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124-490: Circa 425 spp., see text Anolis is a genus of anoles ( US : / ə ˈ n oʊ . l i z / ), iguanian lizards in the family Dactyloidae , native to the Americas . With more than 425 species, it represents the world's most species-rich amniote tetrapod genus, although many of these have been proposed to be moved to other genera, in which case only about 45 Anolis species remain. Previously, it

248-557: A species : see Botanical name and Specific name (zoology) . The rules for the scientific names of organisms are laid down in the nomenclature codes , which allow each species a single unique name that, for animals (including protists ), plants (also including algae and fungi ) and prokaryotes ( bacteria and archaea ), is Latin and binomial in form; this contrasts with common or vernacular names , which are non-standardized, can be non-unique, and typically also vary by country and language of usage. Except for viruses ,

372-527: A French fleet sailing towards Haiti forced General Christophe to withdraw from the east, leaving it in French hands. In 1808, a second revolution against France broke out on the island. Following Napoleon's invasion of Spain, the criollos of Santo Domingo revolted against French rule and, with the aid of the United Kingdom, returned Santo Domingo to Spanish control. France would never regain control of

496-477: A charter was passed in 1501, allowing the import of African slaves by Ferdinand and Isabel. The Spanish believed Africans would be more capable of performing physical labor. From 1519 to 1533, the indigenous uprising known as Enriquillo 's Revolt, after the Taíno cacique who led them, ensued, resulting from escaped African slaves on the island ( maroons ) possibly working with the Taíno people. Precious metals played

620-487: A devastating impact on both mortality and fertility of the Taíno population over the first quarter century. Colonial administrators and Dominican and Hieronymite friars observed that the search for gold and agrarian enslavement through the encomienda system were deciminating the indigenous population. Demographic data from two provinces in 1514 shows a low birth rate, consistent with a 3.5% annual population decline. In 1503, Spaniards began to bring enslaved Africans after

744-450: A different structure with larger rectangular walls and a porch. The Taíno village also had a flat court used for ball games and festivals. Religiously, the Taíno people were polytheists, and their gods were called Zemí. Religious worship and dancing were common, and medicine men or priests also consulted the Zemí for advice in public ceremonies. For food, the Taíno relied on meat and fish as

868-413: A diversity in the recognition of individuals. Sensory systems must be able to effectively pick up on dewlap signals, thus coevolving with the changes in dewlap characteristics. One of the main limitations to these theories is that of gene flow . Population genetic theory says that gene flow can counteract evolutionary adaptations made and prevent signal divergence, due to an influx of abnormal alleles into

992-590: A high payment for compensation to slaveholders who lost their property, and Haiti was saddled with unmanageable debt for decades. By this point, both states were united under Haitian control. However, suppression of the Dominican culture would lead to the Dominican War of Independence and the establishment of the Dominican Republic in 1844. (This is one of the reasons for the tensions between

1116-633: A large role in the history of the island after Columbus's arrival. One of the first inhabitants Columbus came across on this island was "a girl wearing only a gold nose plug". Soon the Taínos were trading pieces of gold for hawk's bells with their cacique declaring the gold came from Cibao . Traveling further east from Navidad, Columbus came across the Yaque del Norte River , which he named Río de Oro (River of Gold) because its "sands abound in gold dust". On Columbus's return during his second voyage, he learned it

1240-696: A largely hunter-gatherer lifestyle. During the 1st millennium BC , the Arawakan-speaking ancestors of the Taino people began to migrate into the Caribbean. Unlike the Archaic peoples, they practiced the intensive production of pottery and agriculture. The earliest evidence of the ancestors of the Taino people on Hispaniola is the Ostionoid culture, which dates to around 600 AD. The Taino represented

1364-651: A later homonym of a validly published name is a nomen illegitimum or nom. illeg. ; for a full list refer to the International Code of Nomenclature for algae, fungi, and plants and the work cited above by Hawksworth, 2010. In place of the "valid taxon" in zoology, the nearest equivalent in botany is " correct name " or "current name" which can, again, differ or change with alternative taxonomic treatments or new information that results in previously accepted genera being combined or split. Prokaryote and virus codes of nomenclature also exist which serve as

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1488-628: A long time and redescribed as new by a range of subsequent workers, or if a range of genera previously considered separate taxa have subsequently been consolidated into one. For example, the World Register of Marine Species presently lists 8 genus-level synonyms for the sperm whale genus Physeter Linnaeus, 1758, and 13 for the bivalve genus Pecten O.F. Müller, 1776. Within the same kingdom, one generic name can apply to one genus only. However, many names have been assigned (usually unintentionally) to two or more different genera. For example,

1612-576: A new colonial elite. The first major slave revolt in the Americas occurred in Santo Domingo during 1521, when enslaved Muslims of the Wolof nation led an uprising in the sugar plantation of admiral Don Diego Colon , son of Christopher Columbus . Many of these insurgents managed to escape where they formed independent maroon communities in the south of the island. Beginning in the 1520s,

1736-493: A primary source for protein. On the island they hunted small mammals, but also snakes, worms, and birds. In lakes and in the sea they were able to catch ducks and turtles. The Taíno also relied on agriculture as a primary food source. The indigenous people of Hispaniola raised crops in a conuco, which is a large mound packed with leaves and fixed crops to prevent erosion. Some common agricultural goods were cassava , maize, squash, beans, peppers, peanuts, cotton, and tobacco, which

1860-409: A reference for designating currently accepted genus names as opposed to others which may be either reduced to synonymy, or, in the case of prokaryotes, relegated to a status of "names without standing in prokaryotic nomenclature". An available (zoological) or validly published (botanical) name that has been historically applied to a genus but is not regarded as the accepted (current/valid) name for

