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108-725: The Seward Peninsula is a remnant of the Bering land bridge , a roughly thousand-mile-wide swath of land connecting Siberia with mainland Alaska during the Pleistocene Ice Age . This land bridge aided in the migration of humans, as well as plant and animal species, from Asia to North America. Excavations at sites such as the Trail Creek Caves and Cape Espenberg in the Bering Land Bridge National Preserve as well as Cape Denbigh to

216-582: A Dene (Athapascan) group, call the Arctic caribou Ɂekwǫ̀ and the boreal woodland caribou tǫdzı . The Gwichʼin (also a Dene group) have over 24 distinct caribou-related words. Reindeer are also called tuttu by the Greenlandic Inuit and hreindýr , sometimes rein , by the Icelanders . The "glacial-interglacial cycles of the upper Pleistocene had a major influence on

324-464: A boreal forest hosting a species assemblage with no modern analogue. These are among the oldest DNA fragments ever sequenced. Carl Linnaeus in 1758 named the Eurasian tundra species Cervus tarandus , the genus Rangifer being credited to Smith, 1827. Rangifer has had a convoluted history because of the similarity in antler architecture (brow tines asymmetrical and often palmate, bez tines,

432-463: A Captain Craycott had brought a live pair from Greenland to England in 1738. He named it Capra groenlandicus , Greenland reindeer. Linnaeus, in the 12th edition of Systema naturae , gave grœnlandicus as a synonym for Cervus tarandus . Borowski disagreed (and again changed the spelling), saying Cervus grönlandicus was morphologically distinct from Eurasian tundra reindeer. Baird placed it under

540-454: A back tine sometimes branched, and branched at the distal end, often palmate). Because of individual variability, early taxonomists were unable to discern consistent patterns among populations, nor could they, examining collections in Europe, appreciate the difference in habitats and the differing function they imposed on antler architecture. Comparative morphometrics, the measurement of skulls,

648-778: A broad, high muzzle to increase the volume of the nasal cavity to warm and moisten the air before it enters the throat and lungs, bez tines set close to the brow tines, distinctive coat patterns, short legs and other adaptations for running long distances, and multiple behaviors suited to tundra, but not to forest (such as synchronized calving and aggregation during rutting and post-calving). As well, many genes, including those for vitamin D metabolism, fat metabolism , retinal development, circadian rhythm , and tolerance to cold temperatures, are found in tundra caribou that are lacking or rudimentary in forest types. For this reason, forest-adapted reindeer and caribou could not survive in tundra or polar deserts . The oldest undoubted Rangifer fossil

756-439: A direct common ancestor , they cannot be Biological specificity#conspecific|conspecific. Similarly, woodland caribou diverged from the ancestors of Arctic caribou before modern barren-ground caribou had evolved, and were more likely related to extinct North American forest reindeer (see Evolution above). Lacking a direct shared ancestor, barren-ground and woodland caribou cannot be conspecific. Molecular data also revealed that

864-611: A forest subspecies, formerly included reindeer west of the Sea of Okhotsk which, however, are indistinguishable genetically from the Jano-Indigirka, East Siberian taiga and Chukotka populations of R. t. sibiricus . Siberian tundra reindeer herds have been in decline but are stable or increasing since 2000. Insular (island) reindeer, classified as the Novaya Zemlya reindeer ( R. t. pearsoni ) occupy several island groups:

972-464: A land bridge from c.  30,000  – c.  11,000  YBP, followed by a Holocene sea-level rise that reopened the strait. Post-glacial rebound has continued to raise some sections of the coast. During the last glacial period , enough of the Earth's water became frozen in the great ice sheets covering North America and Europe to cause a drop in sea levels . For thousands of years

1080-428: A large scale in the world. Both wild and domestic reindeer have been an important source of food, clothing, and shelter for Arctic people from prehistorical times. They are still herded and hunted today. In some traditional Christmas legends, Santa Claus's reindeer pull a sleigh through the night sky to help Santa Claus deliver gifts to good children on Christmas Eve. Names follow international convention before

1188-725: A mtDNA haplotype with Labrador caribou, in the North American lineage (i.e., woodland caribou). Røed et al. (1991) had noted: Among Baffin Island caribou the TFL2 allele was the most common allele (p=0.521), while this allele was absent, or present in very low frequencies, in other caribou populations (Table 1), including the Canadian barren-ground caribou from the Beverly herd. A large genetic difference between Baffin Island caribou and

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1296-816: A new revision of the genus. Abbreviations: The table above includes, as per the recent revision, R. t. caboti (the Labrador caribou (the Eastern Migratory population DU4)), and R. t. terranovae (the Newfoundland caribou (the Newfoundland population DU5)), which molecular analyses have shown to be of North American (i.e., woodland caribou) lineage; and four mountain ecotypes now known to be of distant Beringia - Eurasia lineage (see Taxonomy above). The scientific name Tarandus rangifer buskensis Millais, 1915 (the Busk Mountains reindeer)

1404-439: A predator-avoidance strategy, which requires large rutting aggregations. Males cannot defend a harem because, while he was busy fighting, they would disappear into the mass of the herd. Males therefore tend individual females; their fights are infrequent and brief. Their antlers are thin, beams round in cross-section, sweep back and then forward with a cluster of branches at the top; these are designed more for visual stimulation of

1512-480: A rich diversity of grasses and herbs. There were patches of shrub tundra with isolated refugia of larch ( Larix ) and spruce ( Picea ) forests with birch ( Betula ) and alder ( Alnus ) trees. It has been proposed that the largest and most diverse megafaunal community residing in Beringia at this time could only have been sustained in a highly diverse and productive environment. Analysis at Chukotka on

