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115-568: Jurassic and Cretaceous marine transgressions from the Pacific are recorded in the sediments of Neuquén Basin. These marine sediments belong to Cuyo Group , Tordillo Formation , Auquilco Formation and Vaca Muerta . In the Late Cretaceous, conditions in the neighboring Andean orogeny changed. A marine regression occurred and the fold and thrust belts of Malargüe (36°00 S), Chos Malal (37° S) and Agrio (38° S) started to develop in

230-555: A Berriasian–Barremian warm-dry phase, an Aptian–Santonian warm-wet phase, and a Campanian–Maastrichtian cool-dry phase. As in the Cenozoic, the 400,000 year eccentricity cycle was the dominant orbital cycle governing carbon flux between different reservoirs and influencing global climate. The location of the Intertropical Convergence Zone (ITCZ) was roughly the same as in the present. The cooling trend of

345-483: A boom of dinosaurian evolution on land as the continents began to separate from each other (Nyasasaurus from 243 to 210 million years ago, approximately 235–30 ma, some of them separated into Sauropodomorphs, Theropods and Herrerasaurids), as well as the first pterosaurs . During the Late Triassic, some advanced cynodonts gave rise to the first Mammaliaformes . All this climatic change, however, resulted in

460-594: A coating. This contrasts with the earth's current flora, in which the dominant land plants in terms of number of species are angiosperms . The earliest members of the genus Ginkgo first appeared during the Middle Jurassic. This genus is represented today by a single species, Ginkgo biloba . Modern conifer groups began to radiate during the Jurassic. Bennettitales , an extinct group of gymnosperms with foliage superficially resembling that of cycads gained

575-600: A decline in diversity of sauropods, stegosaurs, and other high-browsing groups, with sauropods particularly scarce in North America. Some island-hopping dinosaurs, such as Eustreptospondylus , evolved to cope with the coastal shallows and small islands of ancient Europe. Other dinosaurs rose up to fill the empty space that the Jurassic-Cretaceous extinction left behind, such as Carcharodontosaurus and Spinosaurus . Seasons came back into effect and

690-477: A few million years before the Triassic–Jurassic extinction event. Sea levels began to rise during the Jurassic, probably caused by an increase in seafloor spreading . The formation of new crust beneath the surface displaced ocean waters by as much as 200 m (656 ft) above today's sea level, flooding coastal areas. Furthermore, Pangaea began to rift into smaller divisions, creating new shoreline around

805-615: A geologic signature associated with the mass extinction that lies between the Mesozoic and Cenozoic Eras . The Cretaceous as a separate period was first defined by Belgian geologist Jean d'Omalius d'Halloy in 1822 as the Terrain Crétacé , using strata in the Paris Basin and named for the extensive beds of chalk ( calcium carbonate deposited by the shells of marine invertebrates , principally coccoliths ), found in

920-483: A global distribution during the Late Triassic, and represented one of the most common groups of Mesozoic seed plants. Flowering plants radiated during the early Cretaceous, first in the tropics , but the even temperature gradient allowed them to spread toward the poles throughout the period. By the end of the Cretaceous, angiosperms dominated tree floras in many areas, although some evidence suggests that biomass

1035-671: A large die-out known as the Triassic–Jurassic extinction event, in which many archosaurs (excluding pterosaurs, dinosaurs and crocodylomorphs ), most synapsids , and almost all large amphibians became extinct, as well as 34% of marine life, in the Earth's fourth mass extinction event. The cause is debatable; flood basalt eruptions at the Central Atlantic magmatic province is cited as one possible cause. The Jurassic ranges from 200 million years to 145 million years ago and features three major epochs: The Early Jurassic,

1150-615: A straight shell, flourished in the seas along with reef-building rudist clams. Inoceramids were also particularly notable among Cretaceous bivalves, and they have been used to identify major biotic turnovers such as at the Turonian-Coniacian boundary. Predatory gastropods with drilling habits were widespread. Globotruncanid foraminifera and echinoderms such as sea urchins and starfish (sea stars) thrived. Ostracods were abundant in Cretaceous marine settings; ostracod species characterised by high male sexual investment had

1265-435: A variety of niches, beginning in the mid-Triassic 4 million to 6 million years after the extinction, and not fully proliferated until 30 million years after the extinction. Animal life was then dominated by various archosaurs: dinosaurs , pterosaurs, and aquatic reptiles such as ichthyosaurs, plesiosaurs, and mosasaurs . The climatic changes of the late Jurassic and Cretaceous favored further adaptive radiation. The Jurassic

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1380-564: A ~0.6 °C increase in temperature. The latter warming interval, occurring at the very end of the Cretaceous, was triggered by the activity of the Deccan Traps. The LKEPCI lasted into the Late Palaeocene , when it gave way to another supergreenhouse interval. The production of large quantities of magma, variously attributed to mantle plumes or to extensional tectonics , further pushed sea levels up, so that large areas of

1495-458: Is a comprehensive overview of described formations: The Neuquén Basin is an important oil and gas producing basin in Argentina. Production started in 1918 and accumulated to 928 thousand cubic metres (5.84 million barrels) of oil equivalent in 2004, representing 45% of the oil production in Argentina and 61% of its gas production. The basin is also important for unconventionals, with

1610-619: Is found in England, northern France, the low countries , northern Germany , Denmark and in the subsurface of the southern part of the North Sea . Chalk is not easily consolidated and the Chalk Group still consists of loose sediments in many places. The group also has other limestones and arenites . Among the fossils it contains are sea urchins , belemnites , ammonites and sea reptiles such as Mosasaurus . In southern Europe,

