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117-606: O. rosmarus rosmarus O. rosmarus divergens O. rosmarus laptevi (debated) The walrus ( Odobenus rosmarus ) is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere . It is the only extant species in the family Odobenidae and genus Odobenus . This species is subdivided into two subspecies :
234-661: A CITES Appendix 3 listing. In October 2017, the Center for Biological Diversity announced they would sue the U.S. Fish and Wildlife Service to force it to classify the Pacific Walrus as a threatened or endangered species. In 1952, walruses in Svalbard were nearly gone due to ivory hunting over a 300 years period, but the Norwegian government banned their commercial hunting and the walruses began to repopulate. By 2018,
351-472: A Germanic language , and it has been attributed largely to either Dutch or Old Norse . Its first part is thought to derive from a word such as Old Norse hvalr ('whale') and the second part has been hypothesized to come from the Old Norse word hross ('horse'). For example, the Old Norse word hrosshvalr means 'horse-whale' and is thought to have been passed in an inverted form to both Dutch and
468-700: A monophyletic origin. A 2021 genetic study found that pinnipeds are more closely related to musteloids. Pinnipeds split from other caniforms 50 million years ago ( mya ) during the Eocene . The earliest fossils of pinnipeds date back to the Late Oligocene . Fossil animals representing basal lineages include Puijila , of the Early Miocene in Arctic Canada. It resembled a modern otter, but shows evidence of quadrupedal swimming—retaining
585-543: A pitch accent . In Modern Greek, all vowels and consonants are short. Many vowels and diphthongs once pronounced distinctly are pronounced as /i/ ( iotacism ). Some of the stops and glides in diphthongs have become fricatives , and the pitch accent has changed to a stress accent . Many of the changes took place in the Koine Greek period. The writing system of Modern Greek, however, does not reflect all pronunciation changes. The examples below represent Attic Greek in
702-440: A diverse and opportunistic diet, feeding on more than 60 genera of marine organisms, including shrimp , crabs, priapulids , spoon worms , tube worms , soft corals , tunicates , sea cucumbers , various mollusks (such as snails , octopuses , and squid ), some types of slow-moving fish , and even parts of other pinnipeds. However, it prefers benthic bivalve mollusks , especially clams , for which it forages by grazing along
819-615: A few, such as the leopard seal , feed on large vertebrates, such as penguins and other seals. Walruses are specialized for feeding on bottom-dwelling mollusks . Male pinnipeds typically mate with more than one female ( polygyny ), though the degree of polygyny varies with the species. The males of land-breeding species tend to mate with a greater number of females than those of ice breeding species . Male pinniped strategies for reproductive success vary between defending females, defending territories that attract females and performing ritual displays or lek mating . Pups are typically born in
936-525: A fifth major dialect group, or it is Mycenaean Greek overlaid by Doric, with a non-Greek native influence. Regarding the speech of the ancient Macedonians diverse theories have been put forward, but the epigraphic activity and the archaeological discoveries in the Greek region of Macedonia during the last decades has brought to light documents, among which the first texts written in Macedonian , such as
1053-506: A form of aquatic locomotion that led to those employed by modern pinnipeds. Potamotherium , which lived in the same period in Europe, was similar to Puijila but more aquatic. The braincase of Potamotherium shows evidence that it used its whiskers to hunt, like modern seals. Both Puijila and Potamotherium fossils have been found in lake deposits, suggesting that seal ancestors were originally adapted for fresh water. Enaliarctos ,
1170-419: A fossil species of late Oligocene /early Miocene (24–22 mya) California , closely resembled modern pinnipeds; it was adapted to an aquatic life with flippers and a flexible spine. Its teeth were more like land predators in that they were more adapted for shearing . Its hind-flippers may have allowed it to walk on land, and it probably did not leave coastal areas as much as its modern relatives. Enaliarctos
1287-586: A layer of fat, or blubber , under the skin to keep warm in cold water, and, other than the walrus, all species are covered in fur. Although pinnipeds are widespread, most species prefer the colder waters of the Northern and Southern Hemispheres. They spend most of their lives in water, but come ashore to mate, give birth, molt or to avoid ocean predators, such as sharks and orcas . Seals mainly live in marine environments but can also be found in fresh water. They feed largely on fish and marine invertebrates ;
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#17327717757571404-565: A peak in the 1970s-1980s, but subsequently, walrus numbers have again declined. Early aerial censuses of Pacific walrus conducted at five-year intervals between 1975 and 1985 estimated populations of above 220,000 in each of the three surveys. In 2006, the population of the Pacific walrus was estimated to be around 129,000 on the basis of an aerial census combined with satellite tracking. There were roughly 200,000 Pacific walruses in 1990. The much smaller population of Atlantic walruses ranges from
1521-550: A prefix /e-/, called the augment . This was probably originally a separate word, meaning something like "then", added because tenses in PIE had primarily aspectual meaning. The augment is added to the indicative of the aorist, imperfect, and pluperfect, but not to any of the other forms of the aorist (no other forms of the imperfect and pluperfect exist). The two kinds of augment in Greek are syllabic and quantitative. The syllabic augment
1638-553: A significant portion (about 42%) of struck and lost animals. Several hundred are removed annually around Greenland. The sustainability of these levels of harvest is difficult to determine given uncertain population estimates and parameters such as fecundity and mortality . The Boone and Crockett Big Game Record book has entries for Atlantic and Pacific walrus. The recorded largest tusks are just over 30 inches and 37 inches long respectively. The effects of global climate change are another element of concern. The extent and thickness of