1620-549: A scathing review by Ian McTaggart-Cowan in 1962. Most authorities continued to consider all or most subspecies valid; some were quite distinct. In his chapter in the authoritative 2005 reference work Mammal Species of the World , referenced by the American Society of Mammalogists , English zoologist Peter Grubb agreed with Valerius Geist , a specialist on large mammals, that these subspecies were valid (i.e., before

1728-772: A very interesting history. Allen (1902) named it as a distinct species, R. granti , from the "western end of Alaska Peninsula , opposite Popoff Island " and noting that: Rangifer granti is a representative of the Barren Ground group of Caribou, which includes R. arcticus of the Arctic Coast and R. granlandicus of Greenland. It is not closely related to R. stonei of the Kenai Peninsula, from which it differs not only in its very much smaller size, but in important cranial characters and in coloration. ...The external and cranial differences between R. granti and

1836-515: A vital role in the subsistence lifestyles of many peninsula residents and ease travel, hunting, and fishing. Most peninsula rivers have at least a small yearly run of several varieties of salmon , as well as Dolly Varden trout , Arctic grayling , whitefish of various species, northern pike , and burbot . Most rivers on the Seward Peninsula freeze in mid-October; spring break-up usually occurs in mid- to late May. The Seward Peninsula

1944-808: A wildlife rehabilitation center in Canada, caribou were considered extirpated from the contiguous United States . The Committee on the Status of Endangered Wildlife in Canada (COSEWIC) classified both the Southern Mountain population DU9 ( R. t. montanus ) and the Central Mountain population DU8 ( R. t. fortidens ) as Endangered and the Northern Mountain population DU7 ( R. t. osborni ) as Threatened. Some species and subspecies are rare and three subspecies have already become extinct:

2052-520: Is a lava field formed roughly 1,000 to 2,000 years ago, which covers roughly 88 sq mi (230 km). Several geothermal hot springs are located throughout the peninsula, including Serpentine Hot Springs , Pilgrim Hot Springs , Granite Mountain, Elim, Clear Creek and Lava Creek. The Seward Peninsula has several rivers. The largest include the Koyuk , Kuzitrin , Niukluk , Fish , Tubuktilik , Kiwalik , Buckland and Agiupuk rivers. These play

2160-619: Is a species of deer with circumpolar distribution , native to Arctic , subarctic , tundra , boreal , and mountainous regions of Northern Europe, Siberia, and North America. It is the only representative of the genus Rangifer . More recent studies suggest the splitting of reindeer and caribou into six distinct species over their range. Reindeer occur in both migratory and sedentary populations, and their herd sizes vary greatly in different regions. The tundra subspecies are adapted for extreme cold, and some are adapted for long-distance migration. Reindeer vary greatly in size and color from

2268-580: Is from Omsk , Russia, dated to 2.1-1.8 Ma. The oldest North American Rangifer fossil is from the Yukon , 1.6 million years before present (BP). A fossil skull fragment from Süßenborn, Germany, R. arcticus stadelmanni , (which is probably misnamed) with "rather thin and cylinder-shaped" antlers, dates to the Middle Pleistocene (Günz) Period, 680,000-620,000 BP. Rangifer fossils become increasingly frequent in circumpolar deposits beginning with

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2376-475: Is necessary with R. montanus or with any of the woodland forms." Osgood and Murie (1935), agreeing with granti ' s close relationship with the barren-ground caribou, brought it under R. arcticus as a subspecies, R. t. granti . Anderson (1946) and Banfield (1961), based on statistical analysis of cranial, dental and other characters, agreed. But Banfield (1961) also synonymized Alaska's large R. stonei with other mountain caribou of British Columbia and

2484-664: Is often seen as more objective than description of differences of color or antler patterns, but actually confounds genetic variance with epistatic and statistical variance as well as compounded environment-based variance. For example, woodland caribou males, rutting in boreal forest where only a few females can be found, collect harems and defend them against other males, for which they have short, straight, strong, much-branched antlers, beams flattened in cross-section, designed for combat — and not too large, so as not to impede them in forested winter ranges. By contrast, modern tundra caribou (see Evolution above) have synchronized calving as

2592-480: Is the westernmost limit of distribution for the black spruce , Picea mariana , a dominant overstory species of the region. Alaska's reindeer herding was concentrated on the Seward Peninsula ever since the first shipment of reindeer were imported there from eastern Siberia in 1892. It was believed that migrating caribou could be prevented from mingling with the domesticated reindeer on the Peninsula because of

2700-605: The Aleutians and islands in the Bering Sea at the close of the nineteenth century indicated that a past land connection might lie beneath the shallow waters between Alaska and Chukotka . The underlying mechanism was first thought to be tectonics, but by 1930 changes in the ice mass balance, leading to global sea-level fluctuations were viewed as the cause of the Bering land bridge. In 1937, Eric Hultén proposed that around

2808-709: The Anadyr River ), and Arctodus simus , American badger , American kiang -like equids, Bootherium and Camelops in North America, with the existence of Homotherium being disputed in Late Pleistocene Siberia. The lack of mastodon and Megalonyx has been attributed to their inhabitation of Alaska and the Yukon being limited to interglacials. However, ground sloth eDNA has potentially been recovered from Siberia. The peopling of

2916-406: The Bering land bridge , that was up to 1,000 km (620 mi) wide at its greatest extent and which covered an area as large as British Columbia and Alberta together, totaling about 1.6 million km (620,000 sq mi), allowing biological dispersal to occur between Asia and North America. Today, the only land that is visible from the central part of the Bering land bridge are

3024-679: The Diomede Islands , the Pribilof Islands of St. Paul and St. George, St. Lawrence Island , St. Matthew Island , and King Island . It is believed that a small human population of at most a few thousand arrived in Beringia from eastern Siberia during the Last Glacial Maximum before expanding into the settlement of the Americas sometime after 16,500 years Before Present (YBP). This would have occurred as

