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105-554: Founded in 1872 as a private concern by Anton Dohrn , in 1982 the Stazione Zoologica came under the supervision and control of the Ministero dell'Università e della Ricerca Scientifica e Tecnologica (Ministry of Universities and Scientific and Technological Research) as a National Institute. Dohrn's idea was to establish an international scientific community provided with laboratory space, equipment, research material and

210-442: A common ancestor is now generally accepted and considered a fundamental scientific concept. In a joint publication with Alfred Russel Wallace , he introduced his scientific theory that this branching pattern of evolution resulted from a process he called natural selection , in which the struggle for existence has a similar effect to the artificial selection involved in selective breeding . Darwin has been described as one of

315-451: A genealogical branching of a single evolutionary tree , in which "It is absurd to talk of one animal being higher than another", thereby discarding Lamarck's idea of independent lineages progressing to higher forms. While developing this intensive study of transmutation, Darwin became mired in more work. Still rewriting his Journal , he took on editing and publishing the expert reports on his collections, and with Henslow's help obtained

420-562: A 230-page "Essay", to be expanded with his research results if he died prematurely. In November, the anonymously published sensational best-seller Vestiges of the Natural History of Creation brought wide interest in transmutation. Darwin scorned its amateurish geology and zoology, but carefully reviewed his own arguments. Controversy erupted, and it continued to sell well despite contemptuous dismissal by scientists. Darwin completed his third geological book in 1846. He now renewed

525-845: A Bachelor of Arts degree as the first step towards becoming an Anglican country parson . Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying. During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens ' Illustrations of British entomology (1829–1932). Through Fox, Darwin became

630-673: A Treasury grant of £1,000 to sponsor this multi-volume Zoology of the Voyage of H.M.S. Beagle , a sum equivalent to about £115,000 in 2021. He stretched the funding to include his planned books on geology, and agreed to unrealistic dates with the publisher. As the Victorian era began, Darwin pressed on with writing his Journal , and in August 1837 began correcting printer's proofs . As Darwin worked under pressure, his health suffered. On 20 September, he had "an uncomfortable palpitation of

735-591: A beginning to each crop of species". Darwin's book The Structure and Distribution of Coral Reefs on his theory of atoll formation was published in May 1842 after more than three years of work, and he then wrote his first "pencil sketch" of his theory of natural selection. To escape the pressures of London, the family moved to rural Down House in Kent in September. On 11 January 1844, Darwin mentioned his theorising to

840-547: A book on geology. When they reached Brazil, Darwin was delighted by the tropical forest , but detested the sight of slavery there , and disputed this issue with FitzRoy. The survey continued to the south in Patagonia . They stopped at Bahía Blanca , and in cliffs near Punta Alta Darwin made a major find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like

945-718: A break and went "geologising" in Scotland. He visited Glen Roy in glorious weather to see the parallel "roads" cut into the hillsides at three heights. He later published his view that these were marine-raised beaches, but then had to accept that they were shorelines of a proglacial lake . Fully recuperated, he returned to Shrewsbury in July 1838. Used to jotting down daily notes on animal breeding, he scrawled rambling thoughts about marriage, career and prospects on two scraps of paper, one with columns headed "Marry" and "Not Marry" . Advantages under "Marry" included "constant companion and

1050-471: A calm spell. On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell 's Principles of Geology , which set out uniformitarian concepts of land slowly rising or falling over immense periods, and Darwin saw things Lyell's way, theorising and thinking of writing

1155-562: A close friend and follower of botany professor John Stevens Henslow . He met other leading parson-naturalists who saw scientific work as religious natural theology , becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley 's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for

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1260-412: A copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during

1365-596: A fascination and expertise in marine invertebrates, dating back to his student days with Grant, by dissecting and classifying the barnacles he had collected on the voyage, enjoying observing beautiful structures and thinking about comparisons with allied structures. In 1847, Hooker read the "Essay" and sent notes that provided Darwin with the calm critical feedback that he needed, but would not commit himself and questioned Darwin's opposition to continuing acts of creation . In an attempt to improve his chronic ill health, Darwin went in 1849 to Dr. James Gully 's Malvern spa and

1470-529: A fortnight mapping strata in Wales. After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth . He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS  Beagle with captain Robert FitzRoy , a position for a gentleman rather than "a mere collector". The ship

