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Puno ( Aymara and Quechua : Punu ) is a city in southeastern Peru , located on the shore of Lake Titicaca . It is the capital city of the Puno Region and the Puno Province with a population of approximately 140,839 (2015 estimate). The city was established in 1668 by viceroy Pedro Antonio Fernández de Castro as capital of the province of Paucarcolla with the name San Juan Bautista de Puno. The name was later changed to San Carlos de Puno, in honor of king Charles II of Spain . Puno has several churches dating back from the colonial period; they were built to service the Spanish population and evangelize the Quechua people .

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101-498: Puno is an important agricultural and livestock region; important livestock are llamas and alpacas , which graze on its immense plateaus and plains . Much of the city economy relies on the black market, fueled by cheap goods smuggled in from Bolivia . Puno is served by the Inca Manco Capac International Airport in nearby Juliaca . Puno is situated between the shores of Lake Titicaca and

202-430: A meat and pack animal by Andean cultures since the pre-Columbian era . Llamas are social animals and live with others as a herd . Their wool is soft and contains only a small amount of lanolin . Llamas can learn simple tasks after a few repetitions. When using a pack, they can carry about 25 to 30% of their body weight for 8 to 13 km (5–8 miles ). The name llama (in the past also spelled "lama" or "glama")

303-434: A better idea of when the cria is expected can be determined. Hand mating is the most efficient method, but it requires the most work on the part of the human involved. A male and female llama are put into the same pen, and mating is monitored. They are then separated and re-mated every other day until one refuses the mating. Usually, one can get in two matings using this method, though some stud males routinely refuse to mate

404-445: A broad denuded area as a piedmont deposit by the rivers which issued from the mountains. Since then, it has been more or less dissected by the erosion of valleys. The central section of the plains thus presents a marked contrast to the northern section. While the northern section owes its smoothness to the removal of local gravels and sands from a formerly uneven surface by the action of degrading rivers and their inflowing tributaries,

505-602: A broad stretch of country underlain by nearly horizontal strata extending westward from the 97th meridian west to the base of the Rocky Mountains , a distance of 300 to 500 mi (480 to 800 km). It extends northward from the Mexican boundary far into Canada. Although the altitude of the plains increases gradually from 600 ft (180 m) or 1,200 ft (370 m) on the east to 4,000–5,000 ft (1,200–1,500 m) or 6,000 ft (1,800 m) near

606-434: A completely or partially domesticated state. Many are also descended from ancestors previously domesticated, a state that tends to produce a certain amount of variation from the original type. The four forms commonly distinguished by the inhabitants of South America are recognized as distinct species, though there are difficulties in defining their distinctive characteristics. These are: The llama and alpaca are only known in

707-523: A female more than once. The separation presumably helps to keep the sperm count high for each mating and also helps to keep the condition of the female llama's reproductive tract more sound. If the mating is unsuccessful within two to three weeks, the female is mated again. Options for feeding llamas are quite wide; various commercial and farm-based feeds are available. The major determining factors include feed cost, availability, nutrient balance and energy density required. Young, actively growing llamas require

808-456: A good fossil record. Camel-like animals have been traced back through early Miocene forms from the thoroughly differentiated, modern species. Their characteristics became more general, and they lost those that distinguished them as camelids; hence, they were classified as ancestral artiodactyls. No fossils of these earlier forms have been found in the Old World , indicating that North America

909-603: A gradual transition, this rainfall line may be taken to divide the drier plains from the moister prairies. However, in Canada the eastern boundary of the plains is well defined by the presence of the Canadian Shield to the northeast. The plains (within the United States) may be described in northern, intermediate, central and southern sections, in relation to certain peculiar features. In Canada, no such division

1010-509: A greater concentration of nutrients than mature animals because of their smaller digestive tract capacities. Llamas that are well-socialized and trained to halter and lead after weaning are very friendly and pleasant to be around. They are extremely curious, and most will approach people easily. However, llamas that are bottle-fed or over-socialized and over-handled as youth will become extremely difficult to handle when mature, when they will begin to treat humans as they treat each other, which

1111-400: A group), consist of the vicuña ( Vicugna vicugna , prev. Lama vicugna ), guanaco ( Lama guanicoe ), Suri alpaca , and Huacaya alpaca ( Vicugna pacos , prev. Lama guanicoe pacos ), and the domestic llama ( Lama glama ). Guanacos and vicuñas live in the wild, while llamas and alpacas exist only as domesticated animals. Although early writers compared llamas to sheep , their similarity to

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1212-697: A height of 1.7 to 1.8 m (5 ft 7 in to 5 ft 11 in) at the top of the head and can weigh between 130 and 272 kg (287 and 600 lb). At maturity, males can weigh 94.74 kg, while females can weigh 102.27 kg. At birth, a baby llama (called a cria ) can weigh between 9 and 14 kg (20 and 31 lb). Llamas typically live for 15 to 25 years, with some individuals surviving 30 years or more. The following characteristics apply especially to llamas. Dentition of adults: incisors ⁠ 1 / 3 ⁠ canines ⁠ 1 / 1 ⁠ , premolars ⁠ 2 / 2 ⁠ , molars ⁠ 3 / 3 ⁠ ; total 32. In