3132-594: The Early Pleistocene (2 million years ago) Kap Kobenhavn Formation of northern Greenland identified preserved DNA fragments of Rangifer , identified as basal but potentially ancestral to modern reindeer. This suggests that reindeer have inhabited Greenland since at least the Early Pleistocene. Around this time, northern Greenland was 11–19 °C (20–34 °F) warmer than the Holocene , with

3240-540: The Last Glacial Maximum , when ice sheets began advancing from 33,000   YBP and reached their maximum limits 26,500   YBP. Deglaciation commenced in the Northern Hemisphere approximately 19,000   YBP and in Antarctica approximately 14,500 years   YBP, which is consistent with evidence that glacial meltwater was the primary source for an abrupt rise in sea level 14,500   YBP and

3348-738: The Last Glacial Period until the present day. In the non-forested mountains of central Norway, such as Jotunheimen , it is still possible to find remains of stone-built trapping pits , guiding fences and bow rests, built especially for hunting reindeer. These can, with some certainty, be dated to the Migration Period , although it is not unlikely that they have been in use since the Stone Age . Cave paintings by ancient Europeans include both tundra and forest types of reindeer. A 2022 study of ancient environmental DNA from

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3456-801: The Late Miocene , 8.7–9.6 million years ago. Rangifer "evolved as a mountain deer, ...exploiting the subalpine and alpine meadows...". Rangifer originated in the Late Pliocene and diversified in the Early Pleistocene , a 2+ million-year period of multiple glacier advances and retreats. Several named Rangifer fossils in Eurasia and North America predate the evolution of modern tundra reindeer. Archaeologists distinguish "modern" tundra reindeer and barren-ground caribou from primitive forms – living and extinct – that did not have adaptations to extreme cold and to long-distance migration. They include

3564-568: The Mi'kmaq qalipu , meaning "snow shoveler", and refers to its habit of pawing through the snow for food. Because of its importance to many cultures, Rangifer and some of its species and subspecies have names in many languages. Inuvialuit of the western Canadian Arctic and Inuit of the eastern Canadian Arctic, who speak different dialects of the Inuit languages , both call the barren-ground caribou tuktu . The Wekʼèezhìi ( Tłı̨chǫ ) people,

3672-663: The Novaya Zemlya Archipelago (about 5,000 animals at last count, but most of these are either domestic reindeer or domestic-wild hybrids), the New Siberia Archipelago (about 10,000 to 15,000), and Wrangel Island (200 to 300 feral domestic reindeer). What was once the second largest herd is the migratory Labrador caribou ( R. t. caboti ) George River herd in Canada, with former variations between 28,000 and 385,000. As of January 2018, there are fewer than 9,000 animals estimated to be left in

3780-746: The Purchase of Alaska from Russia in 1867. 65°20′N 164°15′W  /  65.333°N 164.250°W  / 65.333; -164.250 Bering land bridge Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia ; on the east by the Mackenzie River in Canada ; on the north by 72° north latitude in the Chukchi Sea ; and on

3888-681: The Queen Charlotte Islands caribou ( R. t. dawsoni ) from western Canada, the Sakhalin reindeer ( R. t. setoni ) from Sakhalin and the East Greenland caribou from eastern Greenland, although some authorities believe that the latter, R. t. eogroenlandicus Degerbøl, 1957, is a junior synonym of the Peary caribou. Historically, the range of the sedentary boreal woodland caribou covered more than half of Canada and into

3996-529: The Riss glaciations , the second youngest of the Pleistocene Epoch, roughly 300,000–130,000 BP. By the 4-Würm period (110,000–70,000 to 12,000–10,000 BP), its European range was extensive, supplying a major food source for prehistoric Europeans. North American fossils outside of Beringia that predate the Last Glacial Maximum (LGM) are of Rancholabrean age (240,000–11,000 years BP) and occur along

4104-817: The Russian mainland. In August 2011 Russia announced an ambitious project to construct a rail tunnel under the Bering Strait, linking the Seward Peninsula in Alaska with the Chukchi Peninsula in Russia. If completed, the project would cost an estimated US$ 65 billion and would be the world's longest tunnel at 64 mi (103 km) long. The peninsula was named after William H. Seward , the United States Secretary of State who negotiated

4212-881: The Scandinavian mountains and R. t. sibiricus across Siberia) and east ( R. t. arcticus in the North American Barrenlands) when rising seas isolated them. Likewise in North America, DNA analysis shows that woodland caribou ( R. caribou ) diverged from primitive ancestors of tundra / barren-ground caribou not during the LGM, 26,000–19,000 years ago, as previously assumed, but in the Middle Pleistocene around 357,000 years ago. At that time, modern tundra caribou had not even evolved. Woodland caribou are likely more related to extinct North American forest caribou than to barren-ground caribou. For example,

4320-474: The Sámi word raingo . Carl Linnaeus chose the word tarandus as the specific epithet, making reference to Ulisse Aldrovandi 's Quadrupedum omnium bisulcorum historia fol. 859–863, Cap. 30: De Tarando (1621). However, Aldrovandi and Conrad Gessner thought that rangifer and tarandus were two separate animals. In any case, the tarandos name goes back to Aristotle and Theophrastus . The use of

4428-864: The polar desert of the high Arctic Archipelago and Grant's caribou ( R. t. granti also called the Porcupine caribou ) lives in the western end of the Alaska Peninsula and the adjacent islands; the other four subspecies, Osborn's caribou ( R. t. osborni ), Stone's caribou ( R. t. stonei ), the Rocky Mountain caribou ( R. t. fortidens ) and the Selkirk Mountains caribou ( R. t. montanus ) are all montane . The extinct insular Queen Charlotte Islands caribou ( R. t. dawsoni ), lived on Graham Island in Haida Gwaii (formerly known as

