Satawal is a solitary coral atoll of one island with about 500 people on just over 1 km located in the Caroline Islands in the Pacific Ocean . It forms a legislative district in Yap State in the Federated States of Micronesia . Satawal is the easternmost island in the Yap island group and is located approximately 70 kilometers (43 mi) east of Lamotrek .
112-470: The island, which measures 2 kilometers (1.2 mi) long northeast-southwest, is up to 0.8 kilometers (0.50 mi) wide and sits atop a small platform-like reef with a narrow fringing reef. The total land area is 1.3 square kilometers (0.50 sq mi), and is thickly wooded with coconut and breadfruit trees. As there are no anchorages for large boats, Satawal is seldom visited by outsiders. Administratively Piagailoe Atoll , located 71 kilometers to
224-430: A fiber , called coir , which has many traditional and commercial uses. Both the exocarp and the mesocarp make up the "husk" of the coconut, while the endocarp makes up the hard coconut "shell". The endocarp is around 4 millimetres ( 1 ⁄ 8 inch) thick and has three distinctive germination pores ( micropyles ) on the distal end. Two of the pores are plugged (the "eyes"), while one is functional. The interior of
336-407: A clustering habit results. Exclusively sympodial genera include many of the rattans , Guihaia , and Rhapis . Several palm genera have both solitary and clustering members. Palms which are usually solitary may grow in clusters and vice versa. Palms have large, evergreen leaves that are either palmately ('fan-leaved') or pinnately ('feather-leaved') compound and spirally arranged at the top of
448-463: A large amount of the human diet and several other human uses, both by absolute amount produced and by number of species domesticated . This is far higher than almost any other plant family, sixth out of domesticated crops in the human diet, and first in total economic value produced – sharing the top spot with the Poaceae and Fabaceae . These human uses have also spread many Arecaceae species around
560-500: A shorter postharvest storage life of around two to three weeks at temperatures of 12 to 15 °C (54 to 59 °F) or up to 2 months at 0 to 1.5 °C (32.0 to 34.7 °F). In comparison, mature coconuts with the husk intact can be stored for three to five months at normal room temperature . Unlike some other plants, the palm tree has neither a taproot nor root hairs , but has a fibrous root system . The root system consists of an abundance of thin roots that grow outward from
672-409: A single domesticated population. Philippine and Malayan dwarf coconuts diverged early into two distinct types. They usually remain genetically isolated when introduced to new regions, making it possible to trace their origins. Numerous other dwarf cultivars also developed as the initial dwarf cultivar was introduced to other regions and hybridized with various tall cultivars. The origin of dwarf varieties
784-663: A species that had started to be dispersed by humans and grown in plantations. Niu vai endocarp fragments have been recovered in archaeological sites in the St. Matthias Islands of the Bismarck Archipelago . The fragments are dated to approximately 1000 BCE, suggesting that cultivation and artificial selection of coconuts were already practiced by the Austronesian Lapita people . Coconuts can also be broadly divided into two general types based on habit:
896-695: A thinner husk and a larger amount of endosperm. Domesticated coconuts also contain more coconut water . These two forms are referred to by the Samoan terms niu kafa for the elongated wild coconuts, and niu vai for the rounded domesticated Pacific coconuts. A full-sized coconut fruit weighs about 1.4 kilograms (3 pounds 1 ounce). Coconuts sold domestically in coconut-producing countries are typically not de-husked. Especially immature coconuts (6 to 8 months from flowering) are sold for coconut water and softer jelly-like coconut meat (known as "green coconuts", "young coconuts", or "water coconuts"), where
1008-637: A variety of different habitats. Their diversity is highest in wet, lowland forests. South America , the Caribbean , and areas of the South Pacific and southern Asia are regions of concentration. Colombia may have the highest number of palm species in one country. There are some palms that are also native to desert areas such as the Arabian Peninsula and parts of northwestern Mexico. Only about 130 palm species naturally grow entirely beyond
1120-691: A variety of products for furnishing and decoration. The coconut has cultural and religious significance in certain societies, particularly in the Austronesian cultures of the Western Pacific where it is featured in their mythologies, songs, and oral traditions. The fall of its mature fruit has led to a preoccupation with death by coconut . It also had ceremonial importance in pre-colonial animistic religions. It has also acquired religious significance in South Asian cultures, where it
1232-609: A water source, and in desert oases such as the date palm . A few palms are adapted to extremely basic lime soils, while others are similarly adapted to extreme potassium deficiency and toxicity of heavy metals in serpentine soils . Palms are a monophyletic group of plants, meaning the group consists of a common ancestor and all its descendants. Extensive taxonomic research on palms began with botanist H.E. Moore , who organized palms into 15 major groups based mostly on general morphological characteristics. The following classification, proposed by N.W. Uhl and J. Dransfield in 1987,
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#17327754540641344-425: Is Southeast Asia , which contain the tall cultivars that are genetically closest to dwarf coconuts. Sequencing of the genome of the tall and dwarf varieties revealed that they diverged 2 to 8 million years ago and that the dwarf variety arose through alterations in genes involved in the metabolism of the plant hormone gibberellin . Another ancestral variety is the niu leka of Polynesia (sometimes called
1456-508: Is a revision of Moore's classification that organizes palms into 6 subfamilies. A few general traits of each subfamily are listed below. The Phytelephantoideae is the sixth subfamily of Arecaceae in N.W. Uhl and J. Dransfield's 1987 classification. Members of this group have distinct monopodial flower clusters. Other distinct features include a gynoecium with five to 10 joined carpels, and flowers with more than three parts per whorl. Fruits are multiple-seeded and have multiple parts. From