1984-528: A stark decrease from the country's 60% forest cover in 1925. The country has been significantly deforested over the last 50 years, resulting in the desertification of many portions of Haitian territory. Haiti's poor citizens use cooking fires often, and this is a major culprit behind the nation's loss of trees. Haitians use trees as fuel either by burning the wood directly, or by first turning it into charcoal in ovens. Seventy-one percent of all fuel consumed in Haiti

2108-427: A taxon; however, the names published in suppressed works are made unavailable via the relevant Opinion dealing with the work in question. In botany, similar concepts exist but with different labels. The botanical equivalent of zoology's "available name" is a validly published name . An invalidly published name is a nomen invalidum or nom. inval. ; a rejected name is a nomen rejiciendum or nom. rej. ;

2232-497: A time when European demand for sugar was high. Slavery kept costs low and profit was maximized. It was an important port in the Americas for goods and products flowing to and from France and Europe. European colonists often died young due to tropical fevers, as well as from violent slave resistance in the late eighteenth century. In 1791, during the French Revolution , a major slave revolt broke out on Saint-Domingue. When

2356-455: A total of c. 520,000 published names (including synonyms) as at end 2019, increasing at some 2,500 published generic names per year. "Official" registers of taxon names at all ranks, including genera, exist for a few groups only such as viruses and prokaryotes, while for others there are compendia with no "official" standing such as Index Fungorum for fungi, Index Nominum Algarum and AlgaeBase for algae, Index Nominum Genericorum and

2480-404: A way to threaten other males in the area. During intermale fights, dewlaps are flashed. More currently, many studies have been done on the dewlap as a function for species recognition , with focus on the relationship of the contrast between dewlap color and environment. The dewlap comes in a variety of colors, including yellow, blue, and red. It was previously believed that the color of the dewlap

2604-672: Is a Hispanophone nation of approximately 11.3 million people. Spanish is spoken by essentially all Dominicans as a primary language. Roman Catholicism is the official and dominant religion and some Evangelicalism and Protestant churches and The Church of Jesus Christ and minority religions such as African religions, Afro-American religions, African diaspora religions , Haitian Vodou , Dominican Vodou , Dominican Santeria , Congos Del Espiritu Santo , Dominican Protestants , Pentecostals , Judaism , Islam and Baháʼí Faith , Hinduism , Buddhism , Unitarian Universalism , Jehovah's Witnesses , Pentecostalism and others also exist. Haiti

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2728-421: Is a Creole-speaking nation of roughly 11.7 million people. Although French is spoken as a primary language by the educated and wealthy minority, virtually the entire population speaks Haitian Creole , one of several French-derived creole languages. Roman Catholicism is the dominant religion , practiced by more than half the population, although in some cases in combination with Haitian Vodou faith. Another 25% of

2852-612: Is discouraged by both the International Code of Zoological Nomenclature and the International Code of Nomenclature for algae, fungi, and plants , there are some five thousand such names in use in more than one kingdom. For instance, A list of generic homonyms (with their authorities), including both available (validly published) and selected unavailable names, has been compiled by the Interim Register of Marine and Nonmarine Genera (IRMNG). The type genus forms

2976-403: Is generally little rainfall outside hurricane season from August to October, and droughts are by no means uncommon when hurricanes do not come. On the northern coast, in contrast, rainfall may peak between December and February, though some rain falls in all months of the year. Annual amounts typically range from 1,700 to 2,000 millimetres (67 to 79 in) on the northern coastal lowlands; there

3100-464: Is most present in male anoles. The dewlap is extended by means of the hyoid muscles in the throat, and can be flashed in a "pulse" pattern where the flap is extended repeatedly, or a "moving flag" pattern, where the lizard flashes it continuously while bobbing up and down. The coloration of the dewlap is caused by two pigments, pterins and carotenoids . Pterin pigments are compounds synthesized from guanine , whereas carotenoids are pigments acquired from

3224-518: Is much greater, around 3,100 millimetres (120 in) per year, but with a similar pattern to that observed in the central lowlands. The variations of temperature depend on altitude and are much less marked than rainfall variations in the island. Lowland Hispaniola is generally more hot and humid, with temperatures averaging 28 °C (82 °F). with high humidity during the daytime, and around 20 °C (68 °F) at night. At higher altitudes, temperatures fall steadily, so that frosts occur during

3348-481: Is no compelling evidence for an outbreak. The natives had no acquired immunity to European diseases, including smallpox . By May 1519, as many as one-third of the remaining Taínos had died. In the century following the Spanish arrival on Hispaniola, the Taíno population fell by up to 95% of the population, out of a pre-contact population estimated from tens of thousands to 8,000,000. Many authors have described

3472-489: Is probably much more in the Cordillera Septentrional , though no data exist. The interior of Hispaniola, along with the southeastern coast centered around Santo Domingo, typically receives around 1,400 millimetres (55 in) per year, with a distinct season from May to October. Usually, this wet season has two peaks: one around May, the other around the hurricane season. In the interior highlands, rainfall

3596-460: Is somewhat arbitrary. Although all species within a genus are supposed to be "similar", there are no objective criteria for grouping species into genera. There is much debate among zoologists about whether enormous, species-rich genera should be maintained, as it is extremely difficult to come up with identification keys or even character sets that distinguish all species. Hence, many taxonomists argue in favor of breaking down large genera. For instance,

3720-416: Is still heated debate over the population of Taíno people on the island of Hispaniola in 1492, but estimates range from no more than a few tens of thousands, according to a 2020 genetic analysis, to upwards of 750,000. A Taíno home consisted of a circular building with woven straw and palm leaves as covering. Most individuals slept in fashioned hammocks, but grass beds were also used. The cacique lived in

3844-474: Is the type species , and the generic name is permanently associated with the type specimen of its type species. Should the specimen turn out to be assignable to another genus, the generic name linked to it becomes a junior synonym and the remaining taxa in the former genus need to be reassessed. In zoological usage, taxonomic names, including those of genera, are classified as "available" or "unavailable". Available names are those published in accordance with