1725-468: Is set by the Permian–Triassic extinction event , during which it has been estimated that up to 90-96% of marine species became extinct although those approximations have been brought into question with some paleontologists estimating the actual numbers as low as 81%. It is also known as the "Great Dying" because it is considered the largest mass extinction in the Earth's history. The upper boundary of

1840-537: Is thought that a large meteor smashed into earth 66 million years ago, creating the Chicxulub Crater in an event known as the K-Pg Extinction (formerly K-T), the fifth and most recent mass extinction event, in which 75% of life became extinct, including all non-avian dinosaurs. Compared to the vigorous convergent plate mountain-building of the late Paleozoic, Mesozoic tectonic deformation

1955-711: The Indian subcontinent , which collided with the Asian plate during the Cenozoic, giving rise to the Himalayas . The Triassic was generally dry, a trend that began in the late Carboniferous , and highly seasonal, especially in the interior of Pangaea. Low sea levels may have also exacerbated temperature extremes. With its high specific heat capacity , water acts as a temperature-stabilizing heat reservoir, and land areas near large bodies of water—especially oceans—experience less variation in temperature. Because much of Pangaea's land

2070-713: The Mancos Shale of western North America. These shales are an important source rock for oil and gas , for example in the subsurface of the North Sea. In northwestern Europe, chalk deposits from the Upper Cretaceous are characteristic for the Chalk Group , which forms the white cliffs of Dover on the south coast of England and similar cliffs on the French Normandian coast. The group

2185-711: The North American Cordillera , as the Nevadan orogeny was followed by the Sevier and Laramide orogenies . Gondwana had begun to break up during the Jurassic Period, but its fragmentation accelerated during the Cretaceous and was largely complete by the end of the period. South America , Antarctica , and Australia rifted away from Africa (though India and Madagascar remained attached to each other until around 80 million years ago); thus,

2300-561: The Permian Period allowed for the radiation of many new lifeforms. In particular, the extinction of the large herbivorous pareiasaurs and carnivorous gorgonopsians left those ecological niches empty. Some were filled by the surviving cynodonts and dicynodonts , the latter of which subsequently became extinct. Recent research indicates that it took much longer for the reestablishment of complex ecosystems with high biodiversity, complex food webs, and specialized animals in

2415-609: The Pliocene and Pleistocene consuming the western fringes of the Neuquén Basin. The Huincul basement high or Huincul ridge ( Spanish : dorsal de Huincul ) is a geological structure that divides Neuquén Basin in two parts. The basement high is one of the most studied features of Neuquén Basin given its importance for hydrocarbon exploration and exploitation. All over the basement high runs an approximate length of 250 kilometres (160 mi). There have been proposals on

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2530-553: The Selli Event . Early Aptian tropical sea surface temperatures (SSTs) were 27–32 °C, based on TEX 86 measurements from the equatorial Pacific. During the Aptian, Milankovitch cycles governed the occurrence of anoxic events by modulating the intensity of the hydrological cycle and terrestrial runoff. The early Aptian was also notable for its millennial scale hyperarid events in the mid-latitudes of Asia. The BAWI itself

2645-728: The Turonian Age, based on isotopic evidence. However, this has subsequently been suggested to be the result of inconsistent isotopic proxies, with evidence of polar rainforests during this time interval at 82° S. Rafting by ice of stones into marine environments occurred during much of the Cretaceous, but evidence of deposition directly from glaciers is limited to the Early Cretaceous of the Eromanga Basin in southern Australia . Flowering plants (angiosperms) make up around 90% of living plant species today. Prior to

2760-614: The dinosaurs ; a hot greenhouse climate; and the tectonic break-up of Pangaea . The Mesozoic is the middle of the three eras since complex life evolved : the Paleozoic , the Mesozoic, and the Cenozoic . The era began in the wake of the Permian–Triassic extinction event , the largest mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event , another mass extinction whose victims included

2875-411: The non-avian dinosaurs , pterosaurs , mosasaurs , and plesiosaurs . The Mesozoic was a time of significant tectonic, climatic, and evolutionary activity. The supercontinent Pangaea began to break apart into separate landmasses. The climate of the Mesozoic was varied, alternating between warming and cooling periods. Overall, however, the Earth was hotter than it is today. Dinosaurs first appeared in

2990-483: The tuatara ) disappeared from North America and Europe after the Early Cretaceous , and were absent from North Africa and northern South America by the early Late Cretaceous . The cause of the decline of Rhynchocephalia remains unclear, but has often been suggested to be due to competition with advanced lizards and mammals. They appear to have remained diverse in high-latitude southern South America during

3105-999: The Albian regularly expanded northward in tandem with expansions of subtropical high pressure belts. The Cedar Mountain Formation's Soap Wash flora indicates a mean annual temperature of between 19 and 26 °C in Utah at the Albian-Cenomanian boundary. Tropical SSTs during the Cenomanian-Turonian Thermal Maximum were at least 30 °C, though one study estimated them as high as between 33 and 42 °C. An intermediate estimate of ~33-34 °C has also been given. Meanwhile, deep ocean temperatures were as much as 15 to 20 °C (27 to 36 °F) warmer than today's; one study estimated that deep ocean temperatures were between 12 and 20 °C during

3220-617: The Andes and did so until Eocene times. This meant an advance of the Andean orogenic deformation since the Late Cretaceous that made the western part of Neuquén Basin to stack in the Malargüe and Agrio fold and thrust belts . This caused a shift in deposition from Pacific to Early Atlantic . In the south of Mendoza Province, the Guañacos fold and thrust belt (36.5° S) appeared and grew in

3335-458: The Atlantic seaway, which has grown continually larger until today. The further separation of the continents gave opportunity for the diversification of new dinosaurs. The Cretaceous is the longest period of the Mesozoic, but has only two epochs: Early and Late Cretaceous. The Early Cretaceous spans from 145 to 100 million years ago. The Early Cretaceous saw the expansion of seaways and