1313-503: A kush (lying down) position, similar to big cats and canines, which is unusual in a large animal. They mate for an extended time (20–45 minutes), also unusual in a large animal. The gestation period of a llama is 11.5 months (350 days). Dams (female llamas) do not lick off their babies, as they have an attached tongue that does not reach outside of the mouth more than 13 millimetres ( 1 ⁄ 2 inch). Rather, they will nuzzle and hum to their newborns. A cria (from Spanish for "baby")

1414-399: A large animal. Female llamas are induced ovulators . Through mating, the female releases an egg and is often fertilized on the first attempt. Female llamas do not go into estrus ("heat"). Like humans, llama males and females mature sexually at different rates. Females reach puberty at about 12 months old; males do not become sexually mature until around three years of age. Llamas mate in

1515-423: A larger brain cavity and orbits and less-developed cranial ridges due to its smaller size. The nasal bones are shorter and broader and are joined by the premaxilla. Vertebrae : The ears are rather long and slightly curved inward, characteristically known as "banana" shaped. There is no dorsal hump. The feet are narrow, the toes being more separated than in the camels, each having a distinct plantar pad. The tail

1616-621: A llama's purchase cost and annual maintenance. Although not every llama is suited to the job, most are a viable, nonlethal alternative for reducing predation, requiring no training and little care. Llamas have a fine undercoat, which can be used for handicrafts and garments . The coarser outer guard hair is used for rugs, wall hangings, and lead ropes. The fiber comes in many colors, ranging from white or grey to reddish-brown, brown, dark brown, and black. Doctors and researchers have determined that llamas possess antibodies that are well-suited to treat certain diseases. Scientists have been studying

1717-436: A strange noise or feels threatened, an alarm call - a loud, shrill sound that rhythmically rises and falls - is sent out, and all others become alert. They will often hum to each other as a form of communication. The llama's groaning noises or going "mwa" (/mwaʰ/) is often a sign of fear or anger. Unhappy or agitated llamas will lay their ears back, while ears being perked upwards is a sign of happiness or curiosity. An "orgle"

1818-594: A west-northwest direction in what is now Oklahoma and Texas which is now known as the De Soto Trail. The Spanish thought that the Great Plains were the location of the mythological Quivira and Cíbola , a place said to be rich in gold. People in the southwest began to acquire horses in the 16th century by trading or stealing them from Spanish colonists in New Mexico. As horse culture moved northward,

1919-404: Is about 500 mi (800 km) east to west and 2,000 mi (3,200 km) north to south. Much of the region was home to American bison herds until they were hunted to near extinction during the mid/late-19th century. It has an area of approximately 500,000 sq mi (1,300,000 km ). Current thinking regarding the geographic boundaries of the Great Plains is shown by this map at

2020-405: Is about 8.4 °C (47.1 °F), and the weather never gets overly warm. During the winter months from June to August, night-time temperatures usually drop below 0 °C (32 °F). At this high altitude, the rays of the sun are very strong. Most of the annual precipitation falls during the southern hemisphere summer, with the winter months being very dry. Music and dance are typical parts of

2121-531: Is characterized by bouts of spitting, kicking and neck wrestling. Llamas are now utilized as certified therapy animals in nursing homes and hospitals. Rojo the Llama , located in the Pacific Northwest was certified in 2008. The Mayo Clinic says animal-assisted therapy can reduce pain, depression, anxiety, and fatigue. This type of therapy is growing in popularity, and several organizations throughout

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2222-426: Is considered a continuation of the birthing patterns observed in the wild. Their crias are up and standing, walking, and attempting to suckle within the first hour after birth. Crias are partially fed with llama milk that is lower in fat and salt and higher in phosphorus and calcium than cow or goat milk. A female llama will only produce about 60 millilitres (2 US fluid ounces) of milk at a time when she gives milk, so

2323-526: Is extraordinarily smooth. It is very dry, except for occasional shallow and temporary water sheets after rains. Llano is separated from the plains on the north by the mature consequent valley of the Canadian River , and from the mountains on the west by the broad and probably mature valley of the Pecos River . On the east, it is strongly undercut by the retrogressive erosion of the headwaters of

2424-643: Is generally more expensive but not always more valuable. Alpacas tend to have a more consistent color throughout the body. The most apparent visual difference between llamas and camels is that camels have a humps and llamas do not. Llamas are not ruminants , pseudo-ruminants, or modified ruminants. They do have a complex three-compartment stomach that allows them to digest lower quality, high cellulose foods. The stomach compartments allow for fermentation of tricky foodstuffs, followed by regurgitation and re-chewing. Ruminants (cows, sheep, goats) have four compartments, whereas llamas have only three stomach compartments:

2525-436: Is instinctively very effective in preventing predation. Some llamas bond more quickly to sheep or goats if introduced just before lambing . Many sheep and goat producers indicate a special bond quickly develops between lambs and their guard llama, and the llama is particularly protective of the lambs. Using llamas as guards has reduced the losses to predators for many producers. The value of the livestock saved each year exceeds

