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47-519: The Midland Oak is an oak tree that grows in Leamington Spa , Warwickshire , near the Lillington boundary, at the junction of Lillington Road and Lillington Avenue. A plaque nearby unveiled in 1988 states that the old tree, the original centuries old Midland Oak was reputed to mark the centre of England . The present tree was planted about 1988, grown from an acorn saved in 1967 when

94-667: A common history of pre-Proto-Germanic speakers throughout the Nordic Bronze Age . The Proto-Germanic language developed in southern Scandinavia (Denmark, south Sweden and southern Norway) and the northern-most part of Germany in Schleswig Holstein and northern Lower Saxony, the Urheimat (original home) of the Germanic tribes. It is possible that Indo-European speakers first arrived in southern Scandinavia with

141-432: A dialect of Proto-Indo-European that had lost its laryngeals and had five long and six short vowels as well as one or two overlong vowels. The consonant system was still that of PIE minus palatovelars and laryngeals, but the loss of syllabic resonants already made the language markedly different from PIE proper. Mutual intelligibility might have still existed with other descendants of PIE, but it would have been strained, and

188-475: A larger scope of linguistic developments, spanning the Nordic Bronze Age and Pre-Roman Iron Age in Northern Europe (second to first millennia BC) to include "Pre-Germanic" (PreGmc), "Early Proto-Germanic" (EPGmc) and "Late Proto-Germanic" (LPGmc). While Proto-Germanic refers only to the reconstruction of the most recent common ancestor of Germanic languages, the Germanic parent language refers to

235-555: A late stage. The early stage includes the stress fixation and resulting "spontaneous vowel-shifts" while the late stage is defined by ten complex rules governing changes of both vowels and consonants. By 250 BC Proto-Germanic had branched into five groups of Germanic: two each in the West and the North and one in the East. The following changes are known or presumed to have occurred in

282-633: A narrow belt along the Himalayas to cover mainland and island Southeast Asia as far as Sumatra , Java , Borneo , and Palawan . Finally, oaks of multiple sections ( Cyclobalanopsis , Ilex , Cerris , Quercus and related genera like Lithocarpus and Castanopsis ) extend across east Asia including China, Korea, and Japan. Potential records of Quercus have been reported from Late Cretaceous deposits in North America and East Asia. These are not considered definitive, as macrofossils older than

329-485: A range of flavours, colours, and aromas. The spongy bark of the cork oak is used to make traditional wine bottle corks. Almost a third of oak species are threatened with extinction due to climate change, invasive pests , and habitat loss . In culture, the oak tree is a symbol of strength and serves as the national tree of many countries. In Indo-European and related religions, the oak is associated with thunder gods . Individual oak trees of cultural significance include

376-508: A single oak tree produces both staminate ('male') flowers in the form of catkins , and small pistillate ('female') flowers, meaning that the trees are monoecious . The fruit is a nut called an acorn , borne in a cup-like structure known as a cupule ; each acorn usually contains one seed and takes 6–18 months to mature, depending on the species. The acorns and leaves contain tannic acid , which helps to guard against fungi and insects. There are some 500 extant species of oaks. Trees in

423-557: A small area of the west of Cuba ; in Mesoamerica it occurs mainly above 1000 metres. The genus crossed the isthmus of Panama when the northern and southern continents came together and is present as one species, Q. humboldtii , above 1000 metres in Colombia. The oaks of North America are of many sections ( Protobalanus , Lobatae , Ponticae , Quercus , and Virentes ) along with related genera such as Notholithocarpus . In

470-535: Is a hardwood tree or shrub in the genus Quercus of the beech family . They have spirally arranged leaves, often with lobed edges, and a nut called an acorn , borne within a cup. The genus is widely distributed in the Northern Hemisphere; it includes some 500 species, both deciduous and evergreen . Fossil oaks date back to the Middle Eocene . Molecular phylogeny shows that the genus