1575-532: A freed black slave who had accompanied Charles Waterton in the South American rainforest . In Darwin's second year at the university, he joined the Plinian Society , a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant 's investigations of

1680-473: A friend in old age ... better than a dog anyhow", against points such as "less money for books" and "terrible loss of time". Having decided in favour of marriage, he discussed it with his father, then went to visit his cousin Emma on 29 July. At this time he did not get around to proposing, but against his father's advice, he mentioned his ideas on transmutation. He married Emma on 29 January 1839 and they were

1785-530: A giant capybara . The armour fragments were actually from Glyptodon , a huge armadillo-like creature, as Darwin had initially thought. These extinct creatures were related to living species in South America. In mid-December, Darwin took lodgings in Cambridge to arrange expert classification of his collections, and prepare his own research for publication. Questions of how to combine his diary into

1890-556: A giant version of the armour on local armadillos . From a jaw and tooth he identified the gigantic Megatherium , then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest. In Patagonia, Darwin came to wrongly believe the territory was devoid of reptiles. On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political and anthropological insights into both native and colonial people at

1995-506: A great name among the Naturalists of Europe". On 2 October 1836, Beagle anchored at Falmouth , Cornwall. Darwin promptly made the long coach journey to Shrewsbury to visit his home and see relatives. He then hurried to Cambridge to see Henslow, who advised him on finding available naturalists to catalogue Darwin's animal collections and to take on the botanical specimens. Darwin's father organised investments, enabling his son to be

2100-581: A library. This was supported and funded by the German Government , Thomas Henry Huxley , Charles Darwin , Francis Balfour and Charles Lyell among others. Dohrn provided a substantial sum himself. Running costs were paid from income derived from the bench system , the sale of scientific journals and specimens and the income from the public aquarium. This system was an important innovations in management of research and it worked. When Anton Dohrn died in 1909 more than 2,200 scientists from Europe and

2205-467: A marine zoological station (one was later established as Observatoire Oceanologique de Villefranche ). The use of marine organisms as models for study had been promoted in Germany by Johannes Peter Müller (1801-1858). The Straits of Messina were known for their richness and had been studied by Spallanzani and August D. Krohn in the past. In Messina, Dohrn and Miklouho-Maclay conceived a plan to cover

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2310-515: A minor role to natural selection, and it was not until the emergence of the modern evolutionary synthesis from the 1930s to the 1950s that a broad consensus developed in which natural selection was the basic mechanism of evolution. Darwin's scientific discovery is the unifying theory of the life sciences , explaining the diversity of life . Darwin was born in Shrewsbury , Shropshire, on 12 February 1809, at his family's home, The Mount . He

2415-514: A paper on fossil insects Zur Kenntniss der Insecten in den Primärformationen . In this he describes Eugereon boeckingi ( Palaeodictyoptera ). He became a privatdozent at Jena in 1868. Dohrn's ideas of zoology changed in the summer 1862 when he returned to study at Jena , where Ernst Haeckel introduced him to Darwin 's work and theories. Dohrn became a fervent defender of Darwin's theory of evolution by natural selection. He saw comparative anatomy, physiology and embryology as means to determine

2520-597: A principle of succession of functions in 1875 on how one organ could become the basis for the evolution of another of an entirely different function. He was an elected International Member of the American Philosophical Society . The institution transitioned from a private one to a public Italian organization through the subsequent management by his son Reinhard Dohrn . Dohrn was born in Stettin (Szczecin) , Prussian Province of Pomerania , into

2625-539: A programmer of laws. Darwin stayed with his freethinking brother Erasmus, part of this Whig circle and a close friend of the writer Harriet Martineau , who promoted the Malthusianism that underpinned the controversial Whig Poor Law reforms to stop welfare from causing overpopulation and more poverty. As a Unitarian, she welcomed the radical implications of transmutation of species , promoted by Grant and younger surgeons influenced by Geoffroy . Transmutation

2730-543: A research institute of international importance in a large university town that itself had a strong international element. After a visiting a newly opened aquarium in Berlin, the Berliner Aquarium Unter den Linden he decided that charging the general public to visit an aquarium might earn the laboratory enough money to pay a salary for a permanent assistant. Naples, with a population of 500,000 inhabitants,

2835-591: A sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs round them grew to form atolls . On the geologically new Galápagos Islands , Darwin looked for evidence attaching wildlife to an older "centre of creation", and found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia,

