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113-886: In biogeography , the Neotropic or Neotropical realm is one of the eight terrestrial realms . This realm includes South America, Central America , the Caribbean Islands , and southern North America. In Mexico, the Yucatán Peninsula and southern lowlands, and most of the east and west coastlines, including the southern tip of the Baja California Peninsula are Neotropical. In the United States southern Florida and coastal Central Florida are considered Neotropical. The realm also includes temperate southern South America. In contrast,

226-679: A warlike tribe that expanded along the Tapajós river and its tributaries and were feared by neighboring tribes. In the early 19th century, the Munduruku were pacified and subjugated by the Brazilians. During the Amazon rubber boom it is estimated that diseases brought by immigrants, such as typhus and malaria , killed 40,000 native Amazonians. In the 1950s, Brazilian explorer and defender of indigenous people, Cândido Rondon , supported

339-476: A combination of historical factors such as: speciation , extinction , continental drift , and glaciation . Through observing the geographic distribution of species, we can see associated variations in sea level , river routes, habitat, and river capture . Additionally, this science considers the geographic constraints of landmass areas and isolation, as well as the available ecosystem energy supplies. Over periods of ecological changes, biogeography includes

452-721: A complex civilization was flourishing along the Amazon in the 1540s. The Pre-Columbian agriculture in the Amazon Basin was sufficiently advanced to support prosperous and populous societies. It is believed that civilization was later devastated by the spread of diseases from Europe, such as smallpox . This civilization was investigated by the British explorer Percy Fawcett in the early twentieth century. The results of his expeditions were inconclusive, and he disappeared mysteriously on his last trip. His name for this lost civilization

565-713: A diverse array of indigenous peoples , who to varying degrees persist in their autonomous and traditional cultures and subsistence within this environment. The number of these peoples who are as yet relatively untouched by external influences continues to decline significantly, however, along with the near-exponential expansion of urbanization , roads, pastoralism and forest industries which encroach on their customary lands and environment. Nevertheless, amidst these declining circumstances this vast "reservoir" of human diversity continues to survive, albeit much depleted. In South America alone, some 350–400 indigenous languages and dialects are still living (down from an estimated 1,500 at

678-421: A global scale. GIS can show certain processes on the earth's surface like whale locations, sea surface temperatures , and bathymetry. Current scientists also use coral reefs to delve into the history of biogeography through the fossilized reefs. Two global information systems are either dedicated to, or have strong focus on, biogeography (in the form of the spatial location of observations of organisms), namely

791-419: A great impact on Charles Darwin , who was inspired to consider species adaptations and evolution after learning about botanical geography. De Candolle was the first to describe the differences between the small-scale and large-scale distribution patterns of organisms around the globe. Several additional scientists contributed new theories to further develop the concept of biogeography. Charles Lyell developed

904-749: A habitat and species of organisms describe the ecological application of biogeography. Historical biogeography describes the long-term, evolutionary periods of time for broader classifications of organisms. Early scientists, beginning with Carl Linnaeus , contributed to the development of biogeography as a science. The scientific theory of biogeography grows out of the work of Alexander von Humboldt (1769–1859), Francisco Jose de Caldas (1768–1816), Hewett Cottrell Watson (1804–1881), Alphonse de Candolle (1806–1893), Alfred Russel Wallace (1823–1913), Philip Lutley Sclater (1829–1913) and other biologists and explorers. The patterns of species distribution across geographical areas can usually be explained through

1017-422: A hazard. Among the largest predatory creatures are the black caiman , jaguar , cougar , and anaconda . In the river, electric eels can produce an electric shock that can stun or kill, while piranha are known to bite and injure humans. Various species of poison dart frogs secrete lipophilic alkaloid toxins through their flesh. There are also numerous parasites and disease vectors. Vampire bats dwell in

1130-413: A number of methods have been developed to produce arguably more complete "predictive" or "modelled" distributions for species based on their associated environmental or other preferences (such as availability of food or other habitat requirements); this approach is known as either Environmental niche modelling (ENM) or Species distribution modelling (SDM). Depending on the reliability of the source data and

1243-987: A result of tectonic uplift (or subsidence ), natural damming created by a landslide , or headward or lateral erosion of the watershed between adjacent basins. Biogeography is a synthetic science, related to geography , biology , soil science , geology , climatology , ecology and evolution . Some fundamental concepts in biogeography include: The study of comparative biogeography can follow two main lines of investigation: There are many types of biogeographic units used in biogeographic regionalisation schemes, as there are many criteria ( species composition , physiognomy , ecological aspects) and hierarchization schemes: biogeographic realms (ecozones), bioregions ( sensu stricto ), ecoregions , zoogeographical regions , floristic regions , vegetation types, biomes , etc. The terms biogeographic unit, biogeographic area can be used for these categories, regardless of rank. In 2008, an International Code of Area Nomenclature

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1356-492: A whole suite of predictor variables for biogeographic analysis, including satellite imaging and processing of the Earth. Two main types of satellite imaging that are important within modern biogeography are Global Production Efficiency Model (GLO-PEM) and Geographic Information Systems (GIS). GLO-PEM uses satellite-imaging gives "repetitive, spatially contiguous, and time specific observations of vegetation". These observations are on