517-471: Is divided into Old World and New World clades , but many oak species hybridise freely, making the genus's history difficult to resolve. Ecologically, oaks are keystone species in habitats from Mediterranean semi-desert to subtropical rainforest . They live in association with many kinds of fungi including truffles . Oaks support more than 950 species of caterpillar , many kinds of gall wasp which form distinctive galls , roundish woody lumps such as

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564-452: Is from Old English ac (seen in placenames such as Acton , from ac + tun , "oak village" ), which in turn is from Proto-Germanic *aiks , "oak". Oaks are hardwood ( dicotyledonous ) trees, deciduous or evergreen, with spirally arranged leaves , often with lobate margins ; some have serrated leaves or entire leaves with smooth margins. Many deciduous species are marcescent , not dropping dead leaves until spring. In spring,

611-807: Is largely a matter of convention. The first coherent text recorded in a Germanic language is the Gothic Bible , written in the later fourth century in the East Germanic variety of the Thervingi Gothic Christians , who had escaped persecution by moving from Scythia to Moesia in 348. Early West Germanic text is available from the fifth century, beginning with the Frankish Bergakker runic inscription . The evolution of Proto-Germanic from its ancestral forms, beginning with its ancestor Proto-Indo-European , began with

658-413: Is termed Pre-Proto-Germanic . Whether it is to be included under a wider meaning of Proto-Germanic is a matter of usage. Winfred P. Lehmann regarded Jacob Grimm 's "First Germanic Sound Shift", or Grimm's law, and Verner's law , (which pertained mainly to consonants and were considered for many decades to have generated Proto-Germanic) as pre-Proto-Germanic and held that the "upper boundary" (that is,

705-695: Is the reconstructed proto-language of the Germanic branch of the Indo-European languages . Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic branches during the fifth century BC to fifth century AD: West Germanic , East Germanic and North Germanic . The latter of these remained in contact with the others over a considerable time, especially with the Ingvaeonic languages (including English ), which arose from West Germanic dialects, and had remained in contact with

752-564: Is the question of what specific tree, in the tree model of language evolution, best explains the paths of descent of all the members of a language family from a common language, or proto-language (at the root of the tree) to the attested languages (at the leaves of the tree). The Germanic languages form a tree with Proto-Germanic at its root that is a branch of the Indo-European tree, which in turn has Proto-Indo-European at its root. Borrowing of lexical items from contact languages makes

799-548: Is typical not of Germanic but Celtic languages. Another is * walhaz 'foreigner; Celt' from the Celtic tribal name Volcae with k → h and o → a . Other likely Celtic loans include * ambahtaz 'servant', * brunjǭ 'mailshirt', * gīslaz 'hostage', * īsarną 'iron', * lēkijaz 'healer', * laudą 'lead', * Rīnaz 'Rhine', and * tūnaz, tūną 'fortified enclosure'. These loans would likely have been borrowed during

846-677: The Corded Ware culture in the mid-3rd millennium BC, developing into the Nordic Bronze Age cultures by the early second millennium BC. According to Mallory, Germanicists "generally agree" that the Urheimat ('original homeland') of the Proto-Germanic language, the ancestral idiom of all attested Germanic dialects, was primarily situated in an area corresponding to the extent of the Jastorf culture . Early Germanic expansion in

893-503: The Funnelbeaker culture , but the sound change in the Germanic languages known as Grimm's law points to a non-substratic development away from other branches of Indo-European. Proto-Germanic itself was likely spoken after c. 500 BC, and Proto-Norse , from the second century AD and later, is still quite close to reconstructed Proto-Germanic, but other common innovations separating Germanic from Proto-Indo-European suggest

940-637: The Norse . A defining feature of Proto-Germanic is the completion of the process described by Grimm's law , a set of sound changes that occurred between its status as a dialect of Proto-Indo-European and its gradual divergence into a separate language. The end of the Common Germanic period is reached with the beginning of the Migration Period in the fourth century AD. The alternative term " Germanic parent language " may be used to include