2940-517: A self-funded gentleman scientist , and an excited Darwin went around the London institutions being fêted and seeking experts to describe the collections. British zoologists at the time had a huge backlog of work, due to natural history collecting being encouraged throughout the British Empire, and there was a danger of specimens just being left in storage. Charles Lyell eagerly met Darwin for

3045-579: A series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. Darwin published his theory of evolution with compelling evidence in his 1859 book On the Origin of Species . By the 1870s, the scientific community and a majority of the educated public had accepted evolution as a fact . However, many initially favoured competing explanations that gave only

3150-719: A similarity between farmers picking the best stock in selective breeding, and a Malthusian Nature selecting from chance variants so that "every part of newly acquired structure is fully practical and perfected", thinking this comparison "a beautiful part of my theory". He later called his theory natural selection , an analogy with what he termed the "artificial selection" of selective breeding. On 11 November, he returned to Maer and proposed to Emma, once more telling her his ideas. She accepted, then in exchanges of loving letters she showed how she valued his openness in sharing their differences, while expressing her strong Unitarian beliefs and concerns that his honest doubts might separate them in

3255-468: A time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its view of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species. Three Fuegians on board, who had been seized during

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3360-427: A wealthy middle-class family. His grandfather, Heinrich Dohrn, had been a wine and spice merchant , and had made the family fortune by trading in sugar and a director of the sugar refinery Pommersche Provinzial Zuckersiederei. This wealth and advice from Alexander von Humboldt , allowed Anton's father, Carl August (1806-1892), to devote himself to his various hobbies; travelling, folk music and insects. Anton,

3465-606: A zoologist. Dohrn received an early education which was based on humanist ideals. He went to study medicine and zoology and Koenigsberg, Bonn, Jena and Berlin. was initially interested in Hemiptera . In 1859, he published Beitrag zur Kenntniss der Harpactoridae in Entomologische Zeitung and the more important Catalogus hemipterorum . He gained his doctorate from Breslau in November 1865 with his thesis "On

3570-471: Is known as a Malthusian catastrophe . Darwin was well-prepared to compare this to Augustin de Candolle 's "warring of the species" of plants and the struggle for existence among wildlife, explaining how numbers of a species kept roughly stable. As species always breed beyond available resources, favourable variations would make organisms better at surviving and passing the variations on to their offspring, while unfavourable variations would be lost. He wrote that

3675-602: Is still unrivalled in Europe today. The first librarian was Emil Schoebel. The station maintained a high level of technical services. Ernst Abbe (1840–1905) of the Zeiss factory, a close friend, supplied sets of Zeiss instruments at low prices, thus bringing Zeiss equipment, sometimes improved, to the attention of the international scientific community. Assistants and guests collaborated in improving section-cutting and staining. For collecting there were several crewed boats, including

3780-913: The Fisheries Research Services Marine Laboratory ( Aberdeen , 1899), and the Bangor Marine Station ( Queen's University of Belfast , 1903). Dohrn's name has been immortalised in an undersea feature, the Anton Dohrn Seamount , a seamount in the Rockall Trough , to the north-west of the British Isles, which has become known for the great biodiversity which lives on the cold-water coral, Lophelia pertusa , in this region. The Carnegie Institution's Department of Marine Biology laboratory at Dry Tortugas , Florida placed

3885-933: The Hopkins Marine Station , opened in California . In Britain, current marine laboratories that originate from this time include the Dunstaffnage Marine Station (today Scottish Association for Marine Science , 1884), the Gatty Marine Laboratory ( University of St Andrews , 1884), the Marine Biological Association of the United Kingdom ( Plymouth , 1888), the Dove Marine Laboratory ( Newcastle University , 1897),

3990-686: The Linnean Society of London . Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex , followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in

4095-554: The Narrative were resolved at the end of the month when FitzRoy accepted Broderip's advice to make it a separate volume, and Darwin began work on his Journal and Remarks . Darwin's first paper showed that the South American landmass was slowly rising, and with Lyell's enthusiastic backing he read it to the Geological Society of London on 4 January 1837. On the same day, he presented his mammal and bird specimens to

4200-546: The Royal Society 's Royal Medal, and it made his reputation as a biologist . Upon the conclusion of his research, Darwin declared "I hate a barnacle as no man ever did before." In 1854, he became a Fellow of the Linnean Society of London , gaining postal access to its library. He began a major reassessment of his theory of species, and in November realised that divergence in the character of descendants could be explained by them becoming adapted to "diversified places in

