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92-644: See text Parinari is a genus of plant in the family Chrysobalanaceae . Species of genus Parinari are found in Subsaharan Africa from Senegal to Sudan and Kenya and south to Namibia and Natal; in Eastern Madagascar; from Indochina through Indonesia, New Guinea, northern Queensland, and the southwest Pacific; and in Central and South America from Costa Rica to Trinidad and southern Brazil. The oldest fossils of Parinari fruits are from

184-574: A gymnasium and a colonnade ( peripatos ), from which the school acquired the name Peripatetic . Aristotle conducted courses and research at the school for the next twelve years. He often lectured small groups of distinguished students and, along with some of them, such as Theophrastus , Eudemus , and Aristoxenus , Aristotle built a large library which included manuscripts, maps, and museum objects. While in Athens, his wife Pythias died and Aristotle became involved with Herpyllis of Stagira. They had

276-541: A "survival of the fittest" origin of living things and their organs, and ridiculed the idea that accidents could lead to orderly results. To put his views into modern terms, he nowhere says that different species can have a common ancestor , or that one kind can change into another , or that kinds can become extinct . Aristotle did not do experiments in the modern sense. He used the ancient Greek term pepeiramenoi to mean observations, or at most investigative procedures like dissection. In Generation of Animals , he finds

368-533: A complex synthesis of the various philosophies existing prior to him. His teachings and methods of inquiry have had a significant impact across the world, and remain a subject of contemporary philosophical discussion. Aristotle's views profoundly shaped medieval scholarship . The influence of his physical science extended from late antiquity and the Early Middle Ages into the Renaissance , and

460-562: A daughter whom they also named Pythias. In 343/42 BC, Aristotle was invited to Pella by Philip II of Macedon in order to become the tutor to his thirteen-year-old son Alexander ; a choice perhaps influenced by the relationship of Aristotle's family with the Macedonian dynasty. Aristotle taught Alexander at the private school of Mieza , in the gardens of the Nymphs , the royal estate near Pella. Alexander's education probably included

552-438: A fertilized hen's egg of a suitable stage and opens it to see the embryo's heart beating inside. Instead, he practiced a different style of science: systematically gathering data, discovering patterns common to whole groups of animals, and inferring possible causal explanations from these. This style is common in modern biology when large amounts of data become available in a new field, such as genomics . It does not result in

644-497: A fluid such as air. In this system, heavy bodies in steady fall indeed travel faster than light ones (whether friction is ignored, or not ), and they do fall more slowly in a denser medium. Newton's "forced" motion corresponds to Aristotle's "violent" motion with its external agent, but Aristotle's assumption that the agent's effect stops immediately it stops acting (e.g., the ball leaves the thrower's hand) has awkward consequences: he has to suppose that surrounding fluid helps to push

736-455: A house is the bricks, stones, timbers, etc., or whatever constitutes the potential house, while the form of the substance is the actual house, namely 'covering for bodies and chattels' or any other differentia that let us define something as a house. The formula that gives the components is the account of the matter, and the formula that gives the differentia is the account of the form. Like his teacher Plato, Aristotle's philosophy aims at

828-597: A linear scale, and noted various exceptions, such as that sharks had a placenta like the tetrapods. To a modern biologist, the explanation, not available to Aristotle, is convergent evolution . Philosophers of science have generally concluded that Aristotle was not interested in taxonomy, but zoologists who studied this question in the early 21st century think otherwise. He believed that purposive final causes guided all natural processes; this teleological view justified his observed data as an expression of formal design. Aristotle's psychology , given in his treatise On

920-480: A number of subjects, such as ethics and politics , as well as standard literary texts, like Euripides and Homer . It is likely that during Aristotle's time in the Macedonian court, other prominent nobles, like Ptolemy and Cassander , would have occasionally attended his lectures. Aristotle encouraged Alexander toward eastern conquest, and his own attitude towards Persia was strongly ethnocentric . In one famous example, he counsels Alexander to be "a leader to

1012-435: A part of particular things. For example, it is possible that there is no particular good in existence, but "good" is still a proper universal form. Aristotle disagreed with Plato on this point, arguing that all universals are instantiated at some period of time, and that there are no universals that are unattached to existing things. In addition, Aristotle disagreed with Plato about the location of universals. Where Plato spoke of