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4536-520: The tundra , taiga (boreal forest) and south through the Canadian Rocky Mountains . Of the eight subspecies classified by Harding (2022) into the Arctic caribou ( R. arcticus ), the migratory mainland barren-ground caribou of Arctic Alaska and Northern Canada ( R. t. arcticus ), summer in tundra and winter in taiga, a transitional forest zone between boreal forest and tundra; the nomadic Peary caribou ( R. t. pearyi ) lives in

4644-553: The Aleutians and the Bering Strait region were tundra plants that had originally dispersed from a now-submerged plain between Alaska and Chukotka, which he named Beringia after Vitus Bering who had sailed into the strait in 1728. The American arctic geologist David Hopkins redefined Beringia to include portions of Alaska and Northeast Asia. Beringia was later regarded as extending from the Verkhoyansk Mountains in

4752-667: The American glaciers blocking the way southward melted, but before the bridge was covered by the sea about 11,000 YBP. The term Beringia was coined by the Swedish botanist Eric Hultén in 1937, from the Danish-born Russian explorer Vitus Bering . During the ice ages, Beringia, like most of Siberia and all of North and Northeast China , was not glaciated because snowfall was very light . The remains of Late Pleistocene mammals that had been discovered on

4860-734: The Americas began when Paleolithic hunter-gatherers ( Paleo-Indians ) entered North America from the North Asian Mammoth steppe via the Beringia land bridge , which had formed between northeastern Siberia and western Alaska due to the lowering of sea level during the Last Glacial Maximum (26,000 to 19,000 years ago). These populations expanded south of the Laurentide Ice Sheet and spread rapidly southward, occupying both North and South America by 12,000 to 14,000 years ago. The earliest populations in

4968-440: The Americas, before roughly 10,000 years ago, are known as Paleo-Indians . Indigenous peoples of the Americas have been linked to Siberian populations by proposed linguistic factors , the distribution of blood types , and in genetic composition as reflected by molecular data, such as DNA . Around 3,000 years ago, the progenitors of the Yupik peoples settled along both sides of the straits. The governments of Russia and

5076-422: The Arctic caribou. Siberian tundra reindeer herds are also in decline, and Rangifer as a whole is considered to be Vulnerable by the International Union for Conservation of Nature (IUCN). Charles Hamilton Smith is credited with the name Rangifer for the reindeer genus, which Albertus Magnus used in his De animalibus , fol. Liber 22, Cap. 268: "Dicitur Rangyfer quasi ramifer". This word may go back to

5184-626: The Beverly herd was also indicated by eight alleles found in the Beverly herd which were absent from the Baffin Island samples. Jenkins et al. (2018) also reported genetic distinctiveness of Baffin Island caribou from all other barren-ground caribou; its genetic signature was not found on the mainland or on other islands; nor were Beverly herd (the nearest mainly barren-ground caribou) alleles present in Baffin Island caribou, evidence of reproductive isolation. These advances in Rangifer genetics were brought together with previous morphological-based descriptions, ecology, behavior and archaeology to propose

5292-501: The Cenozoic. [REDACTED] Africa [REDACTED] Antarctica [REDACTED] Asia [REDACTED] Australia [REDACTED] Europe [REDACTED] North America [REDACTED] South America [REDACTED] Afro-Eurasia [REDACTED] Americas [REDACTED] Eurasia [REDACTED] Oceania Reindeer See text , traditionally 1, but possibly up to 6 The reindeer or caribou ( Rangifer tarandus )

5400-464: The Eurasian reindeer radiation dates to the large Riss glaciation (347,000 to 128,000 years ago), based on the Norwegian-Svalbard split 225,000 years ago. Finnish forest reindeer ( R. t. fennicus ) likely evolved from Cervus [Rangifer] geuttardi Desmarest, 1822, a reindeer that adapted to forest habitats in Eastern Europe as forests expanded during an interglacial period before the LGM (the Würmian or Weichsel glaciation );. The fossil species geuttardi

5508-399: The George River herd, as reported by the Canadian Broadcasting Corporation . The New York Times reported in April 2018 of the disappearance of the only herd of southern mountain woodland caribou in the contiguous United States , with an expert calling it "functionally extinct" after the herd's size dwindled to a mere three animals. After the last individual, a female, was translocated to

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5616-412: The Greenland caribou ( R. t. groenlandicus ) and the Svalbard reindeer ( R. t. platyrhynchus ), although not closely related to each other, were the most genetically divergent among Rangifer clades; that modern (see Evolution above) Eurasian tundra reindeer ( R. t. tarandus and R. t. sibiricus ) and North American barren-ground caribou ( R. t. arcticus ), although sharing ancestry, were separable at

5724-422: The Last Glacial Maximum. This was followed by a single population of modern wolves expanding out of their Beringia refuge to repopulate the wolf's former range, replacing the remaining Late Pleistocene wolf populations across Eurasia and North America. The extinct pine species Pinus matthewsii has been described from Pliocene sediments in the Yukon areas of the refugium. The existence of fauna endemic to

5832-425: The Mammals of the World Vol. 2: Hoofed Mammals . Most Russian authors also recognized R. t. angustirostris , a forest reindeer from east of Lake Baikal . However, since 1991, many genetic studies have revealed deep divergence between modern tundra reindeer and woodland caribou. Geist (2007) and others continued arguing that the woodland caribou was incorrectly classified, noting that "true woodland caribou,

