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77-679: In addition to monkey puzzle trees, the park is also home to coigüe, ñirre, oak, lenga, orchids, carnivorous plants. The park provides habitat for the mountain lion , the pudú , a small Chilean deer, and Darwin's fox . Among the birds are the Magellanic woodpecker , the Andean tapaculo and the chucao tapaculo . The park features 30 roads and 15 trails that can be explored by car or foot. The National Forest Corporation (Chile) information center and camping area are both found in Pehuenco, where

154-566: A 96.7% plant-based diet) and 2.57 (for Iceland , with 50% meat and fish, 50% plants). These values are comparable to those of non-apex predators such as the anchovy or pig . However, Peter D. Roopnarine criticized Bonhommeau's approach in 2014, arguing that humans are apex predators and that the HTL was based on terrestrial farming where indeed humans have a low trophic level, mainly eating producers (crop plants at level 1) or primary consumers (herbivores at level 2), which as expected places humans at

231-567: A complete picture. However, indirect evidence such as the absence of any discernible predator in an environment is suggestive. Anomalocaris was an aquatic apex predator, in the Cambrian. Its mouthparts are clearly predatory, and there were no larger animals in the seas at that time. Carnivorous theropod dinosaurs including Allosaurus and Tyrannosaurus are theorized to have been apex predators, based on their size, morphology, and dietary needs. A Permian shark, Triodus sessilis ,

308-454: A comprehensive literature review of more than 160 studies on cougar ecology, ecological interactions with 485 other species in cougar-inhabited ecosystems have been shown to involve different areas of interaction, ranging from the use of other species as food sources and prey, fear effects on potential prey, effects from carcass remains left behind, to competitive effects on other predator species in shared habitat. The most common research topic in

385-565: A female reaches estrous again, her offspring must disperse or the male will kill them. Males tend to disperse further than females. One study has shown a high mortality rate among cougars that travel farthest from their maternal range, often due to conflicts with other cougars. In a study area in New Mexico , males dispersed farther than females, traversed large expanses of non-cougar habitat and were probably most responsible for nuclear gene flow between habitat patches. Life expectancy in

462-766: A female's litter can have multiple paternities. Copulation is brief but frequent. Chronic stress can result in low reproductive rates in captivity as well as in the field. Gestation is 82–103 days long. Only females are involved in parenting. Litter size is between one and six cubs, typically two. Caves and other alcoves that offer protection are used as litter dens. Born blind, cubs are completely dependent on their mother at first and begin to be weaned at around three months of age. As they grow, they go out on forays with their mother, first visiting kill sites and, after six months, beginning to hunt small prey on their own. Kitten survival rates are just over one per litter. Juveniles remain with their mothers for one to two years. When

539-484: A foremost crepuscular and nocturnal activity pattern in a ranching area in southern Argentina. Home range sizes and overall cougar abundance depend on terrain, vegetation, and prey abundance. Research suggests a lower limit of 25 km (9.7 sq mi) and upper limit of 1,300 km (500 sq mi) of home range for males. Large male home ranges of 150 to 1,000 km (58 to 386 sq mi) with female ranges half that size. One female adjacent to

616-468: A kill to a preferred spot, covers it with brush, and returns to feed over a period of days. The cougar is generally reported to not be a scavenger , but deer carcasses left exposed for study were scavenged by cougars in California, suggesting more opportunistic behavior. Aside from humans, no species preys upon mature cougars in the wild, although conflicts with other predators or scavengers occur. Of

693-517: A large pack of seven to 11 wolves killing a female cougar and her kittens, while in nearby Sun Valley, Idaho , a 2-year-old male cougar was found dead, apparently killed by a wolf pack. Conversely, one-to-one confrontations tend to be dominated by the cat, and there are various documented accounts where wolves have been ambushed and killed, including adult male specimens. Wolves more broadly affect cougar population dynamics and distribution by dominating territory and prey opportunities, and disrupting

