E and from ~0.42 to ~0.68 during the TOAE, and many scholars conclude this change in osmium isotope ratios evidences the responsibility of this large igneous province for the biotic crises. Mercury anomalies from

10126-562: The last 300 Ma, and possibly the largest of the Phanerozoic. A positive δ C excursion, likely resulting from the mass burial of organic carbon during the anoxic event, is known from the falciferum ammonite zone, chemostratigraphically identifying the TOAE. Large igneous province resulted in increased silicate weathering and an acceleration of the hydrological cycle , as shown by a increased amount of terrestrially derived organic matter found in sedimentary rocks of marine origin during

10248-477: The late Early Jurassic in association with the eruption of the Karoo-Ferrar large igneous provinces , opening the western Indian Ocean and beginning the fragmentation of Gondwana. At the beginning of the Jurassic, North and South America remained connected, but by the beginning of the Late Jurassic they had rifted apart to form the Caribbean Seaway, also known as the Hispanic Corridor, which connected

10370-414: The late Pliensbachian was an icehouse period. These ice sheets are believed to have been thin and stretched into lower latitudes, making them extremely sensitive to temperature changes. A warming trend lasting from the latest Pliensbachian to the earliest Toarcian was interrupted by a "cold snap" in the middle polymorphum zone, equivalent to the tenuicostatum ammonite zone, which was then followed by

10492-472: The late Sinemurian–Pliensbachian before regressing to near present levels by the late Pliensbachian. There seems to have been a gradual rise to a peak of ~75 m above present sea level during the Toarcian. During the latest part of the Toarcian, the sea level again dropped by several tens of metres. It progressively rose from the Aalenian onwards, aside from dips of a few tens of metres in the Bajocian and around

10614-403: The latest Jurassic to earliest Cretaceous, have been suggested to represent the most promising candidates for fixing the Jurassic–Cretaceous boundary In particular, the first appearance Calpionella alpina , co-inciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous. The working definition for the boundary has often been placed as

10736-404: The methane released from ocean sediments was rapidly sequestered, buffering its ability to act as a major positive feedback, and that methane clathrate dissociation occurred too late to have had an appreciable causal impact on the extinction event. Hypothetical release of methane clathrates extremely depleted in heavy carbon isotopes has furthermore been considered unnecessary as an explanation for

10858-718: The middle Oxford Clay, the Stewartby Member , and the upper Oxford Clay, the Weymouth Member . The upper facies is a fossil poor assemblage of calcareous mudstones. Oxford Clay appears at the surface around Oxford , Peterborough and Weymouth and is exposed in many quarries around these areas. The top of the Lower Oxford Clay shows a lithological change, where fissile shale changes to grey mudstone. The Middle and Upper Oxford Clays differ slightly, as they are separated by an argillaceous limestone in

10980-428: The modern species, are known from the Middle Jurassic in the Northern Hemisphere. Several other lineages of ginkgoaleans are known from Jurassic rocks, including Yimaia , Grenana , Nagrenia and Karkenia . These lineages are associated with Ginkgo- like leaves, but are distinguished from living and fossil representatives of Ginkgo by having differently arranged reproductive structures. Umaltolepis from

11102-415: The most dire crises in their evolutionary history. Brachiopod taxa of large size declined significantly in abundance. Uniquely, the brachiopod genus Soaresirhynchia thrived during the later stages of the TOAE due to its low metabolic rate and slow rate of growth, making it a disaster taxon . The species S. bouchardi is known to have been a pioneer species that colonised areas denuded of brachiopods in

11224-478: The northwestern Tethyan region. Ostracods also suffered a major diversity loss, with almost all ostracod clades’ distributions during the time interval corresponding to the serpentinum zone shifting towards higher latitudes to escape intolerably hot conditions near the Equator. Bivalves likewise experienced a significant turnover. The decline of bivalves exhibiting high endemism with narrow geographic ranges