2626-506: Is peculiarly elaborate. Known as the Badlands , it is a minutely dissected form with a relief of a few hundred feet. This is due to several causes: The central section of the Great Plains, between latitudes 42° and 36° , occupying eastern Colorado and western Kansas , is mostly a dissected fluviatile plain. That is, this section was once smoothly covered with a gently sloping plain of gravel and sand that had been spread far forward on

2727-638: Is served by several small institutes of technology, education and other technical or junior college-type facilities. Additionally it is home to the Universidad Nacional del Altiplano de Puno (UNAP), which was founded in 1856. Puno features an subtropical highland climate ( Cwb , according to the Köppen climate classification ). As Puno is located at such a high elevation, it experiences more extreme weather conditions than would be expected for its tropical latitude. The average annual temperature

2828-443: Is short, and the fiber is long, woolly, and soft. In essential structural characteristics, as well as in general appearance and habits, all the animals of this genus very closely resemble each other, so whether they should be considered as belonging to one, two, or more species is a matter of controversy among naturalists . The question is complicated by the circumstances of most individuals who have come under observation, either in

2929-658: Is the Kuntur Wasi viewpoint, which has a large metal sculpture of a condor . There are some 700 steps to climb to reach the sculpture. During the celebrations of the Feast of the " Virgen de la Candelaria " and the Regional Competition of Autochthonous Dances. Puno's access to Lake Titicaca is surrounded by 41 floating islands . To this day, the Uros people maintain and live on these man-made islands, depending on

3030-505: Is the mating sound of a llama or alpaca, made by the sexually aroused male. The sound is reminiscent of gargling but with a more forceful, buzzing edge. Males begin the sound when they become aroused and continue throughout copulation . Using llamas as livestock guards in North America began in the early 1980s, and some sheep producers have used llamas successfully since then. Some would even use them to guard their smaller cousins,

3131-485: Is the name for a baby llama, alpaca , vicuña , or guanaco . Crias are typically born with all the herd's females gathering to protect against the male llamas and potential predators. Llamas give birth standing. Birth is usually quick and problem-free, over in less than 30 minutes. Most births occur between 8 am and noon, during the warmer daylight hours. This may increase cria survival by reducing fatalities due to hypothermia during cold Andean nights. This birthing pattern

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3232-594: Is used in the United States to describe a sub-section of the even more vast Interior Plains physiographic division, which covers much of the interior of North America. It also has currency as a region of human geography , referring to the Plains Indians or the Plains states . In Canada the term is rarely used; Natural Resources Canada , the government department responsible for official mapping, treats

3333-455: Is used: the climatic and vegetation regions are more impactful on human settlement than mere topography, and therefore the region is split into (from north to south), the taiga plains, boreal plains , aspen parkland , and prairie ecoregion regions. The northern section of the Great Plains, north of latitude 44° , includes eastern Montana , eastern Wyoming , most of North Dakota and South Dakota , southwestern Minnesota and portions of

3434-563: The American burying beetle (Nicrophorus americus), Salt Creek Tiger Beetle (Cinidela nevadica lincolniana), Great Plains Giant Tiger Beetle (Amblycheila chylindriformis), Microstylum morosum , Bean leaf beetle (Cerotoma trifurcata), Great Plains Camel Cricket (Daihinia brevipes), and the Great plains spittlebug (Lepyronia gibbosa). Some species in the Great plains have gone extinct in

3535-593: The Arikara , Mandan , Pawnee , and Wichita . Wars with the Ojibwe and Cree peoples pushed the Lakota (Teton Sioux) west onto the Great Plains in the mid- to late-17th century. The Shoshone originated in the western Great Basin and spread north and east into present-day Idaho and Wyoming. By 1500, some Eastern Shoshone had crossed the Rocky Mountains into the Great Plains. After 1750, warfare and pressure from

3636-690: The Commission for Environmental Cooperation , a North American Free Trade Agreement (NAFTA) agency composed of the geographical agencies of the Mexican, American, and Canadian governments, uses the "Great Plains" as an ecoregion synonymous with predominant prairies and grasslands rather than as physiographic region defined by topography. The Great Plains ecoregion includes five sub-regions: Temperate Prairies, West-Central Semi-Arid Prairies, South-Central Semi-Arid Prairies, Texas Louisiana Coastal Plains, and Tamaulipas-Texas Semi-Arid Plain, which overlap or expand upon other Great Plains designations. The region

3737-825: The Department of the Interior . In contrast, U.S. Forest Service , an agency of the U. S. Department of Agriculture , administers the National Forests and National Grasslands, under a multiple-use concept. By law, the U.S. Forest Service must consider all resources, with no single resource emphasized to the detriment of others, including water, soil, grazing, timber harvesting, and minerals (mining and drilling), as well as recreation and conservation of fish and wildlife. Each individual state also administers state lands, typically smaller areas, for various purposes including conservation and recreation. Grasslands are among

