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51-583: The Chiapas catfish ( Lacantunia enigmatica ) is an unusual species of catfish (order Siluriformes) from the Usumacinta River basin in the Mexican state of Chiapas and in Guatemala . It was scientifically described in 2005 and placed in its own family Lacantuniidae . While discovery of an undescribed species of catfish is not uncommon, discovery of a new family of any vertebrate group

102-740: A hydrofoil . Some have a mouth that can expand to a large size and contains no incisiform teeth; catfish generally feed through suction or gulping rather than biting and cutting prey. Some families, though, notably the Loricariidae and Astroblepidae , have a suckermouth that allows them to fasten themselves to objects in fast-moving water. Catfish also have a maxilla reduced to a support for barbels ; this means that they are unable to protrude their mouths as other fish such as carp . Catfish may have up to four pairs of barbels - nasal, maxillary (on each side of mouth), and two pairs of chin barbels, though pairs of barbels may be absent depending on

153-464: A bullhead catfish by the nickname "chucklehead", while in another state or region, that nickname refers to the blue catfish. Representatives of the genus Ictalurus have been introduced into European waters in the hope of obtaining a sporting and food resource, but the European stock of American catfishes has not achieved the dimensions of these fish in their native waters and have only increased

204-472: A catfish weighing 88 kilograms (194 lb) was caught in the River Ebro , Spain, by an 11-year-old British schoolgirl. Order (biology) What does and does not belong to each order is determined by a taxonomist , as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that

255-484: A defense, these spines may be locked into place so that they stick outwards, enabling them to inflict severe wounds. In numerous catfish species, these fin rays can be used to deliver a stinging protein if the fish is irritated; as many as half of all catfish species may be venomous in this fashion, making the Siluriformes overwhelmingly the vertebrate order with the largest number of venomous species. This venom

306-478: A diverse group of ray-finned fish . Named for their prominent barbels , which resemble a cat 's whiskers , catfish range in size and behavior from the three largest species alive, the Mekong giant catfish from Southeast Asia , the wels catfish of Eurasia , and the piraíba of South America , to detritivores (species that eat dead material on the bottom), and even to a tiny parasitic species commonly called

357-495: A juvenile obaichthyid lepisosteiform , possibly a junior synonym of Obaichthys . The authors of the original study still stood by their original conclusion based on the absence of important holostean characters, and noted that it could not be a juvenile, since the bones were completely ossified. The taxonomy of catfish is quickly changing. In a 2007 and 2008 paper, Horabagrus , Phreatobius , and Conorhynchos were not classified under any current catfish families. There

408-843: A rate three times faster than that of the past century. In June 2005, researchers named the newest family of catfish, Lacantuniidae , only the third new family of fish distinguished in the last seventy years, the others being the coelacanth in 1938 and the megamouth shark in 1983. The new species in Lacantuniidae , Lacantunia enigmatica , was found in the Lacantun river in the Mexican state of Chiapas . The higher-level phylogeny of Siluriformes has gone through several recent changes, mainly due to molecular phylogenetic studies. While most studies, both morphological and molecular, agree that catfishes are arranged into three main lineages ,

459-543: A result of long branch attraction , incorrectly placing Loricarioidei as the earliest-branching catfish lineage. When a data filtering method was used to reduce lineage rate heterogeneity (the potential source of bias) on their dataset, a final phylogeny was recovered which showed the Diplomystidae are the earliest-branching catfish, followed by Loricarioidei and Siluroidei as sister lineages, providing both morphological and molecular support for Diplomystidae being

510-427: A taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow consistent naming schemes . Orders of plants , fungi , and algae use the suffix -ales (e.g. Dictyotales ). Orders of birds and fishes use

561-573: Is Phreatobius cisternarum , known to live underground in phreatic habitats. Numerous species from the families Ariidae and Plotosidae , and a few species from among the Aspredinidae and Bagridae , are found in salt water. In the Southern United States, catfish species may be known by a variety of slang names, such as "mud cat", "polliwogs", or "chuckleheads". These nicknames are not standardized, so one area may call