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1104-498: A priori principles. Aristotle's "natural philosophy" spans a wide range of natural phenomena including those now covered by physics, biology and other natural sciences. In Aristotle's terminology, "natural philosophy" is a branch of philosophy examining the phenomena of the natural world, and includes fields that would be regarded today as physics, biology and other natural sciences. Aristotle's work encompassed virtually all facets of intellectual inquiry. Aristotle makes philosophy in

1196-418: A property or a relation to other things. When one looks at an apple, for example, one sees an apple, and one can also analyse a form of an apple. In this distinction, there is a particular apple and a universal form of an apple. Moreover, one can place an apple next to a book, so that one can speak of both the book and apple as being next to each other. Plato argued that there are some universal forms that are not

1288-551: A protective response. The first such plant receptors were identified in rice and in Arabidopsis thaliana . Plants have some of the largest genomes of all organisms. The largest plant genome (in terms of gene number) is that of wheat ( Triticum aestivum ), predicted to encode ≈94,000 genes and thus almost 5 times as many as the human genome . The first plant genome sequenced was that of Arabidopsis thaliana which encodes about 25,500 genes. In terms of sheer DNA sequence,

1380-420: A range of physical and biotic stresses which cause DNA damage , but they can tolerate and repair much of this damage. Plants reproduce to generate offspring, whether sexually , involving gametes , or asexually , involving ordinary growth. Many plants use both mechanisms. When reproducing sexually, plants have complex lifecycles involving alternation of generations . One generation, the sporophyte , which

1472-621: A set of six books called the Organon around 40 BC by Andronicus of Rhodes or others among his followers. The books are: The order of the books (or the teachings from which they are composed) is not certain, but this list was derived from analysis of Aristotle's writings. It goes from the basics, the analysis of simple terms in the Categories, the analysis of propositions and their elementary relations in On Interpretation , to

1564-549: A son whom Aristotle named after his father, Nicomachus . This period in Athens, between 335 and 323 BC, is when Aristotle is believed to have composed many of his philosophical works. He wrote many dialogues, of which only fragments have survived. Those works that have survived are in treatise form and were not, for the most part, intended for widespread publication; they are generally thought to be lecture aids for his students. His most important treatises include Physics , Metaphysics , Nicomachean Ethics , Politics , On

1656-562: A thrown stone, in the Physics (254b10), and "natural motion", such as of a falling object, in On the Heavens (300a20). In violent motion, as soon as the agent stops causing it, the motion stops also: in other words, the natural state of an object is to be at rest, since Aristotle does not address friction . With this understanding, it can be observed that, as Aristotle stated, heavy objects (on

1748-544: A woman with origins from Chalcis , Euboea . Nicomachus was said to have belonged to the medical guild of Asclepiadae and was likely responsible for Aristotle's early interest in biology and medicine. Ancient tradition held that Aristotle's family descended from the legendary physician Asclepius and his son Machaon . Both of Aristotle's parents died when he was still at a young age and Proxenus of Atarneus became his guardian. Although little information about Aristotle's childhood has survived, he probably spent some time in

1840-404: Is diploid (with 2 sets of chromosomes ), gives rise to the next generation, the gametophyte , which is haploid (with one set of chromosomes). Some plants also reproduce asexually via spores . In some non-flowering plants such as mosses, the sexual gametophyte forms most of the visible plant. In seed plants (gymnosperms and flowering plants), the sporophyte forms most of the visible plant, and

1932-425: Is a change where the substrate of the thing that has undergone the change has itself changed. In that particular change he introduces the concept of potentiality ( dynamis ) and actuality ( entelecheia ) in association with the matter and the form. Referring to potentiality, this is what a thing is capable of doing or being acted upon if the conditions are right and it is not prevented by something else. For example,

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2024-454: Is a historical accident: his works on botany have been lost, but two books on plants by his pupil Theophrastus have survived. Aristotle reports on the sea-life visible from observation on Lesbos and the catches of fishermen. He describes the catfish , electric ray , and frogfish in detail, as well as cephalopods such as the octopus and paper nautilus . His description of the hectocotyl arm of cephalopods, used in sexual reproduction,

2116-547: Is a similar process. Structures such as runners enable plants to grow to cover an area, forming a clone . Many plants grow food storage structures such as tubers or bulbs which may each develop into a new plant. Some non-flowering plants, such as many liverworts, mosses and some clubmosses, along with a few flowering plants, grow small clumps of cells called gemmae which can detach and grow. Plants use pattern-recognition receptors to recognize pathogens such as bacteria that cause plant diseases. This recognition triggers