3838-968: The Great Plains toad ( Anaxyrus cognatus ), plains leopard frog ( Lithobates blairi ), and plains spadefoot toad ( Spea bombifrons ). Some species predominately associated with various river basins in the Great Plains include sturgeon chub ( Macrhybopsis gelida ), peppered chub ( Macrhybopsis tetranema ), prairie chub ( Macrhybopsis australis ), western silvery minnow ( Hybognathus argyritis ), plains minnow ( Hybognathus placitus ), smalleye shiner ( Notropis buccula ), Arkansas River shiner ( Notropis girardi ), Red River shiner ( Notropis bairdi ), Topeka shiner ( Notropis topeka ), plains topminnow ( Fundulus sciadicus ), plains killifish ( Fundulus zebrinus ), Red River pupfish ( Cyprinodon rubrofluviatilis ), and Arkansas darter ( Etheostoma cragini ). The great plains also has many invertebrate species living here both alive and extinct such as

3939-498: The Rocky Mountains and in Central America . Some of the fossil llamas were much larger than current forms. Some species remained in North America during the last ice ages. North American llamas are categorized as an extinct genus, Hemiauchenia . Llama-like animals would have been a common sight 25,000 years ago in modern-day California , Texas , New Mexico , Utah , Missouri , and Florida . The camelid lineage has

4040-521: The United States and Canada late in the 20th century; their descendants now number more than 158,000 llamas and 100,000 alpacas. In Aymara mythology, llamas are important beings. The Heavenly Llama is said to drink water from the ocean and urinates as it rains. According to Aymara eschatology , llamas will return to the water springs and ponds where they come from at the end of time. Lamoids, or llamas (as they are more generally known as

4141-521: The bleak and elevated parts of the mountain range bordering the region of perpetual snow , amidst rocks and precipices, occurring in various suitable localities throughout Peru , in the southern part of Ecuador , and as far south as the middle of Bolivia . Its manners very much resemble those of the chamois of the European Alps ; it is as vigilant, wild, and timid. Vicuña fiber is extremely delicate and soft and highly valued for weaving, but

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4242-487: The camel was soon recognized. They were included in the genus Camelus along with alpaca in the Systema Naturae ( 1758 ) of Carl Linnaeus . They were, however, separated by Georges Cuvier in 1800 under the name of lama along with the guanaco . DNA analysis has confirmed that the guanaco is the wild ancestor of the llama, while the vicuña is the wild ancestor of the alpaca; the latter two were placed in

4343-696: The mixed grass prairie , the tallgrass prairie between the Great Lakes and Appalachian Plateau , and the Taiga Plains and Boreal Plains ecozones in Northern Canada . "Great Plains", or Western Plains , is also the ecoregion of the Great Plains or alternatively the western portion of the Great Plains. The Great Plains lie across both the Central United States and Western Canada , encompassing: The term "Great Plains"

4444-556: The Blackfoot, Crow, Lakota, Cheyenne, and Arapaho pushed Eastern Shoshone south and westward. Some of them moved as far south as Texas, emerging as the Comanche by 1700. The first known contact between Europeans and Indians in the Great Plains occurred in what is now Texas, Kansas, and Nebraska from 1540 to 1542 with the arrival of Francisco Vázquez de Coronado , a Spanish conquistador. In that same period, Hernando de Soto crossed

4545-503: The Canadian provinces including southeastern Alberta , southern Saskatchewan and southwestern Manitoba . The strata here are Cretaceous or early Tertiary , lying nearly horizontal. The surface is shown to be a plain of degradation by a gradual ascent here and there to the crest of a ragged escarpment, the escarpment-remnant of a resistant stratum. There are also the occasional lava -capped mesas and dike formed ridges, surmounting

4646-462: The Center for Great Plains Studies, University of Nebraska–Lincoln . This definition, however, is primarily ecological, not physiographic. The Boreal Plains of Western Canada are physiographically the same, but differentiated by their tundra and forest (rather than grassland) appearance. The term "Great Plains", for the region west of about the 96th meridian west and east of the Rocky Mountains ,

4747-557: The Comanche were among the first to commit to a fully mounted nomadic lifestyle. This occurred by the 1730s, when they had acquired enough horses to put all their people on horseback. The real beginning of the horse culture of the plains began with the Pueblo Revolt of 1680 in New Mexico and the capture of thousands of horses and other livestock. In 1683 a Spanish expedition into Texas found horses among Native people. In 1690,

4848-551: The Great Plains and into the valleys and lower elevations of the eastern Rocky Mountains and portions of the American southwest . Other snakes include the plains hog-nosed snake ( Heterodon nasicus ), western milksnake ( Lampropeltis gentilis ), great plains ratsnake ( Pantherophis emoryi ), bullsnake ( Pituophis catenifer sayi ), plains black-headed snake ( Tantilla nigriceps ), plains gartersnake ( Thamnophis radix ), and lined snake ( Tropidoclonion lineatum ). Reptile diversity increases significantly in southern regions of

4949-629: The Great Plains and that included trade networks west to the Rocky Mountains. Mississippians settled the Great Plains at sites now in Oklahoma and South Dakota . Siouan language speakers may have originated in the lower Mississippi River region. They were agriculturalists and may have been part of the Mound Builder civilization during the 9th–12th centuries. Pressure from other Indian tribes, themselves driven west and south by