3450-486: The Cenomanian between the Albian and Turonian. The Cretaceous is divided into Early and Late Cretaceous epochs , or Lower and Upper Cretaceous series . In older literature, the Cretaceous is sometimes divided into three series: Neocomian (lower/early), Gallic (middle) and Senonian (upper/late). A subdivision into 12 stages , all originating from European stratigraphy, is now used worldwide. In many parts of

3565-571: The Cenozoic Era   —   the ichthyosaurs , last remaining temnospondyls ( Koolasuchus ), and nonmammalian cynodonts ( Tritylodontidae )   —   were already extinct millions of years before the event occurred. Coccolithophorids and molluscs , including ammonites , rudists , freshwater snails , and mussels , as well as organisms whose food chain included these shell builders, became extinct or suffered heavy losses. For example, ammonites are thought to have been

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3680-645: The Coniacian through the Maastrichtian. During the Cretaceous, the late- Paleozoic -to-early-Mesozoic supercontinent of Pangaea completed its tectonic breakup into the present-day continents , although their positions were substantially different at the time. As the Atlantic Ocean widened, the convergent-margin mountain building ( orogenies ) that had begun during the Jurassic continued in

3795-574: The Cretaceous is sharply defined, being placed at an iridium -rich layer found worldwide that is believed to be associated with the Chicxulub impact crater , with its boundaries circumscribing parts of the Yucatán Peninsula and extending into the Gulf of Mexico . This layer has been dated at 66.043 Mya. At the end of the Cretaceous, the impact of a large body with the Earth may have been

3910-570: The Cretaceous is usually a marine system consisting of competent limestone beds or incompetent marls . Because the Alpine mountain chains did not yet exist in the Cretaceous, these deposits formed on the southern edge of the European continental shelf , at the margin of the Tethys Ocean . During the Cretaceous, the present North American continent was isolated from the other continents. In

4025-537: The Cretaceous seas. Stagnation of deep sea currents in middle Cretaceous times caused anoxic conditions in the sea water leaving the deposited organic matter undecomposed. Half of the world's petroleum reserves were laid down at this time in the anoxic conditions of what would become the Persian Gulf and the Gulf of Mexico. In many places around the world, dark anoxic shales were formed during this interval, such as

4140-616: The Jurassic, the North Atlantic already opened, leaving a proto-ocean between Europe and North America. From north to south across the continent, the Western Interior Seaway started forming. This inland sea separated the elevated areas of Laramidia in the west and Appalachia in the east. Three dinosaur clades found in Laramidia (troodontids, therizinosaurids and oviraptorosaurs) are absent from Appalachia from

4255-576: The LKEPCI. During this period of relatively cool temperatures, the ITCZ became narrower, while the strength of both summer and winter monsoons in East Asia was directly correlated to atmospheric CO 2 concentrations. Laramidia likewise had a seasonal, monsoonal climate. The Maastrichtian was a time of chaotic, highly variable climate. Two upticks in global temperatures are known to have occurred during

4370-400: The Late Cretaceous declined for poorly understood reasons, though this might be due to tendencies of the fossil record, as their diversity seems to be much higher than previously thought. Birds became increasingly common and diversified into a variety of enantiornithe and ornithurine forms. Though mostly small, marine hesperornithes became relatively large and flightless, adapted to life in

4485-467: The Late Cretaceous, where lizards remained rare, with their remains outnumbering terrestrial lizards 200:1. Choristoderes , a group of freshwater aquatic reptiles that first appeared during the preceding Jurassic, underwent a major evolutionary radiation in Asia during the Early Cretaceous, which represents the high point of choristoderan diversity, including long necked forms such as Hyphalosaurus and

4600-555: The Late Cretaceous-Early Palaeogene Cool Interval (LKEPCI). Tropical SSTs declined from around 35 °C in the early Campanian to around 28 °C in the Maastrichtian. Deep ocean temperatures declined to 9 to 12 °C, though the shallow temperature gradient between tropical and polar seas remained. Regional conditions in the Western Interior Seaway changed little between the MKH and

4715-628: The MKH. Mean annual temperatures at the poles during the MKH exceeded 14 °C. Such hot temperatures during the MKH resulted in a very gentle temperature gradient from the equator to the poles; the latitudinal temperature gradient during the Cenomanian-Turonian Thermal Maximum was 0.54 °C per ° latitude for the Southern Hemisphere and 0.49 °C per ° latitude for the Northern Hemisphere, in contrast to present day values of 1.07 and 0.69 °C per ° latitude for

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4830-526: The MKH. The poles were so warm that ectothermic reptiles were able to inhabit them. Beginning in the Santonian, near the end of the MKH, the global climate began to cool, with this cooling trend continuing across the Campanian. This period of cooling, driven by falling levels of atmospheric carbon dioxide, caused the end of the MKH and the transition into a cooler climatic interval, known formally as

4945-404: The Maastrichtian, bucking the trend of overall cooler temperatures during the LKEPCI. Between 70 and 69 Ma and 66–65 Ma, isotopic ratios indicate elevated atmospheric CO 2 pressures with levels of 1000–1400 ppmV and mean annual temperatures in west Texas between 21 and 23 °C (70 and 73 °F). Atmospheric CO 2 and temperature relations indicate a doubling of pCO 2 was accompanied by

5060-473: The Mesozoic is set at the Cretaceous–Paleogene extinction event (or K–Pg extinction event ), which may have been caused by an asteroid impactor that created Chicxulub Crater on the Yucatán Peninsula . Towards the Late Cretaceous, large volcanic eruptions are also believed to have contributed to the Cretaceous–Paleogene extinction event. Approximately 50% of all genera became extinct, including all of