4305-942: The Zoological Society . The ornithologist John Gould soon announced that the Galápagos birds that Darwin had thought a mixture of blackbirds , " gros-beaks " and finches , were, in fact, twelve separate species of finches . On 17 February, Darwin was elected to the Council of the Geological Society, and Lyell's presidential address presented Owen's findings on Darwin's fossils, stressing geographical continuity of species as supporting his uniformitarian ideas. Early in March, Darwin moved to London to be near this work, joining Lyell's social circle of scientists and experts such as Charles Babbage , who described God as

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4410-521: The first Beagle voyage then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there

4515-519: The marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aborigines "good-humoured & pleasant", their numbers depleted by European settlement. FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed, and the survey supported Darwin's theorising. FitzRoy began writing

4620-653: The motor vessel Anton Dohrn in service in July 1911 for ocean science work. The vessel served in the United States Navy as the patrol vessel Anton Dohrn (SP-1086) from 1917 to 1919. Charles Darwin Charles Robert Darwin ( / ˈ d ɑːr w ɪ n / DAR -win ; 12 February 1809 – 19 April 1882) was an English naturalist , geologist , and biologist , widely known for his contributions to evolutionary biology . His proposition that all species of life have descended from

4725-598: The ordinary degree. Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature , explaining adaptation as God acting through laws of nature . He read John Herschel 's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described

4830-455: The "Stazione Zoologica di Messina", but quickly realized the technical difficulties of studying marine life without a permanent structure and support facilities, such as trained personnel and a library. In 1870 Dohrn decided that Naples would be a better place for his Station. This choice was due to the greater biological richness of the Gulf of Naples and also to the possibility of starting

4935-596: The "final cause of all this wedging, must be to sort out proper structure, & adapt it to changes", so that "One may say there is a force like a hundred thousand wedges trying force into every kind of adapted structure into the gaps of in the economy of nature, or rather forming gaps by thrusting out weaker ones." This would result in the formation of new species. As he later wrote in his Autobiography : In October 1838, that is, fifteen months after I had begun my systematic enquiry, I happened to read for amusement Malthus on Population, and being well prepared to appreciate

5040-716: The 1886 addition. The first assistants were zoologists Nicolaus Kleinenberg and Hugo Eisig and one of the Preparators was Salvatore Lobianco (Lo Bianco) (1860–1910), who wrote The Methods Employed at the Naples Zoological Station for the Preservation of Marine Animals . Others were Dr. Brandt (librarian); Dr. Lang; Dr. Giesbrecht; Petersen (engineer). By 1910 the permanent staff were Professor Dr. Paul Mayer and Dr. Gross, morphology; Dr. Burian, comparative physiology; Dr. Henze, chemistry; Dr. Gast,

5145-725: The Anatomy of Hemiptera". He visited Heligoland along with Ernst Haeckel to examine marine organisms for the first time. Haeckel talked about a zoological station at this time. In 1867 Dohrn visited Millport , Isle of Cumbrae as a guest of David Robertson (1806-1896) who had founded the Millport Biological Station. During this period he became associated with the English scientific establishment through his father's friendship with Henry Tibbats Stainton . He also met Thomas Henry Huxley (1825-1895). In 1866, he published

5250-765: The British Association at Liverpool steps have been taken by Dr. Dohrn to secure the moral assistance of some other scientific bodies, and that the Academy of Belgium has passed a vote acknowledging the great value of the proposed Observatories. Besides this, the Government at Berlin has given instruction to the German Embassy at Florence and to the General Consul at Naples to do everything to secure success to Dr. Dohrn's enterprise. Next October

5355-676: The Stazione Zoologica, and the new way of thinking and funding research are the main legacies of Dohrn. The model was copied a number of times throughout the world. In 1878 Johns Hopkins University founded the Chesapeake Zoological Laboratory, under the direction of W.K. Brooks . Then, in 1888, the Marine Biological Laboratory was founded at Woods Hole and in 1892 the first laboratory on the United States West Coast ,

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5460-611: The Stazione for one year where he or she would find available all that was required to conduct research (laboratory space, animal supply, chemicals, an exceptional library and expert staff). He contributed his own library and obtained donations of books from publishers and authors, including Darwin. These facilities were supplied with no strings attached, in the sense that investigators were completely free to pursue their own projects and ideas. This system worked so that when Anton Dohrn died in 1909 more than 2,200 scientists from Europe and