2208-419: Is also a final cause or end. Then Aristotle proceeds and concludes that the actuality is prior to potentiality in formula, in time and in substantiality. With this definition of the particular substance (i.e., matter and form), Aristotle tries to solve the problem of the unity of the beings, for example, "what is it that makes a man one"? Since, according to Plato there are two Ideas: animal and biped, how then

2300-473: Is known about Aristotle's life. He was born in the city of Stagira in northern Greece during the Classical period . His father, Nicomachus , died when Aristotle was a child, and he was brought up by a guardian. At around eighteen years old, he joined Plato 's Academy in Athens and remained there until the age of thirty seven ( c.  347 BC ). Shortly after Plato died, Aristotle left Athens and, at

2392-496: Is known as botany , a branch of biology . All living things were traditionally placed into one of two groups, plants and animals . This classification dates from Aristotle (384–322 BC), who distinguished different levels of beings in his biology , based on whether living things had a "sensitive soul" or like plants only a "vegetative soul". Theophrastus , Aristotle's student, continued his work in plant taxonomy and classification. Much later, Linnaeus (1707–1778) created

2484-541: Is man a unity? However, according to Aristotle, the potential being (matter) and the actual one (form) are one and the same. Aristotle's immanent realism means his epistemology is based on the study of things that exist or happen in the world, and rises to knowledge of the universal, whereas for Plato epistemology begins with knowledge of universal Forms (or ideas) and descends to knowledge of particular imitations of these. Aristotle uses induction from examples alongside deduction , whereas Plato relies on deduction from

2576-413: Is the fulfilment of the end of the potentiality. Because the end ( telos ) is the principle of every change, and potentiality exists for the sake of the end, actuality, accordingly, is the end. Referring then to the previous example, it can be said that an actuality is when a plant does one of the activities that plants do. For that for the sake of which ( to hou heneka ) a thing is, is its principle, and

2668-686: The Antarctic flora , consisting of algae, mosses, liverworts, lichens, and just two flowering plants, have adapted to the prevailing conditions on that southern continent. Plants are often the dominant physical and structural component of the habitats where they occur. Many of the Earth's biomes are named for the type of vegetation because plants are the dominant organisms in those biomes, such as grassland , savanna , and tropical rainforest . Aristotle Aristotle ( Attic Greek : Ἀριστοτέλης , romanized:  Aristotélēs ; 384–322 BC)

2760-706: The Cretaceous so rapid that Darwin called it an " abominable mystery ". Conifers diversified from the Late Triassic onwards, and became a dominant part of floras in the Jurassic . In 2019, a phylogeny based on genomes and transcriptomes from 1,153 plant species was proposed. The placing of algal groups is supported by phylogenies based on genomes from the Mesostigmatophyceae and Chlorokybophyceae that have since been sequenced. Both

2852-652: The Early Modern period. John Philoponus (in Late antiquity ) and Galileo (in Early modern period ) are said to have shown by experiment that Aristotle's claim that a heavier object falls faster than a lighter object is incorrect. A contrary opinion is given by Carlo Rovelli , who argues that Aristotle's physics of motion is correct within its domain of validity, that of objects in the Earth 's gravitational field immersed in

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2944-525: The Great Comet of 371 BC . Aristotle was one of the first people to record any geological observations. He stated that geological change was too slow to be observed in one person's lifetime. The geologist Charles Lyell noted that Aristotle described such change, including "lakes that had dried up" and "deserts that had become watered by rivers", giving as examples the growth of the Nile delta since

3036-476: The History of Animals in a graded scale of perfection, a nonreligious version of the scala naturae , with man at the top. His system had eleven grades of animal, from highest potential to lowest, expressed in their form at birth: the highest gave live birth to hot and wet creatures, the lowest laid cold, dry mineral-like eggs. Animals came above plants , and these in turn were above minerals. He grouped what

3128-553: The Macedonian capital, making his first connections with the Macedonian monarchy . At the age of seventeen or eighteen, Aristotle moved to Athens to continue his education at Plato's Academy . He became distinguished as a researcher and lecturer, earning for himself the nickname "mind of the school" by his tutor Plato . In Athens, he probably experienced the Eleusinian Mysteries as he wrote when describing

3220-456: The Milky Way was made up of "those stars which are shaded by the earth from the sun's rays," pointing out partly correctly that if "the size of the sun is greater than that of the earth and the distance of the stars from the earth many times greater than that of the sun, then... the sun shines on all the stars and the earth screens none of them." He also wrote descriptions of comets, including