1568-509: Is also cited as evidence that the plant originated in the region. For example, the Polynesian and Melanesian term niu ; Tagalog and Chamorro term niyog ; and the Malay word nyiur or nyior . Other evidence for a Central Indo-Pacific origin is the native range of the coconut crab ; and the higher amounts of C. nucifera -specific insect pests in the region (90%) in comparison to
1680-562: Is also often stated that coconuts can travel 110 days, or 5,000 km (3,000 mi), by sea and still be able to germinate. This figure has been questioned based on the extremely small sample size that forms the basis of the paper that makes this claim. Thor Heyerdahl provides an alternative, and much shorter, estimate based on his first-hand experience crossing the Pacific Ocean on the raft Kon-Tiki : The nuts we had in baskets on deck remained edible and capable of germinating
1792-421: Is consistently warm and humid. Coconut palms require warm conditions for successful growth, and are intolerant of cold weather. Some seasonal variation is tolerated, with good growth where mean summer temperatures are between 28 and 37 °C (82 and 99 °F), and survival as long as winter temperatures are above 4–12 °C (39–54 °F); they will survive brief drops to 0 °C (32 °F). Severe frost
1904-556: Is documented in the Steve Thomas book The Last Navigator , which also treats Mau Piailug's traditional navigation system in some depth. Sanford J. Low produced The Navigators: Pathfinders of the Pacific , a documentary film about Mau Piailug and communal life on Satawal including food preparation, fishing and boat building. On March 18, 2007, Piailug presided over the first Pwo ceremony for navigators on Satawal in 56 years. At
2016-605: Is from Dravidian *aṭ-ay-kkāy ("areca nut"). The suffix -aceae is the feminine plural of the Latin -āceus ("resembling"). Palm originates from Latin palma semantically overlapping with sense of " hand front " (due to similar splayed shape) ultimately from Proto-Indo-European * pl̥h₂meh₂ , a direct descendant folm once existed in Old English. Whether as shrubs, tree-like, or vines, palms have two methods of growth: solitary or clustered. The common representation
2128-577: Is further strengthened by other similar botanical evidence of contact, like the pre-colonial presence of sweet potato in Oceanian cultures. During the colonial era , Pacific coconuts were further introduced to Mexico from the Spanish East Indies via the Manila galleons . In contrast to the Pacific coconuts, Indo-Atlantic coconuts were largely spread by Arab and Persian traders into
2240-617: Is highly tolerant of salinity . It prefers areas with abundant sunlight and regular rainfall (1,500–2,500 mm [59–98 in] annually), which makes colonizing shorelines of the tropics relatively straightforward. Coconuts also need high humidity (at least 70–80%) for optimum growth, which is why they are rarely seen in areas with low humidity. However, they can be found in humid areas with low annual precipitation such as in Karachi , Pakistan , which receives only about 250 mm ( 9 + 3 ⁄ 4 in) of rainfall per year, but
2352-587: Is more typical. Plants are intolerant to cold and prefer copious precipitation and full sunlight. Many insect pests and diseases affect the species and are a nuisance for commercial production. In 2022, about 73% of the world's supply of coconuts was produced by Indonesia , India , and the Philippines . Cocos nucifera is a large palm, growing up to 30 metres (100 feet) tall, with pinnate leaves 4–6 m (13–20 ft) long, and pinnae 60–90 centimetres (2–3 ft) long; old leaves break away cleanly, leaving
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#17327754540642464-467: Is not thought to be indigenous to South America, and the highest genetic diversity is present in Asian Cocos , indicating that at least the modern species Cocos nucifera is native to there . In addition, fossils of potential Cocos ancestors have been recovered from both Colombia and India. In order to resolve this enigma, a 2014 study proposed that the ancestors of Cocos had likely originated on
2576-496: Is often called "anomalous secondary growth". The Arecaceae are notable among monocots for their height and for the size of their seeds, leaves, and inflorescences. Ceroxylon quindiuense , Colombia's national "tree", is the tallest monocot in the world, reaching up to 60 metres (197 ft) tall. The coco de mer ( Lodoicea maldivica ) has the largest seeds of any plant, 40–50 centimetres (16–20 in) in diameter and weighing 15–30 kilograms (33–66 lb) each ( coconuts are
2688-683: Is possible north of subtropical climates, and some higher latitude locales such as Ireland, Scotland, England, and the Pacific Northwest feature a few palms in protected locations and microclimates . In the United States , there are at least 12 native palm species, mostly occurring in the states of the Deep South and Florida . Palms inhabit a variety of ecosystems. More than two-thirds of palm species live in humid moist forests, where some species grow tall enough to form part of
2800-460: Is that of a solitary shoot ending in a crown of leaves. This monopodial character may be exhibited by prostrate, trunkless, and trunk-forming members. Some common palms restricted to solitary growth include Washingtonia and Roystonea . Palms may instead grow in sparse though dense clusters. The trunk develops an axillary bud at a leaf node, usually near the base, from which a new shoot emerges. The new shoot, in turn, produces an axillary bud and
2912-700: Is the Pacific coconut from the Philippines. This, together with their use of the South American sweet potato , suggests that Austronesian peoples may have sailed as far east as the Americas. In the Hawaiian Islands , the coconut is regarded as a Polynesian introduction , first brought to the islands by early Polynesian voyagers (also Austronesians) from their homelands in the southern islands of Polynesia. Specimens have been collected from
3024-790: Is used in rituals of Hinduism . It forms the basis of wedding and worship rituals in Hinduism. It also plays a central role in the Coconut Religion founded in 1963 in Vietnam . Coconuts were first domesticated by the Austronesian peoples in Island Southeast Asia and were spread during the Neolithic via their seaborne migrations as far east as the Pacific Islands , and as far west as Madagascar and