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3968-615: Is the most populous island in the West Indies , and the second-largest by land area , after Cuba . The 76,192-square-kilometre (29,418 sq mi) island is divided into two separate sovereign countries: the Spanish-speaking Dominican Republic (48,445 km (18,705 sq mi) to the east and the French and Haitian Creole –speaking Haiti (27,750 km (10,710 sq mi) to

4092-765: Is under the sovereignty of the Dominican Republic occupying the eastern portion and 27,750 square kilometers (10,710 sq mi) under the sovereignty of Haiti occupying the western portion. The island of Cuba lies 80 kilometers (50 mi) to the west across the Windward Passage; to the southwest lie Jamaica , separated by the Jamaica Channel , the Cayman Islands and Navassa Island ; 190 km (120 mi) . Puerto Rico lies 130 km (81 mi) east of Hispaniola across

4216-428: Is what mattered most in interlizard interactions, but it has since been found that there exists a relationship between habitat light conditions and dewlap color. This means that, rather than the color being of importance, it is the contrast of the dewlap against the background of its environment that best visually signals to other lizards. There have been many methods used to determine this. Persons et. al (1999) found that

4340-427: Is wood or charcoal. Haiti's government began establishing protected areas across the country in 1968. These 26 areas today represent nearly 7 per cent of the country's land and 1.5 per cent of its waters. In the Dominican Republic, the forest cover has increased. In 2003, the Dominican Republic's forest cover had been reduced to 32% of its land area, but by 2011, forest cover had increased to nearly 40%. The success of

4464-541: The Caribbean Sea was raided by increasingly numerous French pirates. In 1541, Spain authorized the construction of Santo Domingo's fortified wall, and in 1560 decided to restrict sea travel to enormous, well-armed convoys. In another move, which would destroy Hispaniola's sugar industry, in 1561 Havana , more strategically located in relation to the Gulf Stream , was selected as the designated stopping point for

4588-477: The French Republic abolished slavery in the colonies on February 4, 1794, it was a European first. The ex-slave army joined forces with France in its war against its European neighbors. In the second 1795 Treaty of Basel (July 22), Spain ceded the eastern two-thirds of the island of Hispaniola, later to become the Dominican Republic. French settlers had begun to colonize some areas in the Spanish side of

4712-621: The International Code of Zoological Nomenclature ; the earliest such name for any taxon (for example, a genus) should then be selected as the " valid " (i.e., current or accepted) name for the taxon in question. Consequently, there will be more available names than valid names at any point in time; which names are currently in use depending on the judgement of taxonomists in either combining taxa described under multiple names, or splitting taxa which may bring available names previously treated as synonyms back into use. "Unavailable" names in zoology comprise names that either were not published according to

4836-824: The International Plant Names Index for plants in general, and ferns through angiosperms, respectively, and Nomenclator Zoologicus and the Index to Organism Names for zoological names. Totals for both "all names" and estimates for "accepted names" as held in the Interim Register of Marine and Nonmarine Genera (IRMNG) are broken down further in the publication by Rees et al., 2020 cited above. The accepted names estimates are as follows, broken down by kingdom: The cited ranges of uncertainty arise because IRMNG lists "uncertain" names (not researched therein) in addition to known "accepted" names;

4960-580: The Mona Passage . The Bahamas and Turks and Caicos Islands lie to the north. Its westernmost point is known as Cap Carcasse. Cuba, Cayman Islands, Navassa Island, Hispaniola, Jamaica, and Puerto Rico are collectively known as the Greater Antilles . Hispaniola is also a part of the Antilles and the West Indies . The island has five major ranges of mountains: The Central Range, known in

5084-554: The Spanish Empire , serving as a headquarters for the further colonial expansion into the Americas . The colony was a meeting point of European explorers, soldiers, and settlers who brought with them the culture, architecture, laws, and traditions of the Old World . Spaniards imposed a harsh regime of forced labor and enslavement of the Taínos, as well as redirection of their food production and labor to Spaniards. This had

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5208-404: The platypus belongs to the genus Ornithorhynchus although George Shaw named it Platypus in 1799 (these two names are thus synonyms ) . However, the name Platypus had already been given to a group of ambrosia beetles by Johann Friedrich Wilhelm Herbst in 1793. A name that means two different things is a homonym . Since beetles and platypuses are both members of the kingdom Animalia,

5332-522: The rain shadow of the mountains in the southern and western portion of the island, and in the Cibao valley in the center-north of the island. The Hispaniolan pine forests occupy the mountainous 15% of the island, above 850 metres (2,790 ft) elevation. The flooded grasslands and savannas ecoregion in the south central region of the island surrounds a chain of lakes and lagoons, the most notable of which are Etang Saumatre and Trou Caïman in Haiti and

5456-425: The Caribbean shows the lizards have changed little in the millions of years. As ectotherms,  Anolis lizards must regulate their body temperature partly through behavioral changes and bask in the sunlight to gain enough heat to become fully active, but lizards cannot behaviorally warm themselves at night when temperatures drop. Because of this, cold tolerance evolves faster than heat tolerance in these lizards. On

5580-453: The Caribs. Christopher Columbus first landed at Hispaniola on December 6, 1492, at a small bay he named San Nicolas, now called Môle-Saint-Nicolas on the north coast of present-day Haiti. He was welcomed in a friendly fashion by the indigenous people known as the Taíno. Trading with the natives yielded more gold than they had come across previously on the other Caribbean islands and Columbus

5704-653: The Dominican Republic as the Cordillera Central , spans the central part of the island, extending from the south coast of the Dominican Republic into northwestern Haiti, where it is known as the Massif du Nord. This mountain range boasts the highest peak in the Antilles, Pico Duarte at 3,101 meters (10,174 ft) above sea level . The Cordillera Septentrional runs parallel to the Central Range across