11346-440: The observed shifts in the carbon isotope record. Other studies contradict and reject the methane hydrate hypothesis, however, concluding that the isotopic record is too incomplete to conclusively attribute the isotopic excursion to methane hydrate dissociation, that carbon isotope ratios in belemnites and bulk carbonates are incongruent with the isotopic signature expected from a massive release of methane clathrates, that much of

11468-468: The ocean enabled high levels of primary productivity to be maintained over the course of the TOAE. Rising sea levels contributed to ocean deoxygenation; as rising sea levels inundated low-lying lands, organic plant matter was transported outwards into the ocean. An alternate model for the development of anoxia is that epicontinental seaways became salinity stratified with strong haloclines , chemoclines , and thermoclines . This caused mineralised carbon on

11590-422: The ocean floor acting as a buffer against large CO 2 emissions. The climate of the Jurassic was generally warmer than that of present, by around 5–10 °C (9–18 °F), with atmospheric carbon dioxide likely about four times higher. Intermittent "cold snap" intervals are known to have occurred during this time period, however, interrupting the otherwise warm greenhouse climate. Forests likely grew near

11712-585: The ocean. This produced exquisitely preserved lagerstätten across the world, such as Ya Ha Tinda, Strawberry Bank, and the Posidonia Shale . As is common during anoxic events, black shale deposition was widespread during the deoxygenation events of the Toarcian. Toarcian anoxia was responsible for the deposition of commercially extracted oil shales, particularly in China. Enhanced hydrological cycling caused clastic sedimentation to accelerate during

11834-476: The oceans, and prompted the development of anoxia, leading to severe biodiversity loss. The biogeochemical crisis is documented by a high amplitude negative carbon isotope excursions, as well as black shale deposition. The Early Toarcian extinction event occurred in two distinct pulses, with the first event being classified by some authors as its own event unrelated to the more extreme second event. The first, more recently identified pulse occurred during

11956-460: The oceans, resulting in large areas of desert and scrubland in the lower latitudes between 40° N and S of the equator. Tropical rainforest and tundra biomes are likely to have been rare or absent. The Jurassic also witnessed the decline of the Pangaean megamonsoon that had characterised the preceding Permian and Triassic periods. Variation in the frequency of wildfire activity in the Jurassic

12078-566: The onset of TOAE are known from pyrites in the Sakahogi and Sakuraguchi-dani localities in Japan, with the Sakahogi site displaying a less extreme but still significant pyritic positive δ S excursion during the PTo-E. Euxinia is further evidenced by enhanced pyrite burial in Zázrivá, Slovakia, enhanced molybdenum burial totalling about 41 Gt of molybdenum, and δ Mo excursions observed in sites in

12200-611: The poles, where they experienced warm summers and cold, sometimes snowy winters; there were unlikely to have been ice sheets given the high summer temperatures that prevented the accumulation of snow, though there may have been mountain glaciers. Dropstones and glendonites in northeastern Siberia during the Early to Middle Jurassic indicate cold winters. The ocean depths were likely 8 °C (14 °F) warmer than present, and coral reefs grew 10° of latitude further north and south. The Intertropical Convergence Zone likely existed over

12322-633: The region more arid. This aridification was interrupted, however, in the spinatus ammonite biozone and across the Pliensbachian-Toarcian boundary itself. The large rise in sea levels resulting from the intense global warming led to the formation of the Laurasian Seaway, which enabled the flow of cool water low in salt content to flow into the Tethys Ocean from the Arctic Ocean . The opening of this seaway may have potentially acted as

12444-648: The region. The GSSP for the base of the Bathonian is Ravin du Bès, Bas-Auran area, Alpes de Haute Provence , France; it was ratified in 2009. The base of the Bathonian is defined by the first appearance of the ammonite Gonolkites convergens , at the base of the Zigzagiceras zigzag ammonite zone . The Callovian is derived from the Latinized name of the village of Kellaways in Wiltshire , England, and