3136-465: Is usually fatal, although they have been known to recover from temperatures of −4 °C (25 °F). Due to this, there are not many coconut palms in California . They may grow but not fruit properly in areas with insufficient warmth or sunlight, such as Bermuda . The conditions required for coconut trees to grow without any care are: The main limiting factor for most locations which satisfy
3248-612: The Americas during the colonial era in the Columbian exchange , but there is evidence of a possible pre-Columbian introduction of Pacific coconuts to Panama by Austronesian sailors. The evolutionary origin of the coconut is under dispute, with theories stating that it may have evolved in Asia , South America, or Pacific islands. Trees can grow up to 30 metres (100 feet) tall and can yield up to 75 fruits per year, though fewer than 30
3360-633: The Caribbean coast of what is now Colombia, and during the Eocene the ancestral Cocos performed a long-distance dispersal across the Atlantic Ocean to North Africa . From here, island-hopping via coral atolls lining the Tethys Sea , potentially boosted by ocean currents at the time, would have proved crucial to dispersal, eventually allowing ancestral coconuts to reach India. The study contended that an adaptation to coral atolls would explain
3472-657: The Comoros . They played a critical role in the long sea voyages of Austronesians by providing a portable source of food and water, as well as providing building materials for Austronesian outrigger boats . Coconuts were also later spread in historic times along the coasts of the Indian and Atlantic Oceans by South Asian , Arab , and European sailors. Based on these separate introductions, coconut populations can still be divided into Pacific coconuts and Indo-Atlantic coconuts, respectively. Coconuts were introduced by Europeans to
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3584-566: The Dasypogonaceae . For a phylogenetic tree of the family, see the list of Arecaceae genera . The Arecaceae were the first modern family of monocots to appear in the fossil record around 80 million years ago (Mya), during the late Cretaceous period. The first modern species, such as Nypa fruticans and Acrocomia aculeata , appeared 69 Mya, as evidenced by fossil Nypa pollen. Palms appear to have undergone an early period of adaptive radiation . By 60 Mya, many of
3696-518: The East African coast. Indo-Atlantic coconuts were also introduced into the Atlantic Ocean by Portuguese ships from their colonies in coastal India and Sri Lanka ; first introduced to coastal West Africa , then onwards into the Caribbean and the east coast of Brazil . All of these introductions are within the last few centuries, relatively recent in comparison to the spread of Pacific coconuts. The coconut palm thrives on sandy soils and
3808-535: The Holocene . But research on them is still ongoing to determine their phylogenetic affinities. Endt & Hayward (1997) have noted their resemblance to members of the South American genus Parajubaea , rather than Cocos , and propose a South American origin. Conran et al. (2015), however, suggests that their diversity in New Zealand indicate that they evolved endemically, rather than being introduced to
3920-512: The Indian Ocean basin, resulting in limited admixture with Pacific coconuts introduced earlier to Madagascar and the Comoros via the ancient Austronesian maritime trade network . Coconuts can be broadly divided into two fruit types – the ancestral niu kafa form with a thick-husked, angular fruit, and the niu vai form with a thin-husked, spherical fruit with a higher proportion of endosperm . The terms are derived from
4032-553: The Maastrichtian – Danian of the early Tertiary (70 to 62 million years ago). C. binoriensis has been claimed by their authors to be the earliest known fossil of Cocos nucifera . Outside of New Zealand and India, only two other regions have reported Cocos -like fossils, namely Australia and Colombia . In Australia, a Cocos -like fossil fruit, measuring 10 cm × 9.5 cm ( 3 + 7 ⁄ 8 in × 3 + 3 ⁄ 4 in), were recovered from
4144-459: The Maypan , King , and Macapuno . These vary by the taste of the coconut water and color of the fruit, as well as other genetic factors. Botanically , the coconut fruit is a drupe , not a true nut . Like other fruits, it has three layers : the exocarp , mesocarp , and endocarp . The exocarp is the glossy outer skin, usually yellow-green to yellow-brown in color. The mesocarp is composed of
4256-479: The Pacific Region . Konan et al. , 2007 explains much resistance with a few alleles at a few microsatellites . They find that ' Vanuatu Tall ' and ' Sri-Lanka Green Dwarf ' are the most resistant while ' West African Tall ' breeds are especially susceptible. Palm tree The Arecaceae ( / ˌ ær ə ˈ k eɪ s i . iː , - ˌ aɪ / ) is a family of perennial , flowering plants in
4368-410: The Samoan language and was adopted into scientific usage by Harries (1978). The niu kafa form is the wild ancestral type, with thick husks to protect the seed, an angular, highly ridged shape to promote buoyancy during ocean dispersal, and a pointed base that allowed fruits to dig into the sand, preventing them from being washed away during germination on a new island. It is the dominant form in
4480-579: The canopy and shorter ones form part of the understory . Some species form pure stands in areas with poor drainage or regular flooding, including Raphia hookeri which is common in coastal freshwater swamps in West Africa. Other palms live in tropical mountain habitats above 1 thousand metres (3 thousand feet), such as those in the genus Ceroxylon native to the Andes . Palms may also live in grasslands and scrublands, usually associated with
4592-407: The genus Cocos . The term " coconut " (or the archaic " cocoanut ") can refer to the whole coconut palm , the seed , or the fruit , which botanically is a drupe , not a nut . They are ubiquitous in coastal tropical regions and are a cultural icon of the tropics . The coconut tree provides food, fuel, cosmetics, folk medicine and building materials, among many other uses. The inner flesh of
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4704-453: The monocot order Arecales . Their growth form can be climbers , shrubs , tree -like and stemless plants, all commonly known as palms . Those having a tree-like form are colloquially called palm trees . Currently, 181 genera with around 2,600 species are known, most of which are restricted to tropical and subtropical climates. Most palms are distinguished by their large, compound, evergreen leaves, known as fronds , arranged at