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612-414: Is a rare event. The Chiapas catfish mainly feeds on crabs , prawns , small fish, and large, tough plant seeds. This catfish is commonly fished in its natural habitat, where it is known as madre de juil , which means "mother of Rhamdia " (a common fish in the area). It first came to the attention of scientists in 1996, when Rócio Rodiles-Hernández, a Mexican ichthyologist conducting a biotic survey on

663-620: Is disagreement on the family status of certain groups; for example, Nelson (2006) lists Auchenoglanididae and Heteropneustidae as separate families, while the All Catfish Species Inventory (ACSI) includes them under other families. FishBase and the Integrated Taxonomic Information System lists Parakysidae as a separate family, while this group is included under Akysidae by both Nelson (2006) and ACSI. Many sources do not list

714-426: Is most similar to Ictaluridae, but lacks some important characteristics to classify this fish into this family or the other two families. Initial study has placed this fish above Diplomystidae , Cetopsidae , and Hypsidoridae , but further classification is unknown. This family probably represents a basal group in which intermediates have disappeared. With recent molecular evidence, Lacantunia has been shown to be

765-481: Is produced by glandular cells in the epidermal tissue covering the spines. In members of the family Plotosidae and of the genus Heteropneustes , this protein is so strong it may hospitalize humans who receive a sting; in Plotosus lineatus , the stings can be lethal. The dorsal- and pectoral-fin spines are two of the most conspicuous features of siluriforms, and differ from those in other fish groups. Despite

816-577: Is usually considered to be the earliest branching catfish lineage and the sister group to the other two lineages, Loricarioidei and Siluroidei. Molecular evidence usually contrasts with this hypothesis, and shows the suborder Loricarioidei as the earliest branching catfish lineage, and sister to a clade that includes the Diplomystidae and Siluroidei; this phylogeny has been obtained in numerous studies based on genetic data. However, it has been suggested that these molecular results are errors as

867-659: The Prodromus Systematis Naturalis Regni Vegetabilis of Augustin Pyramus de Candolle and the Genera Plantarum of Bentham & Hooker, it indicated taxa that are now given the rank of family (see ordo naturalis , ' natural order '). In French botanical publications, from Michel Adanson 's Familles naturelles des plantes (1763) and until the end of the 19th century, the word famille (plural: familles )

918-795: The Cypriniformes (carps and minnows), Characiformes (characins and tetras), Gonorynchiformes (milkfish and beaked salmons) and Gymnotiformes (South American knifefish), a superorder characterized by the Weberian apparatus . Some place Gymnotiformes as a sub-order of Siluriformes; however, this is not as widely accepted. Currently, the Siluriformes are said to be the sister group to the Gymnotiformes, though this has been debated due to more recent molecular evidence. As of 2007 there were about thirty-six extant catfish families, and about 3,093 extant species have been described. This makes

969-795: The Late Cretaceous , including the Andinichthyidae , Vorhisia vulpes and possibly Arius . A potential fossil record is known from the earlier Coniacian - Santonian stages in Niger of West Africa , though this has been considered unreliable, and the putative earliest armored catfish known from the fossil record, Afrocascudo , lived during the Cenomanian age of the Late Cretaceous in Morocco of North Africa ( Kem Kem Group ). The describers of Afrocascudo claimed that

1020-418: The candiru , Vandellia cirrhosa . Neither the armour-plated types nor the naked types have scales. Despite their name, not all catfish have prominent barbels or "whiskers". Members of the Siluriformes order are defined by features of the skull and swimbladder . Catfish are of considerable commercial importance; many of the larger species are farmed or fished for food. Many of the smaller species, particularly

1071-596: The sister group to Claroteidae , an African catfish family. It probably diverged from the claroteids in the Late Cretaceous between 75 and 90 million years ago. The generic name of the fish reflects its distribution in the Lacantún River drainage, flowing through the Montes Azules and Selva Lacandona Biosphere Reserves in Chiapas , Mexico into the Usumacinta River . After its initial discovery in

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1122-521: The Americas. They are the only ostariophysans that have entered freshwater habitats in Madagascar , Australia, and New Guinea . They are found in fresh water/ brackish water environments, though most inhabit shallow, running water. Representatives of at least eight families are hypogean (live underground) with three families that are also troglobitic (inhabiting caves). One such species