1469-415: Is a moist broadleaf tropical rainforest in the Amazon biome that covers most of the Amazon basin of South America . This basin encompasses 7,000,000 km (2,700,000 sq mi), of which 6,000,000 km (2,300,000 sq mi) are covered by the rainforest . This region includes territory belonging to nine nations and 3,344 indigenous territories . The majority of the forest, 60%,

1582-424: Is an alternate view than that of Linnaeus. Buffon's law eventually became a principle of biogeography by explaining how similar environments were habitats for comparable types of organisms. Buffon also studied fossils which led him to believe that the Earth was over tens of thousands of years old, and that humans had not lived there long in comparison to the age of the Earth. Following the period of exploration came

1695-502: Is an integrative field of inquiry that unites concepts and information from ecology , evolutionary biology , taxonomy , geology , physical geography , palaeontology , and climatology . Modern biogeographic research combines information and ideas from many fields, from the physiological and ecological constraints on organismal dispersal to geological and climatological phenomena operating at global spatial scales and evolutionary time frames. The short-term interactions within

1808-811: Is believed that the drainage basin of the Amazon was split along the middle of the continent by the Purus Arch . Water on the eastern side flowed toward the Atlantic , while to the west water flowed toward the Pacific across the Amazonas Basin . As the Andes Mountains rose, however, a large basin was created that enclosed a lake; now known as the Solimões Basin . Within the last 5–10 million years, this accumulating water broke through

1921-526: Is considerable, and areas cleared of forest are visible to the naked eye from outer space. In the 1970s, construction began on the Trans-Amazonian highway . This highway represented a major threat to the Amazon rainforest. The highway still has not been completed, limiting the environmental damage. Between 1991 and 2000, the total area of forest lost in the Amazon rose from 415,000 to 587,000 km (160,000 to 227,000 sq mi), with most of

2034-570: Is endemic to the Neotropic realm, occupying a larger geographic area than any other piranha species. Some fish groups originally unique to the Neotropics include: Examples of other animal groups that are entirely or mainly restricted to the Neotropical region include: According to Simberloff. as of 1984 there were a total of 92,128 species of flowering plants (Angiosperms) in the Neotropics. Plant families endemic and partly subendemic to

2147-426: Is estimated at 356 ± 47 tonnes per hectare. To date, an estimated 438,000 species of plants of economic and social interest have been registered in the region with many more remaining to be discovered or catalogued. The total number of tree species in the region is estimated at 16,000. The green leaf area of plants and trees in the rainforest varies by about 25% as a result of seasonal changes. Leaves expand during

2260-507: Is in Brazil , followed by Peru with 13%, Colombia with 10%, and with minor amounts in Bolivia , Ecuador , French Guiana , Guyana , Suriname , and Venezuela . Four nations have " Amazonas " as the name of one of their first-level administrative regions , and France uses the name " Guiana Amazonian Park " for French Guiana's protected rainforest area. The Amazon represents over half of

2373-404: Is in the geological similarities between varying locations around the globe, the geographic distribution of some fossils (including the mesosaurs ) on various continents, and the jigsaw puzzle shape of the landmasses on Earth. Though Wegener did not know the mechanism of this concept of Continental Drift, this contribution to the study of biogeography was significant in the way that it shed light on

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2486-723: Is mostly covered by tropical moist broadleaf forest , including the vast Amazon rainforest , which stretches from the Andes Mountains to the Atlantic Ocean, and the lowland forests of the Guianas . The bioregion also includes tropical savanna and tropical dry forest ecoregions. The Central Andes lie between the Gulfs of Guayaquil and Penas and thus encompass southern Ecuador, Chile, Peru, western Bolivia, and northwest and western Argentina. Eastern South America includes

2599-473: Is on how the environment and humans affect the distribution of species as well as other manifestations of Life such as species or genetic diversity. Biogeography is being applied to biodiversity conservation and planning, projecting global environmental changes on species and biomes, projecting the spread of infectious diseases, invasive species, and for supporting planning for the establishment of crops. Technological evolving and advances have allowed for generating

2712-448: Is the branch of biogeography that studies the distribution of plants. Zoogeography is the branch that studies distribution of animals. Mycogeography is the branch that studies distribution of fungi, such as mushrooms . Knowledge of spatial variation in the numbers and types of organisms is as vital to us today as it was to our early human ancestors , as we adapt to heterogeneous but geographically predictable environments . Biogeography

2825-571: Is the conversion of forested areas to non-forested areas. The main sources of deforestation in the Amazon are human settlement and the development of the land. In 2022, about 20% of the Amazon rainforest has already been deforested and a further 6% was "highly degraded". Research suggests that upon reaching about 20–25% (hence 0–5% more), the tipping point to flip it into a non-forest ecosystem – degraded savannah – (in eastern, southern and central Amazonia) will be reached. This process of savanisation would take decades to take full effect. Prior to

2938-614: The Age of Enlightenment in Europe, which attempted to explain the patterns of biodiversity observed by Buffon and Linnaeus. At the birth of the 19th century, Alexander von Humboldt, known as the "founder of plant geography", developed the concept of physique generale to demonstrate the unity of science and how species fit together. As one of the first to contribute empirical data to the science of biogeography through his travel as an explorer, he observed differences in climate and vegetation. The Earth

3051-838: The Caatinga xeric shrublands of northeastern Brazil, the broad Cerrado grasslands and savannas of the Brazilian Plateau , and the Pantanal and Chaco grasslands. The diverse Atlantic forests of eastern Brazil are separated from the forests of Amazonia by the Caatinga and Cerrado, and are home to a distinct flora and fauna. North of the Gulf of Guayaquil in Ecuador and Colombia, a series of accreted oceanic terranes (discrete allochthonous fragments) have developed that constitute