5565-533: The United States had worked at Naples and more than 50 tables-per-year had been rented out. It might be said that international scientific collaboration in the modern sense was born at the Stazione, based on quick and free communication of ideas, methods and results. Dohrn suffered from heart problems in his last years and died in Munich on September 26, 1909. He was buried on October 3 at Jena. The success of

5670-580: The United States had worked at Stazione Zoologica and more than 50 tables-per-year had been rented out. "Report of the Committee, consisting of Dr. Anton Dohrn, Professor Rolleston and Mr. P. L. Sclater, appointed for the purpose of promoting the Foundation of Zoological Stations in different parts of the World:—Reporter, Dr. Dohrn [Jena]."-"The Committee beg to report that since the last Meeting of

5775-618: The Wedgwoods were adopting Anglicanism . Robert Darwin, a freethinker , had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury , but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had a taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending

5880-459: The afterlife. While he was house-hunting in London, bouts of illness continued and Emma wrote urging him to get some rest, almost prophetically remarking "So don't be ill any more my dear Charley till I can be with you to nurse you." He found what they called "Macaw Cottage" (because of its gaudy interiors) in Gower Street , then moved his "museum" in over Christmas. On 24 January 1839, Darwin

5985-506: The anatomy and life cycle of marine invertebrates in the Firth of Forth , and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech . One day, Grant praised Lamarck 's evolutionary ideas . Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin

6090-451: The biota of Mediterranean (in 1876 Anton Dohrn added a section of Botany), Charles Darwin had advised Dohrn that establishing a library would be unwise (Groeben, 1982, p. 29). Dohrn argued that availability of all the major published sources was essential. He gave his own books and scientific journals to the Station and persuaded publishers and scientists to donate their publications. The Naples Station's biological reference collection

6195-437: The botanist Joseph Dalton Hooker , writing with melodramatic humour "it is like confessing a murder". Hooker replied, "There may, in my opinion, have been a series of productions on different spots, & also a gradual change of species. I shall be delighted to hear how you think that this change may have taken place, as no presently conceived opinions satisfy me on the subject." By July, Darwin had expanded his "sketch" into

6300-429: The building at Naples will be commenced under the personal superintendence of Dr. Dohrn, who will be accompanied by the assistant architect of the Berlin Aquarium. The contractors agree to finish the building in one year, so that in January 1873 the Aquarium in Naples may be expected to be in working order." British Association for the Advancement of Science Report on the 1871 Meeting in Edinburgh The oldest building of

6405-429: The comparative anatomist St. George Mivart . Dohrn's ideas on functional phylogenetics underlie some of the embryological hypotheses such as the origin of the vertebrate jaw from gill arches. In his original writings he claimed that gill slits had changed into a variety of vertebrate organs. From 1850 to 1852, the zoologist and geologist Carl Vogt had lived in Nice where he tried unsuccessfully to enlist support for

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6510-569: The directorship of Antonio Miralto. In 1987 Gaetano Salvatore, dean of the Medical School of Università Federico II in Napoli, was appointed president of the Stazione Zoologica. After his death in 1997, Professor Giorgio Bernardi was appointed as president. He launched the study of molecular evolution "at the institutional level" completing Dohrn's vision. 40°49′58″N 14°14′09″E  /  40.8327°N 14.2358°E  / 40.8327; 14.2358 Anton Dohrn Felix Anton Dohrn FRS FRSE (29 December 1840 – 26 September 1909)

6615-490: The duties of Secretary of the Geological Society. After initially declining the work, he accepted the post in March 1838. Despite the grind of writing and editing the Beagle reports, Darwin made remarkable progress on transmutation, taking every opportunity to question expert naturalists and, unconventionally, people with practical experience in selective breeding such as farmers and pigeon fanciers . Over time, his research drew on information from his relatives and children,

6720-434: The evolutionary history of various organisms. At that time comparative embryology was the keystone of morphological evolutionary studies, based on Haeckel's recapitulation theory , the idea that an organism during its embryonic development passes through the major stages of the evolutionary past of its species . Morphology became one of the major ways in which zoologists sought to expand and develop Darwinian theory in