5828-399: The Dominican Republic. The island has four distinct ecoregions . The Hispaniolan moist forests ecoregion covers approximately 50% of the island, especially the northern and eastern portions, predominantly in the lowlands but extending up to 2,100 meters (6,900 ft) elevation. The Hispaniolan dry forests ecoregion occupies approximately 20% of the island, lying in the rain shadow of

5952-530: The Dominican forest growth is due to several Dominican government policies and private organizations for the purpose of reforesting, and a strong educational campaign that has resulted in increased awareness by the Dominican people of the importance of forests for their welfare and other forms of life on the island. Hispaniola is the most populous Caribbean island with a combined population of 23 million inhabitants as of July 2023 . The Dominican Republic

6076-622: The Fort of Santo Tomás, present day Jánico , leaving Captain Pedro Margarit in command of 56 men. On 24 March 1495, Columbus with his ally Guacanagarix , embarked on a war of revenge against Caonabo, capturing him and his family while "killing many Indians and capturing others." Afterwards, "every person of fourteen years of age or upward was to pay a large hawk's bell of gold dust", every three months, as "the Spaniards were sure there

6200-420: The French botanist Joseph Pitton de Tournefort (1656–1708) is considered "the founder of the modern concept of genera". The scientific name (or the scientific epithet) of a genus is also called the generic name ; in modern style guides and science, it is always capitalised. It plays a fundamental role in binomial nomenclature , the system of naming organisms , where it is combined with the scientific name of

6324-606: The Italian Peter Martyr d'Anghiera , and the Spaniards Bartolomé de las Casas and Gonzalo Fernández de Oviedo . Based on a comprehensive survey and map prepared by Andrés de Morales in 1508, Martyr reported that the island as a whole was called Quizquella (or Quisqueya ) and Ayiti referred to a rugged mountainous region on the western end of the island. Diego Álvarez Chanca , a physician on Columbus's second voyage, also noted that “Ayiti” or Haïti

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6448-466: The Royal services to suppress their former buccaneer allies. In the 1697 Treaty of Ryswick , Spain formally ceded the western third of the island to France. Saint-Domingue quickly came to overshadow the east in both wealth and population. Nicknamed the "Pearl of the Antilles", it became the most prosperous colony in the West Indies , with a system of human slavery used to grow and harvest sugar cane during

6572-520: The Spanish re-captured Tortuga for the last time . In 1655 the island of Tortuga was reoccupied by the English and French. In 1660 the English appointed a Frenchman as Governor who proclaimed the King of France, set up French colours, and defeated several English attempts to reclaim the island. In 1665, French colonization of the island was officially recognized by King Louis XIV . The French colony

6696-486: The ability to adapt to different areas of the environment in a way where multiple species can coexist effectively. The amount of vegetation in an environment affects the amount of light absorbed. Studies have shown that lighting affects the dewlap’s function as a visual signal. The diversity in vegetation in Anolis lizards’ environments has caused a similar diversity in dewlap morphology, as different species of anoles adapt to

6820-412: The ability to stay submerged for as long as 18 minutes. The Anolis lizards that are less susceptible to predation are those with a dewlap in which both the scales and the exposed skin areas between them match the usual pale gray or whitish of the rest of the ventral surface. The dewlap is a flap of skin found beneath the jaw or throat of Anolis lizards. It can present in a variety of colorations, and

6944-442: The base for higher taxonomic ranks, such as the family name Canidae ("Canids") based on Canis . However, this does not typically ascend more than one or two levels: the order to which dogs and wolves belong is Carnivora ("Carnivores"). The numbers of either accepted, or all published genus names is not known precisely; Rees et al., 2020 estimate that approximately 310,000 accepted names (valid taxa) may exist, out of

7068-400: The clades are recognized as full genera, about 45 species remain in Anolis , with the remaining moved to Audantia (9 species), Chamaelinorops (7 species), Ctenonotus (more than 40 species), Dactyloa ( circa 95 species), Deiroptyx (almost 35 species), Norops (about 190 species), and Xiphosurus (around 15 species). Some of these can be further subdivided. For example, Phenacosaurus

7192-437: The diet. Both cause the red-yellow hues most commonly found in Anolis lizard dewlaps. The function of the dewlap in Anolis lizards has been a topic of debate for centuries. It is thought that the dewlap is flashed as a visual signal for other competing males, or as a courtship signal for single females. It has also been hypothesized that the dewlap serves as a signal for sex recognition. The first study done on dewlap function

7316-408: The different species occupied their various niches in the trees as a " community ". A study of lizard fossils trapped in amber shows that the lizard communities have existed for about 20 million years or more. Four modern ecomorph body types, trunk-crown, trunk-ground, trunk, and twig, are represented in the amber fossils study. Close comparison of the lizard fossils with their descendants alive today in

7440-495: The dominant group on the island during the period of European contact. Each society on the island was a small independent kingdom with a lead known as a cacique . In 1492, which is considered the peak of the Taíno, there were five different kingdoms on the island, the Xaragua, Higuey (Caizcimu), Magua (Huhabo), Ciguayos (Cayabo or Maguana), and Marien (Bainoa). Many distinct Taíno languages also existed in this time period. There

7564-626: The dry season on the highest peaks, where maxima are no higher than 18 °C (64 °F). There are many bird species in Hispaniola , and the island's amphibian species are also diverse. There are many species endemic to the island including insects and other invertebrates, reptiles, amphibians, fishs, birds and mammals (originally animals, native animals) and also (imported animals, introduced animals, not native animals or invasive species) just like farm animals, transport animals, house animals, pets and more. The two endemic terrestrial mammals on

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7688-525: The entire island of Hispaniola from the 1490s until the 17th century, when French pirates began establishing bases on the western side of the island. The official name was La Española , meaning "The Spanish (Island)". It was also called Santo Domingo , after Saint Dominic . The island was called various names by its native people, the Taíno . The Taino had no written language, hence, historical evidence for these names comes through three European historians:

