58-637: Ptolemaia is a genus of extinct Afrotherian mammals from the Oligocene of East Africa. The genus and type species, P. lyonsi , was described by Henry Fairfield Osborn in 1908 from the Jebel Qatrani Formation of Egypts' Fayum Depression . The genus name alludes to the Ptolemaic dynasty of Ancient Greece which ruled over the Egyptian region where Ptolemaia was discovered, while
116-770: A bipedal stance for some behavior, and may walk a few steps bipedally. Pangolins are also good swimmers. Pangolins are insectivorous . Most of their diet consists of various species of ants and termites and may be supplemented by other insects, especially larvae. They are somewhat particular and tend to consume only one or two species of insects, even when many species are available. A pangolin can consume 140 to 200 g (5 to 7 oz) of insects per day. Pangolins are an important regulator of termite populations in their natural habitats. Pangolins have very poor vision . They also lack teeth. They rely heavily on smell and hearing , and they have other physical characteristics to help them eat ants and termites. Their skeletal structure
174-488: A ball when threatened, with its overlapping scales acting as armor , while it protects its face by tucking it under its tail. The scales are sharp, providing extra defense from predators. Pangolins can emit a noxious-smelling chemical from glands near the anus, similar to the spray of a skunk . They have short legs, with sharp claws which they use for burrowing into ant and termite mounds and for climbing. The tongues of pangolins are extremely long, and like those of
232-517: A clade. Additionally, there might be some dental synapomorphies uniting afroinsectiphilians, if not afrotheres as a whole: p4 talonid and trigonid of similar breadth, a prominent p4 hypoconid, presence of a P4 metacone and absence of parastyles on M1–2. Afrotheria is now recognized as one of the three major groups within the Eutheria (containing placental mammals ). Relations within the three cohorts, Afrotheria, Xenarthra , Boreoeutheria , and
290-411: A genetic dysfunction, making them extremely fragile. They are susceptible to diseases such as pneumonia and the development of ulcers in captivity, complications that can lead to an early death. In addition, pangolins rescued from illegal trade often have a higher chance of being infected with parasites such as intestinal worms , further lessening their chance for rehabilitation and reintroduction to
348-575: A high risk of extinction (perhaps related to the large size of many). Species loss within this already small group would comprise a particularly great loss of genetic and evolutionary diversity. The IUCN Afrotheria Specialist Group notes that Afrotheria, as currently reconstructed, includes nearly a third of all mammalian orders currently found in Africa and Madagascar, but only 75 of more than 1,200 mammalian species in those areas. While most extant species assigned to Afrotheria live in Africa, some (such as
406-433: A pangolin is marked by large, hardened, overlapping, plate-like scales, which are soft on newborn pangolins, but harden as the animal matures. They are made of keratin , the same material from which human fingernails and tetrapod claws are made, and are structurally and compositionally very different from the scales of reptiles. The pangolin's scaled body is comparable in appearance to a pine cone . It can curl up into
464-679: A study found that it is "not too late" to establish conservation efforts for Philippine pangolins ( Manis culionensis ), a species that is only found on the island province of Palawan. Taiwan is one of the few conservation grounds for pangolins in the world after the country enacted the 1989 Wildlife Conservation Act. The introduction of Wildlife Rehabilitation Centers in places like Luanshan ( Yanping Township ) in Taitung and Xiulin townships in Hualien became important communities for protecting pangolins and their habitats and has greatly improved
522-618: Is a superorder of placental mammals , the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles , elephant shrews (also known as sengis), otter shrews , tenrecs , aardvarks , hyraxes , elephants , sea cows , and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting
580-419: Is also active by day, while other species of pangolins spend most of the daytime sleeping, curled up into a ball (" volvation "). Arboreal pangolins live in hollow trees, whereas the ground-dwelling species dig tunnels to a depth of 3.5 m (11 ft). Some pangolins walk with their front claws bent under the foot pad, although they use the entire foot pad on their rear limbs. Furthermore, some exhibit