5940-431: The Queen Charlotte Islands). The boreal woodland caribou ( R. t. caribou ), lives in the boreal forest of northeastern Canada: the Labrador or Ungava caribou of northern Quebec and northern Labrador ( R. t. caboti ), and the Newfoundland caribou of Newfoundland ( R. t. terranovae ) have been found to be genetically in the woodland caribou lineage. In Eurasia, both wild and domestic reindeer are distributed across

6048-418: The Reindeer and Caribou, Genus Rangifer (1961), eliminated R. t. caboti (the Labrador caribou), R. t. osborni (Osborn's caribou — from British Columbia ) and R. t. terranovae (the Newfoundland caribou) as invalid and included only barren-ground caribou , renamed as R. t. groenlandicus (formerly R. arcticus ) and woodland caribou as R. t. caribou . However, Banfield made multiple errors, eliciting

6156-413: The Seward Peninsula include the Bendeleben Mountains , Darby Mountains , and York Mountains . The Bendeleben Mountains exhibit evidence of recent faulting in the late Cenozoic, with the majority of tectonic deformation and mountain formation occurring in the Cretaceous, which is attributed to regional tectonic block rotation of the Bering plate in the Arctic. The Lost Jim Lava Flow north of Kuzitrin Lake

6264-467: The Siberian edge of the land bridge indicated that from c.  57,000  – c.  15,000  YBP (MIS 3 to MIS 2) the environment was wetter and colder than the steppe–tundra to the east and west, with warming in parts of Beringia from c.  15,000  YBP. These changes provided the most likely explanation for mammal migrations after c.  15,000  YBP, as the warming provided increased forage for browsers and mixed feeders. At

6372-657: The Siberian forest reindeer ( R. t. valentinae , formerly called the Busk Mountains reindeer ( R. t. buskensis ) by American taxonomists) occupies the Altai and Ural Mountains . Male ("bull") and female ("cow") reindeer can grow antlers annually, although the proportion of females that grow antlers varies greatly between populations. Antlers are typically larger on males. Antler architecture varies by species and subspecies and, together with pelage differences, can often be used to distinguish between species and subspecies (see illustrations in Geist, 1991 and Geist, 1998). About 25,000 mountain reindeer ( R. t. tarandus ) still live in

6480-524: The United States announced a plan to formally establish "a transboundary area of shared Beringian heritage". Among other things this agreement would establish close ties between the Bering Land Bridge National Preserve and the Cape Krusenstern National Monument in the United States and Beringia National Park in Russia. Biogeographical evidence demonstrates previous connections between North America and Asia. Similar dinosaur fossils occur both in Asia and in North America . The dinosaur Saurolophus

6588-509: The Yukon as invalid subspecies of woodland caribou, then R. t. caribou . This left the small, migratory barren-ground caribou of Alaska and the Yukon, including the Porcupine caribou herd, without a name, which Banfield rectified in his 1974 Mammals of Canada by extending to them the name " granti ". The late Valerius Geist (1998), in the only error in his whole illustrious career, re-analyzed Banfield's data with additional specimens found in an unpublished report he cites as "Skal, 1982", but

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6696-499: The Yukon where it was blocked by the Wisconsin glaciation . Therefore, the flora and fauna of Beringia were more related to those of Eurasia rather than North America. Beringia received more moisture and intermittent maritime cloud cover from the north Pacific Ocean than the rest of the Mammoth steppe, including the dry environments on either side of it. This moisture supported a shrub-tundra habitat that provided an ecological refugium for plants and animals. In East Beringia 35,000 YBP,

6804-466: The Yukon. In the driest and coldest periods of the Late Pleistocene, and possibly during the entire Pleistocene, moisture occurred along a north–south gradient with the south receiving the most cloud cover and moisture due to the air-flow from the North Pacific. In the Late Pleistocene, Beringia was a mosaic of biological communities. Commencing from c.  57,000  YBP ( MIS 3), steppe–tundra vegetation dominated large parts of Beringia with

6912-512: The average water depth of the Bering Strait is 40–50 m (130–160 ft); therefore the land bridge opened when the sea level dropped more than 50 m (160 ft) below the current level. A reconstruction of the sea-level history of the region indicated that a seaway existed from c.  135,000  – c.  70,000  YBP, a land bridge from c.  70,000  – c.  60,000  YBP, an intermittent connection from c.  60,000  – c.  30,000  YBP,

7020-539: The beginning of the Holocene , some mesic habitat -adapted species left the refugium and spread westward into what had become tundra-vegetated northern Asia and eastward into northern North America. The latest emergence of the land bridge was c.  70,000 years ago. However, from c.  24,000  – c.  13,000  YBP the Laurentide Ice Sheet fused with the Cordilleran Ice Sheet , which blocked gene flow between Beringia (and Eurasia) and continental North America. The Yukon corridor opened between

7128-412: The bridge was finally inundated around 11,000 YBP. The fossil evidence from many continents points to the extinction of large animals, termed Pleistocene megafauna , near the end of the last glaciation. During the Ice Age a vast, cold and dry Mammoth steppe stretched from the arctic islands southwards to China, and from Spain eastwards across Eurasia and over the Bering land bridge into Alaska and

7236-421: The climate was warmer and wetter. The environmental conditions were not homogenous in Beringia. Recent stable isotope studies of woolly mammoth bone collagen demonstrate that western Beringia ( Siberia ) was colder and drier than eastern Beringia ( Alaska and Yukon ), which was more ecologically diverse. Grey wolves suffered a species-wide population bottleneck (reduction) approximately 25,000 YBP during

7344-426: The dry beds of the English Channel and North Sea , and the dry bed of the South China Sea linked Sumatra , Java , and Borneo to Indochina . The last glacial period , commonly referred to as the "Ice Age", spanned 125,000 –14,500   YBP and was the most recent glacial period within the current ice age , which occurred during the last years of the Pleistocene era. The Ice Age reached its peak during