5050-588: The Great Plains include National Parks and National Monuments, administers by the National Park Service with the responsibility of preserving ecological and historical places and making them available to the public. The U.S. Fish & Wildlife Service manages the National Wildlife Refuges, with the primary responsibility of conserving and protecting fish, wildlife, plants, and habitat in the public trust. Both are agencies of

5151-476: The Great Plains include the swift fox ( Vulpes velox ) and the endangered black-footed ferret ( Mustela nigripes ). The lesser prairie chicken ( Tympanuchus pallidicinctus ) is endemic to the Great Plains and the distribution of the greater prairie chicken ( Tympanuchus cupido ) predominantly occurs in the region, although the latter historically ranged further eastward. The Harris's sparrow ( Zonotrichia querula ) spends winter months in southern areas of

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5252-512: The Great Plains is the most tornado active area in the world and is sometimes referred to as Tornado Alley . The Great Plains are part of the floristic North American Prairies province , which extends from the Rocky Mountains to the Appalachians . Although the American bison ( Bison bison ) historically ranged throughout much of North America (from New York to Oregon and Canada to northern Mexico), they are strongly associated with

5353-751: The Great Plains where they once roamed in immense herds. Pronghorn ( Antilocapra americana ) range into western areas of the region. The black-tailed prairie dog ( Cynomys ludovicianus ) is another iconic species among several rodents that are linked to the region including the thirteen-lined ground squirrel ( Ictidomys tridecemlineatus ), spotted ground squirrel ( Xerospermophilus spilosoma ), Franklin's ground squirrel ( Poliocitellus franklinii ), plains pocket gopher ( Geomys bursarius ), hispid pocket mouse ( Chaetodipus hispidus ), olive-backed pocket mouse ( Perognathus fasciatus ), plains pocket mouse ( Perognathus flavescens ), and plains harvest mouse ( Reithrodontomys montanus ), Two carnivores associated with

5454-488: The Great Plains. The ornate box turtle ( Terrapene ornata ) and great plains skink ( Plestiodon obsoletus ) occur in southern areas. Although few salamanders are strongly associated with region, the western tiger salamander ( Ambystoma mavortium ) ranges through much of the Great Plains and the Rocky Mountains, as does the Rocky Mountain toad ( Anaxyrus w. woodhousi ). Other anurans related to region include

5555-503: The High Plains) is periodically subjected to extended periods of drought ; high winds in the region may then generate devastating dust storms . The eastern Great Plains near the eastern boundary falls in the humid subtropical climate zone in the southern areas, and the northern and central areas fall in the humid continental climate . Many thunderstorms occur in the plains in the spring through summer. The southeastern portion of

5656-583: The Interior Plains as one unit consisting of several related plateaus and plains. There is no region referred to as the "Great Plains" in the Atlas of Canada . In terms of human geography, the term prairie is more commonly used in Canada, and the region is known as the Canadian Prairies , prairie provinces or simply "the prairies". The North American Environmental Atlas , produced by

5757-897: The Puno folklore. The most important dances are the Wifala de Asillo , the Ichu Carnival, the Tuntuna, the Khashua de Capachica , the Machu-tusuj , the Kcajelo , and the Pandilla Puneña . Llama Camelus glama Linnaeus, 1758 The llama ( / ˈ l ɑː m ə / ; Spanish pronunciation: [ˈʎama] or [ˈʝama] ) ( Lama glama ) is a domesticated South American camelid , widely used as

5858-650: The Red, Brazos, and Colorado rivers of Texas and presents a ragged escarpment approximately 500 to 800 ft (150 to 240 m) high, overlooking the central denuded area of that state. There, between the Brazos and Colorado rivers, occurs a series of isolated outliers capped by limestone that underlies both the Llano Uplift on the west and the Grand Prairies escarpment on the east. The southern and narrow part of

5959-594: The Tragulina ( chevrotains ), the Pecora ( ruminants ), and the Whippomorpha ( hippos and cetaceans , which belong to Artiodactyla from a cladistic , if not traditional, standpoint). The Tylopoda have more or less affinity to each of the sister taxa , standing in some respects in a middle position between them, sharing some characteristics from each, but in others showing special modifications not found in any of

6060-553: The United States participate. When correctly reared, llamas spitting at a human is a rare thing. Llamas are very social herd animals, however, and sometimes spit at each other to discipline lower-ranked llamas. A llama's social rank in a herd is never static. They can always move up or down the social ladder by picking small fights. This is usually done between males to see which will become dominant. Their fights are visually dramatic, characterized by spitting, ramming each other with their chests, neck wrestling, and kicking, mainly to knock

6161-537: The afterlife. The Moche of pre-Columbian Peru depicted llamas quite realistically in their ceramics. In the Inca Empire , llamas were the only beasts of burden, and many of the people dominated by the Inca had long traditions of llama herding. For the Inca nobility, the llama was symbolic, and llama figures were often buried with the dead. In South America, llamas are still used as beasts of burden, as well as for