4816-572: The trunk smooth. On fertile soil, a tall coconut palm tree can yield up to 75 fruits per year, but more often yields less than 30. Given proper care and growing conditions, coconut palms produce their first fruit in six to ten years, taking 15 to 20 years to reach peak production. True-to-type dwarf varieties of Pacific coconuts have been cultivated by the Austronesian peoples since ancient times. These varieties were selected for slower growth, sweeter coconut water, and often brightly colored fruits. Many modern varieties are also grown, including
4928-469: The "Compact Dwarfs"). Although it shares similar characteristics to dwarf coconuts (including slow growth), it is genetically distinct and is thus believed to be independently domesticated, likely in Tonga . Other cultivars of niu leka may also exist in other islands of the Pacific, and some are probably descendants of advanced crosses between Compact Dwarfs and Southeast Asian Dwarf types. Coconut fruit in
5040-751: The "Tall" (var. typica ) and "Dwarf" (var. nana ) varieties. The two groups are genetically distinct, with the dwarf variety showing a greater degree of artificial selection for ornamental traits and for early germination and fruiting. The tall variety is outcrossing while dwarf palms are self-pollinating , which has led to a much greater degree of genetic diversity within the tall group. The dwarf coconut cultivars are fully domesticated, in contrast to tall cultivars which display greater diversity in terms of domestication (and lack thereof). The fact that all dwarf coconuts share three genetic markers out of thirteen (which are only present at low frequencies in tall cultivars) makes it likely that they all originate from
5152-479: The 17th century use the name as well. He consulted the catalogue Herbarium Amboinense by Georg Eberhard Rumphius where Rumphius said that coccus was a homonym of coccum and coccus from Greek κόκκος kokkos meaning "grain" or "berry", but Romans identified coccus with " kermes insects "; Rumphius preferred the word cocus as a replacement. However, the word cocus could also mean "cook" like coquus in Latin, so Linnaeus chose Cocos directly from
5264-516: The Americas (20%), and Africa (4%). A study in 2011 identified two highly genetically differentiated subpopulations of coconuts, one originating from Island Southeast Asia (the Pacific group) and the other from the southern margins of the Indian subcontinent (the Indo-Atlantic group). The Pacific group is the only one to display clear genetic and phenotypic indications that they were domesticated; including dwarf habit, self-pollination, and
5376-421: The Americas. They are now almost ubiquitous between 26° N and 26° S except for the interiors of Africa and South America. The 2014 coral atoll origin hypothesis proposed that the coconut had dispersed in an island hopping fashion using the small, sometimes transient, coral atolls. It noted that by using these small atolls, the species could easily island-hop. Over the course of evolutionary time-scales
5488-647: The Ceroxyloideae and Phytelephantoideae. Studies have suggested the lack of a fully resolved hypothesis for the relationships within the family is due to a variety of factors, including difficulties in selecting appropriate outgroups, homoplasy in morphological character states, slow rates of molecular evolution important for the use of standard DNA markers , and character polarization. However, hybridization has been observed among Orbignya and Phoenix species, and using chloroplast DNA in cladistic studies may produce inaccurate results due to maternal inheritance of
5600-517: The Chinchilla Sand Formation dated to the latest Pliocene or basal Pleistocene . Rigby (1995) assigned them to modern Cocos nucifera based on its size. In Colombia, a single Cocos -like fruit was recovered from the middle to late Paleocene Cerrejón Formation . The fruit, however, was compacted in the fossilization process and it was not possible to determine if it had the diagnostic three pores that characterize members of
5712-574: The IUCN, and was published in 1996. The rarest palm known is Hyophorbe amaricaulis . The only living individual remains at the Botanic Gardens of Curepipe in Mauritius . Some pests are specialists to particular taxa . Pests that attack a variety of species of palms include: The palm branch was a symbol of triumph and victory in classical antiquity . The Romans rewarded champions of
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#17327754540645824-678: The Indo-Atlantic coconuts. However, they may have also been partially selected for thicker husks for coir production, which was also important in Austronesian material culture as a source for cordage in building houses and boats. The niu vai form is the domesticated form dominant in Pacific coconuts. They were selected for by the Austronesian peoples for their larger endosperm-to-husk ratio as well as higher coconut water content, making them more useful as food and water reserves for sea voyages. The decreased buoyancy and increased fragility of this spherical, thin-husked fruit would not matter for
5936-526: The Middle East and North Africa. A wide range of common products and foods are derived from palms. In contemporary times, palms are also widely used in landscaping. In many historical cultures, because of their importance as food, palms were symbols for such ideas as victory, peace, and fertility. The word Arecaceae is derived from the word areca with the suffix "-aceae". Areca is derived from Portuguese , via Malayalam അടയ്ക്ക ( aṭaykka ), which
6048-445: The Pacific Ocean and that they could not have dispersed worldwide without human agency. More recently, genomic analysis of cultivated coconut ( C. nucifera L.) has shed light on the movement. However, admixture , the transfer of genetic material, evidently occurred between the two populations. Given that coconuts are ideally suited for inter-island group ocean dispersal, obviously some natural distribution did take place. However,
6160-549: The Pacific unaided. If they were naturally distributed and had been in the Pacific for a thousand years or so, then we would expect the eastern shore of Australia, with its own islands sheltered by the Great Barrier Reef , to have been thick with coconut palms: the currents were directly into, and down along this coast. However, both James Cook and William Bligh (put adrift after the Bounty mutiny ) found no sign of
