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95-552: The rough-skinned newt or roughskin newt ( Taricha granulosa ) is a North American newt known for the strong toxin exuded from its skin. A stocky newt with rounded snout, it ranges from light brown to olive or brownish-black on top, with the underside, including the head, legs, and tail, a contrasting orange to yellow. The skin is granular, but males are smooth-skinned during breeding season. They measure 6 to 9 cm (2.4 to 3.5 in) in snout-to-vent length, and 11 to 18 cm (4.3 to 7.1 in) overall. They are similar to

190-561: A writing system , built huge pyramids and temples , had a complex calendar , and developed the concept of zero around 400 CE. The first recorded European references to North America are in Norse sagas where it is referred to as Vinland . The earliest verifiable instance of pre-Columbian trans-oceanic contact by any European culture with the North America mainland has been dated to around 1000 CE. The site , situated at

285-433: A close interaction between newts and snakes. The whelk predators used their own shell to open the shell of their prey, oftentimes breaking both shells of the predator and prey in the process. This led to the fitness of larger-shelled prey to be higher and then more selected for through generations, however, the predator's population selected for those who were more efficient at opening the larger-shelled prey. This example

380-401: A form of chemical based avoidance behavior to avoid being eaten by predators, mainly the common garter snake. The snakes, after swallowing, digesting, and metabolizing a rough-skinned newt, release a chemical signature. This stimulus can be detected by a nearby newt and trigger an avoidant response, which allows them to minimize predation risks. In this way, newts are able to differentiate whether

475-450: A heightened resistance. The toxin in newts consists of a tradeoff. Each time they release the toxin, they inject themselves with a few milligrams. The TTX becomes concentrated in certain parts of the tissue after passing through cell membranes. As a result of tissue exposure to the toxin, newts have evolved a protection mechanism via a single amino acid substitution to the voltage-gated sodium channel normally affected by TTX. Puffer fishes show

570-416: A mycelium and brought by rain or wind before infecting tubers and leaves. Black colours appear on the plant because of the infection of its cellular system necessary for the multiplication of the oomycete infectious population. The parasite contains virulent-avirulent allelic combinations in several microsatellite loci, likewise the host contains several multiloci resistance genes (or R gene ). That interaction

665-664: A period of mountain building called the Laramide orogeny , between 80 and 55 mya. The formation of the Isthmus of Panama that connected the continent to South America arguably occurred approximately 12 to 15 mya, and the Great Lakes (as well as many other northern freshwater lakes and rivers) were carved by receding glaciers about 10,000 years ago. North America is the source of much of what humanity knows about geologic time periods. The geographic area that would later become

760-590: A powerful earthquake killed seven people in 2009. Volcanic eruptions are common in the region. In 1968, the Arenal Volcano , in Costa Rica, erupted and killed 87 people. Fertile soils from weathered volcanic lavas have made it possible to sustain dense populations in agriculturally productive highland areas. Central America has many mountain ranges ; the longest are the Sierra Madre de Chiapas ,

855-408: A similar amino acid sequence that allows them to survive from their own toxin exposure. Predation on newts by T. sirtalis also shows evidence that tetrodotoxin may serve as protection of eggs by the mother. While TTX is mainly located in the glands of the skin, the rough-skinned newt, as well as some other amphibians also possesses TTX in the ovaries and eggs. The higher the skin toxin levels were in

950-407: A snake is resistant or sensitive to the toxin in order to avoid being preyed upon. However, newts do not avoid the corpses of a recently digested newt that has been left to decompose. This behavior is unlike salamanders that have been documented in avoiding other injured salamanders. Parasites include the trematode Halipegus occidualis , the adult form of which may infest the newt's esophagus and

1045-540: A variety of reactions, including curiosity, trading, cooperation, resignation, and resistance. The indigenous population declined substantially following European arrival, primarily due to the introduction of Eurasian diseases, such as smallpox , to which the indigenous peoples lacked immunity, and because of violent conflicts with Europeans. Indigenous culture changed significantly and their affiliation with political and cultural groups also changed. Several linguistic groups died out , and others changed quite quickly. On

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1140-665: Is a very large continent that extends from north of the Arctic Circle to south of the Tropic of Cancer . Greenland, along with the Canadian Shield , is tundra with average temperatures ranging from 10 to 20 °C (50 to 68 °F), but central Greenland is composed of a very large ice sheet. This tundra radiates throughout Canada, but its border ends near the Rocky Mountains (but still contains Alaska) and at