7812-479: The flora in this naturally protected area consists of 621 species of vascular plants, of which 153 are highly endemic to La Hispaniola. The most prominent endemic species of flora that abound in the area are Ebano Verde (green ebony), Magnolia pallescens, a highly endangered hardwood. Recent in-depth studies of satellite imagery and environmental analysis regarding forest classification conclude that Haiti actually has approximately 30% tree cover; this is, nevertheless,

7936-486: The following year when Columbus brought 1,300 men to Hispaniola in November 1493 with the intention of establishing a permanent settlement. They found the encampment at Navidad had been destroyed and all the crewmen left behind killed by the natives. Columbus decided to sail east in search of a better site to found a new settlement. In January 1494 they established La Isabela in present-day Dominican Republic . In 1496,

8060-446: The form "author, year" in zoology, and "standard abbreviated author name" in botany. Thus in the examples above, the genus Canis would be cited in full as " Canis Linnaeus, 1758" (zoological usage), while Hibiscus , also first established by Linnaeus but in 1753, is simply " Hibiscus L." (botanical usage). Each genus should have a designated type , although in practice there is a backlog of older names without one. In zoology, this

8184-402: The four Greater Antillean Islands about 50 million years ago, Anolis lizards spread on each island to occupy niches in the island's trees. Some living in the tree canopy area, others low on the tree trunk near the ground; others in the mid-trunk area, others on twigs. Each new species developed its own distinct body type, called an ecomorph , adapted to the tree niche where it lived. Together,

8308-737: The generic name (or its abbreviated form) still forms the leading portion of the scientific name, for example, Canis lupus lupus for the Eurasian wolf subspecies, or as a botanical example, Hibiscus arnottianus ssp. immaculatus . Also, as visible in the above examples, the Latinised portions of the scientific names of genera and their included species (and infraspecies, where applicable) are, by convention, written in italics . The scientific names of virus species are descriptive, not binomial in form, and may or may not incorporate an indication of their containing genus; for example,

8432-470: The gold mines of Miguel Díaz and Francisco de Garay in 1504, as pit mines became royal mines for Ferdinand II of Aragon , who reserved the best mines for himself, though placers were open to private prospectors. King Ferdinand kept 967 natives in the San Cristóbal mining area, supervised by salaried miners. Under the royal governor Nicolás de Ovando , the indigenous people were forced to work in

8556-492: The gold mines. By 1503, the Spanish Crown legalized the allocation of private grants of indigenous labor to particular Spaniards for mining through the encomienda system. Once the indigenous were forced into mining far from their home villages, they suffered hunger and other difficult conditions. By 1508, the Taíno population of about 400,000 was reduced to 60,000, and by 1514, only 26,334 remained. About half resided in

8680-426: The government of Philip III ordered all inhabitants of Hispaniola to move close to Santo Domingo, to fight against piracy. Rather than secure the island, his action meant that French, English, and Dutch pirates established their own bases on the less populated north and west coasts of the island. In 1625, French and English pirates arrived on the island of Tortuga , just off the northwest coast of Hispaniola, which

8804-526: The highest point in Haiti, at 2,680 meters (8,790 ft) above sea level. A depression runs parallel to the southern range, between the southern range and the Chaîne des Matheux-Sierra de Neiba. It is known as the Plaine du Cul-de-Sac in Haiti, and Haiti's capital Port-au-Prince lies at its western end. The depression is home to a chain of salt lakes , including Lake Azuei in Haiti and Lake Enriquillo in

8928-432: The idea that a newly defined genus should fulfill these three criteria to be descriptively useful: Moreover, genera should be composed of phylogenetic units of the same kind as other (analogous) genera. The term "genus" comes from Latin genus , a noun form cognate with gignere ('to bear; to give birth to'). The Swedish taxonomist Carl Linnaeus popularized its use in his 1753 Species Plantarum , but

9052-466: The island are the Hispaniolan hutia ( Plagiodontia aedium ) and the Hispaniolan solenodon ( Solenodon paradoxus ). There are also many avian species on the island, with six endemic genera ( Calyptophilus , Dulus , Nesoctites , Phaenicophilus , Xenoligea and Microligea ). More than half of the original distribution of its ecoregions has been lost due to habitat destruction impacting

9176-457: The island of Hispaniola , both high-altitude and low-altitude lizard populations exist, and the thermal conditions at high and low elevations differ significantly. High-altitude lizards have shifted their ecological niche to boulder environments, where warming themselves is easier, and they show changes in the shape of limbs and skull that make them better adapted to these environments. To escape dangers, species that evolved near water have adapted

9300-671: The island, and after some 12 years of Spanish dominion, the leaders in Santo Domingo revolted again, and eastern Hispaniola was declared independent as the Republic of Spanish Haiti in 1821. Fearing the influence of a society of slaves that had successfully revolted against their owners, the United States and European powers refused to recognize Haiti, the second republic in the Western Hemisphere . France demanded

9424-633: The largest component, with 23,236 ± 5,379 accepted genus names, of which 20,845 ± 4,494 are angiosperms (superclass Angiospermae). By comparison, the 2018 annual edition of the Catalogue of Life (estimated >90% complete, for extant species in the main) contains currently 175,363 "accepted" genus names for 1,744,204 living and 59,284 extinct species, also including genus names only (no species) for some groups. The number of species in genera varies considerably among taxonomic groups. For instance, among (non-avian) reptiles , which have about 1180 genera,

9548-505: The largest gulf of the Antilles. The southern range begins in the southwesternmost Dominican Republic as the Sierra de Bahoruco , and extends west into Haiti as the Massif de la Selle and the Massif de la Hotte , which form the mountainous spine of Haiti's southern peninsula. Pic de la Selle is the highest peak in the southern range, the third highest peak in the Antilles and consequently