770-628: A level slightly above 2. Roopnarine instead calculated the position of humans in two marine ecosystems, a Caribbean coral reef and the Benguela system near South Africa. In these systems, humans mainly eat predatory fish and have a fractional trophic level of 4.65 and 4.5, respectively, which in Roopnarine's view makes those humans apex predators. In 2021, Miki Ben-Dor and colleagues compared human biology to that of animals at various trophic levels. Using metrics as diverse as tool use and acidity of

847-424: A powerful leap onto the back of its prey and a suffocating neck bite. The cougar can break the neck of some of its smaller prey with a strong bite and momentum bearing the animal to the ground. Kills are generally estimated around one large ungulate every two weeks. The period shrinks for females raising young, and may be as short as one kill every three days when cubs are nearly mature around 15 months. The cat drags

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924-481: A puma habituates to humans or is in a condition of severe starvation. Attacks are most frequent during late spring and summer when juvenile cougars leave their mothers and search for new territory. Apex predator An apex predator , also known as a top predator or superpredator , is a predator at the top of a food chain , without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics , meaning that they occupy

1001-531: A reduction in the population of sperm whales , apex predators with a fractional trophic level of 4.7, by hunting has caused an increase in the population of the large squid, with trophic level over 4 (carnivores that eat other carnivores). This effect, called mesopredator release , occurs in terrestrial and marine ecosystems; for instance, in North America, the ranges of all apex carnivores have contracted whereas those of 60% of mesopredators have grown in

1078-873: A tree. The cougar has the most extensive range of any wild land animal in the Americas, spanning 110 degrees of latitude from the Yukon in Canada to the southern Andes in Chile. The species was extirpated from eastern North America, aside from Florida, but they may be recolonizing their former range and isolated populations have been documented east of their contemporary ranges in both the Midwestern US and Canada. The cougar lives in all forest types, lowland and mountainous deserts, and in open areas with little vegetation up to an elevation of 5,800 m (19,000 ft). In

1155-614: Is a large cat native to the Americas. It inhabits North , Central and South America , making it the most widely distributed wild, terrestrial mammal in the Western Hemisphere , and one of the most widespread in the world. Its range spans the Yukon , British Columbia and Alberta provinces of Canada , the Rocky Mountains and areas in the western United States . Further south, its range extends through Mexico to

1232-467: Is a learned behavior and they do not generally recognize humans as prey. In a 10-year study in New Mexico of wild cougars who were not habituated to humans, the animals did not exhibit threatening behavior to researchers who approached closely (median distance=18.5 m; 61 feet) except in 6% of cases; 14 ⁄ 16 of those were females with cubs. Attacks on people, livestock, and pets may occur when

1309-473: Is a mostly solitary animal. Only mothers and kittens live in groups, with adults meeting rarely. While generally loners, cougars will reciprocally share kills and seem to organize themselves into small communities defined by the territories of dominant males. Cats within these areas socialize more frequently with each other than with outsiders. In the vicinity of a cattle ranch in northern Mexico, cougars exhibited nocturnal activity that overlapped foremost with

1386-413: Is a partial skull from the late Calabrian ( Ensenadan ) age. The head of the cougar is round, and the ears are erect. Its powerful forequarters, neck, and jaw serve to grasp and hold large prey. It has four retractile claws on its hind paws and five on its forepaws, of which one is a dewclaw . The larger front feet and claws are adaptations for clutching prey. Cougars are slender and agile members of

1463-559: Is afforded protection under the Endangered Species Act . The Texas Mountain Lion Conservation Project was launched in 2009 and aimed at raising local people's awareness of the status and ecological role of the cougar and mitigating conflict between landowners and cougars. The cougar is threatened by habitat loss , habitat fragmentation , and depletion of its prey base due to poaching . Hunting

1540-596: Is also sometimes used in the United States. The first use of puma in English dates to 1777, introduced from Spanish from the Quechua language . In the western United States and Canada, it is also called "mountain lion", a name first used in writing in 1858. Other names include "panther" (although it does not belong to the genus Panthera ) and "catamount" (meaning "cat of the mountains"). Felis concolor