1235-524: Is an ancient craton which forms the geologic core of North America; it formed between 1.5 and 1.0 billion years ago during the Proterozoic eon. The Canadian Shield is the largest exposure of this craton. From the Late Paleozoic to Early Mesozoic eras, North America was joined with the other modern-day continents as part of the supercontinent Pangaea , with Eurasia to its east. One of

1330-419: Is an excellent example of asymmetrical arms race because while the prey is evolving a physical trait, the predators are adapting in a much different way. Floodplain death adders eat three types of frogs: one nontoxic, one producing mucus when taken by the predator, and the highly toxic frogs, however, the snakes have also found if they wait to consume their toxic prey the potency decreases. In this specific case,

1425-485: Is called gene-for-gene relationship and is, in general, widespread in plant diseases. Expression of genetic patterns in the two species is a combination of resistance and virulence characteristics in order to have the best survival rate. Bats have evolved to use echolocation to detect and catch their prey. Moths have in turn evolved to detect the echolocation calls of hunting bats, and evoke evasive flight maneuvers, or reply with their own ultrasonic clicks to confuse

1520-546: Is in present-day Panama at the Darien Gap on the Colombia -Panama border, placing almost all of Panama within North America. Alternatively, some geologists physiographically locate its southern limit at the Isthmus of Tehuantepec , Mexico, with Central America extending southeastward to South America from this point. The Caribbean islands, or West Indies, are considered part of North America. The continental coastline

1615-648: Is long and irregular. The Gulf of Mexico is the largest body of water indenting the continent, followed by Hudson Bay . Others include the Gulf of Saint Lawrence and the Gulf of California . Before the Central American isthmus formed, the region had been underwater. The islands of the West Indies delineate a submerged former land bridge , which had connected North and South America via what are now Florida and Venezuela . There are several islands off

1710-459: Is much larger than a sodium ion, it acts like a cork in a bottle and prevents the flow of sodium. The reverse binding to sodium channels in nerve cells blocks electrical signals necessary for conducting nerve impulses. This inhibition of firing action potentials has the effect of inducing paralysis and death by asphyxiation. Throughout much of the newt's range, the common garter snake ( Thamnophis sirtalis ) has been observed to exhibit resistance to

1805-559: Is needed to kill any other conceivable predator. Some newts secrete enough toxins to kill several adult humans. It appears that in some areas, the common garter snake has surpassed the newt in the evolutionary arms race by developing such a strong resistance to the toxin that the newt is unable to compete with its production of the toxin. There has been phylogenetic evidence that indicates elevated resistance to TTX has originated independently and only in certain species of garter snakes. The resistance has evolved in at least two unrelated species in

1900-522: Is no longer valid, as specimens that look similar to T.g.m. have been found in areas of Alaska as well. Many newts produce toxins from skin glands as a defense against predation , but the toxins of the genus Taricha are particularly potent. An acrid smell radiates from the newt, which acts as a warning for animals to stay away. Toxicity is generally experienced only if the newt is ingested, although some individuals have been reported to experience skin irritation after dermal contact, particularly if

1995-561: Is part of North America geographically. In a geologic sense, Bermuda is not part of the Americas, but an oceanic island that was formed on the fissure of the Mid-Atlantic Ridge over 100 million years ago (mya). The nearest landmass to it is Cape Hatteras , North Carolina . However, Bermuda is often thought of as part of North America, especially given its historical, political and cultural ties to Virginia and other parts of

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2090-707: Is the Morrison Formation of the western U.S. Canada is geologically one of the oldest regions in the world, with more than half of the region consisting of Precambrian rocks that have been above sea level since the beginning of the Palaeozoic era. Canada's mineral resources are diverse and extensive. Across the Canadian Shield and in the north there are large iron, nickel, zinc , copper, gold, lead, molybdenum , and uranium reserves. Large diamond concentrations have been recently developed in

2185-527: Is the humid subtropical climate . This area has the wettest cities in the contiguous U.S. , with annual precipitation reaching 67 in (1,700 mm) in Mobile, Alabama . Stretching from the borders of the humid continental and subtropical climates, and going west to the Sierra Nevada , south to the southern tip of Durango , north to the border with tundra climate, the steppe / desert climates are

2280-729: The Americas at least 20,000 years ago, but various evidence points to possibly earlier dates. The Paleo-Indian period in North America followed the Last Glacial Period, and lasted until about 10,000 years ago when the Archaic period began. The classic stage followed the Archaic period, and lasted from approximately the 6th to 13th centuries. Beginning in 1000 AD, the Norse were the first Europeans to begin exploring and ultimately colonizing areas of North America. In 1492,