9672-432: The lighting conditions in their environment. The ability for a lizard to signal effectively also means it is able to defend its territory and attract mates more effectively, making it a good competitor. The relationship between background contrast and visual signals also suggests that there exists a coevolution between the signals and sensory systems of Anolis lizards. The environmental diversity of Anolis habitats causes

9796-459: The lizard genus Anolis has been suggested to be broken down into 8 or so different genera which would bring its ~400 species to smaller, more manageable subsets. Hispaniola Hispaniola ( / ˌ h ɪ s p ə n ˈ j oʊ l ə / , also UK : /- p æ n ˈ -/ ) is an island between Cuba and Puerto Rico in the Greater Antilles of the Caribbean . Hispaniola

9920-443: The local fauna and some of the original animals either threat, threatened with extinction or totally extinct, because of climate change or because they have been hunted by humans or their habitats have been felled or changed for some reasons or have become some of the animals have been threatened by (introduced animals, not native animals or invasive species) or there are fighting for space to survive and perhaps some animals that feed on

10044-734: The location within the vegetation where they forage (such as in the crown of trees vs. the trunk vs. underlying shrubs). These divergences in habitat are accompanied by morphological changes primarily related to moving on the substrate diameter they most frequently encounter, with twig ecomorphs having short limbs, while trunk ecomorphs have long limbs. In addition, these patterns repeat on numerous islands, with animals in similar habitats converging on similar body forms repeatedly. This demonstrates adaptive radiation can actually be predictable based on habitat encountered, and experimental introductions onto formerly lizard-free islands have proven Anolis evolution can be predicted. After appearing on each of

10168-467: The merchant flotas , which had a royal monopoly on commerce with the Americas. In 1564, the island's main inland cities Santiago de los Caballeros and Concepción de la Vega were destroyed by an earthquake. In the 1560s, English privateers joined the French in regularly raiding Spanish shipping in the Americas. By the early 17th century, Hispaniola and its nearby islands (notably Tortuga ) became regular stopping points for Caribbean pirates . In 1606,

10292-495: The mining towns of Concepción, Santiago, Santo Domingo, and Buenaventura. The repartimiento of 1514 accelerated emigration of the Spanish colonists, coupled with the exhaustion of the mines. The first documented outbreak of smallpox , previously an Eastern hemisphere disease, occurred on Hispaniola in December 1518 among enslaved African miners. Some scholars speculate that European diseases arrived before this date, but there

10416-403: The most (>300) have only 1 species, ~360 have between 2 and 4 species, 260 have 5–10 species, ~200 have 11–50 species, and only 27 genera have more than 50 species. However, some insect genera such as the bee genera Lasioglossum and Andrena have over 1000 species each. The largest flowering plant genus, Astragalus , contains over 3,000 species. Which species are assigned to a genus

10540-563: The most frequently used term in English-speaking countries for the island in scientific and cartographic works. In 1918, the United States occupation government, led by Harry Shepard Knapp , obliged the use of the name Hispaniola on the island, and recommended the use of that name to the National Geographic Society . The name "Haïti" was adopted by Haitian revolutionary Jean-Jacques Dessalines in 1804, as

10664-456: The mountains in the southern and western portion of the island and in the Cibao valley in the center-north of the island. The Hispaniolan pine forests occupy the mountainous 15% of the island, above 850 metres (2,790 ft) elevation. The flooded grasslands and savannas ecoregion in the south central region of the island surrounds a chain of lakes and lagoons in which the most notable include that of Lake Azuei and Trou Caïman in Haiti and

10788-428: The name could not be used for both. Johann Friedrich Blumenbach published the replacement name Ornithorhynchus in 1800. However, a genus in one kingdom is allowed to bear a scientific name that is in use as a generic name (or the name of a taxon in another rank) in a kingdom that is governed by a different nomenclature code. Names with the same form but applying to different taxa are called "homonyms". Although this

10912-473: The name to Española , and when Peter Martyr detailed his account of the island in Latin, he rendered its name as Hispaniola . Due to Taíno, Spanish and French influences on the island, historically the whole island was often referred to as Haïti , Hayti , Santo Domingo , or Saint-Domingue . Martyr's literary work was translated into English and French soon after being written, the name Hispaniola became

11036-613: The nearby Lake Enriquillo in the Dominican Republic, which is not only the lowest point of the island, but also the lowest point for an island country. Hispaniola's climate shows considerable variation due to its diverse mountainous topography, and is the most varied island of all the Antilles . Except in the Northern Hemisphere summer season, the predominant winds over Hispaniola are the northeast trade winds . As in Jamaica and Cuba, these winds deposit their moisture on

11160-494: The nearby Lake Enriquillo in the Dominican Republic. In Haiti , deforestation has long been cited by scientists as a source of ecological crisis; the timber industry dates back to French colonial rule. Haiti has seen a dramatic reduction of forests due to the excessive and increasing use of charcoal as fuel for cooking. Various media outlets have suggested that the country has just 2% forest cover , but this has not been substantiated by research. Also extremely important are

11284-445: The new population. This causes a genetic homogenization and challenges the idea that dewlap morphology in Anolis lizards and their sensory systems have coevolved. Genus The composition of a genus is determined by taxonomists . The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including

11408-805: The northern end of the Dominican Republic, extending into the Atlantic Ocean as the Samaná Peninsula . The Cordillera Central and Cordillera Septentrional are separated by the lowlands of the Cibao Valley and the Atlantic coastal plains, which extend westward into Haiti as the Plaine du Nord (Northern Plain). The lowest of the ranges is the Cordillera Oriental, in the eastern part of the country. The Sierra de Neiba rises in

11532-403: The northern mountains, and create a distinct rain shadow on the southern coast, where some areas receive as little as 400 millimetres (16 in) of rainfall, and have semi-arid climates . Annual rainfall under 600 millimetres (24 in) also occurs on the southern coast of Haiti's northwest peninsula and in the central Azúa region of the Plaine du Cul-de-Sac. In these regions, moreover, there