638-430: Is called the gizzard , and it is also covered in keratinous spines. These spines further aid in the grinding up and digestion of the pangolin's prey. Some species, such as the tree pangolin , use their strong, prehensile tails to hang from tree branches and strip away bark from the trunk, exposing insect nests inside. Pangolins are solitary and meet only to have sex, with mating typically taking place at night after
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#1732772157532696-408: Is found in the comparative anatomical literature for the idea that at least part of this group forms a clade . The Paleocene genus Ocepeia , which is the most completely-known Paleocene African mammal and the oldest afrotherian known from a complete skull, shares similarities with both Paenungulata and Afroinsectiphilia, and may help to characterize the ancestral body type of afrotherians. Since
754-473: Is reported in Nigeria , where the animal is on the verge of extinction due to overexploitation . The overexploitation comes from hunting pangolins for game meat and the reduction of their forest habitats due to deforestation caused by timber harvesting . The pangolin are hunted as game meat for both medicinal purposes and food consumption. The nucleic acid sequence of a specific receptor-binding domain of
812-707: Is sturdy and they have strong front legs used for tearing into termite mounds. They use their powerful front claws to dig into trees, soil, and vegetation to find prey, then proceed to use their long tongues to probe inside the insect tunnels and to retrieve their prey. The structure of their tongue and stomach is key to aiding pangolins in obtaining and digesting insects. Their saliva is sticky, causing ants and termites to stick to their long tongues when they are hunting through insect tunnels. Without teeth, pangolins cannot also chew; but while foraging , they ingest small stones ( gastroliths ), which accumulate in their stomachs to help to grind up ants. This part of their stomach
870-472: The IUCN , while three are classified as critically endangered . All pangolin species are currently listed under Appendix I of CITES which prohibits international trade, except when the product is intended for non-commercial purposes and a permit has been granted. China had been the main destination country for pangolins until 2018, where it was surpassed by Vietnam. In 2019, Vietnam was reported to have seized
928-616: The Malay word pengguling meaning "one who rolls up" from guling or giling "to roll"; it was used for the Sunda pangolin ( Manis javanica ). However, the modern name is tenggiling . In Javanese , it is terenggiling ; and in the Philippine languages , it is goling , tanggiling , or balintong (with the same meaning). In ancient India, according to Aelian , it was known as the phattáges (φαττάγης). The physical appearance of
986-806: The Song dynasty , the notion of penetrating and clearing blockages was emphasized in the Taiping sheng hui fan ("Formulas from Benevolent Sages Compiled During the Era of Peace and Tranquility"), compiled by Wang Huaiyin in 992. In the 21st century, the main uses of pangolin scales are quackery practices based on unproven claims the scales dissolve blood clots , promote blood circulation, or help lactating women secrete milk. The supposed health effects of pangolin meat and scales claimed by folk medicine practitioners are based on their consumption of ants, long tongues, and protective scales. The official pharmacopoeia of
1044-502: The carnivorans , with which they form a clade , the Ferae . Palaeanodonts are even closer relatives to pangolins, being classified with pangolins in the clade Pholidotamorpha . The split between carnivorans and pangolins is estimated to have occurred 79.47 Ma (million years) ago. The first dichotomy in the phylogeny of extant Manidae separates Asian pangolins ( Manis ) from African pangolins ( Smutsia and Phataginus ). Within
1102-456: The giant anteater and the tube-lipped nectar bat , the root of the tongue is not attached to the hyoid bone but is in the thorax between the sternum and the trachea . Large pangolins can extend their tongues as much as 40 cm (16 in), with a diameter of only about 0.5 cm ( 1 ⁄ 5 in). Most pangolins are nocturnal animals which use their well-developed sense of smell to find insects. The long-tailed pangolin