987-643: The Paleogene , and possibly from before the Eocene are mostly poorly preserved without critical features for certain identification. Amongst the oldest unequivocal records of Quercus are pollen from Austria, dating to the Paleocene -Eocene boundary, around 55 million years ago. The oldest records of Quercus in North America are from Oregon , dating to the Middle Eocene, around 44 million years ago, with

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1034-681: The Pre-Roman Iron Age (fifth to first centuries BC) placed Proto-Germanic speakers in contact with the Continental Celtic La Tène horizon . A number of Celtic loanwords in Proto-Germanic have been identified. By the first century AD, Germanic expansion reached the Danube and the Upper Rhine in the south and the Germanic peoples first entered the historical record . At about the same time, extending east of

1081-790: The Royal Oak in Britain, the Charter Oak in the United States, and the Guernica Oak in the Basque Country . The generic name Quercus is Latin for "oak", derived from Proto-Indo-European *kwerkwu- , "oak", which is also the origin of the name " fir ", another important or sacred tree in Indo-European culture. The word "cork", for the bark of the cork oak , similarly derives from Quercus . The common name "oak"

1128-621: The Vistula ( Oksywie culture , Przeworsk culture ), Germanic speakers came into contact with early Slavic cultures, as reflected in early Germanic loans in Proto-Slavic . By the third century, Late Proto-Germanic speakers had expanded over significant distance, from the Rhine to the Dniepr spanning about 1,200 km (700 mi). The period marks the breakup of Late Proto-Germanic and

1175-436: The oak apple , and a large number of pests and diseases. Oak leaves and acorns contain enough tannin to be toxic to cattle, but pigs are able to digest them safely. Oak timber is strong and hard, and has found many uses in construction and furniture-making. The bark was traditionally used for tanning leather . Wine barrels are made of oak; these are used for aging alcoholic beverages such as sherry and whisky , giving them

1222-484: The "lower boundary" was the dropping of final -a or -e in unstressed syllables; for example, post-PIE * wóyd-e > Gothic wait , 'knows'. Elmer H. Antonsen agreed with Lehmann about the upper boundary but later found runic evidence that the -a was not dropped: ékwakraz … wraita , 'I, Wakraz, … wrote (this)'. He says: "We must therefore search for a new lower boundary for Proto-Germanic." Antonsen's own scheme divides Proto-Germanic into an early stage and

1269-556: The 2nd century CE, as well as the non-runic Negau helmet inscription, dated to the 2nd century BCE), and in Roman Empire -era transcriptions of individual words (notably in Tacitus ' Germania , c. AD 90 ). Proto-Germanic developed out of pre-Proto-Germanic during the Pre-Roman Iron Age of Northern Europe. According to the Germanic substrate hypothesis , it may have been influenced by non-Indo-European cultures, such as

1316-429: The Germanic subfamily exhibited a less treelike behaviour, as some of its characteristics were acquired from neighbours early in its evolution rather than from its direct ancestors. The internal diversification of West Germanic developed in an especially non-treelike manner. Proto-Germanic is generally agreed to have begun about 500 BC. Its hypothetical ancestor between the end of Proto-Indo-European and 500 BC

1363-706: The Old World, oaks of section Quercus extend across the whole of Europe including European Russia apart from the far north, and north Africa (north of the Sahara) from Morocco to Libya. In Mediterranean Europe, they are joined by oaks of the sections Cerris and Ilex , which extend across Turkey, the Middle East, Iran, Afghanistan and Pakistan , while section Ponticae is endemic to the western Caucasus in Turkey and Georgia . Oaks of section Cyclobalanopsis extend in

1410-489: The beginning of the (historiographically recorded) Germanic migrations . The earliest available complete sentences in a Germanic language are variably dated to the 2nd century AD, around 300 AD or the first century AD in runic inscriptions (such as the Tune Runestone ). The language of these sentences is known as Proto-Norse , although the delineation of Late Common Germanic from Proto-Norse at about that time