1755-442: A single ancestor, or diphyletic , recent genetic evidence suggests all three descended from a caniform ancestor most closely related to modern bears. Recent multigene analysis indicates the odobenids and otariids diverged from the phocids about 20–26 million years ago, while the odobenids and the otariids separated 15–20 million years ago. Odobenidae was once a highly diverse and widespread family, including at least twenty species in
1872-522: A single walrus was sighted on the shores of the Baltic Sea - at Rügen Island, Germany , Mielno , Poland and Skälder Bay , Sweden . In July 2022, there was a report of a lost, starving walrus (nicknamed as Stena ) in the coastal waters of the towns of Hamina and Kotka in Kymenlaakso , Finland , that, despite rescue attempts, died of starvation when the rescuers tried to transport it to
1989-542: A strong Northwest Greek influence, and can in some respects be considered a transitional dialect, as exemplified in the poems of the Boeotian poet Pindar who wrote in Doric with a small Aeolic admixture. Thessalian likewise had come under Northwest Greek influence, though to a lesser degree. Pamphylian Greek , spoken in a small area on the southwestern coast of Anatolia and little preserved in inscriptions, may be either
2106-456: A substantial breeding population occurred on isolated Sable Island , 100 mi (160 km) southeast of Nova Scotia and 500 mi (800 km) due east of Portland, Maine , until the early Colonial period. Abundant walrus remains have also been recovered from the southern North Sea dating to the Eemian interglacial period, when that region would have been submerged as it is today, unlike
2223-710: A vital art form. According to Adolf Erik Nordenskiöld , European hunters and Arctic explorers found walrus meat not particularly tasty, and only ate it in case of necessity; however walrus tongue was a delicacy. Walrus hunts are regulated by resource managers in Russia , the United States, Canada, and Greenland , and by representatives of the respective hunting communities. An estimated four to seven thousand Pacific walruses are harvested in Alaska and in Russia, including
2340-510: A vowel or /n s r/ ; final stops were lost, as in γάλα "milk", compared with γάλακτος "of milk" (genitive). Ancient Greek of the classical period also differed in both the inventory and distribution of original PIE phonemes due to numerous sound changes, notably the following: The pronunciation of Ancient Greek was very different from that of Modern Greek . Ancient Greek had long and short vowels ; many diphthongs ; double and single consonants; voiced, voiceless, and aspirated stops ; and
2457-433: A year before weaning, but the young can spend up to five years with the mothers. Walrus milk contains higher amounts of fats and protein compared to land animals but lower compared to phocid seals . This lower fat content in turn causes a slower growth rate among calves and a longer nursing investment for their mothers. Because ovulation is suppressed until the calf is weaned, females give birth at most every two years, leaving
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#17327717757572574-418: Is added to stems beginning with consonants, and simply prefixes e (stems beginning with r , however, add er ). The quantitative augment is added to stems beginning with vowels, and involves lengthening the vowel: Some verbs augment irregularly; the most common variation is e → ei . The irregularity can be explained diachronically by the loss of s between vowels, or that of the letter w , which affected
2691-402: Is because of their massive ankle bones and flatter heels. In water, true seals rely on the side-to-side motion of their hind-flippers and lower body to move forward. The phocid's skull has thickened mastoids , puffed up entotympanic bones , nasal bones with a pointed tip in the back and a non-existent supraorbital foramen. The hip has a more converse ilium . A 2006 molecular study supports
2808-448: Is considered by some linguists to have been closely related to Greek . Among Indo-European branches with living descendants, Greek is often argued to have the closest genetic ties with Armenian (see also Graeco-Armenian ) and Indo-Iranian languages (see Graeco-Aryan ). Ancient Greek differs from Proto-Indo-European (PIE) and other Indo-European languages in certain ways. In phonotactics , ancient Greek words could end only in
2925-552: Is estimated to have been at least 2,300 kg (5,000 lb). The Atlantic subspecies weighs about 10–20% less than the Pacific subspecies. Male Atlantic walrus weigh an average of 900 kg (2,000 lb). The Atlantic walrus also tends to have relatively shorter tusks and somewhat more of a flattened snout . Females weigh about two-thirds as much as males, with the Atlantic females averaging 560 kg (1,230 lb), sometimes weighing as little as 400 kg (880 lb), and
3042-410: Is highly variable, they generally have relatively few teeth other than the tusks. The maximal number of teeth is 38 with dentition formula: 3.1.4.2 3.1.3.2 , but over half of the teeth are rudimentary and occur with less than 50% frequency, such that a typical dentition includes only 18 teeth 1.1.3.0 0.1.3.0 Surrounding the tusks is a broad mat of stiff bristles ("mystacial vibrissae "), giving
3159-411: Is little evidence to prove this. Due to its great size and tusks, the walrus has only two natural predators: the orca and the polar bear . The walrus does not, however, comprise a significant component of either of these predators' diets. Both the orca and the polar bear are also most likely to prey on walrus calves. The polar bear often hunts the walrus by rushing at beached aggregations and consuming
3276-406: Is more like that of true seals, relying less on flippers and more on sinuous whole body movements. Also like phocids, it lacks external ears. The extraocular muscles of the walrus are well-developed. This and its lack of orbital roof allow it to protrude its eyes and see in both a frontal and dorsal direction. However, vision in this species appears to be more suited for short-range. While this
3393-485: Is the "old Sea Vitch—the big, ugly, bloated, pimpled, fat-necked, long-tusked walrus of the North Pacific, who has no manners except when he is asleep". [REDACTED] Pinniped Pinnipeds (pronounced / ˈ p ɪ n ɪ ˌ p ɛ d z / ), commonly known as seals , are a widely distributed and diverse clade of carnivorous , fin -footed, semiaquatic , mostly marine mammals . They comprise