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6262-464: The alpaca. They are used most commonly in the western regions of the United States , where larger predators, such as coyotes and feral dogs, are prevalent. Typically, a single gelding (castrated male) is used. Research suggests using multiple guard llamas is not as effective as one. Multiple males tend to bond with one another rather than with the livestock and may ignore the flock. A gelded male of two years of age bonds closely with its new charges and

6363-653: The area of the ancient Great Plains for thousands to millions of years. The vast majority of these animals became extinct in North America at the end of the Pleistocene (around 13,000 years ago). A number of significant fossil sites are located in the Great Plains including Agate Fossil Beds National Monument ( Nebraska ), Ashfall Fossil Beds ( Nebraska ), Clayton Lake State Park ( New Mexico ), Dinosaur Valley State Park ( Texas ), Hudson-Meng Bison Kill (Nebraska), Makoshika State Park (Montana), and The Mammoth Site ( South Dakota ). Public and protected lands in

6464-471: The central section, it is for the most part a dissected fluviatile plain. However, the lower lands which surround it on all sides place it in such strong relief that it stands up as a table-land, known from the time of Mexican occupation as the Llano Estacado . It measures roughly 150 mi (240 km) east-west and 400 mi (640 km) north-south. It is of very irregular outline, narrowing to

6565-417: The cria must frequently suckle to receive the nutrients it requires. In harem mating , the male is left with females most of the year. For field mating , a female is turned into a field with a male llama and left there for some time. This is the easiest method in terms of labor but the least useful in predicting a likely birth date. An ultrasound test can be performed, and together with the exposure dates,

6666-425: The domestic state and are variable in size and of many colors, often white, brown, or piebald. Some are grey or black. The guanaco and vicuña are wild. The guanaco is endangered; it has a nearly uniform light-brown color, passing into white below. The guanaco and vicuña certainly differ: The vicuña is more petite, more slender in its proportions, and has a shorter head than the guanaco. The vicuña lives in herds on

6767-531: The encroachment of European settlers as well as economic incentives such as the fur trade, alongside the arrival of the horse and firearms from Europe pushed multiple tribes onto the Great Plains. Among those to have lived on the Great Plains were the Blackfoot , Crow , Sioux , Cheyenne , Arapaho , Comanche , and others. Eastern portions of the Great Plains were inhabited by tribes who lived at Etzanoa and in semi-permanent villages of earth lodges, such as

6868-433: The general level by 500 ft (150 m) or more and manifestly demonstrating the widespread erosion of the surrounding plains. All these reliefs are more plentiful towards the mountains in central Montana. The peneplain is no longer in the cycle of erosion that witnessed its production. It appears to have suffered a regional uplift or increase in elevation, for the upper Missouri River and its branches no longer flow on

6969-581: The genus Vicugna . The genera Lama and Vicugna are, with the two species of true camels, the sole existing representatives of a very distinct section of the Artiodactyla or even-toed ungulates, called Tylopoda , or "bump-footed", from the peculiar bumps on the soles of their feet. The Tylopoda consists of a single family, the Camelidae, and shares the order Artiodactyla with the Suina ( pigs ),

7070-760: The great plains like the Rocky Mountain Locust (Melanoplus spretus). During the Cretaceous Period (145–66 million years ago), the Great Plains were covered by a shallow inland sea called the Western Interior Seaway . However, during the Late Cretaceous to the Paleocene (65–55 million years ago), the seaway had begun to recede, leaving behind thick marine deposits and a relatively flat terrain which

7171-620: The lake for their survival, and are a large tourist destination. Dragon boat racing, an old tradition in Puno, is a very popular activity amongst tourists. Puno is the first major hub in the constant migration of indigenous peoples of the Andes to the larger cities of Peru. It is the largest city in the Southern Altiplano and is the recipient of new residents from surrounding smaller agricultural communities of people seeking better opportunities for education and employment. As such, Puno

7272-501: The largest group. They rise like a large island from the sea, occupying an oval area of about 100 mi (160 km) north-south by 50 mi (80 km) east-west. At Black Elk Peak , they reach an altitude of 7,216 ft (2,199 m) and have an effective relief over the plains of 2,000 or 3,000 ft (610 or 910 m) This mountain mass is of flat-arched, dome-like structure, now well dissected by radiating consequent streams. The weaker uppermost strata have been eroded down to

7373-468: The least protected biomes. Humans have converted much of the prairies for agricultural purposes or to create pastures. Several of the protected lands in the region are centered around aberrant and uncharacteristic features of the region, such as mountains, outcrops, and canyons (e.g. Devil's Tower National Monument , Wind Cave National Park , Scotts Bluff National Monument ), and as splendid and worthy as they are, they are not primarily focused on conserving

7474-426: The level of the plains where their upturned edges are evenly truncated. The next following harder strata have been sufficiently eroded to disclose the core of underlying igneous and metamorphic crystalline rocks in about half of the domed area. In the intermediate section of the plains, between latitudes 44° and 42° , including southern South Dakota and northern Nebraska , the erosion of certain large districts