1617-400: Is apparently correlated with the presence of other predators, prey species, livestock and humans. It is an ambush predator that pursues a wide variety of prey. Ungulates , particularly deer , are its primary prey, but it also hunts rodents . It is territorial and lives at low population densities. Individual home ranges depend on terrain, vegetation and abundance of prey. While large, it

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1694-560: Is believed to have originated in Asia about 11 million years ago ( Mya ). Taxonomic research on felids remains partial, and much of what is known about their evolutionary history is based on mitochondrial DNA analysis. Significant confidence intervals exist with suggested dates. In the latest genomic study of the Felidae, the common ancestor of today's Leopardus , Lynx , Puma , Prionailurus , and Felis lineages migrated across

1771-403: Is legal in the western United States. In Florida, heavy traffic causes frequent accidents involving cougars. Highways are a major barrier to the dispersal of cougars. The cougar populations in California are becoming fragmented with the increase in human population and infrastructure growth in the state. Human–wildlife conflict in proximity of 5 km (1.9 sq mi) of cougar habitat

1848-664: Is less. Whereas the size of cougars tends to increase as much as distance from the equator increases, which crosses the northern portion of South America, jaguars are generally smaller north of the Amazon River in South America and larger south of it. For example, while South American jaguars are comparatively large, and may exceed 90 kg (200 lb), North American jaguars in Mexico's Chamela-Cuixmala Biosphere Reserve weigh approximately 50 kg (110 lb), about

1925-565: Is lower in areas closer to the equator . A survey of North America research found 68% of prey items were ungulates, especially deer. Only the Florida panther showed variation, often preferring feral hogs and armadillos . Cougars have been known to prey on introduced gemsbok populations in New Mexico . One individual cougar was recorded as hunting 29 gemsbok, which made up 58% of its recorded kills. Most gemsbok kills were neonates, but some adults were also known to have been taken. Elsewhere in

2002-587: Is not always the dominant apex predator in its range, yielding prey to other predators. It is reclusive and mostly avoids people. Fatal attacks on humans are rare but increased in North America as more people entered cougar habitat and built farms. The cougar is listed as Least Concern on the IUCN Red List . Intensive hunting following European colonization of the Americas and ongoing human development into cougar habitat has caused populations to decline in most parts of its historical range. In particular,

2079-452: Is open to the public all year round. This La Araucanía Region location article is a stub . You can help Misplaced Pages by expanding it . This article related to a protected area in South America is a stub . You can help Misplaced Pages by expanding it . Mountain lion Also see text The cougar ( Puma concolor ) ( / ˈ k uː ɡ ər / , KOO-gər ), also known as the panther , mountain lion , catamount and puma ,

2156-507: Is pronounced in areas with a median human density of 32.48 inhabitants/km (84.1 inhabitants/sq mi) and a median livestock population density of 5.3 heads/km (14 heads/sq mi). Conflict is generally lower in areas more than 16.1 km (10.0 mi) away from roads and 27.8 km (17.3 mi) away from settlements. Due to the expanding human population , cougar ranges increasingly overlap with areas inhabited by humans. Attacks on humans are very rare, as cougar prey recognition

2233-827: Is shrinking the sea ice of the Arctic, forcing polar bears to fast on land for increasingly long periods. Dramatic changes in the Greater Yellowstone Ecosystem were recorded after the gray wolf , both an apex predator and a keystone species (one with a large effect on its ecosystem), was reintroduced to Yellowstone National Park in 1995 as a conservation measure. Elk , the wolves' primary prey, became less abundant and changed their behavior, freeing riparian zones from constant grazing and allowing willows , aspens , and cottonwoods to flourish, creating habitats for beaver , moose , and scores of other species. In addition to their effect on prey species,