5175-484: The Mesozoic) ended with the Cretaceous–Paleogene extinction event , a large mass extinction in which many groups, including non-avian dinosaurs, pterosaurs , and large marine reptiles , died out, widely thought to have been caused by the impact of a large asteroid that formed the Chicxulub crater in the Gulf of Mexico. The end of the Cretaceous is defined by the abrupt Cretaceous–Paleogene boundary (K–Pg boundary),

5290-583: The Mesozoic, but would remain small—less than 15 kg (33 lb)—until the Cenozoic. Flowering plants appeared in the Early Cretaceous and would rapidly diversify through the end of the era, replacing conifers and other gymnosperms ( sensu lato ), such as ginkgoales , cycads and bennettitales as the dominant group of plants. The phrase "Age of Reptiles" was introduced by the 19th century paleontologist Gideon Mantell who viewed it as dominated by diapsids such as Iguanodon , Megalosaurus , Plesiosaurus , and Pterodactylus . The current name

5405-474: The Mid-Triassic, and became the dominant terrestrial vertebrates in the Late Triassic or Early Jurassic, occupying this position for about 150 or 135 million years until their demise at the end of the Cretaceous. Archaic birds appeared in the Jurassic, having evolved from a branch of theropod dinosaurs, then true toothless birds appeared in the Cretaceous. The first mammals also appeared during

5520-561: The Middle Jurassic, and the Late Jurassic. The Early Jurassic spans from 200 to 175 million years ago. The climate was tropical and much more humid than the Triassic, as a result of the large seas appearing between the land masses. In the oceans, plesiosaurs , ichthyosaurs and ammonites were abundant. On land, dinosaurs and other archosaurs staked their claim as the dominant race, with theropods such as Dilophosaurus at

5635-541: The South Atlantic and Indian Oceans were newly formed. Such active rifting lifted great undersea mountain chains along the welts, raising eustatic sea levels worldwide. To the north of Africa the Tethys Sea continued to narrow. During most of the Late Cretaceous, North America would be divided in two by the Western Interior Seaway , a large interior sea, separating Laramidia to the west and Appalachia to

5750-570: The Southern and Northern hemispheres, respectively. This meant weaker global winds, which drive the ocean currents, and resulted in less upwelling and more stagnant oceans than today. This is evidenced by widespread black shale deposition and frequent anoxic events . Tropical SSTs during the late Albian most likely averaged around 30 °C. Despite this high SST, seawater was not hypersaline at this time, as this would have required significantly higher temperatures still. On land, arid zones in

5865-474: The Tethys Ocean. Temperatures continued to increase, then began to stabilize. Humidity also increased with the proximity of water, and deserts retreated. The climate of the Cretaceous is less certain and more widely disputed. Probably, higher levels of carbon dioxide in the atmosphere are thought to have almost eliminated the north–south temperature gradient : temperatures were about the same across

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5980-576: The Tethys to the Arctic Ocean and enabling biotic exchange between the two oceans. At the peak of the Cretaceous transgression , one-third of Earth's present land area was submerged. The Cretaceous is justly famous for its chalk ; indeed, more chalk formed in the Cretaceous than in any other period in the Phanerozoic . Mid-ocean ridge activity—or rather, the circulation of seawater through

6095-717: The Vaca Muerta and Los Molles formations being major shale gas producers. Source rock formations are predominantly the Vaca Muerta, and to a lesser extent the Agrio and Los Molles Formations . Reservoir rocks comprise the Mulichinco and Chachao Formations . Deeper reservoirs are the Lotena and Barda Negra Formations . Regional seal rocks are the evaporites of the Auquilco and Huitrín Formations , with local seals

6210-411: The Vaca Muerta, Agrio and Catriel Formations . Cretaceous The Cretaceous ( IPA : / k r ɪ ˈ t eɪ ʃ ə s / krih- TAY -shəss ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era , as well as the longest. At around 79   million years, it is the ninth and longest geological period of

6325-425: The ancestors of modern-day birds also diversified. They inhabited every continent, and were even found in cold polar latitudes. Pterosaurs were common in the early and middle Cretaceous, but as the Cretaceous proceeded they declined for poorly understood reasons (once thought to be due to competition with early birds , but now it is understood avian adaptive radiation is not consistent with pterosaur decline ). By

6440-406: The beginnings of the breakup of Pangaea and the opening of the Tethys Ocean . Ecosystems had recovered from the Permian extinction. Algae, sponge, corals, and crustaceans all had recovered, and new aquatic reptiles evolved, such as ichthyosaurs and nothosaurs . On land, pine forests flourished, as did groups of insects such as mosquitoes and fruit flies. Reptiles began to get bigger and bigger, and

6555-408: The continental crust were covered with shallow seas. The Tethys Sea connecting the tropical oceans east to west also helped to warm the global climate. Warm-adapted plant fossils are known from localities as far north as Alaska and Greenland , while dinosaur fossils have been found within 15 degrees of the Cretaceous south pole . It was suggested that there was Antarctic marine glaciation in

6670-431: The current level (about 21%) throughout the Mesozoic, some concluding they were lower in the Triassic and part of the Jurassic but higher in the Cretaceous, and some concluding they were higher throughout most or all of the Triassic, Jurassic and Cretaceous. The dominant land plant species of the time were gymnosperms , which are vascular, cone-bearing, non-flowering plants such as conifers that produce seeds without

6785-550: The dramatic rifting of the supercontinent Pangaea, which gradually split into a northern continent, Laurasia , and a southern continent, Gondwana . This created the passive continental margin that characterizes most of the Atlantic coastline (such as along the U.S. East Coast ) today. By the end of the era, the continents had rifted into nearly their present forms, though not their present positions. Laurasia became North America and Eurasia , while Gondwana split into South America , Africa , Australia , Antarctica and