12566-458: The rough, uneven bathymetry in the northward limb of this gyre, oxic bottom waters had relatively few impediments to diffuse into the southwestern Tethys, which spared it from the far greater prevalence of anoxia and euxinia that characterised the northern Tethys. The Panthalassan deep water site of Sakahogi was mainly anoxic-ferruginous across the interval spanning the late Pliensbachian to the TOAE, but transient sulphidic conditions did occur during

12688-498: The seafloor to be recycled back into the photic zone, driving widespread primary productivity and in turn anoxia. The freshening of the Arctic Ocean by way of melting of Northern Hemisphere ice caps was a likely trigger of such stratification and a slowdown of global thermohaline circulation. Stratification also occurred due to the freshening of surface water caused by an enhanced water cycle. Rising seawater temperatures amidst

12810-573: The three series of von Buch in the Swabian Jura into six subdivisions defined by ammonites and other fossils. The German palaeontologist Albert Oppel in his studies between 1856 and 1858 altered d'Orbigny's original scheme and further subdivided the stages into biostratigraphic zones, based primarily on ammonites. Most of the modern stages of the Jurassic were formalized at the Colloque du Jurassique à Luxembourg in 1962. The Jurassic Period

12932-598: The time, such as the Bächental bituminous marls, though its occurrence in areas like Greece has been cited as evidence of its global nature. The negative δ C shift is also known from the Arabian Peninsula , the Ordos Basin , and the Neuquén Basin . The negative δ C excursion has been found to be up to -8% in bulk organic and carbonate carbon, although analysis of compound specific biomarkers suggests

13054-405: The wake of the extinction event, many derived clades of ornithischians, sauropods, and theropods emerged, with most of these post-extinction clades greatly increasing in size relative to dinosaurs before the TOAE. Eusauropods were propelled to ecological dominance after their survival of the Toarcian cataclysm. Megalosaurids experienced a diversification event in the latter part of the Toarcian that

13176-407: The world's major landmasses were coalesced into the supercontinent Pangaea , which during the Early Jurassic began to break up into northern supercontinent Laurasia and the southern supercontinent Gondwana . The rifting between North America and Africa was the first to initiate, beginning in the early Jurassic, associated with the emplacement of the Central Atlantic Magmatic Province . During

13298-533: The yew family ( Taxaceae ) is Palaeotaxus rediviva , from the Hettangian of Sweden, suggested to be closely related to the living Austrotaxus , while Marskea jurassica from the Middle Jurassic of Yorkshire, England and material from the Callovian–Oxfordian Daohugou Bed in China are thought to be closely related to Amentotaxus , with the latter material assigned to the modern genus, indicating that Taxaceae had substantially diversified by

13420-575: Was a global oceanic anoxic event , representing possibly the most extreme case of widespread ocean deoxygenation in the entire Phanerozoic eon. In addition to the PTo-E and TOAE, there were multiple other, smaller extinction pulses within this span of time. Occurring during the supergreenhouse climate of the Early Toarcian Thermal Maximum (ETTM), the Early Toarcian extinction was associated with large igneous province volcanism, which elevated global temperatures, acidified

13542-419: Was a negative feedback loop retarding the otherwise pronounced warming and may have caused global cooling in the aftermath of the TOAE. In anoxic and euxinic marine basins in Europe, organic carbon burial rates increased by ~500%. Furthermore, anoxia was not limited to oceans; large lakes also experienced oxygen depletion and black shale deposition. Euxinia occurred in the northwestern Tethys Ocean during

13664-474: Was first defined and introduced into scientific literature by Alcide d'Orbigny in 1842. It takes its name from the French town of Semur-en-Auxois , near Dijon . The original definition of Sinemurian included what is now the Hettangian. The GSSP of the Sinemurian is located at a cliff face north of the hamlet of East Quantoxhead , 6 kilometres east of Watchet , Somerset , England , within the Blue Lias , and