6825-713: The family butler, neighbours, colonists and former shipmates. He included mankind in his speculations from the outset, and on seeing an orangutan in the zoo on 28 March 1838 noted its childlike behaviour. The strain took a toll, and by June he was being laid up for days on end with stomach problems, headaches and heart symptoms. For the rest of his life, he was repeatedly incapacitated with episodes of stomach pains, vomiting, severe boils , palpitations, trembling and other symptoms, particularly during times of stress, such as attending meetings or making social visits. The cause of Darwin's illness remained unknown, and attempts at treatment had only ephemeral success. On 23 June, he took

6930-467: The first time on 29 October and soon introduced him to the up-and-coming anatomist Richard Owen , who had the facilities of the Royal College of Surgeons to work on the fossil bones collected by Darwin. Owen's surprising results included other gigantic extinct ground sloths as well as the Megatherium Darwin had identified, a near complete skeleton of the unknown Scelidotherium and a hippopotamus -sized rodent -like skull named Toxodon resembling

7035-399: The globe with a network of zoological research stations, analogous to railway stations, where scientists could stop, collect material, make observations and conduct experiments, before moving on to the next station. In 1859-60 Haeckel and his students described 144 new species of Radiolaria collected in the waters. For a while Messina came to be called the "Mecca of the German Privat-Dozent." It

7140-520: The heart", so his doctors urged him to "knock off all work" and live in the country for a few weeks. After visiting Shrewsbury, he joined his Wedgwood relatives at Maer Hall , Staffordshire, but found them too eager for tales of his travels to give him much rest. His charming, intelligent, and cultured cousin Emma Wedgwood , nine months older than Darwin, was nursing his invalid aunt. His uncle Josiah pointed out an area of ground where cinders had disappeared under loam and suggested that this might have been

7245-472: The highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt 's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick 's geology course, then on 4 August travelled with him to spend

7350-405: The international status of the Stazione and to guarantee its economic and hence political independence and freedom of research , Dohrn introduced a series of innovative measures to finance his project. Firstly, the rental of work and research space (the "Bench system"), for an annual fee universities, governments, scientific institutions, private foundations or individuals could send one scientist to

7455-405: The latter years of the 19th century. Dohrn chose to become a "Darwinian morphologist". Dohrn received his doctorate in 1865 at Breslau under Eduard Grube , and his Habilitation in 1868 at Jena with Rudolf Virchow , Ernst Haeckel and Carl Gegenbaur . The study subjects were medicine and zoology , and his Jena monograph was Studien zur Embryologie der Arthropoden . From 1868 to 1870, he

7560-538: The most influential figures in human history and was honoured by burial in Westminster Abbey . Darwin's early interest in nature led him to neglect his medical education at the University of Edinburgh ; instead, he helped to investigate marine invertebrates . His studies at the University of Cambridge 's Christ's College from 1828 to 1831 encouraged his passion for natural science . However, it

7665-429: The museum; Hermann Linden, secretary; Sig. Santorelli, Preparator. Zoologists and morphologists were the first guests of the new Institute. Included were Heinrich Wilhelm Gottfried von Waldeyer-Hartz , Francis Balfour , Ray Lankester , August Weismann , Giovanni Battista Grassi , Antonio della Valle , Oskar Schmidt , Ambrosius Hubrecht (Professor of Utrecht University, an embryologist). In 1897 Ida Henrietta Hyde

7770-503: The nearby Anglican Shrewsbury School as a boarder . Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat the poor of Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone ,

7875-485: The official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history. In Cape Town , South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries,

7980-496: The origin of species". Without telling Darwin, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society , and reported in magazines, including The Athenaeum . Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have

8085-522: The parents of ten children, seven of whom survived to adulthood. Continuing his research in London, Darwin's wide reading now included the sixth edition of Malthus 's An Essay on the Principle of Population . On 28 September 1838, he noted its assertion that human "population, when unchecked, goes on doubling itself every twenty-five years, or increases in a geometrical ratio", a geometric progression so that population soon exceeds food supply in what

8190-573: The possibility that "one species does change into another" to explain the geographical distribution of living species such as the rheas, and extinct ones such as the strange extinct mammal Macrauchenia , which resembled a giant guanaco , a llama relative. Around mid-July, he recorded in his "B" notebook his thoughts on lifespan and variation across generations – explaining the variations he had observed in Galápagos tortoises , mockingbirds, and rheas. He sketched branching descent, and then