7575-484: The line that divides the Great Plains into an area that receives 20 in (510 mm) or more of rainfall per year and an area that receives less than 20 in (510 mm). In this context, the High Plains, as well as Southern Alberta , south-western Saskatchewan and Eastern Montana are mainly semi arid steppe land and are generally characterised by rangeland or marginal farmland . The region (especially

7676-456: The lower jaw, the three incisors are long, spatulate, and procumbent; the outer ones are the smallest. Next to these is a curved, suberect canine, followed after an interval by an isolated minute and often deciduous simple conical premolar; then a contiguous series of one premolar and three molars, which differ from those of Camelus in having a small accessory column at the anterior outer edge. The skull generally resembles that of Camelus , with

7777-760: The more common "prairie". The Great Plains are the westernmost portion of the vast North American Interior Plains , which extend east to the Appalachian Plateau . The United States Geological Survey divides the Great Plains in the United States into ten physiographic subdivisions: Further to this can be added Canadian physiographic sub-regions such as the Alberta Plain, Cypress Hills , Manitoba Escarpment (eastward), Manitoba Plain, Missouri Coteau (shared), Rocky Mountain Foothills (eastward), and Saskatchewan Plain. The Great Plains consist of

7878-413: The mountains surrounding the city. There is less than two miles of flat land between the shores and the foothills, which has caused the growing city to continue to expand upwards onto the hillsides. As a result, the town's less developed and poorest areas, which are high on the hillsides, often have very steep streets, which are generally unpaved and cannot be accessed by automobile. Up one of these streets

7979-442: The mountains, the local relief is generally small. The semi-arid climate excludes tree growth and opens far-reaching views. The plains are by no means a simple unit. They are of diverse structure and of various stages of erosional development. They are occasionally interrupted by buttes and escarpments . They are frequently broken by valleys. Yet on the whole, a broadly extended surface of moderate relief so often prevails that

8080-436: The name, Great Plains, for the region as a whole is well-deserved. The western boundary of the plains is usually well-defined by the abrupt ascent of the mountains. The eastern boundary of the plains (in the United States) is more climatic than topographic . The line of 20 in (510 mm) of annual rainfall trends a little east of northward near the 97th meridian. If a boundary must be drawn where nature presents only

8181-429: The other off balance. The females are usually only seen spitting to control other herd members. One may determine how agitated the llama is by the materials in the spit. The more irritated the llama is, the further back into each of the three stomach compartments it will try to draw materials from for its spit. While the social structure might constantly change, they live as a family and care for each other. If one notices

8282-409: The other taxa. The 19th-century discoveries of a vast and previously unexpected extinct Paleogene fauna of North America, as interpreted by paleontologists Joseph Leidy , Edward Drinker Cope , and Othniel Charles Marsh , aided understanding of the early history of this family. Llamas were not always confined to South America ; abundant llama-like remains were found in Pleistocene deposits in

8383-434: The plains and prairies. United States: Canada: the Great Plains biome is found to be at the brink of collapse due to woody plant encroachment , with 62% of Northern American grassland lost to date. The first Peoples ( Paleo-Indians ) arrived on the Great Plains thousands of years ago. The introduction of corn around 800 CE allowed the development of the mound-building Mississippian culture along rivers that crossed

8484-539: The production of fiber and meat . The Inca deity Urcuchillay was depicted in the form of a multicolored llama. Great Plains The Great Plains are a broad expanse of flatland in North America . The region is located just to the east of the Rocky Mountains , much of it covered in prairie , steppe , and grassland . They are the western part of the Interior Plains , which include

8585-416: The quantity that each animal produces is small. Alpacas are primarily descended from wild vicuña ancestors. In contrast, domesticated llamas are descended primarily from wild guanaco ancestors, although a considerable amount of hybridization between the two species has occurred. Differential characteristics between llamas and alpacas include the llama's larger size, longer head, and curved ears. Alpaca fiber

8686-647: The region. Other species migrate from the south in the spring and spend their breeding season on the plains, including the white-faced ibis ( Plegadis chihi ), mountain plover ( Charadrius montanus ), marbled godwit ( Limosa fedoa ), Sprague's pipit ( Anthus spragueii ), Cassin's sparrow ( Peucaea cassinii ), Baird's sparrow ( Centronyx bairdii ), lark bunting ( Calamospiza melanocorys ), chestnut-collared longspur ( Calcarius ornatus ), thick-billed longspur or McCown's longspur ( Rhynchophanes mccownii ), and dickcissel ( Spiza americana ). The prairie rattlesnake ( Crotalus viridis ) ranges throughout much of

8787-474: The river from a better graded preglacial valley by the Pleistocene ice sheet . Here, the ice sheet overspread the plains from the moderately elevated Canadian highlands far on the north-east, instead of from the much higher mountains nearby on the west. The present altitude of the plains near the mountain base is 4,000 ft (1,200 m). The northern plains are interrupted by several small mountain areas. The Black Hills, chiefly in western South Dakota, are