3164-654: The Cretaceous–Paleogene extinction event altered local flora and fauna. Much later, about two to three million years ago, South America was joined with North America by the formation of the Isthmus of Panama , which allowed a biotic exchange between the two continents, the Great American Interchange . South American species like the ancestors of the Virginia opossum ( Didelphis virginiana ) and

3277-532: The Eocene era (from 56 million years to 33.9 million years ago). It appeared following a global reduction of tropical temperatures when the Atlantic Ocean had widened sufficiently to provide a warm, moist climate to the Amazon basin. The rainforest has been in existence for at least 55 million years, and most of the region remained free of savanna -type biomes at least until the current ice age when

3390-713: The Global Biodiversity Information Facility (GBIF: 2.57 billion species occurrence records reported as at August 2023) and, for marine species only, the Ocean Biodiversity Information System (OBIS, originally the Ocean Biogeographic Information System : 116 million species occurrence records reported as at August 2023), while at a national scale, similar compilations of species occurrence records also exist such as

3503-470: The Great American Interchange , an important biogeographical event. The Neotropic includes more tropical rainforest ( tropical and subtropical moist broadleaf forests ) than any other realm, extending from southern Mexico through Central America and northern South America to southern Brazil, including the vast Amazon rainforest . These rainforest ecoregions are one of the most important reserves of biodiversity on Earth. These rainforests are also home to

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3616-581: The Villas-Bôas brothers ' campaign, which faced strong opposition from the government and the ranchers of Mato Grosso and led to the establishment of the first Brazilian National Park for indigenous people along the Xingu River in 1961. In 1961, British explorer Richard Mason was killed by an uncontacted Amazon tribe known as the Panará . The Matsés made their first permanent contact with

3729-547: The armadillo moved into North America, and North Americans like the ancestors of South America's camelids , including the llama ( Lama glama ), moved south. The long-term effect of the exchange was the extinction of many South American species, mostly by outcompetition by northern species. The Neotropical realm has 31 endemic bird families, which is over twice the number of any other realm. They include tanagers , rheas , tinamous , curassows , antbirds , ovenbirds , toucans , and seriemas . Bird families originally unique to

3842-434: The release of the carbon contained within the vegetation, which could accelerate global warming . Amazonian evergreen forests account for about 10% of the world's terrestrial primary productivity and 10% of the carbon stores in ecosystems – of the order of 1.1 × 10 metric tonnes of carbon. Amazonian forests are estimated to have accumulated 0.62 ± 0.37 tons of carbon per hectare per year between 1975 and 1996. In 2021 it

3955-689: The temperate rain forests of the Valdivian temperate rain forests and Magellanic subpolar forests ecoregions, and the Juan Fernández Islands and Desventuradas Islands , are a refuge for the ancient Antarctic flora , which includes trees like the southern beech ( Nothofagus ), podocarps , the alerce ( Fitzroya cupressoides ), and Araucaria pines like the monkey-puzzle tree ( Araucaria araucana ). These rainforests are endangered by extensive logging and their replacement by fast-growing non-native pines and eucalyptus . South America

4068-469: The 2000s. Oil companies have to set up their operations by opening new roads through the forests, which often contributes to deforestation in the region. 9.4% of the territory of the Amazon is affected by oil fields. Mining is also a major driver of deforestation. 17% of the area of the Amazon Rainforest is affected by mining. The transition to solar and wind energy, digitalization, raised

4181-479: The 36 volume Histoire Naturelle, générale et particulière , in which he argued that varying geographical regions would have different forms of life. This was inspired by his observations comparing the Old and New World, as he determined distinct variations of species from the two regions. Buffon believed there was a single species creation event, and that different regions of the world were homes for varying species, which

4294-456: The Amazon basin, Orinoco basin, and Guianas ) with an exceptionally low (flat) topographic relief, the many waterways have had a highly reticulated history over geological time . In such a context, stream capture is an important factor affecting the evolution and distribution of freshwater organisms. Stream capture occurs when an upstream portion of one river drainage is diverted to the downstream portion of an adjacent basin. This can happen as

4407-461: The Amazon in 1492. By 1900, the native indigenous population had fallen to 1 million and by the early 1980s it was less than 200,000. The first European to travel the length of the Amazon River was Francisco de Orellana in 1542. The BBC's Unnatural Histories presents evidence that Orellana, rather than exaggerating his claims as previously thought, was correct in his observations that

4520-526: The Amazon rainforest are probably the result of centuries of human management, rather than naturally occurring as has previously been supposed. In the region of the Xingu tribe, remains of some of these large settlements in the middle of the Amazon forest were found in 2003 by Michael Heckenberger and colleagues of the University of Florida . Among those were evidence of roads, bridges and large plazas. In

4633-727: The Amazon rainforest comes from the Bodélé depression in Northern Chad in the Sahara desert. The dust contains phosphorus , important for plant growth. The yearly Sahara dust replaces the equivalent amount of phosphorus washed away yearly in Amazon soil from rains and floods. NASA's CALIPSO satellite has measured the amount of dust transported by wind from the Sahara to the Amazon: an average of 182 million tons of dust are windblown out of