2310-570: The Amazon Rainforest and the southern Andes Mountains in Patagonia . It is an adaptable generalist species , occurring in most American habitat types. It prefers habitats with dense underbrush and rocky areas for stalking but also lives in open areas. The cougar is largely solitary. Its activity pattern varies from diurnality and cathemerality to crepuscularity and nocturnality between protected and non-protected areas, and

2387-703: The Bering land bridge into the Americas 8.0 to 8.5 million years ago. The lineages subsequently diverged in that order. North American felids then invaded South America 2–4   Mya as part of the Great American Interchange , following the formation of the Isthmus of Panama . The cheetah lineage is suggested by some studies to have diverged from the Puma lineage in the Americas and migrated back to Asia and Africa, while other research suggests

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2464-473: The Cambrian period when animals such as Anomalocaris and Timorebestia dominated the seas. Humans have for many centuries interacted with other apex predators including the wolf , birds of prey , and cormorants to hunt game animals, birds, and fish respectively. More recently, humans have started interacting with apex predators in new ways. These include interactions via ecotourism , such as with

2541-598: The Cockscomb Basin of Belize were nocturnal but avoided each other. In a protected cloud forest in the central Andes of Colombia, cougars were active from late afternoon to shortly before sunrise and sometimes during noon and early afternoon. In protected areas of the Madidi - Tambopata Landscape in Bolivia and Peru, cougars were active throughout the day but with a tendency to nocturnal activity that overlapped with

2618-457: The Cordillera de Nahuelbuta . At sundown, you can enjoy the most sublime postcard view Nahuelbuta has to offer: a sky marked by red and pink hues framed by hundreds of thousand-year-old monkey puzzle trees. The administration of the park is located in Pehuenco, which is about 42 km from Angol . There are 10 camping sites here, with picnic tables, fire areas, rustic toilets and water. It

2695-460: The Felidae . They are the fourth largest cat species worldwide; adults stand about 60 to 90 cm (24 to 35 in) tall at the shoulders. Adult males are around 2.4 m (7 ft 10 in) long from nose to tail tip, and females average 2.05 m (6 ft 9 in), with overall ranges between 1.50 to 2.75 m (4 ft 11 in to 9 ft 0 in) nose to tail suggested for

2772-820: The San Andres Mountains was found with a big range of 215 km (83 sq mi), necessitated by poor prey abundance. Research has shown cougar abundances from 0.5 animals to as many as seven per 100 km (39 sq mi). Male home ranges include or overlap with females but, at least where studied, not with those of other males. The home ranges of females overlap slightly. Males create scrapes composed of leaves and duff with their hind feet, and mark them with urine and sometimes feces . When males encounter each other, they vocalize and may engage in violent conflict if neither backs down. Cougars communicate with various vocalizations. Aggressive sounds include growls, spits, snarls, and hisses. During

2849-884: The Santa Ana Mountains , it prefers steep canyons, escarpments, rim rocks and dense brush. In Mexico, it was recorded in the Sierra de San Carlos . In the Yucatán Peninsula , it inhabits secondary and semi- deciduous forests in El Eden Ecological Reserve . In El Salvador , it was recorded in the lower montane forest in Montecristo National Park and in a river basin in the Morazán Department above 700 m (2,300 ft) in 2019. In Colombia , it

2926-618: The eastern cougar population is considered to be mostly locally extinct in eastern North America since the early 20th century, with the exception of the isolated Florida panther subpopulation. The word cougar is borrowed from the Portuguese çuçuarana , via French; it was originally derived from the Tupi language . A current form in Brazil is suçuarana . In the 17th century, Georg Marcgrave named it cuguacu ara . Marcgrave's rendering

3003-476: The tiger shark , and through rewilding efforts, such as the proposed reintroduction of the Iberian lynx . Apex predators affect prey species' population dynamics and populations of other predators, both in aquatic and terrestrial ecosystems. Non-native predatory fish, for instance, have sometimes devastated formerly dominant predators. A lake manipulation study found that when the non-native smallmouth bass