1457-498: The dates of borrowings and sound laws are not precisely known, it is not possible to use loans to establish absolute or calendar chronology. Most loans from Celtic appear to have been made before or during the Germanic Sound Shift . For instance, one specimen * rīks 'ruler' was borrowed from Celtic * rīxs 'king' (stem * rīg- ), with g → k . It is clearly not native because PIE * ē → ī

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1504-414: The development of a separate common way of speech among some geographically nearby speakers of a prior language and ended with the dispersion of the proto-language speakers into distinct populations with mostly independent speech habits. Between the two points, many sound changes occurred. Phylogeny as applied to historical linguistics involves the evolutionary descent of languages. The phylogeny problem

1551-432: The earlier boundary) was the fixing of the accent, or stress, on the root syllable of a word, typically on the first syllable. Proto-Indo-European had featured a moveable pitch-accent consisting of "an alternation of high and low tones" as well as stress of position determined by a set of rules based on the lengths of a word's syllables. The fixation of the stress led to sound changes in unstressed syllables. For Lehmann,

1598-480: The entire journey that the dialect of Proto-Indo-European that would become Proto-Germanic underwent through the millennia. The Proto-Germanic language is not directly attested by any complete surviving texts; it has been reconstructed using the comparative method . However, there is fragmentary direct attestation of (late) Proto-Germanic in early runic inscriptions (specifically the Vimose inscriptions , dated to

1645-496: The erosion of unstressed syllables, which would continue in its descendants. The final stage of the language included the remaining development until the breakup into dialects and, most notably, featured the development of nasal vowels and the start of umlaut , another characteristic Germanic feature. Loans into Proto-Germanic from other (known) languages or from Proto-Germanic into other languages can be dated relative to each other by which Germanic sound laws have acted on them. Since

1692-404: The genus Quercus consisted of Old World and New World clades. The entire genome of Quercus robur (the pedunculate oak) has been sequenced , revealing an array of mutations that may underlie the evolution of longevity and disease resistance in oaks. In addition, hundreds of oak species have been compared (at RAD-seq loci), allowing a detailed phylogeny to be constructed. However,

1739-641: The genus are often large and slow-growing; Q. alba can reach an age of 600 years, a diameter of 13 feet (4.0 m) and a height of 145 feet (44 m). The Granit oak in Bulgaria, a Q. robur specimen, has an estimated age of 1637 years, making it the oldest oak in Europe. The Wi'aaSal tree, a live oak in the reservation of the Pechanga Band of Indians , California, is at least 1000 years old, and might be as much as 2000 years old, which would make it

1786-1569: The high signal of introgressive hybridization (the transfer of genetic material by repeated backcrossing with hybrid offspring) in the genus has made it difficult to resolve an unambiguous, unitary history of oaks. The phylogeny from Hipp et al. 2019 is: CTB lineage [REDACTED] Cyclobalanoides Glauca [REDACTED] Acuta [REDACTED] Semiserrata East Asian Cerris [REDACTED] West Eurasian Cerris [REDACTED] Early-diverging Ilex [REDACTED] East Asian Ilex [REDACTED] Himalaya-Mediterranean [REDACTED] Himalayan subalpine [REDACTED] Agrifoliae [REDACTED] Palustres [REDACTED] Coccineae (Rubrae) [REDACTED] Phellos (Laurifoliae) [REDACTED] Texas red oaks [REDACTED] Erythromexicana [REDACTED] [REDACTED] [REDACTED] [REDACTED] Dumosae [REDACTED] Prinoids [REDACTED] Albae [REDACTED] Roburoids [REDACTED] Stellatae [REDACTED] Texas white oaks [REDACTED] Leucomexicana [REDACTED] Proto-Germanic language Caucasus East Asia Eastern Europe Northern Europe Pontic Steppe Northern/Eastern Steppe Europe South Asia Steppe Europe Caucasus India Indo-Aryans Iranians East Asia Europe East Asia Europe Indo-Aryan Iranian Indo-Aryan Iranian Others European Proto-Germanic (abbreviated PGmc ; also called Common Germanic )