6900-463: The earliest crown group birds. Acanthomorph fish, the most diverse group of modern vertebrates, appeared in aquatic habitats around the middle of the Cretaceous. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth by the end of the Cretaceous, coincident with the decline and extinction of previously widespread gymnosperm groups. The Cretaceous (along with

7015-544: The early and mid-Cretaceous (becoming extinct during the late Cretaceous Cenomanian-Turonian anoxic event ), plesiosaurs throughout the entire period, and mosasaurs appearing in the Late Cretaceous. Sea turtles in the form of Cheloniidae and Panchelonioidea lived during the period and survived the extinction event. Panchelonioidea is today represented by a single species; the leatherback sea turtle . The Hesperornithiformes were flightless, marine diving birds that swam like grebes . Baculites , an ammonite genus with

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7130-437: The east, then receded late in the period, leaving thick marine deposits sandwiched between coal beds. Bivalve palaeobiogeography also indicates that Africa was split in half by a shallow sea during the Coniacian and Santonian, connecting the Tethys with the South Atlantic by way of the central Sahara and Central Africa, which were then underwater. Yet another shallow seaway ran between what is now Norway and Greenland, connecting

7245-535: The end of the AACS, which ended around 111 Ma with the Paquier/Urbino Thermal Maximum, giving way to the Mid-Cretaceous Hothouse (MKH), which lasted from the early Albian until the early Campanian. Faster rates of seafloor spreading and entry of carbon dioxide into the atmosphere are believed to have initiated this period of extreme warmth, along with high flood basalt activity. The MKH was punctuated by multiple thermal maxima of extreme warmth. The Leenhardt Thermal Event (LTE) occurred around 110 Ma, followed shortly by

7360-448: The end of the Cretaceous. The high sea level and warm climate of the Cretaceous meant large areas of the continents were covered by warm, shallow seas, providing habitat for many marine organisms. The Cretaceous was named for the extensive chalk deposits of this age in Europe, but in many parts of the world, the deposits from the Cretaceous are of marine limestone , a rock type that is formed under warm, shallow marine conditions. Due to

7475-411: The end of the period only three highly specialized families remained; Pteranodontidae , Nyctosauridae , and Azhdarchidae . The Liaoning lagerstätte ( Yixian Formation ) in China is an important site, full of preserved remains of numerous types of small dinosaurs, birds and mammals, that provides a glimpse of life in the Early Cretaceous. The coelurosaur dinosaurs found there represent types of

7590-407: The enlarged ridges—enriched the oceans in calcium ; this made the oceans more saturated, as well as increased the bioavailability of the element for calcareous nanoplankton . These widespread carbonates and other sedimentary deposits make the Cretaceous rock record especially fine. Famous formations from North America include the rich marine fossils of Kansas 's Smoky Hill Chalk Member and

7705-519: The entire Phanerozoic . The name is derived from the Latin creta , ' chalk ', which is abundant in the latter half of the period. It is usually abbreviated K , for its German translation Kreide . The Cretaceous was a period with a relatively warm climate , resulting in high eustatic sea levels that created numerous shallow inland seas . These oceans and seas were populated with now- extinct marine reptiles , ammonites , and rudists , while dinosaurs continued to dominate on land. The world

7820-528: The extinction fed on insects , larvae , worms , and snails, which in turn fed on dead plant and animal matter. Scientists theorise that these organisms survived the collapse of plant-based food chains because they fed on detritus . In stream communities , few groups of animals became extinct. Stream communities rely less on food from living plants and more on detritus that washes in from land. This particular ecological niche buffered them from extinction. Similar, but more complex patterns have been found in

7935-467: The fern prairies, chased by many new predators such as Allosaurus . Conifer forests made up a large portion of the forests. In the oceans, plesiosaurs were quite common, and ichthyosaurs flourished. This epoch was the peak of the reptiles. The Late Jurassic spans from 163 to 145 million years ago. During this epoch, the first avialans , such as Archaeopteryx , evolved from small coelurosaurian dinosaurs. The increase in sea levels opened up

8050-402: The first crocodilians and dinosaurs evolved, which sparked competition with the large amphibians that had previously ruled the freshwater world, respectively mammal-like reptiles on land. Following the bloom of the Middle Triassic, the Late Triassic, from 237 to 201 million years ago, featured frequent heat spells and moderate precipitation (10–20 inches per year). The recent warming led to

8165-505: The first records of the gharial-like Neochoristodera , which appear to have evolved in the regional absence of aquatic neosuchian crocodyliformes. During the Late Cretaceous the neochoristodere Champsosaurus was widely distributed across western North America. Due to the extreme climatic warmth in the Arctic, choristoderans were able to colonise it too during the Late Cretaceous. In the seas, rays , modern sharks and teleosts became common. Marine reptiles included ichthyosaurs in

8280-487: The food web. In the oceans, mosasaurs ruled, filling the role of the ichthyosaurs, which, after declining, had disappeared in the Cenomanian-Turonian boundary event . Though pliosaurs had gone extinct in the same event, long-necked plesiosaurs such as Elasmosaurus continued to thrive. Flowering plants, possibly appearing as far back as the Triassic, became truly dominant for the first time. Pterosaurs in

8395-492: The fossil record. The Late Cretaceous spans from 100 to 66 million years ago. The Late Cretaceous featured a cooling trend that would continue in the Cenozoic Era. Eventually, tropics were restricted to the equator and areas beyond the tropic lines experienced extreme seasonal changes in weather. Dinosaurs still thrived, as new taxa such as Tyrannosaurus , Ankylosaurus , Triceratops and hadrosaurs dominated