6272-604: The Portuguese word coco instead. The specific name nucifera is derived from the Latin words nux (nut) and fera (bearing), for 'nut-bearing'. Coconuts have a nearly cosmopolitan distribution due to human cultivation and dispersal. However, their original distribution was in the Central Indo-Pacific , in the regions of Maritime Southeast Asia and Melanesia . Modern genetic studies have identified
6384-580: The Rathayatra carts which carry the deities of Krishna and his family members in the cart festival of Jagganath Puri in India are marked with the emblem of a palm tree. Specifically it is the symbol of Krishna's brother, Baladeva. In 1840, the American geologist Edward Hitchcock (1793–1864) published the first tree-like paleontology chart in his Elementary Geology , with two separate trees of life for
6496-623: The Taiwanese Austronesian languages makes it likely that the Austronesian coconut culture developed only after Austronesians started colonizing the Philippine islands . The importance of the coconut in Austronesian cultures is evidenced by shared terminology of even very specific parts and uses of coconuts, which were carried outwards from the Philippines during the Austronesian migrations. Indo-Atlantic type coconuts were also later spread by Arab and South Asian traders along
6608-399: The ancestral traits of tall habits and elongated triangular fruits. The coconut played a critical role in the migrations of the Austronesian peoples. They provided a portable source of both food and water, allowing Austronesians to survive long sea voyages to colonize new islands as well as establish long-range trade routes. Based on linguistic evidence, the absence of words for coconut in
6720-530: The area. The sale of seeds to nurseries and collectors is another threat, as the seeds of popular palms are sometimes harvested directly from the wild. In 2006, at least 100 palm species were considered endangered, and nine species have been reported as recently extinct. However, several factors make palm conservation more difficult. Palms live in almost every type of warm habitat and have tremendous morphological diversity. Most palm seeds lose viability quickly, and they cannot be preserved in low temperatures because
6832-407: The base. The stamens generally number six, with filaments that may be separate, attached to each other, or attached to the pistil at the base. The fruit is usually a single- seeded drupe (sometimes berry-like) but some genera (e.g., Salacca ) may contain two or more seeds in each fruit. Like all monocots , palms do not have the ability to increase the width of a stem ( secondary growth ) via
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#17327754540646944-414: The center of origin of coconuts as being the Central Indo-Pacific , the region between western Southeast Asia and Melanesia , where it shows greatest genetic diversity. Their cultivation and spread was closely tied to the early migrations of the Austronesian peoples who carried coconuts as canoe plants to the islands they settled. The similarities of the local names in the Austronesian region
7056-471: The chloroplast DNA. Chemical and molecular data from non-organelle DNA, for example, could be more effective for studying palm phylogeny. Recently, nuclear genomes and transcriptomes have been used to reconstruct the phylogeny of palms. This has revealed, for example, that a whole-genome duplication event occurred early in the evolution of the Arecaceae lineage, that was not experienced by its sister clade,
7168-514: The coconut shell that resemble facial features. Coco and coconut apparently came from 1521 encounters by Portuguese and Spanish explorers with Pacific Islanders , with the coconut shell reminding them of a ghost or witch in Portuguese folklore called coco (also côca ). In the West it was originally called nux indica , a name used by Marco Polo in 1280 while in Sumatra . He took
7280-490: The coconuts in the Americas are genetically closest related to the coconuts in the Philippines , and not to any other nearby coconut populations (including Polynesia ). Such an origin indicates that the coconuts were not introduced naturally, such as by sea currents. The researchers concluded that it was brought by early Austronesian sailors to the Americas from at least 2,250 BP, and may be proof of pre-Columbian contact between Austronesian cultures and South American cultures. It
7392-503: The cold kills the embryo. Using botanical gardens for conservation also presents problems, since they can rarely house more than a few plants of any species or truly imitate the natural setting. There is also the risk that cross-pollination can lead to hybrid species. The Palm Specialist Group of the World Conservation Union (IUCN) began in 1984, and has performed a series of three studies to find basic information on
7504-748: The control of the Empire of Japan after World War I , and was subsequently administered under the South Seas Mandate . Following World War II , the island came under the control of the United States of America and was administered as part of the Trust Territory of the Pacific Islands from 1947, and became part of the Federated States of Micronesia from 1979. On March 18, 1994, the freighter Oceanus ventured out of
7616-729: The deserts, it also created a more amenable habitat for the people to live in by providing shade and protection from the desert winds. An indication of the importance of palms in ancient times is that they are mentioned more than 30 times in the Bible , and at least 22 times in the Quran . The Torah also references the "70 date palm trees", which symbolize the 70 aspects of Torah that are revealed to those who "eat of its fruit." Arecaceae have great economic importance, including coconut products, oils, dates, palm syrup , ivory nuts, carnauba wax, rattan cane, raffia, and palm wood . This family supplies
7728-568: The early domestication of Pacific coconuts by the Austronesian peoples in maritime Southeast Asia during the Austronesian expansion (c. 3000 to 1500 BCE). Although archaeological remains dating to 1000 to 500 BCE also suggest that the Indo-Atlantic coconuts were also later independently cultivated by the Dravidian peoples , only Pacific coconuts show clear signs of domestication traits like dwarf habits, self-pollination, and rounded fruits. Indo-Atlantic coconuts, in contrast, all have