3080-466: The United States. Establishing wildlife corridors and protecting sufficient range areas are critical for the sustainability of cougar populations. Research simulations showed that it faces a low extinction risk in areas larger than 2,200 km (850 sq mi). Between one and four new individuals entering a population per decade markedly increases persistence, thus highlighting the importance of habitat corridors. The Florida panther population

3157-552: The activity of calves . In a nature reserve in central Mexico, the activity of cougars was crepuscular and nocturnal, overlapping largely with the activity of the nine-banded armadillo ( Dasypus novemcinctus ). Cougars in the montane Abra-Tanchipa Biosphere Reserve in southeastern Mexico displayed a cathemeral activity pattern. Data from 12 years of camera trapping in the Pacific slope and Talamanca Cordillera of Costa Rica showed cougars as cathemeral. Both cougars and jaguars in

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3234-907: The activity of main prey species. During an 8-year-long study in a modified landscape in southeastern Brazil, male cougars were primarily nocturnal, but females were active at night and day. Cougars were diurnal in the Brazilian Pantanal , but crepuscular and nocturnal in protected areas in the Cerrado , Caatinga and ecotone biomes. Cougars in the Atlantic Forest were active throughout the day but displayed peak activity during early mornings in protected areas and crepuscular and nocturnal activity in less protected areas. In central Argentina, cougars were active day and night in protected areas but were active immediately after sunset and before sunrise outside protected areas. Cougars displayed

3311-736: The authors of Mammal Species of the World recognized the following six subspecies in 2005: In 2006, the Florida panther was still referred to as a distinct subspecies P. c. coryi in research works. As of 2017 , the Cat Classification Taskforce of the Cat Specialist Group recognizes only two subspecies as valid : Lynx Cheetah A. jubatus [REDACTED] Cougar [REDACTED] Jaguarundi H. yagouaroundi [REDACTED] Felis Otocolobus Prionailurus The family Felidae

3388-539: The average female in the same region averages about 42 kg (93 lb). On average, adult male cougars in British Columbia weigh 56.7 kg (125 lb) and adult females 45.4 kg (100 lb), though several male cougars in British Columbia weighed between 86.4 and 95.5 kg (190 and 211 lb). Depending on the locality, cougars can be smaller or bigger than jaguars but are less muscular and not as powerfully built, so on average, their weight

3465-434: The cats take prey more frequently and spend less time feeding on each kill. Unlike several subordinate predators from other ecosystems, cougars do not appear to exploit spatial or temporal refuges to avoid competitors. The gray wolf and the cougar compete more directly for prey, mostly in winter. Packs of wolves can steal cougars' kills, and there are some documented cases of cougars being killed by them. One report describes

3542-557: The cheetah diverged in the Old World itself. A high level of genetic similarity has been found among North American cougar populations, suggesting they are all fairly recent descendants of a small ancestral group. Culver et al. propose the original North American cougar population was extirpated during the Pleistocene extinctions some 10,000 years ago, when other large mammals, such as Smilodon , also disappeared. North America

3619-596: The consequences of both controlling prey density and restricting smaller predators, and may be capable of self-regulation. They are central to the functioning of ecosystems, the regulation of disease, and the maintenance of biodiversity. When introduced to subarctic islands, for example, Arctic foxes ' predation of seabirds has been shown to turn grassland into the tundra . Such wide-ranging effects on lower levels of an ecosystem are termed trophic cascades . The removal of top-level predators, often through human agency, can cause or disrupt trophic cascades. For example,

3696-511: The cougar diet. Learned, individual prey recognition was observed, as some cougars rarely killed bighorn sheep, while others relied heavily on the species. In the Central and South American cougar range area, the ratio of deer in the diet declines. Small to mid-sized mammals, including large rodents such as the capybara , are preferred. Ungulates accounted for only 35% of prey items in one survey, about half that of North America. Competition with