8510-503: The genus Berriasella , but its use as a stratigraphic indicator has been questioned, as its first appearance does not correlate with that of C. alpina . The boundary is officially considered by the International Commission on Stratigraphy to be approximately 145 million years ago, but other estimates have been proposed based on U-Pb geochronology, ranging as young as 140 million years ago. The upper boundary of

8625-625: The group Maniraptora , which includes modern birds and their closest non-avian relatives, such as dromaeosaurs , oviraptorosaurs , therizinosaurs , troodontids along with other avialans . Fossils of these dinosaurs from the Liaoning lagerstätte are notable for the presence of hair-like feathers . Insects diversified during the Cretaceous, and the oldest known ants , termites and some lepidopterans , akin to butterflies and moths , appeared. Aphids , grasshoppers and gall wasps appeared. Rhynchocephalians (which today only includes

8740-412: The high sea level, there was extensive space for such sedimentation . Because of the relatively young age and great thickness of the system, Cretaceous rocks are evident in many areas worldwide. Chalk is a rock type characteristic for (but not restricted to) the Cretaceous. It consists of coccoliths , microscopically small calcite skeletons of coccolithophores , a type of algae that prospered in

8855-551: The highest rates of extinction and turnover. Thylacocephala , a class of crustaceans, went extinct in the Late Cretaceous. The first radiation of the diatoms (generally siliceous shelled, rather than calcareous ) in the oceans occurred during the Cretaceous; freshwater diatoms did not appear until the Miocene . Calcareous nannoplankton were important components of the marine microbiota and important as biostratigraphic markers and recorders of environmental change. The Cretaceous

8970-658: The last epoch of the Jurassic, the Tithonian, continued into the Berriasian, the first age of the Cretaceous. The North Atlantic seaway opened and enabled the flow of cool water from the Boreal Ocean into the Tethys. There is evidence that snowfalls were common in the higher latitudes during this age, and the tropics became wetter than during the Triassic and Jurassic. Glaciation was restricted to high- latitude mountains, though seasonal snow may have existed farther from

9085-512: The late Valanginian (~ 134 million years ago) found in Israel and Italy, initially at low abundance. Molecular clock estimates conflict with fossil estimates, suggesting the diversification of crown-group angiosperms during the Late Triassic or the Jurassic, but such estimates are difficult to reconcile with the heavily sampled pollen record and the distinctive tricolpate to tricolporoidate (triple grooved) pollen of eudicot angiosperms. Among

9200-465: The late Cretaceous, and all else that depended on them suffered, as well. Herbivorous animals, which depended on plants and plankton as their food, died out as their food sources became scarce; consequently, the top predators , such as Tyrannosaurus rex , also perished. Yet only three major groups of tetrapods disappeared completely; the nonavian dinosaurs , the plesiosaurs and the pterosaurs . The other Cretaceous groups that did not survive into

9315-574: The latest Albian. Approximately 94 Ma, the Cenomanian-Turonian Thermal Maximum occurred, with this hyperthermal being the most extreme hothouse interval of the Cretaceous and being associated with a sea level highstand. Temperatures cooled down slightly over the next few million years, but then another thermal maximum, the Coniacian Thermal Maximum, happened, with this thermal event being dated to around 87 Ma. Atmospheric CO 2 levels may have varied by thousands of ppm throughout

9430-711: The l’Arboudeyesse Thermal Event (ATE) a million years later. Following these two hyperthermals was the Amadeus Thermal Maximum around 106 Ma, during the middle Albian. Then, around a million years after that, occurred the Petite Verol Thermal Event (PVTE). Afterwards, around 102.5 Ma, the Event 6 Thermal Event (EV6) took place; this event was itself followed by the Breistroffer Thermal Maximum around 101 Ma, during

9545-517: The middle Cretaceous, becoming the dominant group of land plants by the end of the period, coincident with the decline of previously dominant groups such as conifers. The oldest known fossils of grasses are from the Albian , with the family having diversified into modern groups by the end of the Cretaceous. The oldest large angiosperm trees are known from the Turonian (c. 90 Mya) of New Jersey, with

9660-470: The most promising candidates for fixing the Jurassic–Cretaceous boundary. In particular, the first appearance Calpionella alpina , coinciding with the base of the eponymous Alpina subzone, has been proposed as the definition of the base of the Cretaceous. The working definition for the boundary has often been placed as the first appearance of the ammonite Strambergella jacobi , formerly placed in

9775-519: The nature of this structure. In the 1970s and 1980s it was proposed by that it was a transpressive fault zone . In 2009, Pángaro described it as being made up of inverted half-grabens . Unconformities have been registered in the basin and were dated at 98, 117, 123, 129, 134 and 154 Ma. The basin contains many stratigraphic units from the Triassic onwards, with large regional variations from east to west and north to south, often described as different formations that are laterally equivalent, this list

9890-541: The non- avian dinosaurs. The Triassic ranges roughly from 252 million to 201 million years ago, preceding the Jurassic Period. The period is bracketed between the Permian–Triassic extinction event and the Triassic–Jurassic extinction event , two of the " big five ", and it is divided into three major epochs: Early, Middle, and Late Triassic. The Early Triassic, about 252 to 247 million years ago,

10005-824: The oceans. Extinction was more severe among animals living in the water column than among animals living on or in the seafloor. Animals in the water column are almost entirely dependent on primary production from living phytoplankton, while animals living on or in the ocean floor feed on detritus or can switch to detritus feeding. The largest air-breathing survivors of the event, crocodilians and champsosaurs , were semiaquatic and had access to detritus. Modern crocodilians can live as scavengers and can survive for months without food and go into hibernation when conditions are unfavorable, and their young are small, grow slowly, and feed largely on invertebrates and dead organisms or fragments of organisms for their first few years. These characteristics have been linked to crocodilian survival at