7840-404: The edible solid endosperm (the "coconut meat" or "coconut flesh") which hardens over time. The small cylindrical embryo is embedded in the solid endosperm directly below the functional pore of the endosperm. During germination, the embryo pushes out of the functional pore and forms a haustorium (the coconut sprout ) inside the central cavity. The haustorium absorbs the solid endosperm to nourish
7952-412: The endocarp is hollow and is lined with a thin brown seed coat around 0.2 mm ( 1 ⁄ 64 in) thick. The endocarp is initially filled with a multinucleate liquid endosperm (the coconut water ). As development continues, cellular layers of endosperm deposit along the walls of the endocarp up to 11 mm ( 3 ⁄ 8 in) thick, starting at the distal end. They eventually form
8064-531: The event five native Hawaiians and eleven others were inducted into Pwo as master navigators. The Polynesian Voyaging Society presented Piailug a canoe, the Alingano Maisu , as a gift for his key role in reviving traditional wayfinding navigation in Hawaii . Coconut The coconut tree ( Cocos nucifera ) is a member of the palm tree family ( Arecaceae ) and the only living species of
8176-460: The flesh, charcoal from the hard shell, and coir from the fibrous husk . Dried coconut flesh is called copra , and the oil and milk derived from it are commonly used in cooking – frying in particular – as well as in soaps and cosmetics . Sweet coconut sap can be made into drinks or fermented into palm wine or coconut vinegar . The hard shells, fibrous husks and long pinnate leaves can be used as material to make
8288-673: The games and celebrated military successes with palm branches. Early Christians used the palm branch to symbolize the victory of the faithful over enemies of the soul, as in the Palm Sunday festival celebrating the triumphal entry of Jesus Christ into Jerusalem . In Judaism , the palm represents peace and plenty, and is one of the Four Species of Sukkot ; the palm may also symbolize the Tree of Life in Kabbalah . The canopies of
8400-500: The habitat, and those with small habitat ranges are most vulnerable to them. The harvesting of heart of palm, a delicacy in salads, also poses a threat because it is derived from the palm's apical meristem , a vital part of the palm that cannot be regrown (except in domesticated varieties, e.g. of peach palm ). The use of rattan palms in furniture has caused a major population decrease in these species that has negatively affected local and international markets, as well as biodiversity in
8512-432: The history of the Middle East and North Africa. In the text "Date Palm Products" (1993), W.H. Barreveld wrote: One could go as far as to say that, had the date palm not existed, the expansion of the human race into the hot and barren parts of the "old" world would have been much more restricted. The date palm not only provided a concentrated energy food, which could be easily stored and carried along on long journeys across
8624-660: The islands by long-distance dispersal. In west-central India, numerous fossils of Cocos -like fruits, leaves, and stems have been recovered from the Deccan Traps . They include morphotaxa like Palmoxylon sundaran , Palmoxylon insignae , and Palmocarpon cocoides . Cocos -like fossils of fruits include Cocos intertrappeansis , Cocos pantii , and Cocos sahnii . They also include fossil fruits that have been tentatively identified as modern Cocos nucifera . These include two specimens named Cocos palaeonucifera and Cocos binoriensis , both dated by their authors to
8736-513: The locations of the admixture events are limited to Madagascar and coastal east Africa, and exclude the Seychelles . This pattern coincides with the known trade routes of Austronesian sailors. Additionally, a genetically distinct subpopulation of coconut on the Pacific coast of Latin America has undergone a genetic bottleneck resulting from a founder effect ; however, its ancestral population
8848-589: The main shipping channel when its captain attempted to peek at topless Satawalese women. The freighter ran aground on the nearby Wenimong coral reef , the primary source of food for the islanders, and 13,000 square meters of the reef were ravaged. The freighter's insurer, the North of England P&I Association , ended up paying US$ 2 million in compensation to the Satawalese. The island is best known for its preservation of traditional navigational techniques without
8960-407: The mature seed, as well as the coconut milk extracted from it, forms a regular part of the diets of many people in the tropics and subtropics . Coconuts are distinct from other fruits because their endosperm contains a large quantity of an almost clear liquid, called " coconut water " or "coconut juice". Mature, ripe coconuts can be used as edible seeds, or processed for oil and plant milk from
9072-694: The modern phylogenomic data, the Phytelephantoideae are tribe in the Ceroxyloideae subfamily. Currently, few extensive phylogenetic studies of the Arecaceae exist. In 1997, Baker et al. explored subfamily and tribe relationships using chloroplast DNA from 60 genera from all subfamilies and tribes. The results strongly showed the Calamoideae are monophyletic, and Ceroxyloideae and Coryphoideae are paraphyletic. The relationships of Arecoideae are uncertain, but they are possibly related to
9184-660: The modern, specialized genera of palms appeared and became widespread and common, much more widespread than their range today. Because palms separated from the monocots earlier than other families, they developed more intrafamilial specialization and diversity. By tracing back these diverse characteristics of palms to the basic structures of monocots, palms may be valuable in studying monocot evolution. Several species of palms have been identified from flowers preserved in amber, including Palaeoraphe dominicana and Roystonea palaea . Fossil evidence of them can also be found in samples of petrified palmwood . The relationship between
9296-544: The northwest, belongs to Satawal municipality. The native language is Satawalese , a Chuukic language closely related to Woleaian , and the entire population of the island numbers approximately 500. Although located in Yap State, the people of Satawal are more closely related, culturally and linguistically, to those of Chuuk . The Satawalese primarily subsist on fishing and some agriculture (coconuts, breadfruit, taro ). They build small thatch houses for sleeping and use