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4746-532: The Amazon rainforest could become unsustainable under conditions of severely reduced rainfall and increased temperatures, leading to an almost complete loss of rainforest cover in the basin by 2100., and severe economic, natural capital and ecosystem services impacts of not averting the tipping point. However, simulations of Amazon basin climate change across many different models are not consistent in their estimation of any rainfall response, ranging from weak increases to strong decreases. The result indicates that

4859-460: The Amazon rainforest, rather than being a pristine wilderness , has been shaped by man for at least 11,000 years through practices such as forest gardening and terra preta . Terra preta is found over large areas in the Amazon forest; and is now widely accepted as a product of indigenous soil management . The development of this fertile soil allowed agriculture and silviculture in the previously hostile environment; meaning that large portions of

4972-423: The Amazon region at least 11,200 years ago. Subsequent development led to late-prehistoric settlements along the periphery of the forest by AD 1250, which induced alterations in the forest cover . For a long time, it was thought that the Amazon rainforest was never more than sparsely populated, as it was impossible to sustain a large population through agriculture given the poor soil. Archeologist Betty Meggers

5085-578: The Amazon region. In August 2019 there were a record number of fires. Deforestation in the Brazilian Amazon rose more than 88% in June 2019 compared with the same month in 2018. The increased area of fire-impacted forest coincided with a relaxation of environmental regulations from the Brazilian government. Notably, before those regulations were put in place in 2008 the fire-impacted area

5198-787: The Amazonas, there has been fighting and wars between the neighboring tribes of the Jivaro . Several tribes of the Jivaroan group, including the Shuar , practised headhunting for trophies and headshrinking . The accounts of missionaries to the area in the borderlands between Brazil and Venezuela have recounted constant infighting in the Yanomami tribes. More than a third of the Yanomamo males, on average, died from warfare. The Munduruku were

5311-603: The Baudo, or Coastal, Mountains and the Cordillera Occidental. The Orinoco is a region of humid forested broadleaf forest and wetland primarily comprising the drainage basin for the Orinoco River and other adjacent lowland forested areas. This region includes most of Venezuela and parts of Colombia, as well as Trinidad and Tobago . The temperate forest ecoregions of southwestern South America, including

5424-597: The Gómez Farias Region, Tamaulipas, Mexico , which has been described as "ground-breaking" and "a classic treatise in historical biogeography". Martin applied several disciplines including ecology , botany , climatology , geology , and Pleistocene dispersal routes to examine the herpetofauna of a relatively small and largely undisturbed area, but ecologically complex, situated on the threshold of temperate – tropical (nearctic and neotropical) regions, including semiarid lowlands at 70 meters elevation and

5537-619: The Indian Ocean was much narrower than it is today, and that South America was closer to the Antarctic, one would be hard pressed to explain the presence of many "ancient" lineages of perching birds in Africa, as well as the mainly South American distribution of the suboscines . Paleobiogeography also helps constrain hypotheses on the timing of biogeographic events such as vicariance and geodispersal , and provides unique information on

5650-493: The Neotropic include Bromeliaceae , Cannaceae and Heliconiaceae . Plant species with economic importance originally unique to the Neotropic include: Biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time . Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude , elevation , isolation and habitat area . Phytogeography

5763-652: The Neotropical Floristic Kingdom excludes southernmost South America, which instead is placed in the Antarctic kingdom . The Neotropic is delimited by similarities in fauna or flora . Its fauna and flora are distinct from the Nearctic realm (which includes most of North America) because of the long separation of the two continents. The formation of the Isthmus of Panama joined the two continents two to three million years ago, precipitating

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5876-451: The Neotropics include hummingbirds (family Trochilidae) and wrens (family Troglodytidae). Mammal groups originally unique to the Neotropics include: The Neotropical realm has 63 endemic fish families and subfamilies, which is more than any other realm. Neotropical fishes include more than 5,700 species, and represent at least 66 distinct lineages in continental freshwaters (Albert and Reis, 2011). The well-known red-bellied piranha

5989-452: The Origin of Species were devoted to geographical distribution. The first discoveries that contributed to the development of biogeography as a science began in the mid-18th century, as Europeans explored the world and described the biodiversity of life. During the 18th century most views on the world were shaped around religion and for many natural theologists, the bible. Carl Linnaeus , in

6102-631: The Purus Arch, joining the easterly flow toward the Atlantic. There is evidence that there have been significant changes in the Amazon rainforest vegetation over the last 21,000 years through the last glacial maximum (LGM) and subsequent deglaciation. Analyses of sediment deposits from Amazon basin paleolakes and the Amazon Fan indicate that rainfall in the basin during the LGM was lower than for

6215-545: The Sahara each year (some dust falls into the Atlantic), 15% of which of falls over the Amazon basin (22 million tons of it consisting of phosphorus). CALIPSO uses a laser range finder to scan the Earth's atmosphere for the vertical distribution of dust and other aerosols. and regularly tracks the Sahara-Amazon dust plume. CALIPSO has measured variations in the dust amounts transported – an 86 percent drop between

6328-482: The Theory of Continental Drift in 1912, though it was not widely accepted until the 1960s. This theory was revolutionary because it changed the way that everyone thought about species and their distribution around the globe. The theory explained how continents were formerly joined in one large landmass, Pangea , and slowly drifted apart due to the movement of the plates below Earth's surface. The evidence for this theory