3312-508: The Physics (215a25), Aristotle effectively states a quantitative law, that the speed, v, of a falling body is proportional (say, with constant c) to its weight, W, and inversely proportional to the density, ρ, of the fluid in which it is falling:; Aristotle implies that in a vacuum the speed of fall would become infinite, and concludes from this apparent absurdity that a vacuum is not possible. Opinions have varied on whether Aristotle intended to state quantitative laws. Henri Carteron held

3404-457: The carpels or ovaries , which develop into fruits that contain seeds . Fruits may be dispersed whole, or they may split open and the seeds dispersed individually. Plants reproduce asexually by growing any of a wide variety of structures capable of growing into new plants. At the simplest, plants such as mosses or liverworts may be broken into pieces, each of which may regrow into whole plants. The propagation of flowering plants by cuttings

3496-623: The eukaryotes that form the kingdom Plantae ; they are predominantly photosynthetic . This means that they obtain their energy from sunlight , using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll . Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular , except for some green algae. Historically, as in Aristotle's biology ,

3588-412: The universal . Aristotle's ontology places the universal ( katholou ) in particulars ( kath' hekaston ), things in the world, whereas for Plato the universal is a separately existing form which actual things imitate. For Aristotle, "form" is still what phenomena are based on, but is "instantiated" in a particular substance. Plato argued that all things have a universal form , which could be either

3680-1003: The "chlorophyte algae" and the "streptophyte algae" are treated as paraphyletic (vertical bars beside phylogenetic tree diagram) in this analysis, as the land plants arose from within those groups. The classification of Bryophyta is supported both by Puttick et al. 2018, and by phylogenies involving the hornwort genomes that have also since been sequenced. Rhodophyta [REDACTED] Glaucophyta [REDACTED] Chlorophyta [REDACTED] Prasinococcales   Mesostigmatophyceae Chlorokybophyceae Spirotaenia [REDACTED] Klebsormidiales [REDACTED] Chara [REDACTED] Coleochaetales [REDACTED] Hornworts [REDACTED] Liverworts [REDACTED] Mosses [REDACTED] Lycophytes [REDACTED] [REDACTED] Gymnosperms [REDACTED] Angiosperms [REDACTED] Plant cells have distinctive features that other eukaryotic cells (such as those of animals) lack. These include

3772-519: The "extreme view" that Aristotle's concept of force was basically qualitative, but other authors reject this. Archimedes corrected Aristotle's theory that bodies move towards their natural resting places; metal boats can float if they displace enough water ; floating depends in Archimedes' scheme on the mass and volume of the object, not, as Aristotle thought, its elementary composition. Aristotle's writings on motion remained influential until

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3864-527: The Colonies , were composed by the philosopher for the young prince. After Philip II's assassination in 336 BC, Aristotle returned to Athens for the second and final time a year later. As a metic , Aristotle could not own property in Athens and thus rented a building known as the Lyceum (named after the sacred grove of Apollo Lykeios ), in which he established his own school. The building included

3956-610: The Greeks and a despot to the barbarians". Alexander's education under the guardianship of Aristotle likely lasted for only a few years, as at around the age of sixteen he returned to Pella and was appointed regent of Macedon by his father Philip. During this time, Aristotle is said to have gifted Alexander an annotated copy of the Iliad , which reportedly became one of Alexander's most prized possessions. Scholars speculate that two of Aristotle's now lost works, On kingship and On behalf of

4048-771: The Persians, and philosophical questions, like the definition of braveness. A widespread speculation in antiquity suggested that Aristotle played a role in Alexander's death, but the only evidence of this is an unlikely claim made some six years after the death. Following Alexander's death, anti-Macedonian sentiment in Athens was rekindled. In 322 BC, Demophilus and Eurymedon the Hierophant reportedly denounced Aristotle for impiety, prompting him to flee to his mother's family estate in Chalcis, Euboea , at which occasion he

4140-491: The Soul and Poetics . Aristotle studied and made significant contributions to "logic, metaphysics, mathematics, physics, biology, botany, ethics, politics, agriculture, medicine, dance, and theatre." While Alexander deeply admired Aristotle, near the end of his life, the two men became estranged having diverging opinions over issues, like the optimal administration of city-states, the treatment of conquered populations, such as