2375-567: The Bering Strait during the Early-Middle Holocene . Prior to the arrival of European explorers and colonists in North America, the natives of North America were divided into many different polities, ranging from small bands of a few families to large empires. They lived in several culture areas , which roughly correspond to geographic and biological zones that defined the representative cultures and lifestyles of

2470-600: The British Columbia Coast , Western Canada , and Northern Canada . In the U.S., they include New England , the Mid-Atlantic , South Atlantic states , East North Central states , West North Central states , East South Central states , West South Central states , Mountain states , and Pacific states . The Great Lakes region and the Pacific Northwest include areas in both Canada and

2565-913: The Caribbean Plate , whereas the Juan de Fuca Plate and Cocos Plate border the North American Plate on its western frontier. The continent can be divided into four great regions (each of which contains many subregions): the Great Plains stretching from the Gulf of Mexico to the Canadian Arctic ; the geologically young, mountainous west, including the Rocky Mountains , the Great Basin , California , and Alaska ;

2660-716: The Caribbean Sea , and to the west and south by the Pacific Ocean . The region includes the Bahamas , Bermuda , Canada , the Caribbean , Central America , Clipperton Island , Greenland , Mexico , Saint Pierre and Miquelon , Turks and Caicos Islands , and the United States . North America covers an area of about 24,709,000 square kilometers (9,540,000 square miles), representing approximately 16.5% of

2755-553: The Cascade Mountains , though occasionally are found (and considered exotic, and possibly artificially introduced) as far as Montana. One isolated population lives in several ponds just north of Moscow, Idaho , and was most likely introduced. A number of subspecies have been defined based on local variants, but only two subspecies have wider recognition: It is now believed that the Taricha granulosa mazamae subspecies

2850-568: The Cordillera Isabelia , and the Cordillera de Talamanca . Between the mountain ranges lie fertile valleys that are suitable for the people; in fact, most of the population of Honduras, Costa Rica, and Guatemala live in valleys. Valleys are also suitable for the production of coffee, beans, and other crops. The indigenous peoples of the Americas have many creation myths , based on which they assert that they have been present on

2945-502: The Earth 's land area and 4.8% of its total surface area. It is the third-largest continent by size after Asia and Africa , and the fourth-largest continent by population after Asia, Africa, and Europe . As of 2021 , North America's population was estimated as over 592 million people in 23 independent states , or about 7.5% of the world's population. In human geography , the terms "North America" and "North American" can refer to Canada,

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3040-465: The East Coast to eastern North Dakota , and stretching down to Kansas , is the humid continental climate featuring intense seasons, with a large amount of annual precipitation, with places like New York City averaging 50 in (1,300 mm). Starting at the southern border of the humid continental climate and stretching to the Gulf of Mexico (whilst encompassing the eastern half of Texas)

3135-620: The Inuit of the high Arctic arrived in North America much later than other native groups, evidenced by the disappearance of Dorset culture artifacts from the archaeological record and their replacement by the Thule people . During the thousands of years of native habitation on the continent, cultures changed and shifted. One of the oldest yet discovered is the Clovis culture (c. 9550–9050 BCE) in modern New Mexico . Later groups include

3230-719: The Mississippian culture and related Mound building cultures, found in the Mississippi River valley and the Pueblo culture of what is now the Four Corners . The more southern cultural groups of North America were responsible for the domestication of many common crops now used around the world, such as tomatoes, squash , and maize . As a result of the development of agriculture in the south, many other cultural advances were made there. The Mayans developed

3325-745: The Red Queen's hypothesis suggested by Leigh Van Valen in 1973. The Bintje potato is derived from a cross between Munstersen and Fransen potato varieties. It was created in the Netherlands in the early 20th century and now is mainly cultivated in the North of France and Belgium. The oomycete Phytophthora infestans is responsible for the potato blight, in particular during the European famine in 1840. Zoospores (mobile spores, characteristics of oomycetes) are liberated by zoosporangia provided from

3420-488: The first official name given to Mexico. North America includes several regions and subregions, each of which have their own respective cultural, economic, and geographic regions. Economic regions include several regions formalized in 20th- and 21st-century trade agreements, including NAFTA between Canada , Mexico , and the United States , and CAFTA between Central America , the Dominican Republic , and