9408-562: The nuts along this 2,000 km (1,200 mi) stretch when he needed water for his crew. Nor were there coconuts on the east side of the African coast until Vasco da Gama , nor in the Caribbean when first visited by Christopher Columbus . They were commonly carried by Spanish ships as a source of fresh water. These provide substantial circumstantial evidence that deliberate Austronesian voyagers were involved in carrying coconuts across
9520-528: The original coloration of the fruit is more aesthetically pleasing. Whole mature coconuts (11 to 13 months from flowering) sold for export, however, typically have the husk removed to reduce weight and volume for transport. This results in the naked coconut "shell" with three pores more familiar in countries where coconuts are not grown locally. De-husked coconuts typically weigh around 750 to 850 grams (1 lb 10 oz to 1 lb 14 oz). De-husked coconuts are also easier for consumers to open, but have
9632-536: The palm (vivipary) was important. Coconuts today can be grouped into two highly genetically distinct subpopulations: the Indo-Atlantic group originating from southern India and nearby regions (including Sri Lanka , the Laccadives , and the Maldives ); and the Pacific group originating from the region between maritime Southeast Asia and Melanesia . Linguistic, archaeological, and genetic evidence all point to
9744-455: The plant near the surface. Only a few of the roots penetrate deep into the soil for stability. This type of root system is known as fibrous or adventitious, and is a characteristic of grass species. Other types of large trees produce a single downward-growing tap root with a number of feeder roots growing from it. 2,000–4,000 adventitious roots may grow, each about 1 cm ( 1 ⁄ 2 in) large. Decayed roots are replaced regularly as
9856-420: The prehistoric and modern distributions of Cocos , would have provided the necessary evolutionary pressures, and would account for morphological factors such as a thick husk to protect against ocean degradation and provide a moist medium in which to germinate on sparse atolls. The name coconut is derived from the 16th-century Portuguese word coco , meaning 'head' or 'skull' after the three indentations on
9968-434: The rainfall and temperature requirements is canopy growth, except those locations near coastlines, where the sandy soil and salt spray limit the growth of most other trees. Wild coconuts are naturally restricted to coastal areas in sandy, saline soils. The fruit is adapted for ocean dispersal. Coconuts could not reach inland locations without human intervention (to carry seednuts, plant seedlings, etc.) and early germination on
10080-493: The round " niu vai " fruit morphology with larger endosperm-to-husk ratios. The distribution of the Pacific coconuts correspond to regions settled by Austronesian voyagers indicating that its spread was largely the result of human introductions. It is most strikingly displayed in Madagascar , an island settled by Austronesian sailors at around 2000 to 1500 BP . The coconut populations on the island show genetic admixture between
10192-399: The same kind of vascular cambium found in non-monocot woody plants . This explains the cylindrical shape of the trunk (almost constant diameter) that is often seen in palms, unlike in ring -forming trees. However, many palms, like some other monocots, do have secondary growth, although because it does not arise from a single vascular cambium producing xylem inwards and phloem outwards, it
10304-457: The sea as far north as Norway (but it is not known where they entered the water). They have been found in the Caribbean and the Atlantic coasts of Africa and South America for less than 500 years (the Caribbean native inhabitants do not have a dialect term for them, but use the Portuguese name), but evidence of their presence on the Pacific coast of South America antedates Columbus's arrival in
10416-486: The second largest). Raffia palms ( Raphia spp.) have the largest leaves of any plant, up to 25 metres (82 ft) long and 3 metres (10 ft) wide. The Corypha species have the largest inflorescence of any plant, up to 7.5 metres (25 ft) tall and containing millions of small flowers. Calamus stems can reach 200 metres (656 ft) in length. Most palms are native to tropical and subtropical climates. Palms thrive in moist and hot climates but can be found in
10528-431: The seedling. Coconut fruits have two distinctive forms depending on § domestication . Wild coconuts feature an elongated triangular fruit with a thicker husk and a smaller amount of endosperm. These allow the fruits to be more buoyant and make it easier for them to lodge into sandy shorelines, making their shape ideal for ocean dispersal. Domesticated Pacific coconuts, on the other hand, are rounded in shape with
10640-458: The shifting atolls would have shortened the paths of colonization, meaning that any one coconut would not have to travel very far to find new land. Coconuts are susceptible to the phytoplasma disease, lethal yellowing . One recently selected cultivar , the 'Maypan' , has been bred for resistance to this disease. Yellowing diseases affect plantations in Africa, India, Mexico, the Caribbean and
10752-485: The status of palms in the wild, use of wild palms, and palms under cultivation. Two projects on palm conservation and use supported by the World Wildlife Fund took place from 1985 to 1990 and 1986–1991, in the American tropics and southeast Asia, respectively. Both studies produced copious new data and publications on palms. Preparation of a global action plan for palm conservation began in 1991, supported by
10864-411: The stem. The leaves have a tubular sheath at the base that usually splits open on one side at maturity. The inflorescence is a spadix or spike surrounded by one or more bracts or spathes that become woody at maturity. The flowers are generally small and white, radially symmetric, and can be either uni- or bisexual. The sepals and petals usually number three each and may be distinct or joined at
10976-482: The subfamilies is shown in the following cladogram: Calamoideae Nypoideae Coryphoideae Ceroxyloideae Arecoideae Evidence for cultivation of the date palm by Mesopotamians and other Middle Eastern peoples exists from more than 5,000 years ago, in the form of date wood, pits for storing dates, and other remains of the date palm in Mesopotamian sites. The date palm had a significant effect on