1173-577: The Lacantún, it was found to also inhabit other parts of the Usumacinta River basin, extending its range into northwestern Guatemala. With the description of the new family, four catfish families are now found in this region. These fish inhabit deep river channels and pools, often but not always with rocks and strong eddy currents. Few specimens were taken in stream mouths. Key variations that differentiate this genus from all other families are

1224-696: The Latin suffix -iformes meaning 'having the form of' (e.g. Passeriformes ), but orders of mammals and invertebrates are not so consistent (e.g. Artiodactyla , Actiniaria , Primates ). For some clades covered by the International Code of Zoological Nomenclature , several additional classifications are sometimes used, although not all of these are officially recognized. In their 1997 classification of mammals , McKenna and Bell used two extra levels between superorder and order: grandorder and mirorder . Michael Novacek (1986) inserted them at

1275-538: The ariid catfish, where the young retain yolk sacs late into juvenile stages, and many pimelodids, which may have elongated barbels and fin filaments or coloration patterns. Sexual dimorphism is reported in about half of all families of catfish. The modification of the anal fin into an intromittent organ (in internal fertilizers) as well as accessory structures of the reproductive apparatus (in both internal and external fertilizers) have been described in species belonging to 11 different families. Catfish have one of

1326-471: The armor is formed solely by expanded vertebral processes that form plates. Finally, the lateral armor of doradids , Sisor , and hoplomyzontines consists of hypertrophied lateral line ossicles with dorsal and ventral lamina . All catfish other than members of the Malapteruridae ( electric catfish ), possess a strong, hollow, bony, leading spine-like ray on their dorsal and pectoral fins . As

1377-518: The catfish order the second or third most diverse vertebrate order; in fact, one out of every twenty vertebrate species is a catfish. Catfish are believed to have a Gondwanan origin primarily centered around South America, as the most basal living catfish groups are known from there. The earliest known definitive members lived in the Americas from the Campanian to Maastrichtian stages of

1428-598: The corner of the mouth, and a rounded caudal fin. This fish has four pairs of barbels. The maximum length of this species reaches 42.7 centimetres (16.8 in) SL . It is depressed (flattened) near the head and compressed (thin) near the tail. The head is blunt and rounded, and the mouth is usually not subterminal (down-turned). The adipose fin is large and thick. This fish has dorsal and pectoral fin spines. Catfish Extant families: Extinct family: Catfish (or catfishes ; order Siluriformes / s ɪ ˈ lj ʊər ɪ f ɔːr m iː z / or Nematognathi ) are

1479-1533: The earliest branching catfish. Below is a list of family relationships by different authors. Lacantuniidae is included in the Sullivan scheme based on recent evidence that places it sister to Claroteidae . Phylogeny of living Siluriformes based on 2017 and extinct families based on Nelson, Grande & Wilson 2016. † Andinichthyidae Nematogenyidae [REDACTED] Trichomycteridae [REDACTED] Callichthyidae [REDACTED] Astroblepidae [REDACTED] Loricariidae [REDACTED] Diplomystidae † Bachmanniidae † Hypsidoridae Cetopsidae [REDACTED] Siluridae [REDACTED] Pangasiidae Mochokidae [REDACTED] Claroteidae Plotosidae Ictaluridae [REDACTED] Clariidae [REDACTED] Ailiidae Sisoridae [REDACTED] Bagridae [REDACTED] Aspredinidae Doradidae [REDACTED] Auchenipteridae [REDACTED] Heptapteridae [REDACTED] Pseudopimelodidae [REDACTED] Pimelodidae [REDACTED] Unassigned families: Extant catfish species live inland or in coastal waters of every continent except Antarctica . Catfish have inhabited all continents at one time or another. They are most diverse in tropical South America, Asia, and Africa, with one family native to North America and one family in Europe. More than half of all catfish species live in

1530-429: The ecological pressure on native European fauna . Walking catfish have also been introduced in the freshwater areas of Florida, with the voracious catfish becoming a major alien pest there. Flathead catfish , Pylodictis olivaris , is also a North American pest on Atlantic slope drainages. Pterygoplichthys species, released by aquarium fishkeepers, have also established feral populations in many warm waters around