3773-431: The cougar, 32 cougar zoological specimens were described and proposed as subspecies until the late 1980s. Genetic analysis of cougar mitochondrial DNA indicates that many of these are too similar to be recognized as distinct at a molecular level but that only six phylogeographic groups exist. The Florida panther samples showed a low microsatellite variation, possibly due to inbreeding . Following this research,

3850-424: The cougar. The cougar has been listed as Least Concern on the IUCN Red List since 2008. However, it is also listed on CITES Appendix II . Hunting it is prohibited in California, Costa Rica , Honduras , Nicaragua , Guatemala , Panama , Venezuela , Colombia, French Guiana , Suriname , Bolivia , Brazil, Chile, Paraguay , Uruguay and most of Argentina. Hunting is regulated in Canada, Mexico, Peru , and

3927-548: The counties of Norfolk , Cumbria , and Northumberland in England, and Aberdeenshire in Scotland as part of the rewilding movement. The reintroduction of large predators is controversial, in part because of concern among farmers for their livestock . Conservationists such as Paul Lister propose instead to allow wolves and bears to hunt their prey in a "managed environment" on large fenced reserves; however, this undermines

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4004-659: The feline's behavior. Preliminary research in Yellowstone , for instance, has shown displacement of the cougar by wolves. One researcher in Oregon noted: "When there is a pack around, cougars are not comfortable around their kills or raising kittens [...] A lot of times a big cougar will kill a wolf, but the pack phenomenon changes the table." Both species are capable of killing mid-sized predators, such as bobcats , Canada lynxes , wolverines and coyotes , and tend to suppress their numbers. Although cougars can kill coyotes,

4081-971: The form of wolves, and in turn with domestic dogs , for 40,000 years; this collaboration may have helped modern humans to outcompete the Neanderthals . Humans still hunt with dogs , which have often been bred as gun dogs to point to , flush out , or retrieve prey . The Portuguese Water Dog was used to drive fish into nets. Several breeds of dog have been used to chase large prey such as deer and wolves. Eagles and falcons , which are apex predators, are used in falconry , hunting birds or mammals. Tethered cormorants , also top predators, have been used to catch fish . Ecotourism sometimes relies on apex predators to attract business. Tour operators may in consequence decide to intervene in ecosystems, for example by providing food to attract predators to areas that can conveniently be visited. This in turn can have effects on predator population and therefore on

4158-403: The highest trophic levels . Food chains are often far shorter on land, usually limited to being secondary consumers – for example, wolves prey mostly upon large herbivores (primary consumers), which eat plants (primary producers). The apex predator concept is applied in wildlife management , conservation , and ecotourism . Apex predators have a long evolutionary history, dating at least to

4235-863: The large predators in Yellowstone National Park – the grizzly and black bears , gray wolf and cougar – the massive grizzly bear appears dominant, often (though not always) able to drive a gray wolf pack, black bear or cougar off their kills. One study found that grizzlies and American black bears visited 24% of cougar kills in Yellowstone and Glacier National Parks , usurping 10% of carcasses. Bears gained up to 113%, and cougars lost up to 26% of their daily energy requirements from these encounters. In Colorado and California, black bears were found to visit 48% and 77% of kills, respectively. In general, cougars are subordinate to black bears when it comes to killing, and when bears are most active,

4312-429: The larger jaguar in South America has been suggested for the decline in the size of prey items. In Central or North America, the cougar and jaguar share the same prey, depending on its abundance. Other listed prey species of the cougar include mice , porcupines , American beavers , raccoons , hares , guanacoes , peccaries , vicuñas , rheas and wild turkeys . Birds and small reptiles are sometimes preyed upon in

4389-429: The latter have been documented attempting to prey on cougar cubs. The cougar and jaguar share overlapping territory in the southern portion of its range. The jaguar tends to take the larger prey where ranges overlap, reducing both the cougar's potential size and the likelihood of direct competition between the two cats. Cougars appear better than jaguars at exploiting a broader prey niche and smaller prey. The cougar