11088-576: The term from the Arabs, who called it جوز هندي jawz hindī , translating to 'Indian nut'. Thenga , its Tamil / Malayalam name, was used in the detailed description of coconut found in Itinerario by Ludovico di Varthema published in 1510 and also in the later Hortus Indicus Malabaricus . Carl Linnaeus first wanted to name the coconut genus Coccus from latinizing the Portuguese word coco , because he saw works by other botanists in middle of
11200-484: The top of an unbranched stem, except for the Hyphaene genus, who has branched palms. However, palms exhibit an enormous diversity in physical characteristics and inhabit nearly every type of habitat within their range, from rainforests to deserts . Palms are among the best known and most extensively cultivated plant families. They have been important to humans throughout much of history, especially in regions like
11312-551: The tree grows new ones. The palm produces both the female and male flowers on the same inflorescence ; thus, the palm is monoecious . However, there is some evidence that it may be polygamomonoecious and may occasionally have bisexual flowers. The female flower is much larger than the male flower. Flowering occurs continuously. Coconut palms are believed to be largely cross- pollinated , although most dwarf varieties are self-pollinating. The evolutionary history and fossil distribution of Cocos nucifera and other members of
11424-596: The trees were carried off by the water and drowned. Eight survivors from the island came to the Marianas in April of 1849 to permanently live there. There were survivors who were still on the island but could not leave for lack of boats. They also planned on leaving the island permanently. As with all of the Caroline Islands, sovereignty passed from Spain to the Empire of Germany in 1899. The island came under
11536-716: The tribe Cocoseae is more ambiguous than modern-day dispersal and distribution, with its ultimate origin and pre-human dispersal still unclear. There are currently two major viewpoints on the origins of the genus Cocos , one in the Indo-Pacific, and another in South America. The vast majority of Cocos -like fossils have been recovered generally from only two regions in the world: New Zealand and west-central India . However, like most palm fossils, Cocos -like fossils are still putative, as they are usually difficult to identify. The earliest Cocos -like fossil to be found
11648-484: The tribe Cocoseae . Nevertheless, Gomez-Navarro et al. (2009), assigned it to Cocos based on the size and the ridged shape of the fruit. Further complicating measures to determine the evolutionary history of Cocos is the genetic diversity present within C. nucifera as well as its relatedness to other palms. Phylogenetic evidence supports the closest relatives of Cocos being either Syagrus or Attalea , both of which are found in South America. However, Cocos
11760-647: The tropics, mostly in humid lowland subtropical climates, in highlands in southern Asia, and along the rim lands of the Mediterranean Sea . The northernmost native palm is Chamaerops humilis , which reaches 44°N latitude along the coast of Liguria , Italy . In the southern hemisphere, the southernmost palm is the Rhopalostylis sapida , which reaches 44°S on the Chatham Islands where an oceanic climate prevails. Cultivation of palms
11872-466: The trunks of breadfruit trees for boat-building. Cultural forms primarily revolve around dance and story-telling, and an alcoholic beverage known as tuba (a palm wine ) is brewed from fermented sap of the coconut flower spike. In 1849 a earthquake struck which caused flooding. This flood led to the island being submerged with the exception of trees people climbed onto. The island was submerged for several hours. People in certain cases who were hanging on
11984-476: The two subpopulations indicating that Pacific coconuts were first brought by the Austronesian settlers, which then interbred with the later Indo-Atlantic coconuts brought by Europeans from India. Genetic studies of coconuts have also confirmed pre-Columbian populations of coconuts in Panama . However, it is not native and displays a genetic bottleneck resulting from a founder effect . A study in 2008 showed that
12096-540: The use of instruments, based on indigenous astronomical and maritime concepts. Despite its small population, Satawal has continued to produce ocean-going canoes and expert navigators versed in these traditions. The best-known of the Satawal master navigators ( paliuw ), Mau Piailug , served as mentor and teacher to the founding members of the Polynesian Voyaging Society . The daily life of Satawal
12208-545: The whole way to Polynesia . But we had laid about half among the special provisions below deck, with the waves washing around them. Every single one of these was ruined by the sea water. And no coconut can float over the sea faster than a balsa raft moves with the wind behind it. He also notes that several of the nuts began to germinate by the time they had been ten weeks at sea, precluding an unassisted journey of 100 days or more. Drift models based on wind and ocean currents have shown that coconuts could not have drifted across
12320-422: The wild is light, buoyant, and highly water resistant. It is claimed that they evolved to disperse significant distances via marine currents . However, it can also be argued that the placement of the vulnerable eye of the nut (down when floating), and the site of the coir cushion are better positioned to ensure that the water-filled nut does not fracture when dropping on rocky ground, rather than for flotation. It
12432-426: The world. Along with dates mentioned above, members of the palm family with human uses are numerous: Like many other plants, palms have been threatened by human intervention and exploitation . The greatest risk to palms is destruction of habitat , especially in the tropical forests , due to urbanization , wood-chipping , mining , and conversion to farmland . Palms rarely reproduce after such great changes in
12544-460: Was Cocos zeylandica , a fossil species described as small fruits, around 3.5 cm ( 1 + 1 ⁄ 2 in) × 1.3 to 2.5 cm ( 1 ⁄ 2 to 1 in) in size, recovered from the Miocene (~23 to 5.3 million years ago) of New Zealand in 1926. Since then, numerous other fossils of similar fruits were recovered throughout New Zealand from the Eocene , Oligocene , and possibly
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