6441-603: The Theory of Uniformitarianism after studying fossils. This theory explained how the world was not created by one sole catastrophic event, but instead from numerous creation events and locations. Uniformitarianism also introduced the idea that the Earth was actually significantly older than was previously accepted. Using this knowledge, Lyell concluded that it was possible for species to go extinct. Since he noted that Earth's climate changes, he realized that species distribution must also change accordingly. Lyell argued that climate changes complemented vegetation changes, thus connecting

6554-639: The U.K. National Biodiversity Network , the Atlas of Living Australia , and many others. In the case of the oceans, in 2017 Costello et al. analyzed the distribution of 65,000 species of marine animals and plants as then documented in OBIS, and used the results to distinguish 30 distinct marine realms, split between continental-shelf and offshore deep-sea areas. Since it is self evident that compilations of species occurrence records cannot cover with any completeness, areas that have received either limited or no sampling,

6667-414: The anticipated effects of climate change can also be used to show potential changes in species distributions that may occur in the future based on such scenarios. Paleobiogeography goes one step further to include paleogeographic data and considerations of plate tectonics . Using molecular analyses and corroborated by fossils , it has been possible to demonstrate that perching birds evolved first in

6780-407: The area and so the lower the yield per hectare becomes. So despite the popular perception, there has been no economical advantage for Brazil from logging rainforest zones and converting these to pastoral fields. The needs of soy farmers have been used to justify many of the controversial transportation projects that are currently developing in the Amazon. The first two highways successfully opened up

6893-421: The biotic and abiotic features of the Earth in his book, Cosmos . Augustin de Candolle contributed to the field of biogeography as he observed species competition and the several differences that influenced the discovery of the diversity of life. He was a Swiss botanist and created the first Laws of Botanical Nomenclature in his work, Prodromus. He discussed plant distribution and his theories eventually had

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7006-409: The climate was drier and savanna more widespread. Following the Cretaceous–Paleogene extinction event , the extinction of the dinosaurs and the wetter climate may have allowed the tropical rainforest to spread out across the continent. From 66 to 34 Mya , the rainforest extended as far south as 45° . Climate fluctuations during the last 34 million years have allowed savanna regions to expand into

7119-549: The demand for cassiterite (the main ore of tin used also for financing gold mining ), manganese and copper , which attracrted many illegal miners to the Amazon. This led to deforestation, different environmental and social problems. Hydropower also creates significant problems in the Amazon. Such activities are defined by the World Rainforest Movement as " Green extractivism ". The European Union–Mercosur free trade agreement , which would form one of

7232-420: The development of the fields of conservation biology and landscape ecology . Classic biogeography has been expanded by the development of molecular systematics , creating a new discipline known as phylogeography . This development allowed scientists to test theories about the origin and dispersal of populations, such as island endemics . For example, while classic biogeographers were able to speculate about

7345-421: The dry season when sunlight is at a maximum, then undergo abscission in the cloudy wet season. These changes provide a balance of carbon between photosynthesis and respiration. Each hectare of the Amazon rainforest contains around 1 billion of invertebrates . The amount of species per hectare in the Amazon rainforest can be presented in the next table: The rainforest contains several species that can pose

7458-591: The early 1960s, access to the forest's interior was highly restricted, and the forest remained basically intact. Farms established during the 1960s were based on crop cultivation and the slash and burn method. However, the colonists were unable to manage their fields and the crops because of the loss of soil fertility and weed invasion. The soils in the Amazon are productive for just a short period of time, so farmers are constantly moving to new areas and clearing more land. These farming practices led to deforestation and caused extensive environmental damage. Deforestation

7571-504: The environmental surroundings to varying species. This largely influenced Charles Darwin in his development of the theory of evolution. Charles Darwin was a natural theologist who studied around the world, and most importantly in the Galapagos Islands . Darwin introduced the idea of natural selection, as he theorized against previously accepted ideas that species were static or unchanging. His contributions to biogeography and

7684-597: The factors affecting organism distribution, and to predict future trends in organism distribution. Often mathematical models and GIS are employed to solve ecological problems that have a spatial aspect to them. Biogeography is most keenly observed on the world's islands . These habitats are often much more manageable areas of study because they are more condensed than larger ecosystems on the mainland. Islands are also ideal locations because they allow scientists to look at habitats that new invasive species have only recently colonized and can observe how they disperse throughout

7797-458: The formation of regional biotas. For example, data from species-level phylogenetic and biogeographic studies tell us that the Amazonian teleost fauna accumulated in increments over a period of tens of millions of years, principally by means of allopatric speciation, and in an arena extending over most of the area of tropical South America (Albert & Reis 2011). In other words, unlike some of

7910-425: The former Lifemapper project at the University of Kansas (now continued as a part of BiotaPhy ) and AquaMaps , which as at 2023 contain modelled distributions for around 200,000 terrestrial, and 33,000 species of teleosts , marine mammals and invertebrates, respectively. One advantage of ENM/SDM is that in addition to showing current (or even past) modelled distributions, insertion of changed parameters such as

8023-655: The geographical distribution of organisms around the globe. Alfred Russel Wallace studied the distribution of flora and fauna in the Amazon Basin and the Malay Archipelago in the mid-19th century. His research was essential to the further development of biogeography, and he was later nicknamed the "father of Biogeography". Wallace conducted fieldwork researching the habits, breeding and migration tendencies, and feeding behavior of thousands of species. He studied butterfly and bird distributions in comparison to