7452-415: The early taxonomists. Similarly, working on museum collections where skins were often faded and in poor states of preservation, early taxonomists could not readily perceive differences in coat patterns that are consistent within a subspecies, but variable among them. Geist calls these "nuptial" characteristics: sexually selected characters that are highly conserved and diagnostic among subspecies. Towards

7560-652: The end of the 19th century, national museums began sending out biological exploration expeditions and collections accumulated. Taxonomists, usually working for the museums, began naming subspecies more rigorously, based on statistical differences in detailed cranial, dental and skeletal measurements than antlers and pelage, supplemented by better knowledge of differences in ecology and behavior. From 1898 to 1937, mammalogists named 12 new species (other than barren-ground and woodland, which had been named earlier) of caribou in Canada and Alaska, and three new species and nine new subspecies in Eurasia, each properly described according to

7668-437: The evolution" of Rangifer species and other Arctic and sub-Arctic species. Isolation of tundra-adapted species Rangifer in Last Glacial Maximum refugia during the last glacial – the Wisconsin glaciation in North America and the Weichselian glaciation in Eurasia – shaped "intraspecific genetic variability " particularly between the North American and Eurasian parts of the Arctic . Reindeer / caribou ( Rangifer ) are in

7776-495: The evolving rules of zoological nomenclature, with type localities designated and type specimens deposited in museums (see table in Species and subspecies below). In the mid-20th century, as definitions of "species" evolved, mammalogists in Europe and North America made all Rangifer species conspecific with R. tarandus , and synonymized most of the subspecies. Alexander William Francis Banfield 's often-cited A Revision of

7884-465: The extinct caribou Torontoceros [Rangifer] hypogaeus , had features (robust and short pedicles, smooth antler surface, and high position of second tine) that relate it to forest caribou. Humans started hunting reindeer in both the Mesolithic and Neolithic Periods, and humans are today the main predator in many areas. Norway and Greenland have unbroken traditions of hunting wild reindeer from

7992-540: The females. Their bez tines are set low, just above the brow tine, which is vertically flattened to protect the eyes while the buck "threshes" low brush, a courtship display. The low bez tines help the wide flat brow tines dig craters in the hard-packed tundra snow for forage, for which reason brow tines are often called "shovels" in North America and "ice tines" in Europe. The differences in antler architecture reflect fundamental differences in ecology and behavior, and in turn deep divisions in ancestry that were not apparent to

8100-770: The four western Canadian montane ecotypes are not woodland caribou: they share a common ancestor with modern barren-ground caribou / tundra reindeer, but distantly, having diverged > 60,000 years ago — before the modern ecotypes had evolved their cold- and darkness-adapted physiologies and mass-migration and aggregation behaviors (see Evolution above). Before Banfield (1961), taxonomists using cranial, dental and skeletal measurements had unequivocally allied these western montane ecotypes with barren-ground caribou, naming them (as in Osgood 1909 Murie, 1935 and Anderson 1946, among others) R. t. stonei , R. t. montanus , R. t. fortidens and R. t. osborni , respectively, and this phylogeny

8208-619: The fringes of the Rocky Mountain and Laurentide ice sheets as far south as northern Alabama ; and in Sangamonian deposits (~100,000 years BP) from western Canada. A R. t. pearyi -sized caribou occupied Greenland before and after the LGM and persisted in a relict enclave in northeastern Greenland until it went extinct about 1900 (see discussion of R. t. eogroenlandicus below). Archaeological excavations showed that larger barren-ground-sized caribou appeared in western Greenland about 4,000 years ago. The late Valerius Geist (1998) dates

8316-536: The genus Rangifer as R. grœnlandicus . It went back and forth as a full species or subspecies of the barren-ground caribou ( R. arcticus ) or a subspecies of the tundra reindeer ( R. tarandus ), but always as the Greenland reindeer / caribou. Taxonomists consistently documented morphological differences between Greenland and other caribou / reindeer in cranial measurements, dentition, antler architecture, etc. Then Banfield (1961) in his famously flawed revision, gave

8424-496: The geography of the peninsula, thereby avoiding loss of reindeer that might wander off with caribou. However, in 1997 the domesticated reindeer joined the Western Arctic Caribou Herd on their summer migration and disappeared. Cape Prince of Wales , the westernmost point on the mainland of the Americas, is on the western tip. The cape is only 51 mi (82 km) from Cape Dezhnev , the closest point on

8532-485: The larger caribou that appeared in Greenland 4,000 years ago originated from Baffin Island (itself unique; see Taxonomy above), a reconstruction of LGM glacial retreat and caribou advance (Yannic et al. 2013) shows colonization by NAL lineage caribou more likely. Their PCA and tree diagrams show Greenland caribou clustering outside of the Beringian-Eurasian lineage. The scientific name R. t. granti has

8640-649: The mountains of Norway, notably in Hardangervidda . In Sweden there are approximately 250,000 reindeer in herds managed by Sámi villages. Russia manages 19 herds of Siberian tundra reindeer ( R. t. sibiricus ) that total about 940,000. The Taimyr herd of Siberian tundra reindeer is the largest wild reindeer herd in the world, varying between 400,000 and 1,000,000; it is a metapopulation consisting of several subpopulations — some of which are phenotypically different — with different migration routes and calving areas. The Kamchatkan reindeer ( R. t. phylarchus ),

8748-550: The name groenlandicus to all the barren-ground caribou in North America, Greenland included, because groenlandicus pre-dates Richardson's R. arctus . However, because genetic data shows the Greenland caribou to be the most distantly related of any caribou to all the others (genetic distance, FST = 44%, whereas most cervid (deer family) species have a genetic distance of 2% to 5% )--as well as behavioral and morphological differences—a recent revision returned it to species status as R. groenlandicus . Although it has been assumed that