8295-520: The replacement of extinct species by others" as "a natural in contradistinction to a miraculous process". When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on

8400-489: The station to visiting scientists in September 1873, and to the general public in January 1874. In 1875 Dohrn published Der Ursprung der Wirbelthiere und das Princip des Functionswechsels: Genealogische Skizzen which proposed the "change of function" theory of the origin of vertebrates. He suggested that an organ could have secondary functions in addition to their primary function and that the subsidiary functions could become primary over evolutionary time. In order to promote

8505-747: The steamers "Johannes Muller" and "Francis Balfour". An engineer and assisting machinists maintained the aquarium and a trained mechanic made instruments for experimental investigations. The aquarium was constructed by William Alford Lloyd [1] . Dohrn had met Lloyd in 1866 in Hamburg . Anton Dohrn followed Goethe in considering arts and science inseparable. He was devoted to music and in 1873 employed Hans von Marées (1837–1887) and Adolf von Hildebrand to enhance Stazione Zoologica with art works. Anton Dohrn's son Reinhard Dohrn (1880–1962) continued his father's work from 1909. Peter Dohrn (1917–2007) acted as director between 1954 and 1967. From 1967 to 1976 SZN

8610-408: The struggle for existence which everywhere goes on from long-continued observation of the habits of animals and plants, it at once struck me that under these circumstances favourable variations would tend to be preserved, and unfavourable ones to be destroyed. The result of this would be the formation of new species. Here, then, I had at last got a theory by which to work... By mid-December, Darwin saw

8715-409: The systematics of the taxa. He wrote his first examination of the species in 1846, but did not formally describe it until 1854. FitzRoy's long delayed Narrative was published in May 1839. Darwin's Journal and Remarks got good reviews as the third volume, and on 15 August it was published on its own. Early in 1842, Darwin wrote about his ideas to Charles Lyell, who noted that his ally "denies seeing

8820-420: The voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations, and at intervals during the voyage his specimens were sent to Cambridge together with letters including

8925-421: The voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting the immediate joint submission of both their theories to

9030-403: The work of earthworms , inspiring "a new & important theory" on their role in soil formation , which Darwin presented at the Geological Society on 1 November 1837. His Journal was printed and ready for publication by the end of February 1838, as was the first volume of the Narrative , but FitzRoy was still working hard to finish his own volume. William Whewell pushed Darwin to take on

9135-454: The youngest son, read zoology and medicine at various German universities ( Königsberg , Bonn , Jena and Berlin ). The attitudes of his father, a liberal one with interest in people, literature, music ( Felix Mendelssohn was his godfather ) and science was to have a key role in Dohrn's own work as a promoter of scientific collaboration. His brother Heinrich Wolfgang Ludwig Dohrn was also

9240-426: The zoological station was opened in 1874. A second building connected to the western end of the first by bridges was added in 1886 and a third was built in 1906 for the new science of comparative physiology. In 1910 the 1874 building was occupied by the public aquarium and the library only, the department for collecting and preserving organisms as well as the individual laboratories for zoologists having been relocated in

9345-707: Was elected a Fellow of the Royal Society (FRS). On 29 January, Darwin and Emma Wedgwood were married at Maer in an Anglican ceremony arranged to suit the Unitarians, then immediately caught the train to London and their new home. Darwin now had the framework of his theory of natural selection "by which to work", as his "prime hobby". His research included extensive experimental selective breeding of plants and animals, finding evidence that species were not fixed and investigating many detailed ideas to refine and substantiate his theory. For fifteen years this work

9450-510: Was Miklouho-Maclay who introduced Dohrn to the Russian-Polish de Baranowski family. In 1874 Dohrn married their eldest daughter, sixteen-year-old Maria Baranowska. They had four children Boguslaw, Wolfgang, Harald and Reinhard. Due to her linguistic abilities, Maria became a successful translator . Dohrn realised how useful it would be for scientists to arrive at a location and find a ready to use laboratory. Dohrn rented two rooms for

9555-480: Was a Docent in zoology at Jena. During these times, he worked several times at facilities located by the sea: Heligoland alongside Haeckel in 1865, Hamburg in 1866, Millport , Scotland with David Robertson in 1867 and 1868 and moved to Messina , Italy, during the winter of 1868 together with his friend and colleague Nicholas Miklouho-Maclay to work on the marine life of the Straits of Messina . In 1870 Dohrn