8136-518: The highest amount of dust transported in 2007 and the lowest in 2011. This is possibly causing by rainfall variations is the Sahel , a strip of semi-arid land on the southern border of the Sahara.. Amazon phosphorus also comes as smoke due to biomass burning in Africa. Wet tropical forests are the most species-rich biome , and tropical forests in the Americas are consistently more species rich than

8249-411: The importance of environmental and geographic similarities or differences as a result of climate and other pressures on the planet. Importantly, late in his career Wegener recognised that testing his theory required measurement of continental movement rather than inference from fossils species distributions. In 1958 paleontologist Paul S. Martin published A Biogeography of Reptiles and Amphibians in

8362-559: The island and change it. They can then apply their understanding to similar but more complex mainland habitats. Islands are very diverse in their biomes , ranging from the tropical to arctic climates. This diversity in habitat allows for a wide range of species study in different parts of the world. One scientist who recognized the importance of these geographic locations was Charles Darwin , who remarked in his journal "The Zoology of Archipelagoes will be well worth examination". Two chapters in On

8475-400: The lost forest becoming pasture for cattle. Seventy percent of formerly forested land in the Amazon, and 91% of land deforested since 1970, have been used for livestock pasture . Currently, Brazil is the largest global producer of soybeans . New research however, conducted by Leydimere Oliveira et al., has shown that the more rainforest is logged in the Amazon, the less precipitation reaches

8588-488: The mid-18th century, improved our classifications of organisms through the exploration of undiscovered territories by his students and disciples. When he noticed that species were not as perpetual as he believed, he developed the Mountain Explanation to explain the distribution of biodiversity; when Noah's ark landed on Mount Ararat and the waters receded, the animals dispersed throughout different elevations on

8701-515: The most important and consequential developments in biogeography has been to show how multiple organisms, including mammals like monkeys and reptiles like squamates , overcame barriers such as large oceans that many biogeographers formerly believed were impossible to cross. See also Oceanic dispersal . Biogeography now incorporates many different fields including but not limited to physical geography, geology, botany and plant biology, zoology, general biology, and modelling. A biogeographer's main focus

8814-505: The mountain. This showed different species in different climates proving species were not constant. Linnaeus' findings set a basis for ecological biogeography. Through his strong beliefs in Christianity, he was inspired to classify the living world, which then gave way to additional accounts of secular views on geographical distribution. He argued that the structure of an animal was very closely related to its physical surroundings. This

8927-683: The nature of the models employed (including the scales for which data are available), maps generated from such models may then provide better representations of the "real" biogeographic distributions of either individual species, groups of species, or biodiversity as a whole, however it should also be borne in mind that historic or recent human activities (such as hunting of great whales , or other human-induced exterminations) may have altered present-day species distributions from their potential "full" ecological footprint. Examples of predictive maps produced by niche modelling methods based on either GBIF (terrestrial) or OBIS (marine, plus some freshwater) data are

9040-476: The northernmost cloud forest in the western hemisphere at over 2200 meters. The publication of The Theory of Island Biogeography by Robert MacArthur and E.O. Wilson in 1967 showed that the species richness of an area could be predicted in terms of such factors as habitat area, immigration rate and extinction rate. This added to the long-standing interest in island biogeography . The application of island biogeography theory to habitat fragments spurred

9153-583: The origins of species in the Hawaiian Islands , phylogeography allows them to test theories of relatedness between these populations and putative source populations on various continents, notably in Asia and North America . Biogeography continues as a point of study for many life sciences and geography students worldwide, however it may be under different broader titles within institutions such as ecology or evolutionary biology. In recent years, one of

9266-503: The outside world in 1969. Before that date, they were effectively at-war with the Peruvian government. Nine countries share the Amazon basin—most of the rainforest, 58.4%, is contained within the borders of Brazil. The other eight countries are Peru with 12.8%, Bolivia with 7.7%, Colombia with 7.1%, Venezuela with 6.1%, Guyana with 3.1%, Suriname with 2.5%, French Guiana with 1.4% and Ecuador with 1%. The rainforest likely formed during

9379-448: The presence or absence of geographical barriers. His observations led him to conclude that the number of organisms present in a community was dependent on the amount of food resources in the particular habitat. Wallace believed species were dynamic by responding to biotic and abiotic factors. He and Philip Sclater saw biogeography as a source of support for the theory of evolution as they used Darwin's conclusion to explain how biogeography

9492-411: The present, and this was almost certainly associated with reduced moist tropical vegetation cover in the basin. In present day, the Amazon receives approximately 9 feet of rainfall annually. There is a debate, however, over how extensive this reduction was. Some scientists argue that the rainforest was reduced to small, isolated refugia separated by open forest and grassland; other scientists argue that

9605-438: The rainforest and can spread the rabies virus. Malaria , yellow fever and dengue fever can also be contracted in the Amazon region. The biodiversity in the Amazon is becoming increasingly threatened, primarily by habitat loss from deforestation as well as increased frequency of fires. Over 90% of Amazonian plant and vertebrate species (13,000–14,000 in total) may have been impacted to some degree by fires. Deforestation

9718-464: The rainforest and led to increased settlement and deforestation. The mean annual deforestation rate from 2000 to 2005 (22,392 km or 8,646 sq mi per year) was 18% higher than in the previous five years (19,018 km or 7,343 sq mi per year). Although deforestation declined significantly in the Brazilian Amazon between 2004 and 2014, there has been an increase to the present day. Brazil's President, Jair Bolsonaro, has supported