3510-679: The Archaic or Epic period ( c. 800–500 BC ), and the Classical period ( c. 500–300 BC ). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers . It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance . This article primarily contains information about
3627-745: The Caribbean monk seal have become extinct in the past century, while the Mediterranean monk seal and Hawaiian monk seal are ranked as endangered by the International Union for Conservation of Nature . Besides hunting, pinnipeds also face threats from accidental trapping , marine pollution , climate change and conflicts with local people. The name "pinniped" derives from the Latin words pinna ' fin ' and pes, pedis ' foot ' . The common name "seal" originates from
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3744-685: The Chukchi Sea through the relatively narrow Bering Strait . The majority of the population of the Pacific walrus spends its summers north of the Bering Strait in the Chukchi Sea of the Arctic Ocean along the northern coast of eastern Siberia , around Wrangel Island , in the Beaufort Sea along the northern shore of Alaska south to Unimak Island , and in the waters between those locations. Smaller numbers of males summer in
3861-606: The Epic and Classical periods of the language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period ( c. 300 BC ), Ancient Greek was followed by Koine Greek , which is regarded as a separate historical stage, though its earliest form closely resembles Attic Greek , and its latest form approaches Medieval Greek . There were several regional dialects of Ancient Greek; Attic Greek developed into Koine. Ancient Greek
3978-833: The Gulf of Anadyr on the southern coast of the Siberian Chukchi Peninsula , and in Bristol Bay off the southern coast of Alaska, west of the Alaska Peninsula . In the spring and fall, walruses congregate throughout the Bering Strait, reaching from the western coast of Alaska to the Gulf of Anadyr. They winter over in the Bering Sea along the eastern coast of Siberia south to the northern part of
4095-685: The IUCN , while the third is "data deficient". The Pacific walrus is not listed as "depleted" according to the Marine Mammal Protection Act nor as "threatened" or "endangered" under the Endangered Species Act . The Russian Atlantic and Laptev Sea populations are classified as Category 2 (decreasing) and Category 3 (rare) in the Russian Red Book . Global trade in walrus ivory is restricted according to
4212-491: The Kamchatka Peninsula , and along the southern coast of Alaska. A 28,000-year-old fossil walrus was dredged up from the bottom of San Francisco Bay , indicating that Pacific walruses ranged that far south during the last Ice Age . Commercial harvesting reduced the population of the Pacific walrus to between 50,000 and 100,000 in the 1950s–1960s. Limits on commercial hunting allowed the population to increase to
4329-716: The Korkeasaari Zoo for treatment. Walruses prefer shallow shelf regions and forage primarily on the sea floor, often from sea ice platforms. They are not particularly deep divers compared to other pinnipeds; the deepest dives in a study of Atlantic walrus near Svalbard were only 31 ± 17 m (102 ± 56 ft). However, a more recent study recorded dives exceeding 500 metres (1,600 ft) in Smith Sound , between NW Greenland and Arctic Canada – in general, peak dive depth can be expected to depend on prey distribution and seabed depth. The walrus has
4446-990: The Old English word seolh , which is in turn derived from the Proto-Germanic * selkhaz . Bearded seal Hooded seal Ringed seal Baikal seal Caspian seal Spotted seal Harbor seal Grey seal Ribbon seal Harp seal Weddell seal Leopard seal Crabeater seal Ross seal Southern elephant seal Northern elephant seal Mediterranean monk seal Hawaiian monk seal Northern fur seal Steller sea lion California sea lion Galápagos sea lion South American sea lion Australian sea lion New Zealand sea lion Brown fur seal Subantarctic fur seal Antarctic fur seal Guadalupe fur seal Juan Fernández fur seal Antipodean fur seal Galápagos fur seal South American fur seal Walrus The German naturalist Johann Karl Wilhelm Illiger
4563-501: The Pella curse tablet , as Hatzopoulos and other scholars note. Based on the conclusions drawn by several studies and findings such as Pella curse tablet , Emilio Crespo and other scholars suggest that ancient Macedonian was a Northwest Doric dialect , which shares isoglosses with its neighboring Thessalian dialects spoken in northeastern Thessaly . Some have also suggested an Aeolic Greek classification. The Lesbian dialect
4680-442: The aurora borealis is believed to be a special world inhabited by those who died by violence, the changing rays representing deceased souls playing ball with a walrus head. Most of the distinctive 12th-century Lewis Chessmen from northern Europe are carved from walrus ivory, though a few have been found to be made of whales' teeth. Because of its distinctive appearance, great bulk, and immediately recognizable whiskers and tusks,
4797-462: The blastula in suspended development before it implants itself in the uterus. This strategy of delayed implantation , common among pinnipeds, presumably evolved to optimize both the mating season and the birthing season, determined by ecological conditions that promote newborn survival. Calves are born during the spring migration, from April to June. They weigh 45 to 75 kg (99 to 165 lb) at birth and are able to swim. The mothers nurse for over
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4914-472: The continental shelves , spending significant amounts of their lives on the sea ice looking for benthic bivalve molluscs . Walruses are relatively long-lived, social animals, and are considered to be a " keystone species " in the Arctic marine regions. The walrus has played a prominent role in the cultures of many indigenous Arctic peoples , who have hunted it for meat, fat, skin, tusks, and bone. During
5031-603: The epic poems , the Iliad and the Odyssey , and in later poems by other authors. Homeric Greek had significant differences in grammar and pronunciation from Classical Attic and other Classical-era dialects. The origins, early form and development of the Hellenic language family are not well understood because of a lack of contemporaneous evidence. Several theories exist about what Hellenic dialect groups may have existed between