13786-569: Was governed by the 405 kyr eccentricity cycle. Thanks to the breakup of Pangaea, the hydrological cycle during the Jurassic was significantly enhanced. The beginning of the Jurassic was likely marked by a thermal spike corresponding to the Triassic–Jurassic extinction and eruption of the Central Atlantic magmatic province. The first part of the Jurassic was marked by the Early Jurassic Cool Interval between 199 and 183 million years ago. It has been proposed that glaciation

13908-582: Was named by Alcide d'Orbigny in 1852, originally the base at the contact between the Forest Marble Formation and the Cornbrash Formation . However, this boundary was later found to be within the upper part of the Bathonian. The base of the Callovian does not yet have a certified GSSP. The working definition for the base of the Callovian is the first appearance of ammonites belonging to the genus Kepplerites . The Oxfordian

14030-615: Was particularly severe. At Ya Ha Tinda, a replacement of the pre-TOAE bivalve assemblage by a smaller, post-TOAE assemblage occurred, while in the Cleveland Basin , the inoceramid Pseudomytiloides dubius experienced the Lilliput effect . Ammonoids , having already experienced a major morphological bottleneck thanks to the Gibbosus Event, about a million years before the Toarcian extinction, suffered further losses in

14152-519: Was possibly a post-extinction radiation that filled niches vacated by the mass death of the Early Toarcian extinction. Insects may have experienced blooms as fish moved en masse to surface waters to escape anoxia and then died in droves due to limited resources. The volcanogenic extinction event initially impacted terrestrial ecosystems more severely than marine ones. A shift towards a low diversity assemblage of cheirolepid conifers, cycads , and Cerebropollenites -producers adapted for high aridity from

14274-490: Was present in the Northern Hemisphere during both the early Pliensbachian and the latest Pliensbachian. There was a spike in global temperatures of around 4–8 °C (7–14 °F) during the early part of the Toarcian corresponding to the Toarcian Oceanic Anoxic Event and the eruption of the Karoo-Ferrar large igneous provinces in southern Gondwana, with the warm interval extending to the end of

14396-687: Was ratified in 2000. The beginning of the Sinemurian is defined by the first appearance of the ammonite Vermiceras quantoxense . Albert Oppel in 1858 named the Pliensbachian Stage after the hamlet of Pliensbach in the community of Zell unter Aichelberg in the Swabian Alb , near Stuttgart , Germany. The GSSP for the base of the Pliensbachian is found at the Wine Haven locality in Robin Hood's Bay , Yorkshire , England, in

14518-430: Was ratified in 2021. The boundary is defined by the first appearance of ammonites marking the boreal Bauhini Zone and the subboreal Baylei Zone. The Tithonian was introduced in scientific literature by Albert Oppel in 1865. The name Tithonian is unusual in geological stage names because it is derived from Greek mythology rather than a place name. Tithonus was the son of Laomedon of Troy and fell in love with Eos ,

14640-597: Was slow and sluggish, being regularly set back thanks to recurrent episodes of oxygen depletion, which continued for hundreds of thousands of years after the main extinction interval. Evidence from the Cleveland Basin suggests it took ~7 Myr for the marine benthos to recover, on par with the Permian-Triassic extinction event . Many marine invertebrate taxa found in South America migrated through

14762-405: Was the further exacerbation of eutrophication and anoxia. The extreme and rapid global warming at the start of the Toarcian promoted intensification of tropical storms across the globe. The extinction event associated with the TOAE primarily affected marine life as a result the collapse of the carbonate factory. Brachiopods were particularly severely hit, with the TOAE representing one of

14884-582: Was unbelievably catastrophic for corals ; 90.9% of all Tethyan coral species and 49% of all genera were wiped out. Calcareous nannoplankton that lived in the deep photic zone suffered, with the decrease in abundance of the taxon Mitrolithus jansae used as an indicator of shoaling of the oxygen minimum zone in the Tethys and the Hispanic Corridor. Other affected invertebrate groups included echinoderms , radiolarians , dinoflagellates , and foraminifera . Trace fossils , an indicator of bioturbation and ecological diversity, became highly undiverse following

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