1833-545: The history of Proto-Germanic in the wider sense from the end of Proto-Indo-European up to the point that Proto-Germanic began to break into mutually unintelligible dialects. The changes are listed roughly in chronological order, with changes that operate on the outcome of earlier ones appearing later in the list. The stages distinguished and the changes associated with each stage rely heavily on Ringe 2006 , Chapter 3, "The development of Proto-Germanic". Ringe in turn summarizes standard concepts and terminology. This stage began with

1880-519: The largest number of oak species, with approximately 160 species in Mexico, of which 109 are endemic, and about 90 in the United States. The second greatest area of oak diversity is China, with approximately 100 species. In the Americas, Quercus is widespread from Vancouver and Nova Scotia in the south of Canada, south to Mexico and across the whole of the eastern United States. It is present in

1927-474: The old tree, which had died, was cut down. The tree survived the extensive work carried out in 2002, when an underground stream, the Bins Brook , was exposed and an overflow basin was created, to prevent the flooding of nearby houses. 52°18′00″N 1°31′47″W  /  52.300062°N 1.529750°W  / 52.300062; -1.529750 Oak See also List of Quercus species . An oak

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1974-560: The oldest oak in the US. Among the smallest oaks is Q. acuta , the Japanese evergreen oak. It forms a bush or small tree to a height of some 30 feet (9.1 m). The genus Quercus is native to the Northern Hemisphere and includes deciduous and evergreen species extending from cool temperate to tropical latitudes in the Americas , Asia, Europe, and North Africa. North America has

2021-877: The oldest records in Asia from the Middle Eocene of Japan; both forms have affinities to the Cyclobalanopsis group. Quercus forms part, or rather two parts, of the Quercoideae subfamily of the Fagaceae , the beech family. Modern molecular phylogenetics suggests the following relationships: Fagus (beeches) Trigonobalanus (3 evergreen species) Lithocarpus (stone oaks) Chrysolepis (chinquapins) Quercus pro parte Notholithocarpus (tan oak) Quercus pro parte Castanopsis (also called chinquapins) Castanea (chestnuts) Molecular techniques for phylogenetic analysis show that

2068-413: The period marked the definitive break of Germanic from the other Indo-European languages and the beginning of Germanic proper, containing most of the sound changes that are now held to define this branch distinctively. This stage contained various consonant and vowel shifts, the loss of the contrastive accent inherited from PIE for a uniform accent on the first syllable of the word root, and the beginnings of

2115-552: The reduction of the resulting unstressed syllables. By this stage, Germanic had emerged as a distinctive branch and had undergone many of the sound changes that would make its later descendants recognisable as Germanic languages. It had shifted its consonant inventory from a system that was rich in plosives to one containing primarily fricatives, had lost the PIE mobile pitch accent for a predictable stress accent, and had merged two of its vowels. The stress accent had already begun to cause

2162-539: The relative position of the Germanic branch within Indo-European less clear than the positions of the other branches of Indo-European. In the course of the development of historical linguistics, various solutions have been proposed, none certain and all debatable. In the evolutionary history of a language family, philologists consider a genetic "tree model" appropriate only if communities do not remain in effective contact as their languages diverge. Early Indo-European had limited contact between distinct lineages, and, uniquely,

2209-511: The separation of a distinct speech, perhaps while it was still forming part of the Proto-Indo-European dialect continuum. It contained many innovations that were shared with other Indo-European branches to various degrees, probably through areal contacts, and mutual intelligibility with other dialects would have remained for some time. It was nevertheless on its own path, whether dialect or language. This stage began its evolution as

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