5148-465: The extant families Odobenidae (whose only living member is the walrus ), Otariidae (the eared seals: sea lions and fur seals ), and Phocidae (the earless seals, or true seals), with 34 extant species and more than 50 extinct species described from fossils . While seals were historically thought to have descended from two ancestral lines, molecular evidence supports them as a monophyletic group (descended from one ancestor). Pinnipeds belong to
5265-662: The nasal bones , with a large and flattened supraorbital foramen . An extra spine splits the supraspinatous fossa and bronchi that are divided in the front. Otariids consist of two types: sea lions and fur seals ; the latter typically being smaller, with pointier snouts, longer fore-flippers and heavier fur coats . Five genera and seven species (one now extinct) of sea lion are known to exist, while two genera and nine species of fur seal exist. While sea lions and fur seals have historically been considered separate subfamilies (Otariinae and Arctocephalinae respectively), genetic and molecular evidence has refuted this, indicating that
5382-550: The northern fur seal is basal to other otariids and the Australian sea lion and New Zealand sea lion are more closely related to Arctocephalus than to other sea lions. Odobenidae has only one living member: the walrus . This animal is noticeable from its larger size (exceeded only by the elephant seals ), nearly hairless skin, flattened snout and long upper canines , known as tusks . Like otariids, walruses can walk on land with their hind limbs. When moving in water,
5499-488: The orbital wall . The extinct family Desmatophocidae lived 23–10 mya in the North Pacific. They had long skulls that with large orbits, interlocked zygomatic bones and rounded molars and premolars . They also were sexually dimorphic and may have been capable of swimming with both or either pair of flippers. They are grouped with modern pinnipeds, but there is debate as to whether they are more closely related to phocids or to otariids and walruses. The ancestors of
5616-501: The present , future , and imperfect are imperfective in aspect; the aorist , present perfect , pluperfect and future perfect are perfective in aspect. Most tenses display all four moods and three voices, although there is no future subjunctive or imperative. Also, there is no imperfect subjunctive, optative or imperative. The infinitives and participles correspond to the finite combinations of tense, aspect, and voice. The indicative of past tenses adds (conceptually, at least)
5733-542: The suborder Caniformia of the order Carnivora ; their closest living relatives are musteloids ( weasels , raccoons , skunks and red pandas ), having diverged about 50 million years ago. Seals range in size from the 1 m (3 ft 3 in) and 45 kg (100 lb) Baikal seal to the 5 m (16 ft) and 3,200 kg (7,100 lb) southern elephant seal . Several species exhibit sexual dimorphism . They have streamlined bodies and four limbs that are modified into flippers . Though not as fast in
5850-485: The 1 m (3 ft 3 in) and 45 kg (100 lb) Baikal seal to the 5 m (16 ft) and 3,200 kg (7,100 lb) southern elephant seal . Overall, they tend to be larger than other carnivores. Several species have male-biased sexual dimorphism that depends on how polygynous a species is: highly polygynous species like elephant seals are extremely sexually dimorphic, while less polygynous species have males and females that are closer in size, or, in
5967-455: The 18th and 19th centuries, the walrus was heavily exploited by American and European sealers and whalers , leading to the near- extirpation of the Atlantic subspecies. As early as 1871 traditional hunters were expressing concern about the numbers of walrus being hunted by whaling fleets. Commercial walrus harvesting is now outlawed throughout its range, although Chukchi , Yupik and Inuit peoples are permitted to kill small numbers towards
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#17327717757576084-411: The 19th century and the early 20th century, walrus were widely hunted for their blubber , walrus ivory , and meat. The population of walruses dropped rapidly all around the Arctic region. It has rebounded somewhat since, though the populations of Atlantic and Laptev walruses remain fragmented and at low levels compared with the time before human interference. The origin of the word walrus derives from
6201-1031: The 5th century BC. Ancient pronunciation cannot be reconstructed with certainty, but Greek from the period is well documented, and there is little disagreement among linguists as to the general nature of the sounds that the letters represent. /oː/ raised to [uː] , probably by the 4th century BC. Greek, like all of the older Indo-European languages , is highly inflected. It is highly archaic in its preservation of Proto-Indo-European forms. In ancient Greek, nouns (including proper nouns) have five cases ( nominative , genitive , dative , accusative , and vocative ), three genders ( masculine , feminine , and neuter ), and three numbers (singular, dual , and plural ). Verbs have four moods ( indicative , imperative , subjunctive , and optative ) and three voices (active, middle, and passive ), as well as three persons (first, second, and third) and various other forms. Verbs are conjugated through seven combinations of tenses and aspect (generally simply called "tenses"):
6318-495: The Archaic period of ancient Greek (see Homeric Greek for more details): Μῆνιν ἄειδε, θεά, Πηληϊάδεω Ἀχιλῆος οὐλομένην, ἣ μυρί' Ἀχαιοῖς ἄλγε' ἔθηκε, πολλὰς δ' ἰφθίμους ψυχὰς Ἄϊδι προΐαψεν ἡρώων, αὐτοὺς δὲ ἑλώρια τεῦχε κύνεσσιν οἰωνοῖσί τε πᾶσι· Διὸς δ' ἐτελείετο βουλή· ἐξ οὗ δὴ τὰ πρῶτα διαστήτην ἐρίσαντε Ἀτρεΐδης τε ἄναξ ἀνδρῶν καὶ δῖος Ἀχιλλεύς. The beginning of Apology by Plato exemplifies Attic Greek from
6435-501: The Atlantic and Pacific subspecies indicate very restricted gene flow, but relatively recent separation, estimated at 500,000 and 785,000 years ago. These dates coincide with the hypothesis derived from fossils that the walrus evolved from a tropical or subtropical ancestor that became isolated in the Atlantic Ocean and gradually adapted to colder conditions in the Arctic. The modern walrus is mostly known from Arctic regions, but
6552-585: The Atlantic walrus ( O. r. rosmarus ), which lives in the Atlantic Ocean , and the Pacific walrus ( O. r. divergens ), which lives in the Pacific Ocean . Adult walrus are characterised by prominent tusks and whiskers , and considerable bulk: adult males in the Pacific can weigh more than 2,000 kilograms (4,400 pounds) and, among pinnipeds , are exceeded in size only by the two species of elephant seals . Walrus live mostly in shallow waters above