1160-669: The specific epithet lyonsi honors H. G. Lyons , then director of the Egyptian Geological Survey. A second species, P. grangeri , was described in 1987, and named after the early 20th century paleontologist Walter W. Granger . Fossils of P. grangeri are also known from Kenya. This article about prehistoric Afrotheria is a stub . You can help Misplaced Pages by expanding it . Afrotheria See below Afrotheria ( / æ f r oʊ ˈ θ ɪər i ə / from Latin Afro- "of Africa" + theria "wild beast")
1218-564: The spike protein belonging to coronaviruses taken from pangolins was found to be a 99% match with SARS coronavirus 2 ( SARS-CoV-2 ), the virus which causes COVID-19 and is responsible for the COVID-19 pandemic . Researchers in Guangzhou , China, hypothesized that SARS-CoV-2 had originated in bats, and prior to infecting humans, was circulating among pangolins. The illicit Chinese trade of pangolins for use in traditional Chinese medicine
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#17327721575321276-678: The threatened tier. Three ( Manis culionensis , M. pentadactyla and M. javanica ) are critically endangered, three ( Phataginus tricuspis , Manis crassicaudata and Smutsia gigantea ) are endangered and two ( Phataginus tetradactyla and Smutsia temminckii ) are vulnerable on the Red List of Threatened Species of the International Union for Conservation of Nature . The name of order Pholidota comes from Ancient Greek ϕολιδωτός – "clad in scales" from φολίς pholís " scale ". The name "pangolin" comes from
1334-449: The xenarthrans within the invalid taxon Edentata . Continuing work on the molecular and morphological diversity of afrotherian mammals has provided ever increasing support for their common ancestry. The afrotherian clade was originally proposed in 1998 based on analyses of DNA sequence data. However, previous studies had hinted at the close interrelationships among subsets of endemic African mammals; some of these studies date to
1392-734: The 1920s and there were sporadic papers in the 1980s and 1990s. The core of the Afrotheria consists of the Paenungulata , i.e., elephants , sea cows , and hyraxes , a group with a long history among comparative anatomists. Hence, while DNA sequence data have proven essential to infer the existence of the Afrotheria as a whole, and while the Afroinsectiphilia ( insectivoran -grade afrotheres including tenrecs , golden moles , sengis , and aardvarks ) were not recognized as part of Afrotheria without DNA data, some precedent
1450-405: The 1990s, increasing molecular and anatomical data have been applied to the classification of animals. Both types of data support the idea that afrotherian mammals are descended from a single common ancestor to the exclusion of other mammals. On the anatomical side, features shared by most, if not all, afrotheres include high vertebral counts, aspects of placental membrane formation , the shape of
1508-490: The African continent was separated from other major land masses. This idea is consistent with the fossil record of Xenarthra , which is restricted to South America (following recent consensus that Eurotamandua is not a xenarthran ). However, Afrotheria itself does not have a fossil record restricted to Africa, and appears in fact to have evolved in the continent's isolation. More recent, genomic-scale phylogenies favor
1566-756: The Cenozoic, and proboscideans (elephants and their relatives) filled the roles of large herbivores such as hippos, camels, rhinos, and tapirs. The sirenians developed aquatic body plans and started spreading to other parts of the world by water (evolving convergently with the other groups of marine mammals such as cetaceans and pinnipeds ). In addition to their similarity with Laurasian mammals in North America, Europe, and Asia, many afrotheres also exhibit convergent evolution with groups of mammals that evolved and lived exclusively in South America, which
1624-785: The IUCN considered all eight species of pangolin on its Red List of Threatened Species as threatened. The IUCN SSC Pangolin Specialist Group launched a global action plan to conserve pangolins, dubbed "Scaling up Pangolin Conservation", in July 2014. This action plan aims to improve all aspects of pangolin conservation with an added emphasis on combating poaching and trafficking of the animal while educating communities on its importance. Another suggested approach to fighting pangolin (and general wildlife) trafficking consists in "following
1682-605: The Indian elephant and three of the four sirenian species) occur elsewhere; many of these are also endangered. Prior to the Quaternary extinction event , proboscideans were present on every continent of the world except Australia and Antarctica . Hyraxes lived in much of Eurasia as recently as the end of the Pliocene . The extinct afrotherian orders of embrithopods and desmostylians were also once widely distributed. However,