9831-676: The rainforest could be threatened through the 21st century by climate change in addition to deforestation. In 1989, environmentalist C.M. Peters and two colleagues stated there is economic as well as biological incentive to protecting the rainforest. One hectare in the Peruvian Amazon has been calculated to have a value of $ 6820 if intact forest is sustainably harvested for fruits, latex, and timber; $ 1000 if clear-cut for commercial timber (not sustainably harvested); or $ 148 if used as cattle pasture. As indigenous territories continue to be destroyed by deforestation and ecocide (such as in

9944-409: The rainforest remained largely intact but extended less far to the north, south, and east than is seen today. This debate has proved difficult to resolve because the practical limitations of working in the rainforest mean that data sampling is biased away from the center of the Amazon basin, and both explanations are reasonably well supported by the available data. More than 56% of the dust fertilizing

10057-578: The realm are, according to Takhtajan (1978), Hymenophyllopsidaceae , Marcgraviaceae , Caryocaraceae , Pellicieraceae , Quiinaceae , Peridiscaceae , Bixaceae , Cochlospermaceae , Tovariaceae , Lissocarpaceae ( Lissocarpa ), Brunelliaceae , Dulongiaceae , Columelliaceae , Julianiaceae , Picrodendraceae , Goupiaceae , Desfontainiaceae , Plocospermataceae , Tropaeolaceae , Dialypetalanthaceae ( Dialypetalanthus ), Nolanaceae ( Nolana ), Calyceraceae , Heliconiaceae , Cannaceae , Thurniaceae and Cyclanthaceae . Plant families that originated in

10170-691: The realm into bioregions , defined as "geographic clusters of ecoregions that may span several habitat types, but have strong biogeographic affinities, particularly at taxonomic levels higher than the species level (genus, family)." Laurel forest and other cloud forest are subtropical and mild temperate forest , found in areas with high humidity and relatively stable and mild temperatures. Tropical rainforest , tropical and subtropical moist broadleaf forests are highlight in Southern North America, Amazonia, Caribbean, Central America, Northern Andes and Central Andes. The Amazonia bioregion

10283-420: The region of Australia or the adjacent Antarctic (which at that time lay somewhat further north and had a temperate climate). From there, they spread to the other Gondwanan continents and Southeast Asia – the part of Laurasia then closest to their origin of dispersal – in the late Paleogene , before achieving a global distribution in the early Neogene . Not knowing that at the time of dispersal,

10396-452: The region was actually densely populated. The Upano Valley sites in present-day eastern Ecuador predate all known complex Amazonian societies. Some 5 million people may have lived in the Amazon region in AD 1500, divided between dense coastal settlements, such as that at Marajó , and inland dwellers. Based on projections of food production, one estimate suggests over 8 million people living in

10509-550: The region. The name Amazon is said to arise from a war Francisco de Orellana fought with the Tapuyas and other tribes. The women of the tribe fought alongside the men, as was their custom. Orellana derived the name Amazonas from the Amazons of Greek mythology , described by Herodotus and Diodorus . Based on archaeological evidence from an excavation at Caverna da Pedra Pintada , human inhabitants first settled in

10622-650: The region. One in five of all bird species are found in the Amazon rainforest, and one in five of the fish species live in Amazonian rivers and streams. Scientists have described between 96,660 and 128,843 invertebrate species in Brazil alone. The biodiversity of plant species is the highest on Earth with one 2001 study finding a quarter square kilometer (62 acres) of Ecuadorian rainforest supports more than 1,100 tree species. A study in 1999 found one square kilometer (247 acres) of Amazon rainforest can contain about 90,790 tonnes of living plants. The average plant biomass

10735-497: The relaxation of regulations placed on agricultural land. He has used his time in office to allow for more deforestation and more exploitation of the Amazon's rich natural resources. Deforestation reached a 15 year high in 2021. Since the discovery of fossil fuel reservoirs in the Amazon rainforest, oil drilling activity has steadily increased, peaking in the Western Amazon in the 1970s and ushering another drilling boom in

10848-428: The study of plant and animal species in: their past and/or present living refugium habitat ; their interim living sites; and/or their survival locales. As writer David Quammen put it, "...biogeography does more than ask Which species? and Where . It also asks Why? and, what is sometimes more crucial, Why not? ." Modern biogeography often employs the use of Geographic Information Systems (GIS), to understand

10961-416: The theory of evolution were different from those of other explorers of his time, because he developed a mechanism to describe the ways that species changed. His influential ideas include the development of theories regarding the struggle for existence and natural selection. Darwin's theories started a biological segment to biogeography and empirical studies, which enabled future scientists to develop ideas about

11074-478: The time of first European contact ), in about 37 distinct language families and a further number of unclassified and isolate languages . Many of these languages and their cultures are also endangered. Accordingly, conservation in the Neotropical realm is a hot political concern, and raises many arguments about development versus indigenous versus ecological rights and access to or ownership of natural resources . The World Wide Fund for Nature (WWF) subdivides

11187-455: The total area of remaining rainforests on Earth , and comprises the largest and most biodiverse tract of tropical rainforest in the world, with an estimated 390 billion individual trees in about 16,000 species. More than 30 million people of 350 different ethnic groups live in the Amazon, which are subdivided into 9 different national political systems and 3,344 formally acknowledged indigenous territories . Indigenous peoples make up 9% of