1581-472: The ending -anae that was initiated by Armen Takhtajan 's publications from 1966 onwards. The order as a distinct rank of biological classification having its own distinctive name (and not just called a higher genus ( genus summum )) was first introduced by the German botanist Augustus Quirinus Rivinus in his classification of plants that appeared in a series of treatises in the 1690s. Carl Linnaeus

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1632-910: The field of zoology , the Linnaean orders were used more consistently. That is, the orders in the zoology part of the Systema Naturae refer to natural groups. Some of his ordinal names are still in use, e.g. Lepidoptera (moths and butterflies) and Diptera (flies, mosquitoes, midges, and gnats). In virology , the International Committee on Taxonomy of Viruses 's virus classification includes fifteen taxomomic ranks to be applied for viruses , viroids and satellite nucleic acids : realm , subrealm , kingdom , subkingdom, phylum , subphylum , class, subclass, order, suborder, family, subfamily , genus, subgenus , and species. There are currently fourteen viral orders, each ending in

1683-529: The fish's anatomy revealed a number of distinctive characteristics unique to this species and the absence of characters needed to place this fish in any existing catfish family. In a 2005 publication, these three ichthyologists, with Julian M. Humphries of the University of Texas, described this fish as a new species, genus and family of catfish. This fish is not related to any of the families found in its range ( Ariidae , Ictaluridae , or Heptapteridae ). It

1734-439: The former ranges throughout Europe, and the latter is restricted to Greece . Mythology and literature record wels catfish of astounding proportions yet are to be proven scientifically. The typical size of the species is about 1.2–1.6 m (3.9–5.2 ft), and fish more than 2 m (6.6 ft) are rare. However, they are known to exceed 2.5 m (8.2 ft) in length and 100 kg (220 lb) in weight. In July 2009,

1785-493: The genus Corydoras , are important in the aquarium hobby . Many catfish are nocturnal , but others (many Auchenipteridae ) are crepuscular or diurnal (most Loricariidae or Callichthyidae , for example). Molecular evidence suggests that in spite of the great morphological diversity in the order, all catfish form a monophyletic group. Catfish belong to a superorder called the Ostariophysi , which also includes

1836-456: The genus is possible. As far as known for most catfish, features that are often characteristic of species, such as mouth and fin positions, fin shapes, and barbel lengths, show little difference between juveniles and adults. For many species, pigmentation pattern is also similar in juveniles and adults. Thus, juvenile catfish generally resemble and develop smoothly into their adult form without distinct juvenile specializations. Exceptions to this are

1887-404: The largest ranges in size within a single order of bony fish . Many catfish have a maximum length of under 12 cm (4.7 in). Some of the smallest species of the Aspredinidae and Trichomycteridae reach sexual maturity at only 1 cm (0.39 in). The wels catfish , Silurus glanis , and the much smaller related Aristotle's catfish , are the only catfish indigenous to Europe ;

1938-574: The order. In loricarioids and in the Asian genus Sisor , the armor is primarily made up of one or more rows of free dermal plates. Similar plates are found in large specimens of Lithodoras . These plates may be supported by vertebral processes , as in scoloplacids and in Sisor , but the processes never fuse to the plates or form any external armor. By contrast, in the subfamily Doumeinae (family Amphiliidae ) and in hoplomyzontines ( Aspredinidae ),

1989-467: The presence of a Weberian apparatus . Their well-developed Weberian apparatus and reduced gas bladder allow for improved hearing and sound production. Catfish do not have scales ; their bodies are often naked. In some species, their mucus -covered skin is used in cutaneous respiration , where the fish breathes through its skin. In some catfish, the skin is covered in bony plates called scutes ; some form of body armor appears in various ways within

2040-517: The presence of a derived loricariid so early on would indicate the extensive diversification of catfish, or at least loricarioids, prior to the beginning of the Late Cretaceous. As extant loricariids are only known from South America, much of this diversification must have occurred on the supercontinent of West Gondwana prior to its fragmentation into South America and Africa. Britz and colleagues suggested that Afrocascudo instead represents