4232-864: The Viridiplantae, along with the red algae and the glaucophytes , in the clade Archaeplastida . There are about 380,000 known species of plants, of which the majority, some 260,000, produce seeds . They range in size from single cells to the tallest trees . Green plants provide a substantial proportion of the world's molecular oxygen; the sugars they create supply the energy for most of Earth's ecosystems and other organisms , including animals, either eat plants directly or rely on organisms which do so. Grain , fruit , and vegetables are basic human foods and have been domesticated for millennia. People use plants for many purposes , such as building materials , ornaments, writing materials , and, in great variety, for medicines . The scientific study of plants

4324-801: The amount of cytoplasm stays the same. Most plants are multicellular . Plant cells differentiate into multiple cell types, forming tissues such as the vascular tissue with specialized xylem and phloem of leaf veins and stems , and organs with different physiological functions such as roots to absorb water and minerals, stems for support and to transport water and synthesized molecules, leaves for photosynthesis, and flowers for reproduction. Plants photosynthesize , manufacturing food molecules ( sugars ) using energy obtained from light . Plant cells contain chlorophylls inside their chloroplasts, which are green pigments that are used to capture light energy. The end-to-end chemical equation for photosynthesis is: This causes plants to release oxygen into

4416-431: The atmosphere. Green plants provide a substantial proportion of the world's molecular oxygen, alongside the contributions from photosynthetic algae and cyanobacteria. Plants that have secondarily adopted a parasitic lifestyle may lose the genes involved in photosynthesis and the production of chlorophyll. Growth is determined by the interaction of a plant's genome with its physical and biotic environment. Factors of

4508-494: The ball along to make it continue to rise even though the hand is no longer acting on it, resulting in the Medieval theory of impetus . Aristotle suggested that the reason for anything coming about can be attributed to four different types of simultaneously active factors. His term aitia is traditionally translated as "cause", but it does not always refer to temporal sequence; it might be better translated as "explanation", but

4600-467: The basis of the modern system of scientific classification , but retained the animal and plant kingdoms , naming the plant kingdom the Vegetabilia. When the name Plantae or plant is applied to a specific group of organisms or taxa , it usually refers to one of four concepts. From least to most inclusive, these four groupings are: There are about 382,000 accepted species of plants, of which

4692-399: The becoming is for the sake of the end; and the actuality is the end, and it is for the sake of this that the potentiality is acquired. For animals do not see in order that they may have sight, but they have sight that they may see. In summary, the matter used to make a house has potentiality to be a house and both the activity of building and the form of the final house are actualities, which

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4784-574: The broad sense coextensive with reasoning, which he also would describe as "science". However, his use of the term science carries a different meaning than that covered by the term "scientific method". For Aristotle, "all science ( dianoia ) is either practical, poetical or theoretical" ( Metaphysics 1025b25). His practical science includes ethics and politics; his poetical science means the study of fine arts including poetry; his theoretical science covers physics, mathematics and metaphysics. In his On Generation and Corruption , Aristotle related each of

4876-433: The cause of earthquakes was a gas or vapor ( anathymiaseis ) that was trapped inside the earth and trying to escape, following other Greek authors Anaxagoras , Empedocles and Democritus . Aristotle also made many observations about the hydrologic cycle. For example, he made some of the earliest observations about desalination: he observed early – and correctly – that when seawater is heated, freshwater evaporates and that

4968-410: The composite natural ones, the study of nature would be the primary kind of knowledge; but if there is some motionless independent thing, the knowledge of this precedes it and is first philosophy, and it is universal in just this way , because it is first. And it belongs to this sort of philosophy to study being as being, both what it is and what belongs to it just by virtue of being. Aristotle examines

5060-406: The concepts of substance ( ousia ) and essence ( to ti ên einai , "the what it was to be") in his Metaphysics (Book VII), and he concludes that a particular substance is a combination of both matter and form, a philosophical theory called hylomorphism . In Book VIII, he distinguishes the matter of the substance as the substratum , or the stuff of which it is composed. For example, the matter of

5152-399: The data to come to the theory of evolution . Aristotle's writings can seem to modern readers close to implying evolution, but while Aristotle was aware that new mutations or hybridizations could occur, he saw these as rare accidents. For Aristotle, accidents, like heat waves in winter, must be considered distinct from natural causes. He was thus critical of Empedocles's materialist theory of

5244-455: The details of Aristotle's life are not well-established. The biographies written in ancient times are often speculative and historians only agree on a few salient points. Aristotle was born in 384 BC in Stagira , Chalcidice , about 55 km (34 miles) east of modern-day Thessaloniki . He was the son of Nicomachus , the personal physician of King Amyntas of Macedon , and Phaestis,