11300-536: The total population, and 60 of the groups remain largely isolated. Large scale deforestation is occurring in the forest, creating different harmful effects. Economic losses due to deforestation in Brazil could be approximately 7 times higher in comparison to the cost of all commodities produced through deforestation. In 2023, the World Bank published a report proposing a non-deforestation based economic program in

11413-524: The tropics. During the Oligocene , for example, the rainforest spanned a relatively narrow band. It expanded again during the Middle Miocene , then retracted to a mostly inland formation at the last glacial maximum . However, the rainforest still managed to thrive during these glacial periods , allowing for the survival and evolution of a broad diversity of species. During the mid-Eocene , it

11526-447: The well-known insular faunas ( Galapagos finches , Hawaiian drosophilid flies, African rift lake cichlids ), the species-rich Amazonian ichthyofauna is not the result of recent adaptive radiations . For freshwater organisms, landscapes are divided naturally into discrete drainage basins by watersheds , episodically isolated and reunited by erosional processes. In regions like the Amazon Basin (or more generally Greater Amazonia,

11639-614: The wet forests in Africa and Asia. As the largest tract of tropical rainforest in the Americas, the Amazonian rainforests have unparalleled biodiversity . One in ten known species in the world lives in the Amazon rainforest. This constitutes the largest collection of living plants and animal species in the world . The region is home to about 2.5 million insect species , tens of thousands of plants, and some 2,000 birds and mammals . To date, at least 40,000 plant species, 2,200 fishes , 1,294 birds, 427 mammals, 428 amphibians, and 378 reptiles have been scientifically classified in

11752-495: The world's largest free trade areas, has been denounced by environmental activists and indigenous rights campaigners. The fear is that the deal could lead to more deforestation of the Amazon rainforest as it expands market access to Brazilian beef. According to a November 2021 report by Brazil's INPE , based on satellite data , deforestation has increased by 22% over 2020 and is at its highest level since 2006. There were 72,843 fires in Brazil in 2019, with more than half within

11865-404: Was a prominent proponent of this idea, as described in her book Amazonia: Man and Culture in a Counterfeit Paradise . She claimed that a population density of 0.2 inhabitants per square kilometre (0.52/sq mi) is the maximum that can be sustained in the rainforest through hunting, with agriculture needed to host a larger population. However, recent anthropological findings have suggested that

11978-410: Was also larger compared to the regulation period of 2009–2018. As these fire continue to move closer to the heart of the Amazon basin, their impact on biodiversity will only increase in scale, as the cumulative fire-impacted area is correlated with the number of species impacted. Environmentalists are concerned about loss of biodiversity that will result from destruction of the forest , and also about

12091-400: Was divided into regions which he defined as tropical, temperate, and arctic and within these regions there were similar forms of vegetation. This ultimately enabled him to create the isotherm, which allowed scientists to see patterns of life within different climates. He contributed his observations to findings of botanical geography by previous scientists, and sketched this description of both

12204-529: Was important to a George Louis Buffon's rival theory of distribution. Closely after Linnaeus, Georges-Louis Leclerc, Comte de Buffon observed shifts in climate and how species spread across the globe as a result. He was the first to see different groups of organisms in different regions of the world. Buffon saw similarities between some regions which led him to believe that at one point continents were connected and then water separated them and caused differences in species. His hypotheses were described in his work,

12317-485: Was originally part of the supercontinent of Gondwana , which included Africa, Australia, India, New Zealand, and Antarctica, and the Neotropic shares many plant and animal lineages with these other continents, including marsupial mammals and the Antarctic flora . After the final breakup of the Gondwana about 110 million years ago, South America was separated from Africa and drifted north and west. 66 million years ago,

12430-715: Was proposed for biogeography. It achieved limited success; some studies commented favorably on it, but others were much more critical, and it "has not yet gained a significant following". Similarly, a set of rules for paleobiogeography has achieved limited success. In 2000, Westermann suggested that the difficulties in getting formal nomenclatural rules established in this field might be related to "the curious fact that neither paleo- nor neobiogeographers are organized in any formal groupings or societies, nationally (so far as I know) or internationally — an exception among active disciplines." Amazon rainforest The Amazon rainforest , also called Amazon jungle or Amazonia ,

12543-414: Was reported that the Amazon for the first time emitted more greenhouse gases than it absorbed. Though often referenced as producing more than a quarter of the Earth's oxygen, this often stated, but misused statistic actually refers to oxygen turnover. The net contribution of the ecosystem is approximately zero. One computer model of future climate change caused by greenhouse gas emissions shows that

12656-502: Was similar to a record of species inheritance. Key findings, such as the sharp difference in fauna either side of the Wallace Line , and the sharp difference that existed between North and South America prior to their relatively recent faunal interchange , can only be understood in this light. Otherwise, the field of biogeography would be seen as a purely descriptive one. Moving on to the 20th century, Alfred Wegener introduced

12769-451: Was the City of Z . Since the 1970s, numerous geoglyphs have been discovered on deforested land dating between AD 1–1250, furthering claims about Pre-Columbian civilizations. Ondemar Dias is accredited with first discovering the geoglyphs in 1977, and Alceu Ranzi is credited with furthering their discovery after flying over Acre . The BBC's Unnatural Histories presented evidence that

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