2091-430: The recently revised family Anchariidae . The family Horabagridae , including Horabagrus , Pseudeutropius , and Platytropius , is not shown by some authors but presented by others as a true group. Thus, the actual number of families differs between authors. The species count is in constant flux due to taxonomic work as well as description of new species. Between 2003 and 2005, over one hundred species were named,

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2142-507: The relationship among these lineages has been a contentious point in which these studies, performed for example by Rui Diogo , differ. The three main lineages in Siluriformes are the family Diplomystidae , the denticulate catfish suborder Loricarioidei (containing the Neotropical "suckermouth" catfishes), and the suborder Siluroidei, which contains the remaining families of the order. According to morphological data, Diplomystidae

2193-580: The river, collected the first specimens from the Chiapas-Guatemala border. Unable to identify the species, Rodiles involved two American catfish specialists, Dean A. Hendrickson of the University of Texas at Austin and John G. Lundberg of the Academy of Natural Sciences of Philadelphia. A detailed comparative morphological study employing high-resolution, three-dimensional CT-scan imagery of

2244-420: The same position. Michael Benton (2005) inserted them between superorder and magnorder instead. This position was adopted by Systema Naturae 2000 and others. In botany , the ranks of subclass and suborder are secondary ranks pre-defined as respectively above and below the rank of order. Any number of further ranks can be used as long as they are clearly defined. The superorder rank is commonly used, with

2295-421: The shape of the animal's skull , jaw muscles, and gas bladder — which fish use to rise and sink in water. The gas bladder has paired diverticulae , while other catfish families either have no diverticulae or singular diverticulae. Some external characteristics that may help distinguish this fish include nostrils set far apart, the presences of nasal barbels, maxillary barbels placed above the lip distantly from

2346-475: The species. Catfish barbels always occur in pairs. Many larger catfish also have chemoreceptors across their entire bodies, which means they "taste" anything they touch and "smell" any chemicals in the water. "In catfish, gustation plays a primary role in the orientation and location of food". Because their barbels and chemoreception are more important in detecting food, the eyes on catfish are generally small. Like other ostariophysans , they are characterized by

2397-599: The widespread use of the spines for taxonomic and phylogenetic studies the fields have struggled to effectively use the information due to a lack of consistency in the nomenclature, with a general standard for the descriptive anatomy of catfish spines proposed in 2022 to try and resolve this problem. Juvenile catfish, like most fish, have relatively large heads, eyes, and posterior median fins in comparison to larger, more mature individuals. These juveniles can be readily placed in their families, particularly those with highly derived fin or body shapes; in some cases, identification of

2448-578: The word family ( familia ) was assigned to the rank indicated by the French famille , while order ( ordo ) was reserved for a higher rank, for what in the 19th century had often been named a cohors (plural cohortes ). Some of the plant families still retain the names of Linnaean "natural orders" or even the names of pre-Linnaean natural groups recognized by Linnaeus as orders in his natural classification (e.g. Palmae or Labiatae ). Such names are known as descriptive family names. In

2499-428: The world. Most catfish are bottom feeders . In general, they are negatively buoyant , which means that they usually sink rather than float due to a reduced gas bladder and a heavy, bony head. Catfish have a variety of body shapes, though most have a cylindrical body with a flattened ventrum to allow for benthic feeding. A flattened head allows for digging through the substrate, as well as perhaps serving as

2550-551: Was the first to apply it consistently to the division of all three kingdoms of nature (then minerals , plants , and animals ) in his Systema Naturae (1735, 1st. Ed.). For plants, Linnaeus' orders in the Systema Naturae and the Species Plantarum were strictly artificial, introduced to subdivide the artificial classes into more comprehensible smaller groups. When the word ordo was first consistently used for natural units of plants, in 19th-century works such as

2601-561: Was used as a French equivalent for this Latin ordo . This equivalence was explicitly stated in the Alphonse Pyramus de Candolle 's Lois de la nomenclature botanique (1868), the precursor of the currently used International Code of Nomenclature for algae, fungi, and plants . In the first international Rules of botanical nomenclature from the International Botanical Congress of 1905,

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