4466-464: The literature used here was the cougar's diet and its prey's regulation. The cougar is a generalist hypercarnivore . It prefers large mammals such as mule deer , white-tailed deer , elk , moose , mountain goat and bighorn sheep . It opportunistically takes smaller prey such as rodents , lagomorphs , smaller carnivores, birds, and even domestic animals, including pets. The mean weight of cougar vertebrate prey increases with its body weight and

4543-442: The mating season, estrus females produce caterwauls or yowls to attract mates, and males respond with similar vocals. Mothers and offspring keep in contact with whistles, chirps, and mews. Females reach sexual maturity at the age of 18 months to three years and are in estrus for about eight days of a 23-day cycle; the gestation period is approximately 91 days. Both adult males and females may mate with multiple partners, and

4620-474: The most popular route begins. Following it will treat you to over four kilometers of scenic paths before ending at Cerro Piedra del Águila (1,379 meters). The peak offers a terrific view of the park's exuberant and pristine natural wonders, the immensity of the Pacific Ocean to the west, and an impressive chain of Andean volcanoes to the east. Other peaks include Cerro Anay and Alto Nahuelbuta, part of

4697-496: The park. Ecologists have debated whether humans are apex predators. For instance, Sylvain Bonhommeau and colleagues argued in 2013 that across the global food web, a fractional human trophic level (HTL) can be calculated as the mean trophic level of every species in the human diet, weighted by the proportion that that species forms in the diet. This analysis gives an average HTL of 2.21, varying between 2.04 (for Burundi , with

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4774-466: The past two centuries. Because apex predators have powerful effects on other predators, herbivores, and plants, they can be important in nature conservation. Humans have hunted many apex predators close to extinction, but in some parts of the world, these predators are now returning. They are increasingly threatened by climate change . For example, the polar bear requires extensive areas of sea ice to hunt its prey, typically seals, but climate change

4851-537: The same as female cougars. Cougar coloring is plain (hence the Latin concolor ["one color"] in the scientific name) but can vary greatly across individuals and even siblings. The coat is typically tawny, but it ranges from silvery-grey to reddish with lighter patches on the underbody, including the jaws, chin, and throat. Infants are spotted and born with blue eyes and rings on their tails; juveniles are pale, and dark spots remain on their flanks. A leucistic individual

4928-431: The south, but this is rarely recorded in North America. Magellanic penguins ( Spheniscus magellanicus ) constitute the majority of prey items in cougar diet in Patagonia 's Bosques Petrificados de Jaramillo National Park and Monte León National Park . Although capable of sprinting, the cougar is typically an ambush predator . It stalks through brush and trees, across ledges, or other covered spots, before delivering

5005-666: The southwestern United States, they have been recorded to also prey on feral horses in the Great Basin , as well as feral donkeys in the Sonoran and Mojave Deserts . Investigations at Yellowstone National Park showed that elk and mule deer were the cougar's primary prey; the prey base is shared with the park's wolves , with which the cougar competes for resources. A study on winter kills from November to April in Alberta showed that ungulates accounted for greater than 99% of

5082-604: The species in general. Of this length, the tail typically accounts for 63 to 95 cm (25 to 37 in). Males generally weigh 53 to 72 kg (117 to 159 lb). Females typically weigh between 34 and 48 kg (75 and 106 lb). Cougar size is smallest close to the equator and larger towards the poles . The largest recorded cougar, shot in 1901, weighed 105.2 kg (232 lb); claims of 125.2 kg (276 lb) and 118 kg (260 lb) have been reported, though they were probably exaggerated. Male cougars in North America average 62 kg (137 lb), while

5159-507: The stomach, they concluded that humans evolved as apex predators, diversifying their diets in response to the disappearance of most of the megafauna that had once been their primary source of food. Apex predators are thought to have existed since at least the Cambrian period, around 500 million years ago. Extinct species cannot be directly determined to be apex predators as their behavior cannot be observed, and clues to ecological relationships, such as bite marks on bones or shells, do not form