8856-487: The northern arctic areas experienced temperatures 1.5 °C (2.7 °F) degrees warmer than today but the southern sub-Arctic regions were 2 °C (4 °F) degrees cooler. During the LGM 22,000 YBP the average summer temperature was 3–5 °C (5–9 °F) degrees cooler than today, with variations of 2.9 °C (5.2 °F) degrees cooler on the Seward Peninsula to 7.5 °C (13.5 °F) cooler in

8964-626: The northern states of the contiguous United States from Maine to Washington . Boreal woodland caribou have disappeared from most of their original southern range and were designated as Threatened in 2002 by the Committee on the Status of Endangered Wildlife in Canada (COSEWIC). Environment and Climate Change Canada reported in 2011 that there were approximately 34,000 boreal woodland caribou in 51 ranges remaining in Canada (Environment Canada, 2011b), although those numbers included montane populations classified by Harding (2022) into subspecies of

9072-618: The origin of these wolves in eastern Beringia during the Middle Pleistocene . Fossil evidence also indicates an exchange of primates and plants between North America and Asia around 55.8 million years ago. 20 million years ago, evidence in North America shows the last natural interchange of mammalian species. Some, like the ancient saber-toothed cats , have a recurring geographical range: Europe, Africa, Asia, and North America. The pattern of bidirectional flow of biota has been asymmetric, with more plants, animals, and fungi generally migrating from Asia to North America than vice versa throughout

9180-415: The previously named subspecies distributions, without naming them as such, plus some ecotypes. Ecotypes are not phylogenetically based and cannot substitute for taxonomy. Meanwhile, genetic data continued to accumulate, revealing sufficiently deep divisions to easily separate Rangifer back into six previously named species and to resurrect several previously named subspecies. Molecular data showed that

9288-468: The receding ice sheets c.  13,000  YBP, and this once again allowed gene flow between Eurasia and continental North America until the land bridge was finally closed by rising sea levels c.  10,000  YBP. During the Holocene, many mesic-adapted species left the refugium and spread eastward and westward, while at the same time the forest-adapted species spread with the forests up from

9396-537: The recent revision (see Reindeer#Taxonomy below). Reindeer / caribou ( Rangifer ) vary in size from the smallest, the Svalbard reindeer ( R. ( t. ) platyrhynchus ), to the largest, Osborn's caribou ( R. t. osborni ). They also vary in coat color and antler architecture. The North American range of caribou extends from Alaska through the Yukon , the Northwest Territories and Nunavut throughout

9504-483: The recent revision): In North America, R. t. caboti , R. t. caribou , R. t. dawsoni , R. t. groenlandicus , R. t. osborni , R. t. pearyi , and R. t. terranovae ; and in Eurasia, R. t. tarandus , R. t. buskensis (called R. t. valentinae in Europe; see below), R. t. phylarchus , R. t. pearsoni , R. t. sibiricus and R. t. platyrhynchus . These subspecies were retained in the 2011 replacement work Handbook of

9612-495: The respective Siberian and North American portions of Beringia has led to the 'Beringian Gap' hypothesis, wherein an unconfirmed geographic factor blocked migration across the land bridge when it emerged. Beringia did not block the movement of most dry steppe-adapted large species such as saiga antelope, woolly mammoth, and caballid horses. Notable restricted fauna include the woolly rhino in Siberia (which went no further east than

9720-481: The result of an underground steam explosion. The Killeak Lakes and White Fish Lake are also volcanic maar lakes of notable size on the northern Seward Peninsula. Four mountain ranges line the southern side of the peninsula, the most prominent being the Kigluaik (or Sawtooth) Mountains. The highest point in the range and the peninsula is the peak of 4,714 ft (1,437 m) Mount Osborn . Other mountain ranges on

9828-541: The sea floors of many interglacial shallow seas were exposed, including those of the Bering Strait , the Chukchi Sea to the north, and the Bering Sea to the south. Other land bridges around the world have emerged and disappeared in the same way. Around 14,000 years ago, mainland Australia was linked to both New Guinea and Tasmania , the British Isles became an extension of continental Europe via

9936-433: The smallest, the Svalbard reindeer ( R. ( t. ) platyrhynchus ), to the largest, Osborn's caribou ( R. t. osborni ). Although reindeer are quite numerous, some species and subspecies are in decline and considered vulnerable . They are unique among deer (Cervidae) in that females may have antlers , although the prevalence of antlered females varies by subspecies. Reindeer are the only successfully semi-domesticated deer on

10044-728: The south by the tip of the Kamchatka Peninsula . It includes the Chukchi Sea , the Bering Sea , the Bering Strait , the Chukchi and Kamchatka Peninsulas in Russia as well as Alaska in the United States and the Yukon in Canada . The area includes land lying on the North American Plate and Siberian land east of the Chersky Range . At various times, it formed a land bridge referred to as

10152-693: The south have provided insight into the timeline of prehistorical migrations from Asia to the Seward Peninsula. Most of the peninsula is in the Nome Census Area , but part is in the Northwest Arctic Borough . These are the communities on the Seward Peninsula, with 2005 state population estimates: Other locations on the Seward Peninsula include the mining towns of Council , Solomon , Candle, Haycock and Taylor. While still frequented by locals of neighboring communities, there are no longer year round residents in these locations. There

10260-607: The south. The arid-adapted species were reduced to minor habitats or became extinct. Beringia constantly transformed its ecosystem as the changing climate affected the environment, determining which plants and animals were able to survive. The land mass could be a barrier as well as a bridge: during colder periods, glaciers advanced and precipitation levels dropped. During warmer intervals, clouds, rain and snow altered soils and drainage patterns. Fossil remains show that spruce , birch and poplar once grew beyond their northernmost range today, indicating that there were periods when