9660-418: Was a " wren " was in the finch group . Darwin had not labelled the finches by island, but from the notes of others on the ship, including FitzRoy, he allocated species to islands. The two rheas were distinct species, and on 14 March Darwin announced how their distribution changed going southwards. By mid-March 1837, barely six months after his return to England, Darwin was speculating in his Red Notebook on

9765-537: Was a prominent German Darwinist and the founder and first director of the first marine-biological and zoological research station in the world, the Stazione Zoologica in Naples , Italy. The institution became a centre for the exchange of biological ideas through the network of visitors from around the world. He worked on embryology and examined vertebrate origins in terms of functional phylogeny and proposed

9870-472: Was anathema to Anglicans defending social order, but reputable scientists openly discussed the subject, and there was wide interest in John Herschel's letter praising Lyell's approach as a way to find a natural cause of the origin of new species. Gould met Darwin and told him that the Galápagos mockingbirds from different islands were separate species, not just varieties, and what Darwin had thought

9975-489: Was called up to (briefly) serve in the Franco-Prussian War . Dohrn's view of Haeckel's philosophical generalizations changed after he read Immanuel Kant and F. A. Lange 's Geschichte des Materialismus (1866). Among Dohrn's most highly cited theoretical contributions was his so-called "principle of succession of functions". It was derived from pre-existing ideas, especially those that had been proposed by

10080-418: Was his five-year voyage on HMS  Beagle from 1831 to 1836 that truly established Darwin as an eminent geologist. The observations and theories he developed during his voyage supported Charles Lyell 's concept of gradual geological change . Publication of his journal of the voyage made Darwin famous as a popular author. Puzzled by the geographical distribution of wildlife and fossils he collected on

10185-425: Was in the background to his main occupation of writing on geology and publishing expert reports on the Beagle collections, in particular, the barnacles . The impetus of Darwin's barnacle research came from a collection of a barnacle colony from Chile in 1835, which he dubbed Mr. Arthrobalanus . His confusion over the relationship of this species ( Cryptophialus minutus ) to other barnacles caused him to fixate on

10290-766: Was invited to occupy a table at the institute. She went on to fund raise to help establish the American Women's Table at the Stazione Zoologica Anton Dohrn. This table was subsequently held by American women zoologists such as Emily Ray Gregory . The three publications issued by the Station were:- Mittheilungen der Zoologischen Station in Neapel , Zoologischer Jahresbericht a reference journal famous for its rapid publication and accuracy and Fauna und Flora des Golfes von Neapel an inventory of

10395-523: Was led by a Commissario Straordinario. In 1976 following the appointment of Alberto Monroy, an embryologist, as Director a radical reorganisation began. SZN was divided into five parts: Biological Oceanography, Benthic Ecology (at the Villa Dohrn in Ischia), Biochemistry, Cell and Developmental Biology and Neurobiology and. In 1982 SZN became an "Ente pubblico di ricerca (National Research Institute) under

10500-504: Was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England. Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel -beds stranded above high tide. High in the Andes he saw seashells, and several fossil trees that had grown on

10605-427: Was one of the largest and most attractive cities of Europe and also had a considerable flow of tourists (30,000 a year) that could potentially visit the aquarium. Dohrn overcame the doubts of the city authorities and persuaded them to give him, free-of-charge, a plot of land at the sea edge, in the beautiful Villa Comunale on the condition that he promised to build the Stazione Zoologica at his own expense. He opened

10710-524: Was rather bored by Robert Jameson 's natural-history course, which covered geology – including the debate between neptunism and plutonism . He learned the classification of plants and assisted with work on the collections of the University Museum , one of the largest museums in Europe at the time. Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge , in January 1828, to study for

10815-574: Was surprised to find some benefit from hydrotherapy . Then, in 1851, his treasured daughter Annie fell ill, reawakening his fears that his illness might be hereditary. She died the same year after a long series of crises. In eight years of work on barnacles, Darwin's theory helped him to find " homologies " showing that slightly changed body parts served different functions to meet new conditions, and in some genera he found minute males parasitic on hermaphrodites , showing an intermediate stage in evolution of distinct sexes . In 1853, it earned him

10920-482: Was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists . Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794), a poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded. Both families were largely Unitarian , though

11025-446: Was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II , to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution. After delays,

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