1740-613: The People's Republic of China included Chinese pangolin scales as an ingredient in TCM formulations. Pangolins were removed from the pharmacopoeia starting from the first half of 2020. Although pangolin scales have been removed from the list of raw ingredients, the scales are still listed as a key ingredient in various medicines. Pangolin parts are also used for medicinal purposes in other Asian countries such as India, Nepal and Pakistan. In some parts of India and Nepal, locals believe that wearing
1798-431: The ankle bones, the relatively late eruption of the permanent dentition, and undescended testicles remaining in the body near the kidneys. The snout is unusually long and mobile in several Afrotherian species, and this was pointed out as a possible shared-derived character. Studies of genomic data, including millions of aligned nucleotides sampled for a growing number of placental mammals, also support Afrotheria as
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1856-448: The burrow riding on the mother's back. Weaning takes place around three months of age, when the young begin to eat insects in addition to nursing. At two years of age, the offspring are sexually mature and are abandoned by the mother. The order Pholidota was long considered to be the sister taxon to Xenarthra (neotropical anteaters , sloths , and armadillos ), but recent genetic evidence indicates their closest living relatives are
1914-555: The desmostylians have recently been viewed as possible perissodactyls , rather than afrotheres, although this is still controversial; the taxonomic placement of embrithopods is also not clear. Afrotheria is a clade of placental mammals, the stem designation for which is Eutheria . Based on precedent, some clades are junior synonyms and arguably should be replaced. [REDACTED] [REDACTED] Pangolin Pangolins , sometimes known as scaly anteaters , are mammals of
1972-643: The efficacy of those practices, and they have no logical mechanism of action , their popularity still drives the black market for animal body parts, despite concerns about toxicity, transmission of diseases from animals to humans, and species extermination. The ongoing demand for parts as ingredients continues to fuel pangolin poaching, hunting and trading . The first record of pangolin scales occurs in Ben Cao Jinji Zhu ("Variorum of Shennong's Classic of Materia Medica", 500 CE), which recommends pangolin scales for protection against ant bites; burning
2030-480: The fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia. Because Africa was isolated by water, Laurasian groups of mammals such as insectivores , rodents , lagomorphs , carnivorans and ungulates could not reach Africa for much of the early to mid- Cenozoic . Instead, the niches occupied by those groups on
2088-491: The former, Manis pentadactyla is the sister group to a clade comprising M. crassicaudata and M. javanica . Within the latter, a split separates the large terrestrial African pangolins of the genus Smutsia from the small arboreal African pangolins of the genus Phataginus . Asian and African pangolins are thought to have diverged about 41.37 Ma ago. Moreover, the basal position of Manis within Pholidota suggests
2146-501: The genomes of dead Manis javanica and Manis pentadactyla . To refer to both sampling site and hosts, they were named Dongyang pangolin virus (DYPV) and Lishui pangolin virus (LSPV). The DYPV pestivirus was also identified in Amblyomma javanense nymph ticks from a diseased pangolin. Pangolin scales and flesh are used as ingredients for various traditional Chinese medicine preparations. While no scientific evidence exists for
2204-469: The group originated in Eurasia, consistent with their laurasiatherian phylogeny. Pangolins are in high demand in southern China and Vietnam because their scales are believed to have medicinal properties in traditional Chinese and Vietnamese medicine . Their meat is also considered a delicacy. 100,000 are estimated to be trafficked a year to China and Vietnam, amounting to over one million over
2262-448: The hypothesis that Afrotheria and Xenarthra comprise sister taxa at the base of the placental mammal radiation, suggesting an ancient Gondwanan clade of placental mammals. A 2021 morphological study also proposed to render Meridiungulata polyphyletic and recognise most of its clades as part of a group called Sudamericungulata , closely related to hyraxes, while Litopterna remains a sister taxon to Perissodactyla . Relations between