6669-486: The Canadian Arctic, across Greenland , Svalbard , and the western part of Arctic Russia. There are eight hypothetical subpopulations of Atlantic walruses, based largely on their geographical distribution and movements: five west of Greenland and three east of Greenland. The Atlantic walrus once ranged south to Sable Island , off of Nova Scotia ; as late as the 18th century, they could be found in large numbers in
6786-545: The Dorians. The Greeks of this period believed there were three major divisions of all Greek people – Dorians, Aeolians, and Ionians (including Athenians), each with their own defining and distinctive dialects. Allowing for their oversight of Arcadian, an obscure mountain dialect, and Cypriot, far from the center of Greek scholarship, this division of people and language is quite similar to the results of modern archaeological-linguistic investigation. One standard formulation for
6903-789: The Greater Gulf of St. Lawrence region, sometimes in colonies of 7-8,000 individuals. This population was nearly eradicated by commercial harvest; their current numbers, though difficult to estimate, probably remain below 20,000. In April 2006, the Canadian Species at Risk Act listed the populations of northwestern Atlantic walrus in Québec , New Brunswick, Nova Scotia , Newfoundland and Labrador as having been eradicated in Canada. A genetically distinct population existed in Iceland that
7020-655: The North Atlantic, and likely reached the Pacific via the Central American Seaway . Phocines mainly stayed in the Northern Hemisphere, while the monachines diversified southward. The lineages of Otariidae and Odobenidae split around 20 mya. The earliest fossil records of otariids are in North Pacific and dated to around 11 mya. Early fossil genera include Pithanotaria and Thalassoleon . The Callorhinus lineage split
7137-543: The Otarioidea and Phocidea diverged around 25 mya. Phocids are known to have existed for at least 15 million years, and molecular evidence supports a divergence of the Monachinae and Phocinae lineages around this time. The fossil genera Monotherium and Leptophoca of southeastern North America represent the earliest members of Monachinae and Phocinae respectively. Both lineages may have originated in
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#17327717757577254-426: The Pacific female averaging 800 kg (1,800 lb). Length typically ranges from 2.2 to 3.6 m (7 ft 3 in to 11 ft 10 in). Newborn walruses are already quite large, averaging 33 to 85 kg (73 to 187 lb) in weight and 1 to 1.4 m (3 ft 3 in to 4 ft 7 in) in length across both sexes and subspecies. All told, the walrus is the third largest pinniped species, after
7371-531: The Pacific subspecies far exceeds normal dimensions. In 1909, a walrus hide weighing 500 kg (1,100 lb) was collected from an enormous bull in Franz Josef Land , while in August 1910, Jack Woodson shot a 4.9-metre-long (16 ft) walrus, harvesting its 450 kg (1,000 lb) hide. Since a walrus's hide usually accounts for about 20% of its body weight, the total body mass of these two giants
7488-491: The Pacific, but the importance of seals in the walrus diet is under debate. There have been isolated observations of walruses preying on seals up to the size of a 200 kg (440 lb) bearded seal . Rarely, incidents of walruses preying on seabirds, particularly the Brünnich's guillemot ( Uria lomvia ), have been documented. Walruses may occasionally prey on ice-entrapped narwhals and scavenge on whale carcasses but there
7605-598: The angry father throws the daughter from a high cliff and, as she drops into the water, she turns into a walrus . According to various legends, the tusks are formed either by the trails of mucus from the weeping girl or her long braids. This myth is possibly related to the Chukchi myth of the old walrus-headed woman who rules the bottom of the sea, who is in turn linked to the Inuit goddess Sedna . Both in Chukotka and Alaska ,
7722-550: The aorist. Following Homer 's practice, the augment is sometimes not made in poetry , especially epic poetry. The augment sometimes substitutes for reduplication; see below. Almost all forms of the perfect, pluperfect, and future perfect reduplicate the initial syllable of the verb stem. (A few irregular forms of perfect do not reduplicate, whereas a handful of irregular aorists reduplicate.) The three types of reduplication are: Irregular duplication can be understood diachronically. For example, lambanō (root lab ) has
7839-419: The augment when it was word-initial. In verbs with a preposition as a prefix, the augment is placed not at the start of the word, but between the preposition and the original verb. For example, προσ(-)βάλλω (I attack) goes to προσ έ βαλoν in the aorist. However compound verbs consisting of a prefix that is not a preposition retain the augment at the start of the word: αὐτο(-)μολῶ goes to ηὐ τομόλησα in
7956-683: The biological walrus's appetite for bivalve mollusks, oysters, primarily nearshore and intertidal inhabitants, these organisms in fact comprise an insignificant portion of its diet in captivity. The "walrus" in the cryptic song " I Am the Walrus " by the Beatles is a reference to the Lewis Carroll poem. Another appearance of the walrus in literature is in the story "The White Seal" in Rudyard Kipling 's The Jungle Book , where it
8073-455: The body. Seals are unique among carnivorans in that their orbital walls are mostly shaped by the maxilla and are not contained by certain facial bones. Compared to land carnivores, pinnipeds have fewer teeth, which are pointed and cone-shaped. They are adapted for holding onto slippery prey rather than shearing meat like the carnassials of other carnivorans. The walrus has unique tusks which are long upper canines. Pinnipeds range in size from
8190-399: The case of Antarctic seals, females are moderately bigger. Males of sexually dimorphic species also tend to have secondary sex characteristics , such as larger or more prominent heads, necks, chests, crests , noses/ proboscises and canine teeth as well as thicker fur and manes. Though more polygynous species tend to be sexually dimorphic, some evidence suggests that size differences between
8307-445: The complex songs of Weddell seals . The meat, blubber and skin of pinnipeds have traditionally been used by indigenous peoples of the Arctic . Seals have been depicted in various cultures worldwide. They are commonly kept in captivity and are even sometimes trained to perform tricks and tasks. Once relentlessly hunted by commercial industries for their products, seals are now protected by international law. The Japanese sea lion and