3515-458: The 1490s, building cities, putting the resident indigenous populations to work, raising crops for Spanish settlers and panning gold to enrich the Spaniards. Much of the indigenous population died due to disease and overwork, spurring the Spaniards on to claim new lands and peoples. An expedition under the command of Spanish settler, Hernán Cortés , sailed westward in 1519 to what turned out to be

3610-630: The Arctic, making Canada one of the world's largest producers. Throughout the Shield, there are many mining towns extracting these minerals. The largest, and best known, is Sudbury , Ontario. Sudbury is an exception to the normal process of forming minerals in the Shield since there is significant evidence that the Sudbury Basin is an ancient meteorite impact crater . The nearby, but less-known Temagami Magnetic Anomaly has striking similarities to

3705-503: The California newt ( Taricha torosa ) but differ in having smaller eyes, yellow irises, V-shaped tooth patterns, and uniformly dark eyelids. Males can be distinguished from females during breeding season by large swollen vent lobes and cornified toe pads. Habitats of rough-skinned newts are found throughout the Pacific Northwest . Their range extends south to Santa Cruz, California , and north to Alaska . They are uncommon east of

3800-651: The Caribbean islands. France took the western half of Hispaniola and developed Saint-Domingue as a cane sugar producing colony worked by black slave labor. Britain took Barbados and Jamaica , and the Dutch and Danes took islands previously claimed by Spain. Britain did not begin settling on the North American mainland until a hundred years after the first Spanish settlements, since it sought first to control nearby Ireland . The first permanent English settlement

3895-635: The Cod fish", in reference to the abundance of cod fish on the East Coast . Waldseemüller used the Latin version of Vespucci's name, Americus Vespucius, in its feminine form of "America", following the examples of "Europa", "Asia", and "Africa". Americus originated from Medieval Latin Emericus (see Saint Emeric of Hungary ), coming from the Old High German name Emmerich . Map makers later extended

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3990-419: The North America's southeastern coast, Spanish explorer Juan Ponce de León , who had accompanied Columbus's second voyage, visited and named in 1513 La Florida . As the colonial period unfolded, Spain, England, and France appropriated and claimed extensive territories in North America eastern and southern coastlines. Spain established permanent settlements on the Caribbean islands of Hispaniola and Cuba in

4085-479: The Sudbury Basin. Its magnetic anomalies are very similar to the Sudbury Basin, and so it could be a second metal-rich impact crater. The Shield is also covered by vast boreal forests that support an important logging industry. The United States can be divided into twelve main geological provinces: Each province has its own geologic history and unique features. The geology of Alaska is typical of that of

4180-535: The U.S. North America occupies the northern portion of the landmass generally referred to as the New World, the Western Hemisphere , the Americas, or simply America, which, in many countries, is considered a single continent with North America a subcontinent . North America is the third-largest continent by area after Asia and Africa . North America's only land connection to South America

4275-553: The United States has been the source of more varieties of dinosaurs than any other modern country. According to paleontologist Peter Dodson, this is primarily due to stratigraphy, climate and geography, human resources, and history. Much of the Mesozoic Era is represented by exposed outcrops in the many arid regions of the continent. The most significant Late Jurassic dinosaur-bearing fossil deposit in North America

4370-611: The United States, Mexico, and Greenland or, alternatively, Canada, Greenland and the US (Mexico being classified as part of Latin America ) or simply Canada and the US (Greenland being classified as either Arctic or European (due to its political status as a part of Denmark) and Mexico classified as Latin American). It is unknown with certainty how and when first human populations first reached North America. People were known to live in

4465-550: The United States, and Mexico. France , Italy , Portugal , Spain , Romania , Greece , and the countries of Latin America use a six-continent model , with the Americas viewed as a single continent and North America designating a subcontinent comprising Canada , the United States , Mexico, and Saint Pierre and Miquelon (politically part of France), and often including Greenland and Bermuda . North America has historically been known by other names, including Spanish North America, New Spain , and América Septentrional,

4560-579: The United States. North America is divided linguistically and culturally into two primary regions, Anglo-America and Latin America . Anglo-America includes most of North America, Belize , and Caribbean islands with English -speaking populations. There are also regions, including Louisiana and Quebec , with large Francophone populations; in Quebec , French is the official language. . The southern portion of North America includes Central America and non-English speaking Caribbean nations. The north of