11656-551: The official name of independent Saint-Domingue, in tribute to the Amerindian predecessors. It was also adopted as the official name of independent Santo Domingo, as the Republic of Spanish Haiti , a state that existed from November 1821 until its annexation by Haiti in February 1822. The Archaic Age people arrived from mainland Central America or northern South America about 6,000 years ago, and are thought to have practised

11780-402: The probability of a dewlap showing being detected goes up with the contrast of dewlap against the background. They determined this by measuring the amount of times a "positive response" of the lizard's eyes turning towards a flashed dewlap occurred among different background contrasts. Similarly, Leal and Fleishman (2002) found that the light conditions in which a lizard displays its dewlap affects

11904-549: The probability of it being visually detected. They did this by measuring the UV spectral reflectance of dewlaps in Anolis cristatellus lizards using a spectroradiometer , then measuring the spectral sensitivity of the lizards’ retinal responses using electroretinographic (ERG) flicker photometry. Anolis lizards have emerged to be a good example of adaptive radiation . The difference in dewlap morphology among Anolis lizard populations demonstrates this phenomenon. Anolis lizards have

12028-541: The provisions of the ICZN Code, e.g., incorrect original or subsequent spellings, names published only in a thesis, and generic names published after 1930 with no type species indicated. According to "Glossary" section of the zoological Code, suppressed names (per published "Opinions" of the International Commission of Zoological Nomenclature) remain available but cannot be used as the valid name for

12152-400: The rarely mentioned species of Pinguicula casabitoana (a carnivorous plant), Gonocalyx tetraptera, Gesneria sylvicola, Lyonia alaini and Myrcia saliana, as well as palo de viento (Didymopanax tremulus), jaiqui (Bumelia salicifolia), pino criciolio) (pino criciol), sangre de pollo (Mecranium amigdalinum) and palo santo (Alpinia speciosa). According to reports in the Dominican Republic and Haiti,

12276-408: The same plants or animals or just something like that. The island has four distinct ecoregions . The Hispaniolan moist forests ecoregion covers approximately 50% of the island, especially the northern and eastern portions, predominantly in the lowlands but extending up to 2,100 meters (6,900 ft) elevation. The Hispaniolan dry forests ecoregion occupies approximately 20% of the island, lying in

12400-495: The same structural niche and have similarities in their size and shape, but they inhabit different climatic niches with was variability in temperature and openness of the environment. This suggests that thermal physiology is more associated with recently diverged anole lizards. Anolis lizards are some of the best examples of both adaptive radiation and convergent evolution . Populations of lizards on isolated islands diverge to occupy separate ecological niches , mostly in terms of

12524-719: The southwest of the Dominican Republic, and continues northwest into Haiti, parallel to the Cordillera Central, as the Montagnes Noires, Chaîne des Matheux and the Montagnes du Trou d'Eau. The Plateau Central lies between the Massif du Nord and the Montagnes Noires , and the Plaine de l'Artibonite lies between the Montagnes Noires and the Chaîne des Matheux, opening westward toward the Gulf of Gonâve ,

12648-497: The specific name particular to the wolf. A botanical example would be Hibiscus arnottianus , a particular species of the genus Hibiscus native to Hawaii. The specific name is written in lower-case and may be followed by subspecies names in zoology or a variety of infraspecific names in botany . When the generic name is already known from context, it may be shortened to its initial letter, for example, C. lupus in place of Canis lupus . Where species are further subdivided,

12772-412: The standard format for a species name comprises the generic name, indicating the genus to which the species belongs, followed by the specific epithet, which (within that genus) is unique to the species. For example, the gray wolf 's scientific name is Canis lupus , with Canis ( Latin for 'dog') being the generic name shared by the wolf's close relatives and lupus (Latin for 'wolf') being

12896-422: The surviving 7,000 troops in late 1803, the leaders of the revolution declared western Hispaniola the new nation of independent Haiti in early 1804. France continued to rule Spanish Santo Domingo. In 1805, Haitian troops of General Henri Christophe tried to conquer all of Hispaniola. They invaded Santo Domingo and sacked the towns of Santiago de los Caballeros and Moca, killing most of their residents, but news of

13020-403: The taxon is termed a synonym ; some authors also include unavailable names in lists of synonyms as well as available names, such as misspellings, names previously published without fulfilling all of the requirements of the relevant nomenclatural code, and rejected or suppressed names. A particular genus name may have zero to many synonyms, the latter case generally if the genus has been known for

13144-407: The territory. Under Napoleon , France reimposed slavery in most of its Caribbean islands in 1802 and sent an army to bring the island into full control. However, thousands of the French troops succumbed to yellow fever during the summer months, and more than half of the French army died because of disease. After an extremely brutal war with atrocities committed on both sides, the French removed

13268-531: The town of Nueva Isabela was founded. After being destroyed by a hurricane, it was rebuilt on the opposite side of the Ozama River and called Santo Domingo . It is the oldest permanent European settlement in the Americas. The island had an important role in the establishment of Latin American colonies for decades to come. Due to its strategic location, it was the military stronghold of conquistadors of

13392-633: The treatment of Tainos in Hispaniola under the Spanish Empire as genocide. Sugar cane was introduced to Hispaniola by settlers from the Canary Islands , and the first sugar mill in the New World was established in 1516, on Hispaniola. The need for a labor force to meet the growing demands of sugar cane cultivation led to an exponential increase in the importation of slaves over the following two decades. The sugar mill owners soon formed

13516-527: The two countries today). After a brief return to Spanish rule, the Dominican Republic would win its final independence in 1865. Haiti would become one of the poorest countries in the Americas, while the Dominican Republic gradually has developed into one of the largest economies of Central America and the Caribbean . Hispaniola is the second-largest island in the Caribbean (after Cuba), with an area of 76,192 square kilometers (29,418 sq mi), 48,440 square kilometers (18,700 sq mi) of which