5336-488: The development of forests in swampy environments dominated by clubmosses and horsetails, including some as large as trees, and the appearance of early gymnosperms , the first seed plants . The Permo-Triassic extinction event radically changed the structures of communities. This may have set the scene for the evolution of flowering plants in the Triassic (~ 200  million years ago ), with an adaptive radiation in

5428-491: The earliest study of formal logic, and his conception of it was the dominant form of Western logic until 19th-century advances in mathematical logic . Kant stated in the Critique of Pure Reason that with Aristotle, logic reached its completion. Most of Aristotle's work is probably not in its original form, because it was most likely edited by students and later lecturers. The logical works of Aristotle were compiled into

5520-515: The early Miocene of Ethiopia, Panama, and Colombia. The genus is closely related to Neocarya . Parinari can be distinguished from other genera in Chrysobalanaceae by the following characteristics: As of May 2014 The Plant List recognises 42 accepted species (including infraspecific names): This Malpighiales article is a stub . You can help Misplaced Pages by expanding it . Plant See text Plants are

5612-442: The first century AD editor who assembled various small selections of Aristotle's works to create the treatise we know by the name Metaphysics . Aristotle called it "first philosophy", and distinguished it from mathematics and natural science (physics) as the contemplative ( theoretikē ) philosophy which is "theological" and studies the divine. He wrote in his Metaphysics (1026a16): If there were no other independent things besides

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5704-533: The following. Brood size decreases with (adult) body mass, so that an elephant has fewer young (usually just one) per brood than a mouse. Lifespan increases with gestation period , and also with body mass, so that elephants live longer than mice, have a longer period of gestation, and are heavier. As a final example, fecundity decreases with lifespan, so long-lived kinds like elephants have fewer young in total than short-lived kinds like mice. Aristotle distinguished about 500 species of animals , arranging these in

5796-531: The forms as existing separately from the things that participate in them, Aristotle maintained that universals exist within each thing on which each universal is predicated. So, according to Aristotle, the form of apple exists within each apple, rather than in the world of the forms. Concerning the nature of change ( kinesis ) and its causes, as he outlines in his Physics and On Generation and Corruption ( 319b–320a), he distinguishes coming-to-be ( genesis , also translated as 'generation') from: Coming-to-be

5888-515: The four elements proposed earlier by Empedocles , earth , water , air , and fire , to two of the four sensible qualities, hot, cold, wet, and dry. In the Empedoclean scheme, all matter was made of the four elements, in differing proportions. Aristotle's scheme added the heavenly aether , the divine substance of the heavenly spheres , stars and planets. Aristotle describes two kinds of motion: "violent" or "unnatural motion", such as that of

5980-409: The gametophyte is very small. Flowering plants reproduce sexually using flowers, which contain male and female parts: these may be within the same ( hermaphrodite ) flower, on different flowers on the same plant , or on different plants . The stamens create pollen , which produces male gametes that enter the ovule to fertilize the egg cell of the female gametophyte. Fertilization takes place within

6072-583: The great majority, some 283,000, produce seeds . The table below shows some species count estimates of different green plant (Viridiplantae) divisions . About 85–90% of all plants are flowering plants. Several projects are currently attempting to collect records on all plant species in online databases, e.g. the World Flora Online . Plants range in scale from single-celled organisms such as desmids (from 10  micrometres   (μm) across) and picozoa (less than 3 μm across), to

6164-428: The ground, say) require more force to make them move; and objects pushed with greater force move faster. This would imply the equation incorrect in modern physics. Natural motion depends on the element concerned: the aether naturally moves in a circle around the heavens, while the 4 Empedoclean elements move vertically up (like fire, as is observed) or down (like earth) towards their natural resting places. In

6256-639: The head of the Lyceum, who in turn passed them down to Neleus of Scepsis in Asia Minor. There, the papers remained hidden for protection until they were purchased by the collector Apellicon . In the meantime, many copies of Aristotle's major works had already begun to circulate and be used in the Lyceum of Athens, Alexandria , and later in Rome . With the Prior Analytics , Aristotle is credited with

6348-440: The image. According to Aristotle, spontaneity and chance are causes of some things, distinguishable from other types of cause such as simple necessity. Chance as an incidental cause lies in the realm of accidental things , "from what is spontaneous". There is also more a specific kind of chance, which Aristotle names "luck", that only applies to people's moral choices. In astronomy , Aristotle refuted Democritus 's claim that