2320-403: The identity of the placental root, remain somewhat controversial. Afrotheria as a clade has usually been discussed without a Linnaean rank, but has been assigned the rank of cohort, magnorder, and superorder. One reconstruction, which applies the molecular clock , proposes that the oldest split occurred between Afrotheria and the other two some 105 million years ago in the mid-Cretaceous , when
2378-592: The largest volumes of pangolin scales, surpassing Nigeria that year. Pangolins are also hunted and eaten in Ghana and are one of the more popular types of bushmeat , while local healers use the pangolin as a source of traditional medicine. Though pangolins are protected by an international ban on their trade, populations have suffered from illegal trafficking due to beliefs in East Asia that their ground-up scales can stimulate lactation or cure cancer or asthma . In
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2436-431: The male and female pangolin meet near a watering hole. Males are larger than females, weighing up to 40% more. While the mating season is not defined, they typically mate once each year, usually during the summer or autumn. Rather than the males seeking out the females, males mark their location with urine or feces and the females find them. If competition over a female occurs, the males use their tails as clubs to fight for
2494-825: The money" rather than "the animal", which aims to disrupt smugglers' profits by interrupting money flows. Financial intelligence gathering could thus become a key tool in protecting these animals, although this opportunity is often overlooked. In 2018, a Chinese NGO launched the Counting Pangolins movement, calling for joint efforts to save the mammals from trafficking. Wildlife conservation group TRAFFIC has identified 159 smuggling routes used by pangolin traffickers and aims to shut these down. Many attempts have been made to breed pangolins in captivity, but due to their reliance on wide-ranging habitats and very particular diets, these attempts are often unsuccessful. Pangolins have significantly decreased immune responses due to
2552-459: The northern continents were filled by various groups of afrotheres via the process of convergent evolution . The small insectivorous afrotheres such as elephant shrews, golden moles, and tenrecs filled the niches of insectivores , the hyraxes filled the roles of rodents and lagomorphs, the aardvarks filled the roles of various medium size ant-eating mammals (anteaters, armadillos, pangolins, echidnas, numbats, etc.) found on other continents throughout
2610-431: The only known mammals with this feature. Depending on the species, they live in hollow trees or burrows . Pangolins are nocturnal , and their diet consists of mainly ants and termites , which they capture using their long tongues. They tend to be solitary animals, meeting only to mate and produce a litter of one to three offspring, which they raise for about two years. Pangolins superficially resemble armadillos , though
2668-455: The opportunity to mate with her. Gestation periods differ by species, ranging from roughly 70 to 140 days. African pangolin females usually give birth to a single offspring at a time, but the Asiatic species may give birth to from one to three. Weight at birth is 80 to 450 g ( 2 + 3 ⁄ 4 to 15 + 3 ⁄ 4 oz), and the average length is 150 mm (6 in). At
2726-710: The order Pholidota ( / f ɒ l ɪ ˈ d oʊ t ə / ). The one extant family, the Manidae , has three genera: Manis , Phataginus , and Smutsia . Manis comprises four species found in Asia, while Phataginus and Smutsia include two species each, all found in sub-Saharan Africa. These species range in size from 30 to 100 cm (12 to 39 in). Several extinct pangolin species are also known. In September 2023, nine species were reported. Pangolins have large, protective keratin scales, similar in material to fingernails and toenails, covering their skin; they are
2784-605: The past decade, numerous seizures of illegally trafficked pangolin and pangolin meat have taken place in Asia. In one such incident in April 2013, 10,000 kg (22,000 pounds) of pangolin meat were seized from a Chinese vessel that ran aground in the Philippines . In another case in August 2016, an Indonesian man was arrested after police raided his home and found over 650 pangolins in freezers on his property. The same threat