8424-615: The dialect of Sparta ), and Northern Peloponnesus Doric (including Corinthian ). All the groups were represented by colonies beyond Greece proper as well, and these colonies generally developed local characteristics, often under the influence of settlers or neighbors speaking different Greek dialects. After the conquests of Alexander the Great in the late 4th century BC, a new international dialect known as Koine or Common Greek developed, largely based on Attic Greek , but with influence from other dialects. This dialect slowly replaced most of
8541-530: The dialects is: West vs. non-West Greek is the strongest-marked and earliest division, with non-West in subsets of Ionic-Attic (or Attic-Ionic) and Aeolic vs. Arcadocypriot, or Aeolic and Arcado-Cypriot vs. Ionic-Attic. Often non-West is called 'East Greek'. Arcadocypriot apparently descended more closely from the Mycenaean Greek of the Bronze Age. Boeotian Greek had come under
8658-545: The dialects of northern Germany as walros and Walross . An alternative theory is that it comes from the Dutch words wal 'shore' and reus 'giant'. The species name rosmarus is Scandinavian. The Norwegian manuscript Konungs skuggsjá , thought to date from around AD 1240, refers to the walrus as rosmhvalr in Iceland and rostungr in Greenland (walruses were by now extinct in Iceland and Norway, while
8775-567: The divergence of early Greek-like speech from the common Proto-Indo-European language and the Classical period. They have the same general outline but differ in some of the detail. The only attested dialect from this period is Mycenaean Greek , but its relationship to the historical dialects and the historical circumstances of the times imply that the overall groups already existed in some form. Scholars assume that major Ancient Greek period dialect groups developed not later than 1120 BC, at
8892-467: The division of phocids into two monophyletic subfamilies: Monachinae, which consists of elephant seals, monk seals and Antarctic seals ; and Phocinae, which consists of all the rest. One popular hypothesis suggested that pinnipeds are diphyletic (descended from two ancestral lines), with walruses and otariids sharing a recent common ancestor with bears ; and phocids sharing one with Musteloidea . However, morphological and molecular evidence support
9009-550: The earliest, followed by the Eumetopias / Zalophus lineage and then the rest, which colonized the Southern Hemisphere. The earliest fossils of Odobenidae— Prototaria of Japan and Proneotherium of Oregon—date to 18–16 mya. These primitive walruses had normal sized canines and fed on fish instead of mollusks. Later taxa like Gomphotaria , Pontolis and Dusignathus had longer canines on both
9126-621: The end of each summer. Traditional hunters used all parts of the walrus. The meat, often preserved, is an important winter nutrition source; the flippers are fermented and stored as a delicacy until spring; tusks and bone were historically used for tools, as well as material for handicrafts; the oil was rendered for warmth and light; the tough hide made rope and house and boat coverings; and the intestines and gut linings made waterproof parkas. While some of these uses have faded with access to alternative technologies, walrus meat remains an important part of local diets, and tusk carving and engraving remain
9243-459: The increase of stampeding deaths crowding the shorelines of the Chukchi Sea between eastern Russia and western Alaska. Analysis of trends in ice cover published in 2012 indicate that Pacific walrus populations are likely to continue to decline for the foreseeable future, and shift further north, but that careful conservation management might be able to limit these effects. Currently, two of the three walrus subspecies are listed as "least-concern" by
9360-435: The individuals crushed or wounded in the sudden exodus, typically younger or infirm animals. The bears also isolate walruses when they overwinter and are unable to escape a charging bear due to inaccessible diving holes in the ice. However, even an injured walrus is a formidable opponent for a polar bear, and direct attacks are rare. Armed with its ivory tusks, walruses have been known to fatally injure polar bears in battles if
9477-854: The intervening glacial lowstand when the shallow North Sea was dry land. Fossils known from San Francisco , Vancouver , and the Atlantic US coast as far south as North Carolina have been referred to glacial periods. An isolated population in the Laptev Sea was considered by some authorities, including many Russian biologists and the canonical Mammal Species of the World , to be a third subspecies, O. r. laptevi (Chapskii, 1940), but has since been determined to be of Pacific walrus origin. While some outsized Pacific males can weigh as much as 2,000 kg (4,400 lb), most weigh between 800 and 1,700 kg (1,800 and 3,700 lb). An occasional male of
9594-403: The large numbers of organisms actually consumed by the walrus, its foraging has a large peripheral impact on benthic communities. It disturbs ( bioturbates ) the sea floor, releasing nutrients into the water column, encouraging mixing and movement of many organisms and increasing the patchiness of the benthos . Seal tissue has been observed in a fairly significant proportion of walrus stomachs in
9711-498: The largest of any land mammal, both in absolute size and relative to body size. Walruses live to about 20–30 years old in the wild. The males reach sexual maturity as early as seven years, but do not typically mate until fully developed at around 15 years of age. They rut from January through April, decreasing their food intake dramatically. The females begin ovulating as soon as four to six years old. The females are diestrous , coming into heat in late summer and around February, yet
9828-414: The largest tusks typically dominate social groups. Tusks are also used to form and maintain holes in the ice and aid the walrus in climbing out of water onto ice. Tusks were once thought to be used to dig out prey from the seabed, but analyses of abrasion patterns on the tusks indicate they are dragged through the sediment while the upper edge of the snout is used for digging. While the dentition of walruses