8888-466: The rumen, omasum, and abomasum. In addition, the llama (and other camelids) have an extremely long and complex large intestine (colon). The large intestine's role in digestion is to reabsorb water, vitamins, and electrolytes from food waste passing through it. The length of the llama's colon allows it to survive on much less water than other animals. This is a major advantage in arid climates where they live. Llamas have an unusual reproductive cycle for

8989-565: The seaway had once occupied. During the Cenozoic era , specifically about 25 million years ago during the Miocene and Pliocene epochs, the continental climate became favorable to the evolution of grasslands. Existing forest biomes declined and grasslands became much more widespread. The grasslands provided a new niche for mammals, including many ungulates and glires , that switched from browsing diets to grazing diets. Traditionally,

9090-405: The south. Its altitude is 5,500 ft (1,700 m) at the highest western point, nearest the mountains whence its gravels were supplied. From there, it slopes southeastward at a decreasing rate, first about 12 ft (3.7 m), then about 7 ft/mi (1.3 m/km), to its eastern and southern borders, where it is 2,000 ft (610 m) in altitude. Like the High Plains farther north, it

9191-459: The southern section owes its smoothness to the deposition of imported gravels and sands upon a previously uneven surface by the action of aggrading rivers and their outgoing distributaries. The two sections are also alike in that residual eminences still here and there surmount the peneplain of the northern section, while the fluviatile plain of the central section completely buried the pre-existent relief. An exception to this statement must be made for

9292-416: The southwest, close to the mountains in southern Colorado, where some lava-capped mesas ( Mesa de Maya , Raton Mesa ) stand several thousand feet above the general plain level, and thus testify to the widespread erosion of this region before it was aggraded. The southern section of the Great Plains, between latitudes 35.5° and 25.5°, lies in western Texas , eastern New Mexico , and western Oklahoma . Like

9393-588: The spread of grasslands and the development of grazers have been strongly linked. However, an examination of mammalian teeth suggests that it is the open, gritty habitat and not the grass itself which is linked to diet changes in mammals, giving rise to the " grit, not grass " hypothesis. Paleontological finds in the area have yielded bones of mammoths , saber-toothed cats and other ancient animals, as well as dozens of other megafauna (large animals over 100 lb [45 kg]) – such as giant sloths , horses , mastodons , and American lion – that dominated

9494-602: The subdued forms of the Wichita Mountains in Oklahoma , the westernmost member of the Ouachita system. The term "Western Plains" is used to describe the ecoregion of the Great Plains, or alternatively the western portion of the Great Plains. In general, the Great Plains have a wide range of weather, with very cold and harsh winters and very hot and humid summers. Wind speeds are often very high, especially in winter. The 100th meridian roughly corresponds with

9595-464: The surface of the plain, but in well graded, maturely opened valleys, several hundred feet below the general level. A significant exception to the rule of mature valleys occurs, however, in the case of the Missouri, the largest river, which is broken by several falls on hard sandstones about 50 mi (80 km) east of the mountains. This peculiar feature is explained as the result of displacement of

9696-590: The table-land, called the Edwards Plateau , is more dissected than the rest, and falls off to the south in a frayed-out fault scarp. This scarp overlooks the coastal plain of the Rio Grande embayment . The central denuded area, east of the Llano, resembles the east-central section of the plains in exposing older rocks. Between these two similar areas, in the space limited by the Canadian and Red Rivers, rise

9797-519: The upper jaw, a compressed, sharp, pointed laniariform incisor near the hinder edge of the premaxilla is followed in the male at least by a moderate-sized, pointed, curved true canine in the anterior part of the maxilla. The isolated canine-like premolar that follows in the camels is not present. The teeth of the molar series, which are in contact with each other, consist of two very small premolars (the first almost rudimentary) and three broad molars , generally constructed like those of Camelus . In

9898-445: The way llamas might contribute to the fight against coronaviruses, including MERS and SARS-CoV-2 (which causes COVID-19 ). Scholar Alex Chepstow-Lusty has argued that the switch from a hunter-gatherer lifestyle to widespread agriculture was only possible because of the use of llama dung as fertilizer . The Moche people frequently placed llamas and their parts in the burials of important people as offerings or provisions for

9999-645: Was adopted by European settlers from native Peruvians . The ancestors of llamas are thought to have originated on the Great Plains of North America about 40 million years ago and subsequently migrated to South America about three million years ago during the Great American Interchange . By the end of the last ice age (10,000–12,000 years ago), camelids were extinct in North America. As of 2007, there were over seven million llamas and alpacas in South America. Some were imported to

10100-577: Was not generally used before the early 20th century. Nevin Fenneman's 1916 study Physiographic Subdivision of the United States brought the term Great Plains into more widespread usage. Before that the region was almost invariably called the High Plains, in contrast to the lower Prairie Plains of the Midwestern states . Today the term " High Plains " is used for a subregion of the Great Plains. The term still remains little-used in Canada compared to

10201-613: Was the original home of camelids and that the ancestors of Old World camels crossed over via the Bering Land Bridge from North America. The formation of the Isthmus of Panama three million years ago allowed camelids to spread to South America as part of the Great American Interchange , where they evolved further. Meanwhile, North American camelids died out at the end of the Pleistocene . A full-grown llama can reach

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