5236-576: The wider ecosystem. As a result, provisioning of species such as the tiger shark is controversial, but its effects are not well established by empirical evidence. Other affected apex predators include big cats and crocodiles. In some densely populated areas like the British Isles , all the large native predators like the wolf, bear, wolverine and lynx have become extirpated , allowing herbivores such as deer to multiply unchecked except by hunting. In 2015, plans were made to reintroduce lynx to

5313-405: The wild is reported at 8 to 13 years and probably averages 8 to 10; a female of at least 18 years was reported killed by hunters on Vancouver Island . Cougars may live as long as 20 years in captivity. Causes of death in the wild include disability and disease, competition with other cougars, starvation, accidents, and, where allowed, hunting. The feline immunodeficiency virus is well-adapted to

5390-562: The wolves' presence also affected one of the park's vulnerable species , the grizzly bear : emerging from hibernation , having fasted for months, the bears chose to scavenge wolf kills, especially during the autumn as they prepared to hibernate once again. The grizzly bear gives birth during hibernation, so the increased food supply is expected to produce an increase in the number of cubs observed. Dozens of other species, including eagles, ravens, magpies , coyotes, and black bears have also been documented as scavenging from wolf kills within

5467-457: Was discovered containing two amphibians ( Archegosaurus decheni and Cheliderpeton latirostre ), one of which had consumed a fish, Acanthodes bronni , showing that the shark had lived at a trophic level of at least 4. Among more recent fossils, the saber-tooth cats, like Smilodon , are considered to have been apex predators in the Cenozoic . Humans hunted with apex predators in

5544-614: Was recorded in a palm oil plantation close to a riparian forest in the Llanos Basin , and close to water bodies in the Magdalena River Valley . In the human-modified landscape of central Argentina, it inhabits bushland with abundant vegetation cover and prey species. Cougars are an important keystone species in Western Hemisphere ecosystems, linking numerous species at many trophic levels. In

5621-647: Was removed, lake trout , the suppressed native apex predator, diversified its prey selection and increased its trophic level . As a terrestrial example, the badger , an apex predator, preys upon and also competes with the hedgehog , a mesopredator , for food such as insects, small mammals, reptiles, amphibians, and the eggs of ground-nesting birds. Removal of badgers (in a trial investigating bovine tuberculosis ) caused hedgehog densities to more than double. Predators that exert top-down control on organisms in their community are often considered keystone species . Apex predators can have profound effects on ecosystems, as

5698-579: Was reproduced in 1648 by his associate Willem Piso . Cuguacu ara was then adopted by John Ray in 1693. In 1774, Georges-Louis Leclerc, Comte de Buffon converted cuguacu ara to cuguar , which was later modified to "cougar" in English. The cougar holds the Guinness record for the animal with the greatest number of names, with over 40 in English alone. "Puma" is the common name used in Latin America and most parts of Europe. The term puma

5775-521: Was seen in Serra dos Órgãos National Park in Rio de Janeiro in 2013 when it was recorded by a camera trap, indicating that pure white individuals do exist within the species, though they are extremely rare. The cougar has large paws and proportionally the largest hind legs in the Felidae, allowing for great leaping and powerful short sprints. It can leap from the ground up to 5.5 m (18 ft) high into

5852-471: Was the scientific name proposed by Carl Linnaeus in 1771 for a cat with a long tail from Brazil. The specific epithet of the name, "concolor", is Latin for "of uniform color". It was placed in the genus Puma by William Jardine in 1834. This genus is part of the Felinae . The cougar is most closely related to the jaguarundi and the cheetah. Following Linnaeus's first scientific description of

5929-559: Was then repopulated by South American cougars . A coprolite identified as from a cougar was excavated in Argentina's Catamarca Province and dated to 17,002–16,573 years old. It contained Toxascaris leonina eggs. This finding indicates that the cougar and the parasite have existed in South America since at least the Late Pleistocene . The oldest fossil record of a cougar ( Puma concolor ) in South America (Argentina)

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