4655-404: The amount of newt then apply a selective pressure favoring snakes with mutations conferring even greater resistance. This cycle of a predator and prey evolving to one another is sometimes termed an evolutionary arms race because the two species compete in developing adaptations and counter adaptations against each other. This has resulted in the newts producing levels of toxin far in excess of what

4750-595: The anterior of its stomach. [REDACTED] Data related to Taricha granulosa at Wikispecies [REDACTED] Media related to Taricha granulosa at Wikimedia Commons North America North America is a continent in the Northern and Western Hemispheres . North America is bordered to the north by the Arctic Ocean , to the east by the Atlantic Ocean , to the southeast by South America and

4845-547: The area had shifted to the Aztec Empire , whose capital city Tenochtitlan was located further north in the Valley of Mexico . The Aztecs were conquered in 1521 by Hernán Cortés . During the so-called Age of Discovery , Europeans explored overseas and staked claims to various parts of North America, much of which was already settled by indigenous peoples. Upon Europeans' arrival in the " New World ", indigenous peoples had

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4940-433: The asymmetry enabled the snakes to overcome the chemical defenses of the toxic frogs after their death. The results of the study showed that the snake became accustomed to the differences in the frogs by their hold and release timing, always holding the nontoxic, while always releasing the highly toxic frogs, with the frogs that discharge mucus somewhere in between. The snakes would also spend generously more time gaped between

5035-423: The bat has just detected their presence or is actively pursuing them. This allows them to decide whether or not defensive ultrasonic clicks are worth the time and energy expenditure. Rough-skinned newts have skin glands that contain a powerful nerve poison, tetrodotoxin , as an anti-predator adaptation . Throughout much of the newt's range, the common garter snake is resistant to the toxin. While in principle

5130-592: The bat's echolocation. The Arctiidae subfamily of Noctuid moths uniquely respond to bat echolocation in three prevailing hypotheses: startle, sonar jamming, and acoustic aposematic defense. All these differences depend on specific environmental settings and the type of echolocation call; however, these hypotheses are not mutually exclusive and can be used by the same moth for defense. The different defense mechanisms have been shown to be directly responsive to bat echolocation through sympatry studies. In places with spatial or temporal isolation between bats and their prey,

5225-479: The best resistant alleles to survive parasitism. As a consequence, allele frequencies vary through time depending on the size of virulent and resistant populations (fluctuation of genetic selection pressure) and generation time (mutation rate) where some genotypes are preferentially selected thanks to the individual fitness gain. Genetic change accumulation in both populations explains a constant adaptation to have lower fitness costs and avoid extinction in accordance with

5320-401: The case of cheetahs and gazelles, where cheetahs evolve to be better at hunting and killing while gazelles evolve not to hunt and kill, but rather to evade capture. Selective pressure between two species can include host-parasite coevolution . This antagonistic relationship leads to the necessity for the pathogen to have the best virulent alleles to infect the organism and for the host to have

5415-432: The continent maintains recognized regions as well. In contrast to the common definition of North America, which encompasses the whole North American continent, the term "North America" is sometimes used more narrowly to refer only to four nations, Canada, Greenland, Mexico, and the U.S. The U.S. Census Bureau includes Saint Pierre and Miquelon, but excludes Mexico from its definition. The term Northern America refers to

5510-981: The continent's coasts; principally, the Arctic Archipelago, the Bahamas , Turks and Caicos , the Greater and Lesser Antilles , the Aleutian Islands (some of which are in the Eastern Hemisphere proper), the Alexander Archipelago , the many thousand islands of the British Columbia Coast, and Newfoundland. Greenland , a self-governing Danish island, and the world's largest , is on the same tectonic plate (the North American Plate ) and

5605-410: The continent. The vast majority of North America is on the North American Plate. Parts of western Mexico, including Baja California, and of California , including the cities of San Diego , Los Angeles , and Santa Cruz , lie on the eastern edge of the Pacific Plate , with the two plates meeting along the San Andreas Fault . The southernmost portion of the continent and much of the West Indies lie on

5700-604: The cordillera, while the major islands of Hawaii consist of Neogene volcanics erupted over a hot spot . Central America is geologically active with volcanic eruptions and earthquakes occurring from time to time. In 1976 Guatemala was hit by a major earthquake , killing 23,000 people; Managua, the capital of Nicaragua, was devastated by earthquakes in 1931 and 1972, the last one killing about 5,000 people; three earthquakes devastated El Salvador, one in 1986 and two in 2001; one earthquake devastated northern and central Costa Rica in 2009, killing at least 34 people; in Honduras