9945-483: The latter follows the other into the water, where the bear is at a disadvantage. Polar bear–walrus battles are often extremely protracted and exhausting, and bears have been known to break away from the attack after injuring a walrus. Orcas regularly attack walruses, although walruses are believed to have successfully defended themselves via counterattack against the larger cetacean. However, orcas have been observed successfully attacking walruses with few or no injuries. In
10062-465: The males are fertile only around February; the potential fertility of this second period is unknown. Breeding occurs from January to March, peaking in February. Males aggregate in the water around ice-bound groups of estrous females and engage in competitive vocal displays. The females join them and copulate in the water. Gestation lasts 15 to 16 months. The first three to four months are spent with
10179-461: The nearby ice floes) hunting for bivalves . In March 2021, a single walrus, nicknamed Wally the Walrus , was sighted at Valentia Island , Ireland , far south of its typical range, potentially due to having fallen asleep on an iceberg that then drifted south towards Ireland. Days later, a walrus, thought to be the same animal, was spotted on the Pembrokeshire coast, Wales . In June 2022,
10296-508: The older dialects, although the Doric dialect has survived in the Tsakonian language , which is spoken in the region of modern Sparta. Doric has also passed down its aorist terminations into most verbs of Demotic Greek . By about the 6th century AD, the Koine had slowly metamorphosed into Medieval Greek . Phrygian is an extinct Indo-European language of West and Central Anatolia , which
10413-521: The pack ice has reached unusually low levels in several recent years. The walrus relies on this ice while giving birth and aggregating in the reproductive period. Thinner pack ice over the Bering Sea has reduced the amount of resting habitat near optimal feeding grounds. This more widely separates lactating females from their calves, increasing nutritional stress for the young and lower reproductive rates. Reduced coastal sea ice has also been implicated in
10530-487: The perfect stem eilēpha (not * lelēpha ) because it was originally slambanō , with perfect seslēpha , becoming eilēpha through compensatory lengthening. Reduplication is also visible in the present tense stems of certain verbs. These stems add a syllable consisting of the root's initial consonant followed by i . A nasal stop appears after the reduplication in some verbs. The earliest extant examples of ancient Greek writing ( c. 1450 BC ) are in
10647-540: The population had increased to an estimated 5,503 walruses in the Svalbard area. The walrus plays an important role in the religion and folklore of many Arctic peoples. Skin and bone are used in some ceremonies, and the animal appears frequently in legends. For example, in a Chukchi version of the widespread myth of the Raven , in which Raven recovers the sun and the moon from an evil spirit by seducing his daughter,
10764-491: The sea bottom, searching and identifying prey with its sensitive vibrissae and clearing the murky bottoms with jets of water and active flipper movements. The walrus sucks the meat out by sealing its powerful lips to the organism and withdrawing its piston-like tongue rapidly into its mouth, creating a vacuum. The walrus palate is uniquely vaulted, enabling effective suction. The diet of the Pacific walrus consist almost exclusively of benthic invertebrates (97 percent). Aside from
10881-458: The sexes originated due to ecological differences, with polygyny developing later. Ancient Greek Ancient Greek ( Ἑλληνῐκή , Hellēnikḗ ; [hellɛːnikɛ́ː] ) includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek ( c. 1400–1200 BC ), Dark Ages ( c. 1200–800 BC ),
10998-429: The spring and summer months and females bear almost all the responsibility for raising them. Mothers of some species fast and nurse their young for a relatively short period of time while others take foraging trips at sea between nursing bouts. Walruses are known to nurse their young while at sea. Seals produce a number of vocalizations , notably the barks of California sea lions , the gong -like calls of walruses and
11115-467: The subfamilies Imagotariinae, Dusignathinae and Odobeninae. The key distinguishing feature was the development of a squirt/suction feeding mechanism; tusks are a later feature specific to Odobeninae, of which the modern walrus is the last remaining ( relict ) species. Two subspecies of walrus are widely recognized: the Atlantic walrus, O. r. rosmarus (Linnaeus, 1758) and the Pacific walrus, O. r. divergens (Illiger, 1815). Fixed genetic differences between
11232-406: The superfamily Phocoidea. There are 34 extant species of pinnipeds, and more than 50 fossil species of pinnipedimorphs. Otariids are also known as eared seals due to their pinnae . These animals swim mainly using their well-developed fore-flippers. They can also "walk" on land by shifting their hind-flippers forward under the body. The front end of an otariid's frontal bone protrudes between
11349-517: The syllabic script Linear B . Beginning in the 8th century BC, however, the Greek alphabet became standard, albeit with some variation among dialects. Early texts are written in boustrophedon style, but left-to-right became standard during the classic period. Modern editions of ancient Greek texts are usually written with accents and breathing marks , interword spacing , modern punctuation , and sometimes mixed case , but these were all introduced later. The beginning of Homer 's Iliad exemplifies
11466-467: The time of the Dorian invasions —and that their first appearances as precise alphabetic writing began in the 8th century BC. The invasion would not be "Dorian" unless the invaders had some cultural relationship to the historical Dorians . The invasion is known to have displaced population to the later Attic-Ionic regions, who regarded themselves as descendants of the population displaced by or contending with
11583-428: The two elephant seals . Walruses maintain such a high body weight because of the blubber stored underneath their skin. This blubber keeps them warm and the fat provides energy to the walrus. The walrus's body shape shares features with both sea lions ( eared seals : Otariidae) and seals ( true seals : Phocidae). As with otariids, it can turn its rear flippers forward and move on all fours; however, its swimming technique