10368-498: The subfamily Odocoileinae , along with roe deer ( Capreolus ), Eurasian elk / moose ( Alces ), and water deer ( Hydropotes ). These antlered cervids split from the horned ruminants Bos (cattle and yaks), Ovis (sheep) and Capra (goats) about 36 million years ago. The Eurasian clade of Odocoileinae (Capreolini, Hydropotini and Alcini) split from the New World tribes of Capreolinae ( Odocoileini and Rangiferini) in

10476-430: The subspecies level; that Finnish forest reindeer ( R. t. fennicus ) clustered well apart from both wild and domestic tundra reindeer and that boreal woodland caribou ( R. t. caribou ) were separable from all others. Meanwhile, archaeological evidence was accumulating that Eurasian forest reindeer descended from an extinct forest-adapted reindeer and not from tundra reindeer (see Evolution above); since they do not share

10584-626: The terms reindeer and caribou for essentially the same animal can cause confusion, but the ICUN clearly delineates the issue: "Reindeer is the European name for the species of Rangifer, while in North America, Rangifer species are known as Caribou." The word reindeer is an anglicized version of the Old Norse words hreinn ("reindeer") and dýr ("animal") and has nothing to do with reins. The word caribou comes through French, from

10692-499: The tundra and into the taiga. Eurasian mountain reindeer ( R. t. tarandus ) are close to North American caribou genetically and visually, but with sufficient differences to warrant division into two species. The unique, insular Svalbard reindeer inhabits the Svalbard Archipelago . The Finnish forest reindeer ( R. t. fennicus ) is spottily distributed in the coniferous forest zones from Finland to east of Lake Baikal :

10800-947: The uniformly dark, small-maned type with the frontally emphasized, flat-beamed antlers", is "scattered thinly along the southern rim of North American caribou distribution". He affirms that the "true woodland caribou is very rare, in very great difficulties and requires the most urgent of attention." In 2011, noting that the former classifications of Rangifer tarandus , either with prevailing taxonomy on subspecies, designations based on ecotypes , or natural population groupings, failed to capture "the variability of caribou across their range in Canada" needed for effective subspecies conservation and management, COSEWIC developed Designatable Unit (DU) attribution, an adaptation of "evolutionary significant units". The 12 designatable units for caribou in Canada (that is, excluding Alaska and Greenland) based on ecology, behavior and, importantly, genetics (but excluding morphology and archaeology) essentially followed

10908-494: The various forms of the Woodland Caribou are so great in almost every respect that no detailed comparison is necessary. ...According to Mr. Stone, Rangifer granti inhabits the " barren land of Alaska Peninsula, ranging well up into the mountains in summer, but descending to the lower levels in winter, generally feeding on the low flat lands near the coast and in the foothills...As regards cranial characters no comparison

11016-525: The west to the Mackenzie River in the east. The distribution of plants in the genera Erythranthe and Pinus are good examples of this, as very similar genera members are found in Asia and the Americas. During the Pleistocene epoch, global cooling led periodically to the expansion of glaciers and the lowering of sea levels. This created land connections in various regions around the globe. Today,

11124-463: Was "not able to find diagnostic features that could segregate this form from the western barren ground type." But Skal 1982 had included specimens from the eastern end of the Alaska Peninsula and the Kenai Peninsula , the range of the larger Stone's caribou. Later, geneticists comparing barren-ground caribou of Alaska with those of mainland Canada found little difference and they all became

11232-604: Was a United States Coast Guard LORAN station at Port Clarence . The U.S. Air Force operates a radar station at the "Tin City" site, 7 mi (11 km) southeast of Wales. The Seward Peninsula has several distinct geologic features. The Devil Mountain Lakes on the northern portion of the peninsula are the largest maar lakes in the world and part of the Espenberg volcanic field . They were formed over 21,000 years ago as

11340-920: Was confirmed by genetic analysis. DNA also revealed three unnamed clades that, based on genetic distance, genetic divergence and shared vs. private haplotypes and alleles , together with ecological and behavioral differences, may justify separation at the subspecies level: the Atlantic- Gaspésie caribou (COSEWIC DU11), an eastern montane ecotype of the boreal woodland caribou, and the Baffin Island caribou. Neither one of these clades has yet been formally described or named. Jenkins et al. (2012) said that "[Baffin Island] caribou are unique compared to other Barrenground herds, as they do not overwinter in forested habitat, nor do all caribou undertake long seasonal migrations to calving areas." It also shares

11448-512: Was found in both Mongolia and western North America. Relatives of Troodon , Triceratops , and Tyrannosaurus rex all came from Asia. The earliest Canis lupus specimen was a fossil tooth discovered at Old Crow, Yukon , Canada. The specimen was found in sediment dated 1 million YBP, however the geological attribution of this sediment is questioned. Slightly younger specimens were discovered at Cripple Creek Sump, Fairbanks , Alaska, in strata dated 810,000 YBP. Both discoveries point to

11556-406: Was later replaced by R. constantini , which was adapted for grasslands, in a second immigration 19,000–20,000 years ago when the LGM turned its forest habitats into tundra, while fennicus survived in isolation in southwestern Europe. R. constantini was then replaced by modern tundra / barren-ground caribou adapted to extreme cold, probably in Beringia, before dispersing west ( R. t. tarandus in

11664-521: Was selected as the senior synonym to R. t. valentinae Flerov, 1933, in Mammal Species of the World but Russian authors do not recognize Millais and Millais' articles in a hunting travelogue, The Gun at Home and Abroad , seem short of a taxonomic authority. The scientific name groenlandicus is fraught with problems. Edwards (1743) illustrated and claimed to have seen a male specimen ("head of perfect horns...") from Greenland and said that

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