10120-653: The oldest records of Angiosperm macrofossils are Montsechia from the Barremian aged Las Hoyas beds of Spain and Archaefructus from the Barremian-Aptian boundary Yixian Formation in China. Tricolpate pollen distinctive of eudicots first appears in the Late Barremian, while the earliest remains of monocots are known from the Aptian. Flowering plants underwent a rapid radiation beginning during

10235-459: The open sea. Metatherians and primitive eutherian also became common and even produced large and specialised genera such as Didelphodon and Schowalteria . Still, the dominant mammals were multituberculates, cimolodonts in the north and gondwanatheres in the south. At the end of the Cretaceous, the Deccan traps and other volcanic eruptions were poisoning the atmosphere. As this continued, it

10350-437: The planet, and about 10° C higher than today. The circulation of oxygen to the deep ocean may also have been disrupted, preventing the decomposition of large volumes of organic matter, which was eventually deposited as " black shale ". Different studies have come to different conclusions about the amount of oxygen in the atmosphere during different parts of the Mesozoic, with some concluding oxygen levels were lower than

10465-641: The poles got seasonally colder, but some dinosaurs still inhabited the polar forests year round, such as Leaellynasaura and Muttaburrasaurus . The poles were too cold for crocodiles, and became the last stronghold for large amphibians such as Koolasuchus . Pterosaurs got larger as genera such as Tapejara and Ornithocheirus evolved. Mammals continued to expand their range: eutriconodonts produced fairly large, wolverine -like predators such as Repenomamus and Gobiconodon , early therians began to expand into metatherians and eutherians , and cimolodont multituberculates went on to become common in

10580-523: The poles. After the end of the first age, however, temperatures began to increase again, with a number of thermal excursions, such as the middle Valanginian Weissert Thermal Excursion (WTX), which was caused by the Paraná-Etendeka Large Igneous Province's activity. It was followed by the middle Hauterivian Faraoni Thermal Excursion (FTX) and the early Barremian Hauptblatterton Thermal Event (HTE). The HTE marked

10695-412: The principal food of mosasaurs , a group of giant marine lizards related to snakes that became extinct at the boundary. Omnivores , insectivores , and carrion -eaters survived the extinction event, perhaps because of the increased availability of their food sources. At the end of the Cretaceous, there seem to have been no purely herbivorous or carnivorous mammals . Mammals and birds that survived

10810-547: The punctuation mark at the end of a progressive decline in biodiversity during the Maastrichtian age. The result was the extinction of three-quarters of Earth's plant and animal species. The impact created the sharp break known as the K–Pg boundary (formerly known as the K–T boundary). Earth's biodiversity required substantial time to recover from this event, despite the probable existence of an abundance of vacant ecological niches . Despite

10925-744: The rise of angiosperms, during the Jurassic and the Early Cretaceous, the higher flora was dominated by gymnosperm groups, including cycads , conifers , ginkgophytes , gnetophytes and close relatives, as well as the extinct Bennettitales . Other groups of plants included pteridosperms or "seed ferns", a collective term that refers to disparate groups of extinct seed plants with fern-like foliage, including groups such as Corystospermaceae and Caytoniales . The exact origins of angiosperms are uncertain, although molecular evidence suggests that they are not closely related to any living group of gymnosperms. The earliest widely accepted evidence of flowering plants are monosulcate (single-grooved) pollen grains from

11040-451: The severity of the K-Pg extinction event, there were significant variations in the rate of extinction between and within different clades . Species that depended on photosynthesis declined or became extinct as atmospheric particles blocked solar energy . As is the case today, photosynthesizing organisms, such as phytoplankton and land plants , formed the primary part of the food chain in

11155-422: The strong regionality of most biostratigraphic markers, and the lack of any chemostratigraphic events, such as isotope excursions (large sudden changes in ratios of isotopes ) that could be used to define or correlate a boundary. Calpionellids , an enigmatic group of planktonic protists with urn-shaped calcitic tests briefly abundant during the latest Jurassic to earliest Cretaceous, have been suggested as

11270-633: The terrestrial fauna of the late Cretaceous Hell Creek Formation . Other important Cretaceous exposures occur in Europe (e.g., the Weald ) and China (the Yixian Formation ). In the area that is now India, massive lava beds called the Deccan Traps were erupted in the very late Cretaceous and early Paleocene. Palynological evidence indicates the Cretaceous climate had three broad phases:

11385-506: The top of the food chain. The first true crocodiles evolved, pushing the large amphibians to near extinction. All-in-all, archosaurs rose to rule the world. Meanwhile, the first true mammals evolved, remaining relatively small, but spreading widely; the Jurassic Castorocauda , for example, had adaptations for swimming, digging and catching fish. Fruitafossor , from the late Jurassic Period about 150 million years ago,

11500-483: The trunk having a preserved diameter of 1.8 metres (5.9 ft) and an estimated height of 50 metres (160 ft). During the Cretaceous, ferns in the order Polypodiales , which make up 80% of living fern species, would also begin to diversify. On land, mammals were generally small sized, but a very relevant component of the fauna , with cimolodont multituberculates outnumbering dinosaurs in some sites. Neither true marsupials nor placentals existed until

11615-789: The ultimate end of the Tithonian-early Barremian Cool Interval (TEBCI). During this interval, precession was the dominant orbital driver of environmental changes in the Vocontian Basin. For much of the TEBCI, northern Gondwana experienced a monsoonal climate. A shallow thermocline existed in the mid-latitude Tethys. The TEBCI was followed by the Barremian-Aptian Warm Interval (BAWI). This hot climatic interval coincides with Manihiki and Ontong Java Plateau volcanism and with