2842-561: The past decade. This makes them the most trafficked animal in the world. This, coupled with deforestation , has led to a large decrease in the numbers of pangolins. Some species, such as Manis pentadactyla have become commercially extinct in certain ranges as a result of overhunting. In November 2010, pangolins were added to the Zoological Society of London 's list of evolutionarily distinct and endangered mammals. All eight species of pangolin are assessed as threatened by
2900-601: The scales as a cure for people crying hysterically during the night. During the Tang dynasty , a recipe for expelling evil spirits with a formulation of scales, herbs, and minerals appeared in 682, and in 752 CE the idea that pangolin scales could also stimulate milk secretion in lactating women, one of the main uses today, was recommended in the Wai Tai Mi Yao ("Arcane Essentials from the Imperial Library"). In
2958-609: The scales of a pangolin can help prevent pneumonia. Pangolin scales have also been used for medicinal purposes in Malaysia, Indonesia and northern Myanmar. Indigenous people in southern Palawan , Philippines, have held the belief that elders could avoid prostate illnesses by wearing belts made with the scales. As a result of increasing threats to pangolins, mainly in the form of illegal, international trade in pangolin skin, scales, and meat, these species have received increasing conservation attention in recent years. As of January 2020 ,
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#17327721575323016-454: The time of birth, the scales are soft and white. After several days, they harden and darken to resemble those of an adult pangolin. During the vulnerable stage, the mother stays with her offspring in the burrow, nursing it, and wraps her body around it if she senses danger. The young cling to the mother's tail as she moves about, although, in burrowing species, they remain in the burrow for the first 2–4 weeks of life. At one month, they first leave
3074-400: The two are not closely related; they have merely undergone convergent evolution . Pangolins are threatened by poaching (for their meat and scales, which are used in traditional medicine ) and heavy deforestation of their natural habitats, and are the most trafficked mammals in the world. As of January 2020 , there are eight species of pangolin whose conservation status is listed in
3132-602: The various afrotherian orders are still being studied. On the basis of molecular studies, elephants and manatees appear to be related, and likewise elephant shrews and aardvarks. These findings are compatible with the work of earlier anatomists. Procaviidae [REDACTED] Trichechidae [REDACTED] Dugongidae [REDACTED] Elephantidae [REDACTED] Orycteropodidae [REDACTED] Macroscelididae [REDACTED] Chrysochloridae [REDACTED] Potamogalidae [REDACTED] Tenrecidae [REDACTED] Many extant members of Afrotheria appear to have
3190-585: The wild. The idea of farming pangolins to reduce the number being illegally trafficked is being explored with little success. The third Saturday in February is promoted as World Pangolin Day by the conservation NPO Annamiticus. World Pangolin Day has been noted for its effectiveness in generating awareness about pangolins. In 2017, Jackie Chan made a public service announcement called WildAid : Jackie Chan & Pangolins (Kung Fu Pangolin) . In December 2020,
3248-403: Was also an island continent for much of the Cenozoic. The common ancestry of these animals was not recognized until the late 1990s. Historically, the Paenungulata had been linked to the true ungulates (particularly the perissodactyls ); the golden mole, tenrecs, and elephant shrews with the traditional (and polyphyletic/incorrect) taxon Insectivora ; and the aardvarks with the pangolins and
3306-432: Was later proved that the testing which suggested that pangolins were a potential host for the virus was flawed, when genetic analysis showed that the spike protein and its binding to receptors in pangolins had minimal effect from the virus, and therefore were not likely mechanisms for COVID-19 infections in humans. In 2020, two novel RNA viruses distantly related to pestiviruses and coltiviruses have been detected in
3364-452: Was suggested as a vector for human transmission. However, whole-genome comparison found that the pangolin and human coronaviruses share only up to 92% of their RNA. Ecologists worried that the early speculation about pangolins being the source may have led to mass slaughters, endangering them further, which was similar to what happened to Asian palm civets during the SARS outbreak. It
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