5795-514: The cultures of the region commonly reflect Western traditions . However, relatively small parts of North America in Canada, the United States, Mexico, and Central America have indigenous populations that continue adhering to their respective pre-European colonial cultural and linguistic traditions. The Americas were named after the Italian explorer Amerigo Vespucci by German cartographers Martin Waldseemüller and Matthias Ringmann . Vespucci explored South America between 1497 and 1502, and

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5890-434: The degree of loss or regression depends on the amount of evolutionary time and whether or not the moth species has developed secondary uses for hearing. Some bats are known to use clicks at frequencies above or below moths' hearing ranges. This is known as the allotonic frequency hypothesis. It argues that the auditory systems in moths have driven their bat predators to use higher or lower frequency echolocation to circumvent

5985-447: The driest in the U.S. Highland climates cut from north to south of the continent, where subtropical or temperate climates occur just below the tropics, as in central Mexico and Guatemala . Tropical climates appear in the island regions and in the subcontinent's bottleneck, found in countries and states bathed by the Caribbean Sea or to the south of the Gulf of Mexico and the Pacific Ocean . Precipitation patterns vary across

6080-408: The end of the Canadian Shield, near the Great Lakes . Climate west of the Cascade Range is described as being temperate weather with average precipitation 20 inches (510 millimeters). Climate in coastal California is described to be Mediterranean , with average temperatures in cities like San Francisco ranging from 57 to 70 °F (14 to 21 °C) over the course of the year. Stretching from

6175-544: The exploratory voyages of Christopher Columbus led to a transatlantic exchange , including migrations of European settlers during the Age of Discovery and the early modern period . Present-day cultural and ethnic patterns reflect interactions between European colonists, indigenous peoples , enslaved Africans , immigrants from Europe, Asia, and descendants of these respective groups. Europe's colonization in North America led to most North Americans speaking European languages, such as English , Spanish , and French , and

6270-473: The eyes are touched after handling the animal without washing hands. In 1979, a 29-year-old man from Oregon died after ingesting a rough-skinned newt. The newt produces a neurotoxin called tetrodotoxin (TTX), which in this species was formerly called "tarichatoxin". It is the same toxin found in pufferfish and a number of other marine animals. This toxin targets voltage-gated sodium channels via binding to distinct but allosterically coupled sites. Because TTX

6365-408: The female, the higher the toxin level found in the egg. This is evidence that high toxin levels of the skin may, in fact, be under indirect selection. Since egg toxin levels would ultimately increase the survivability from predators, such as the garter snake, of the offspring, egg toxin levels may be under direct selection by mates, which is detectable via skin toxin levels. The rough skinned newt uses

6460-421: The first American inhabitants sailed from Beringia some 13,000 years ago, with widespread habitation of the Americas during the end of the Last Glacial Period , in what is known as the Late Glacial Maximum , around 12,500 years ago. The oldest petroglyphs in North America date from 15,000 to 10,000 years before present. Genetic research and anthropology indicate additional waves of migration from Asia via

6555-399: The genus Thamnophis and at least twice within T. sirtalis . The toxin, when injected into animals, may not kill resistant animals; however, they are normally slowed down by its toxic effects. In snakes, individuals who showed some resistance tended to move slower after TTX injection, while those with less resistance become paralyzed. Newts are not immune to their own toxin; they only have

6650-795: The geographic center of either the 50 states, the conterminous United States, or the North American continent." Nonetheless, there is a 4.6-meter (15 ft) field stone obelisk in Rugby claiming to mark the center. The North American continental pole of inaccessibility is located 1,650 km (1,030 mi) from the nearest coastline, between Allen and Kyle, South Dakota at 43°22′N 101°58′W  /  43.36°N 101.97°W  / 43.36; -101.97  ( Pole of Inaccessibility North America ) . Canada can be divided into roughly seven physiographic divisions: The lower 48 U.S. states can be divided into roughly eight physiographic divisions: Mexico can be divided into roughly fifteen physiographic divisions: North America

6745-419: The indigenous people who lived there, including the bison hunters of the Great Plains and the farmers of Mesoamerica . Native groups also are classified by their language families , which included Athapascan and Uto-Aztecan languages. Indigenous peoples with similar languages did not always share the same material culture , however, and were not necessarily always allies. Anthropologists speculate that