6440-529: The land 1,200  million years ago , but it was not until the Ordovician , around 450  million years ago , that the first land plants appeared, with a level of organisation like that of bryophytes. However, fossils of organisms with a flattened thallus in Precambrian rocks suggest that multicellular freshwater eukaryotes existed over 1000 mya. Primitive land plants began to diversify in

6532-412: The large water-filled central vacuole , chloroplasts , and the strong flexible cell wall , which is outside the cell membrane . Chloroplasts are derived from what was once a symbiosis of a non-photosynthetic cell and photosynthetic cyanobacteria . The cell wall, made mostly of cellulose , allows plant cells to swell up with water without bursting. The vacuole allows the cell to change in size while

6624-515: The largest trees ( megaflora ) such as the conifer Sequoia sempervirens (up to 120 metres (380 ft) tall) and the angiosperm Eucalyptus regnans (up to 100 m (325 ft) tall). The naming of plants is governed by the International Code of Nomenclature for algae, fungi, and plants and the International Code of Nomenclature for Cultivated Plants . The ancestors of land plants evolved in water. An algal scum formed on

6716-665: The late Silurian , around 420  million years ago . Bryophytes, club mosses, and ferns then appear in the fossil record. Early plant anatomy is preserved in cellular detail in an early Devonian fossil assemblage from the Rhynie chert . These early plants were preserved by being petrified in chert formed in silica-rich volcanic hot springs. By the end of the Devonian, most of the basic features of plants today were present, including roots, leaves and secondary wood in trees such as Archaeopteris . The Carboniferous period saw

6808-468: The modern zoologist would call vertebrates as the hotter "animals with blood", and below them the colder invertebrates as "animals without blood". Those with blood were divided into the live-bearing ( mammals ), and the egg-laying ( birds , reptiles , fish ). Those without blood were insects, crustacea (non-shelled – cephalopods, and shelled ) and the hard-shelled molluscs ( bivalves and gastropods ). He recognised that animals did not exactly fit into

6900-404: The oceans are then replenished by the cycle of rainfall and river runoff ("I have proved by experiment that salt water evaporated forms fresh and the vapor does not when it condenses condense into sea water again.") Aristotle was the first person to study biology systematically, and biology forms a large part of his writings. He spent two years observing and describing the zoology of Lesbos and

6992-501: The physical or abiotic environment include temperature , water , light, carbon dioxide , and nutrients in the soil. Biotic factors that affect plant growth include crowding, grazing, beneficial symbiotic bacteria and fungi, and attacks by insects or plant diseases . Frost and dehydration can damage or kill plants. Some plants have antifreeze proteins , heat-shock proteins and sugars in their cytoplasm that enable them to tolerate these stresses . Plants are continuously exposed to

7084-538: The plant kingdom encompassed all living things that were not animals , and included algae and fungi . Definitions have narrowed since then; current definitions exclude the fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants ( hornworts , liverworts , mosses , lycophytes , ferns , conifers and other gymnosperms , and flowering plants ). A definition based on genomes includes

7176-407: The poet Dante called him "the master of those who know". His works contain the earliest known formal study of logic, and were studied by medieval scholars such as Peter Abelard and Jean Buridan . Aristotle's influence on logic continued well into the 19th century. In addition, his ethics , although always influential, gained renewed interest with the modern advent of virtue ethics . In general,

7268-457: The request of Philip II of Macedon , tutored his son Alexander the Great beginning in 343 BC. He established a library in the Lyceum, which helped him to produce many of his hundreds of books on papyrus scrolls . Though Aristotle wrote many treatises and dialogues for publication, only around a third of his original output has survived , none of it intended for publication. Aristotle provided

7360-433: The same certainty as experimental science, but it sets out testable hypotheses and constructs a narrative explanation of what is observed. In this sense, Aristotle's biology is scientific. From the data he collected and documented, Aristotle inferred quite a number of rules relating the life-history features of the live-bearing tetrapods (terrestrial placental mammals) that he studied. Among these correct predictions are

7452-439: The seed of a plant in the soil is potentially ( dynamei ) a plant, and if it is not prevented by something, it will become a plant. Potentially, beings can either 'act' ( poiein ) or 'be acted upon' ( paschein ), which can be either innate or learned. For example, the eyes possess the potentiality of sight (innate – being acted upon), while the capability of playing the flute can be possessed by learning (exercise – acting). Actuality