11730-643: The upper Cretaceous of Western Europe . The name Cretaceous was derived from the Latin creta , meaning chalk . The twofold division of the Cretaceous was implemented by Conybeare and Phillips in 1822. Alcide d'Orbigny in 1840 divided the French Cretaceous into five étages (stages): the Neocomian , Aptian, Albian, Turonian, and Senonian, later adding the Urgonian between Neocomian and Aptian and

11845-628: The very end, but a variety of non-marsupial metatherians and non-placental eutherians had already begun to diversify greatly, ranging as carnivores ( Deltatheroida ), aquatic foragers ( Stagodontidae ) and herbivores ( Schowalteria , Zhelestidae ). Various "archaic" groups like eutriconodonts were common in the Early Cretaceous, but by the Late Cretaceous northern mammalian faunas were dominated by multituberculates and therians , with dryolestoids dominating South America . The apex predators were archosaurian reptiles , especially dinosaurs , which were at their most diverse stage. Avians such as

11960-407: The world, alternative local subdivisions are still in use. From youngest to oldest, the subdivisions of the Cretaceous period are: The lower boundary of the Cretaceous is currently undefined, and the Jurassic–Cretaceous boundary is currently the only system boundary to lack a defined Global Boundary Stratotype Section and Point (GSSP). Placing a GSSP for this boundary has been difficult because of

12075-531: Was about the size of a chipmunk, and its teeth, forelimbs and back suggest that it dug open the nests of social insects (probably termites , as ants had not yet appeared) ; Volaticotherium was able to glide for short distances, such as modern flying squirrels . The first multituberculates such as Rugosodon evolved. The Middle Jurassic spans from 175 to 163 million years ago. During this epoch, dinosaurs flourished as huge herds of sauropods, such as Brachiosaurus and Diplodocus , filled

12190-566: Was also an important interval in the evolution of bioerosion , the production of borings and scrapings in rocks, hardgrounds and shells. Mesozoic The Mesozoic Era is the era of Earth's geological history , lasting from about 252 to 66 million years ago , comprising the Triassic , Jurassic and Cretaceous Periods . It is characterized by the dominance of gymnosperms such as cycads , ginkgoaceae and araucarian conifers, and of archosaurian reptiles such as

12305-928: Was comparatively mild. The sole major Mesozoic orogeny occurred in what is now the Arctic , creating the Innuitian orogeny , the Brooks Range , the Verkhoyansk and Cherskiy Ranges in Siberia, and the Khingan Mountains in Manchuria. This orogeny was related to the opening of the Arctic Ocean and suturing of the North China and Siberian cratons to Asia. In contrast, the era featured

12420-537: Was distant from its shores, temperatures fluctuated greatly, and the interior probably included expansive deserts . Abundant red beds and evaporites such as halite support these conclusions, but some evidence suggests the generally dry climate of the Triassic was punctuated by episodes of increased rainfall. The most important humid episodes were the Carnian Pluvial Event and one in the Rhaetian ,

12535-593: Was dominated by deserts in the interior of the Pangaea supercontinent. The Earth had just witnessed a massive die-off in which 95% of all life became extinct, and the most common vertebrate life on land were Lystrosaurus , labyrinthodonts , and Euparkeria along with many other creatures that managed to survive the Permian extinction. Temnospondyls reached peak diversity during the early Triassic. The Middle Triassic, from 247 to 237 million years ago, featured

12650-593: Was followed by the Aptian-Albian Cold Snap (AACS) that began about 118 Ma. A short, relatively minor ice age may have occurred during this so-called "cold snap", as evidenced by glacial dropstones in the western parts of the Tethys Ocean and the expansion of calcareous nannofossils that dwelt in cold water into lower latitudes. The AACS is associated with an arid period in the Iberian Peninsula . Temperatures increased drastically after

12765-454: Was largely ice-free, although there is some evidence of brief periods of glaciation during the cooler first half, and forests extended to the poles. Many of the dominant taxonomic groups present in modern times can be ultimately traced back to origins in the Cretaceous. During this time, new groups of mammals and birds appeared, including the earliest relatives of placentals & marsupials ( Eutheria and Metatheria respectively), and

12880-515: Was originally described as the "secondary" era, following the "primary" ( Paleozoic ), and preceding the Tertiary . Following the Paleozoic, the Mesozoic extended roughly 186 million years, from 251.902 to 66 million years ago when the Cenozoic Era began. This time frame is separated into three geologic periods . From oldest to youngest: The lower boundary of the Mesozoic

12995-592: Was proposed in 1840 by the British geologist John Phillips (1800–1874). "Mesozoic" literally means 'middle life', deriving from the Greek prefix meso- ( μεσο- 'between') and zōon ( ζῷον 'animal, living being'). In this way, the Mesozoic is comparable to the Cenozoic ( lit.   ' new life ' ) and Paleozoic ('old life') eras as well as the Proterozoic ('earlier life') Eon. The Mesozoic Era

13110-460: Was still dominated by cycads and ferns until after the Cretaceous–Paleogene extinction. Some plant species had distributions that were markedly different from succeeding periods; for example, the Schizeales , a fern order, were skewed to the Northern Hemisphere in the Mesozoic, but are now better represented in the Southern Hemisphere. The extinction of nearly all animal species at the end of

13225-485: Was the height of archosaur diversity, and the first birds and eutherian mammals also appeared. Some have argued that insects diversified in symbiosis with angiosperms, because insect anatomy , especially the mouth parts, seems particularly well-suited for flowering plants. However, all major insect mouth parts preceded angiosperms, and insect diversification actually slowed when they arrived, so their anatomy originally must have been suited for some other purpose. At

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