6840-442: The initial attack. The coevolution is still highly asymmetrical because of the advantage the predators have over their prey. When a species has not been subject to an arms race previously, it may be at a severe disadvantage and face extinction well before it could ever hope to adapt to a new predator, competitor, etc. This should not seem surprising, as one species may have been in evolutionary struggles for millions of years while

6935-468: The land since its creation, but there is no evidence that humans evolved there. The specifics of the initial settlement of the Americas by ancient Asians are subject to ongoing research and discussion. The traditional theory has been that hunters entered the Bering Land Bridge between eastern Siberia and present-day Alaska from 27,000 to 14,000 years ago. A growing viewpoint is that

7030-657: The mainland in Mexico. With local indigenous allies, the Spanish conquered the Aztec empire in central Mexico in 1521. Spain then established permanent cities in Mexico, Central America, and Spanish South America in the sixteenth century. Once Spaniards conquered the high civilization of the Aztecs and Incas, the Caribbean was a backwater of the Spanish empire. Other European powers began to intrude on areas claimed by Spain, including

7125-927: The middle into the main range of the Rockies and the coast ranges in California , Oregon , Washington , and British Columbia , with the Great Basin—a lower area containing smaller ranges and low-lying deserts—in between. The highest peak is Denali in Alaska. The U.S. Geographical Survey (USGS) states that the geographic center of North America is "6 miles [10 km] west of Balta, Pierce County, North Dakota " at about 48°10′N 100°10′W  /  48.167°N 100.167°W  / 48.167; -100.167 , about 24 kilometers (15 mi) from Rugby, North Dakota . The USGS further states that "No marked or monumented point has been established by any government agency as

7220-494: The moth hearing. Barbastelle bats have evolved to use a quieter mode of echolocation, calling at a reduced volume and further reducing the volume of their clicks as they close in on prey moths. The lower volume of clicks reduces the effective successful hunting range, but results in a significantly higher number of moths caught than other, louder bat species. Moths have further evolved the ability to discriminate between high and low echolocation click rates, which indicates whether

7315-411: The moth species hearing mechanism tends to regress. Fullard et al. (2004) compared adventive and endemic Noctiid moth species in a bat-free habitat to ultrasound and found that all of the adventive species reacted to the ultrasound by slowing their flight times, while only one of the endemic species reacted to the ultrasound signal, indicating a loss of hearing over time in the endemic population. However,

7410-402: The name America to North America. In 1538, Gerardus Mercator used the term America on his world map of the entire Western Hemisphere . On his subsequent 1569 map , Mercator called North America "America or New India" ( America sive India Nova ). The Spanish Empire called its territories in North and South America "Las Indias", and the name given to the state body that oversaw the region

7505-451: The newt. Toxin-resistant garter snakes are the only known animals today that can eat a rough-skinned newt and survive. In evolutionary theory, the relationship between the rough-skinned newt and the common garter snake is considered an example of co-evolution . The mutations in the snake's genes that conferred resistance to the toxin have resulted in a selective pressure that favors newts which produce more potent levels of toxin. Increases in

7600-441: The newts producing levels of toxin far in excess of that needed to kill any other predator. In populations where garter snakes and newts live together, higher levels of tetrodotoxin and resistance to it are observed in the two species respectively. Where the species are separated, the toxin levels and resistance are lower. While isolated garter snakes have lower resistance, they still demonstrate an ability to resist low levels of

7695-416: The newts. Successful predation of the rough-skinned newt by the common garter snake is made possible by the ability of individuals in a common garter snake population to gauge whether the newt's level of toxin is too high to feed on. T. sirtalis assays toxin levels of the rough-skinned newt and decides whether or not the levels are manageable by partially swallowing the newt, and either swallowing or releasing

7790-493: The northernmost countries and territories of North America: the U.S., Bermuda , Canada, Greenland, and St. Pierre and Miquelon. Although the term does not refer to a unified region, Middle America includes Mexico, Central America, and the Caribbean. North America's largest countries by land area are Canada and the U.S., both of which have well-defined and recognized subregions. In Canada, these include (from east to west) Atlantic Canada , Central Canada , Canadian Prairies ,

7885-440: The northernmost extent of the island named Newfoundland , has provided unmistakable evidence of Norse settlement. Norse explorer Leif Erikson (c. 970–1020 CE) is thought to have visited the area. Erikson was the first European to make landfall on the continent (excluding Greenland). The Mayan culture was still present in southern Mexico and Guatemala when the Spanish conquistadors arrived, but political dominance in