7544-706: The sights one viewed at the Mysteries, "to experience is to learn" ( παθεĩν μαθεĩν ). Aristotle remained in Athens for nearly twenty years before leaving in 348/47 BC after Plato's death. The traditional story about his departure records that he was disappointed with the Academy's direction after control passed to Plato's nephew Speusippus , although it is possible that the anti-Macedonian sentiments in Athens could have also influenced his decision. Aristotle left with Xenocrates to Assos in Asia Minor , where he

7636-469: The smallest published genome is that of the carnivorous bladderwort ( Utricularia gibba) at 82 Mb (although it still encodes 28,500 genes) while the largest, from the Norway spruce ( Picea abies ), extends over 19.6 Gb (encoding about 28,300 genes). Plants are distributed almost worldwide. While they inhabit several biomes which can be divided into a multitude of ecoregions , only the hardy plants of

7728-756: The study of more complex forms, namely, syllogisms (in the Analytics ) and dialectics (in the Topics and Sophistical Refutations ). The first three treatises form the core of the logical theory stricto sensu : the grammar of the language of logic and the correct rules of reasoning. The Rhetoric is not conventionally included, but it states that it relies on the Topics . What is today called Aristotelian logic with its types of syllogism (methods of logical argument), Aristotle himself would have labelled "analytics". The term "logic" he reserved to mean dialectics . The word "metaphysics" appears to have been coined by

7820-558: The surrounding seas, including in particular the Pyrrha lagoon in the centre of Lesbos. His data in History of Animals , Generation of Animals , Movement of Animals , and Parts of Animals are assembled from his own observations, statements given by people with specialized knowledge, such as beekeepers and fishermen, and less accurate accounts provided by travellers from overseas. His apparent emphasis on animals rather than plants

7912-475: The time of Homer , and "the upheaving of one of the Aeolian islands , previous to a volcanic eruption ."' Meteorologica lends its name to the modern study of meteorology, but its modern usage diverges from the content of Aristotle's ancient treatise on meteors . The ancient Greeks did use the term for a range of atmospheric phenomena, but also for earthquakes and volcanic eruptions. Aristotle proposed that

8004-478: The traditional rendering will be employed here. Aristotle describes experiments in optics using a camera obscura in Problems , book 15. The apparatus consisted of a dark chamber with a small aperture that let light in. With it, he saw that whatever shape he made the hole, the sun's image always remained circular. He also noted that increasing the distance between the aperture and the image surface magnified

8096-543: Was an Ancient Greek philosopher and polymath . His writings cover a broad range of subjects spanning the natural sciences , philosophy , linguistics , economics , politics , psychology , and the arts . As the founder of the Peripatetic school of philosophy in the Lyceum in Athens , he began the wider Aristotelian tradition that followed, which set the groundwork for the development of modern science . Little

8188-413: Was invited by his former fellow student Hermias of Atarneus ; he stayed there for a few years and left around the time of Hermias' death. While at Assos, Aristotle and his colleague Theophrastus did extensive research in botany and marine biology , which they later continued at the near-by island of Lesbos . During this time, Aristotle married Pythias , Hermias's adoptive daughter and niece, and had

8280-798: Was not replaced systematically until the Enlightenment and theories such as classical mechanics were developed. He influenced Judeo-Islamic philosophies during the Middle Ages, as well as Christian theology , especially the Neoplatonism of the Early Church and the scholastic tradition of the Catholic Church . Aristotle was revered among medieval Muslim scholars as "The First Teacher", and among medieval Christians like Thomas Aquinas as simply "The Philosopher", while

8372-523: Was said to have stated "I will not allow the Athenians to sin twice against philosophy" – a reference to Athens's trial and execution of Socrates . He died in Chalcis, Euboea of natural causes later that same year, having named his student Antipater as his chief executor and leaving a will in which he asked to be buried next to his wife. Aristotle left his works to Theophrastus, his successor as

8464-588: Was widely disbelieved until the 19th century. He gives accurate descriptions of the four-chambered fore-stomachs of ruminants , and of the ovoviviparous embryological development of the hound shark . He notes that an animal's structure is well matched to function so birds like the heron (which live in marshes with soft mud and live by catching fish) have a long neck, long legs, and a sharp spear-like beak, whereas ducks that swim have short legs and webbed feet. Darwin , too, noted these sorts of differences between similar kinds of animal, but unlike Aristotle used

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