13640-576: The values quoted are the mean of "accepted" names alone (all "uncertain" names treated as unaccepted) and "accepted + uncertain" names (all "uncertain" names treated as accepted), with the associated range of uncertainty indicating these two extremes. Within Animalia, the largest phylum is Arthropoda , with 151,697 ± 33,160 accepted genus names, of which 114,387 ± 27,654 are insects (class Insecta). Within Plantae, Tracheophyta (vascular plants) make up

13764-429: The virus species " Salmonid herpesvirus 1 ", " Salmonid herpesvirus 2 " and " Salmonid herpesvirus 3 " are all within the genus Salmonivirus ; however, the genus to which the species with the formal names " Everglades virus " and " Ross River virus " are assigned is Alphavirus . As with scientific names at other ranks, in all groups other than viruses, names of genera may be cited with their authorities, typically in

13888-642: The west. The only other divided island in the Caribbean is Saint Martin , which is shared between France ( Saint Martin ) and the Netherlands ( Sint Maarten ). Hispaniola is the site of one of the first European forts in the Americas, La Navidad (1492–1493), as well as the first settlement La Isabela (1493–1500), and the first permanent settlement, the current capital of the Dominican Republic, Santo Domingo (est. 1498). These settlements were founded successively during each of Christopher Columbus 's first three voyages . The Spanish Empire controlled

14012-402: Was by Mertens (1926). The initial assumption was that the dewlap was flashed as a method of sexual selection , and that the males would use it in order to attract females. It is hypothesized that female Anolis lizards are more attracted to males who flash their dewlaps more often, or have more brightly colored dewlaps. This was then challenged by the hypothesis that males flashed their dewlap as

14136-466: Was classified under the family Polychrotidae that contained all the anoles, as well as Polychrus , but recent studies place it in the Dactyloidae. This very large genus displays considerable paraphyly , but phylogenetic analysis suggests a number of subgroups or clades . Whether these clades are best recognized as subgenera within Anolis or separate genera remains a matter of dispute. If

14260-490: Was given the name Saint-Domingue . By 1670 a Welsh privateer named Henry Morgan invited the pirates on the island of Tortuga to set sail under him. They were hired by the French as a striking force that allowed France to have a much stronger hold on the Caribbean region. Consequently, the pirates never really controlled the island and kept Tortuga as a neutral hideout. The capital of the French Colony of Saint-Domingue

14384-460: Was led to believe that much more gold would be found inland. Before he could explore further, his flagship, the Santa Maria , ran aground and sank in the bay on December 24. With only two smaller ships remaining for the voyage home, Columbus built a fortified encampment, La Navidad , on the shore and left behind 21 crewman to await his return the following year. Colonization began in earnest

14508-581: Was made in the cordillera central, which led to a mining boom. By 1501 Columbus's cousin, Giovanni Colombo, had discovered gold near Buenaventura. The deposits were later known as Minas Nuevas. Two major mining areas resulted, one along San Cristobal -Buenaventura, and another in Cibao within the La Vega -Cotuy- Bonao triangle, while Santiago de los Caballeros , Concepción , and Bonao became mining towns. The gold rush of 1500–1508 ensued, and Ovando expropriated

14632-464: Was more gold in the island than the natives had yet found, and were determined to make them dig it out." Gold mining using forced indigenous labor began early on Hispaniola. Miguel Díaz and Francisco de Garay discovered large gold nuggets on the lower Haina River in 1496. These San Cristobal mines were later known as the Minas Viejas mines. Then, in 1499, the first major discovery of gold

14756-526: Was moved from Tortuga to Port-de-Paix on the mainland of Hispaniola in 1676. In 1680, new Acts of Parliament forbade sailing under foreign flags (in opposition to former practice). This was a major legal blow to the Caribbean pirates. Settlements were made in the Treaty of Ratisbon of 1684, signed by the European powers, that put an end to piracy. Most of the pirates after this time were hired out into

14880-507: Was often listed as a full genus in the past, but it is a subclade within Dactyloa ( Dactyloa heteroderma species group). Among the subgroups within Anolis are: In 2011, the green (or Carolina) anole ( Anolis carolinensis ) became the first reptile to have its complete genome published. Closely related, recently diverged anole lizards exhibited more divergence in thermal biology than in morphology. These anole lizards are thought to have

15004-401: Was originally settled by a few Spanish colonists. The pirates were attacked in 1629 by Spanish forces commanded by Don Fadrique de Toledo , who fortified the island, and expelled the French and English. As most of the Spanish army left for the main island of Hispaniola to root out French colonists there, the French returned to Tortuga in 1630 and had constant battles for several decades. In 1654,

15128-436: Was the chief Caonabo who had massacred his settlement at Navidad. While Columbus established a new settlement the village of La Isabela on Jan. 1494, he sent Alonso de Ojeda and 15 men to search for the mines of Cibao. After a six-day journey, Ojeda came across an area containing gold, in which the gold was extracted from streams by the Taíno people. Columbus himself visited the mines of Cibao on 12 March 1494. He constructed

15252-415: Was the easternmost province of the island, an area in the Dominican Republic called " Los Haitises " national park. On the other hand, Oviedo and Las Casas both recorded that the entire island was called Ayiti by the Taíno. When Columbus took possession of the island in 1492, he named it Insula Hispana in Latin and La Isla Española in Spanish, both meaning "the Spanish island". Las Casas shortened

15376-568: Was used as an aspect of social life and religious ceremonies. The Taíno people traveled often and used hollowed canoes with paddles when on the water for fishing or for migration purposes, and upwards of 100 people could fit into a single canoe. The Taíno came frequently in contact with the Caribs , another indigenous tribe. The Taíno people had to defend themselves using bows and arrows with poisoned tips and some war clubs. When Columbus landed on Hispaniola, many Taíno leaders wanted protection from

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