7980-535: The other might never have faced such pressures. This is a common problem in isolated ecosystems such as Australia or the Hawaiian Islands . In Australia, many invasive species , such as cane toads and rabbits , have spread rapidly due to a lack of competition and a lack of adaptations to cane toad bufotenine on the part of potential predators. Introduced species are a major reason why some indigenous species become endangered or even extinct , as

8075-605: The raised but relatively flat plateau of the Canadian Shield in the northeast; and the varied eastern region, which includes the Appalachian Mountains, the coastal plain along the Atlantic seaboard, and the Florida peninsula. Mexico, with its long plateaus and cordilleras , falls largely in the western region, although the eastern coastal plain does extend south along the Gulf. The western mountains are split in

8170-500: The region, and as such rainforest , monsoon , and savanna types can be found, with rains and high temperatures throughout the year. Notable North American fauna include the bison , black bear , jaguar , cougar , prairie dog , turkey , pronghorn , raccoon , coyote , and monarch butterfly . Notable plants that were domesticated in North America include tobacco , maize , squash , tomato , sunflower , blueberry , avocado , cotton , chile pepper , and vanilla . Laurentia

8265-409: The release of the highly toxic frogs than the short gaped time between the release of the frogs that discharge mucus. Therefore, the snakes have a much higher advantage of being able to cope with the different frogs defensive mechanisms, while the frogs could eventually increase the potency of their toxic knowing the snakes would adapt to that change as well, such as the snakes having venom themselves for

8360-487: The results of the formation of Pangaea was the Appalachian Mountains , which formed some 480 mya, making it among the oldest mountain ranges in the world. When Pangaea began to rift around 200 mya, North America became part of Laurasia , before it separated from Eurasia as its own continent during the mid- Cretaceous period. The Rockies and other western mountain ranges began forming around this time from

8455-434: The tetrodotoxin produced in the newt's skin. While in principle the toxin binds to a tube-shaped protein that acts as a sodium channel in the snake's nerve cells, researchers have identified a genetic disposition in several snake populations where the protein is configured in such a way as to hamper or prevent binding of the toxin. In each of these populations, the snakes exhibit resistance to the toxin and successfully prey upon

8550-493: The toxin binds to a tube-shaped protein that acts as a sodium channel in the snake's nerve cells, a mutation in several snake populations configures the protein in such a way as to hamper or prevent binding of the toxin, conferring resistance. In turn, resistance creates a selective pressure that favors newts that produce more toxin. That in its turn imposes a selective pressure favoring snakes with mutations conferring even greater resistance. This evolutionary arms race has resulted in

8645-547: The toxin, suggesting an ancestral predisposition to tetrodotoxin resistance. The lower levels of resistance in separated populations suggest a fitness cost of both toxin production and resistance. Snakes with high levels of tetrodotoxin resistance crawl more slowly than isolated populations of snakes, making them more vulnerable to predation. The same pattern is seen in isolated populations of newts, which have less toxin in their skin. There are geographic hotspots where levels of tetrodotoxin and resistance are extremely high, showing

8740-549: Was called the Council of the Indies . The United Nations and its statistics division recognize North America as including three regions: Northern America, Central America , and the Caribbean . "Northern America" is a distinct term from "North America", excluding Central America, which itself may or may not include Mexico . In the limited context of regional trade agreements, the term is used to reference three nations: Canada,

8835-817: Was in Jamestown, Virginia in 1607, followed by additional colonial establishments on the east coast from present-day Georgia in the south to Massachusetts in the north, forming the Thirteen Colonies of British America. The English did not establish settlements north or east of the St. Lawrence Valley in present-day Canada until after the conclusion of the American Revolutionary War . Britain's early settlements in present-day Canada included St. John's, Newfoundland in 1630 and Halifax, Nova Scotia in 1749. The first permanent French settlement

8930-517: Was in Quebec City, Quebec in 1608 Evolutionary arms race Arms races may be classified as either symmetrical or asymmetrical. In a symmetrical arms race, selection pressure acts on participants in the same direction. An example of this is trees growing taller as a result of competition for light, where the selective advantage for either species is increased height. An asymmetrical arms race involves contrasting selection pressures, such as

9025-473: Was the first European to suggest that the Americas represented a landmass not then known to the Europeans. In 1507, Waldseemüller published a world map, and placed the word "America" on the continent of present-day South America. The continent north of present-day Mexico was then referred to as Parias. On a 1553 world map published by Petrus Apianus , North America was called "Baccalearum", meaning "realm of

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