11700-762: The upper and lower jaw. The familiar long upper tusks developed in the genera Valenictus and Odobenus . The lineage of the modern walrus may have spread from the North Pacific to the North Atlantic through the Caribbean and Central American Seaway 8–5 mya, and then back to the North Pacific via the Arctic 1 mya, or to the Arctic and subsequently the North Atlantic during the Pleistocene . Pinnipeds have streamlined, spindle-shaped bodies with small or non-existent ear flaps, rounded heads, short muzzles, flexible necks, limbs modified into flippers and small tails. The mammary glands and genitals can withdraw into
11817-429: The vibrissae, the walrus is sparsely covered with fur and appears bald. Its skin is highly wrinkled and thick, up to 10 cm (4 in) around the neck and shoulders of males. The blubber layer beneath is up to 15 cm (6 in) thick. Young walruses are deep brown and grow paler and more cinnamon-colored as they age. Old males, in particular, become nearly pink. Because skin blood vessels constrict in cold water,
11934-492: The walrus a characteristic whiskered appearance. There can be 400 to 700 vibrissae in 13 to 15 rows reaching 30 cm (12 in) in length, though in the wild they are often worn to much shorter lengths due to constant use in foraging. The vibrissae are attached to muscles and are supplied with blood and nerves, making them highly sensitive organs capable of differentiating shapes 3 mm ( 1 ⁄ 8 in) thick and 2 mm ( 3 ⁄ 32 in) wide. Aside from
12051-688: The walrus also appears in the popular cultures of peoples with little direct experience with the animal, particularly in English children's literature. Perhaps its best-known appearance is in Lewis Carroll 's whimsical poem " The Walrus and the Carpenter " that appears in his 1871 book Through the Looking-Glass . In the poem, the eponymous antiheroes use trickery to consume a great number of oysters . Although Carroll accurately portrays
12168-419: The walrus can appear almost white when swimming. As a secondary sexual characteristic , males also acquire significant nodules, called "bosses", particularly around the neck and shoulders. The walrus has an air sac under its throat which acts like a flotation bubble and allows it to bob vertically in the water and sleep. The males possess a large baculum (penis bone), up to 63 cm (25 in) in length,
12285-662: The walrus in the Carta Marina in 1539, first referred to the walrus as the ros marus , probably a Latinization of morž , and this was adopted by Linnaeus in his binomial nomenclature . The coincidental similarity between morse and the Latin word morsus ('a bite') supposedly contributed to the walrus's reputation as a "terrible monster". The compound Odobenus comes from odous ( Greek for 'teeth') and baino (Greek for 'walk'), based on observations of walruses using their tusks to pull themselves out of
12402-445: The walrus relies on its hind limbs for locomotion, while its forelimbs are used for steering. Also, it has no outer ears. The epipterygoid of the jaw is well developed and the back of the nasal bones are horizontal. In the feet, the calcaneuses protrude in the middle. Phocids are known as true or "earless" seals. These animals lack outer ears and cannot position their hind-flippers to move on land, making them more cumbersome. This
12519-413: The walrus with the lowest reproductive rate of any pinniped. The rest of the year (late summer and fall), walruses tend to form massive aggregations of tens of thousands of individuals on rocky beaches or outcrops. The migration between the ice and the beach can be long-distance and dramatic. In late spring and summer, for example, several hundred thousand Pacific walruses migrate from the Bering Sea into
12636-704: The water as dolphins , seals are more flexible and agile. Otariids primarily use their front limbs to propel themselves through the water, while phocids and walruses primarily use their hind limbs for this purpose. Otariids and walruses have hind limbs that can be pulled under the body and used as legs on land. By comparison, terrestrial locomotion by phocids is more cumbersome. Otariids have visible external ears, while phocids and walruses lack these. Pinnipeds have well-developed senses—their eyesight and hearing are adapted for both air and water, and they have an advanced tactile system in their whiskers or vibrissae. Some species are well adapted for diving to great depths. They have
12753-587: The water. The term divergens in Latin means 'turning apart', referring to their tusks. The Inuttitut term for the creature is aivik , similar to the Inuktitut word: aiviq ᐊᐃᕕᖅ. The walrus is a mammal in the order Carnivora . It is the sole surviving member of the family Odobenidae, one of three lineages in the suborder Pinnipedia along with true seals ( Phocidae ) and eared seals ( Otariidae ). While there has been some debate as to whether all three lineages are monophyletic , i.e. descended from
12870-971: The word evolved in Greenland). Several place names in Iceland, Greenland and Norway may originate from walrus sites: Hvalfjord, Hvallatrar and Hvalsnes to name some, all being typical walrus breeding grounds. The archaic English word for walrus— morse —is widely thought to have come from the Slavic languages , which in turn borrowed it from Finno-Ugric languages, and ultimately (according to Ante Aikio ) from an unknown Pre-Finno-Ugric substrate language of Northern Europe. Compare морж ( morž ) in Russian, mursu in Finnish , morša in Northern Saami , and morse in French. Olaus Magnus , who depicted
12987-534: The world. In 1989, Annalisa Berta and colleagues proposed the unranked clade Pinnipedimorpha to contain the fossil genus Enaliarctos and modern seals as a sister group . Pinnipeds belong to the order Carnivora and the suborder Caniformia (known as dog-like carnivorans). Of the three extant families, the Otariidae and Odobenidae are grouped in the superfamily Otarioidea, while the Phocidae belong to
13104-480: Was Aeolic. For example, fragments of the works of the poet Sappho from the island of Lesbos are in Aeolian. Most of the dialect sub-groups listed above had further subdivisions, generally equivalent to a city-state and its surrounding territory, or to an island. Doric notably had several intermediate divisions as well, into Island Doric (including Cretan Doric ), Southern Peloponnesus Doric (including Laconian ,
13221-452: Was a pluricentric language , divided into many dialects. The main dialect groups are Attic and Ionic , Aeolic , Arcadocypriot , and Doric , many of them with several subdivisions. Some dialects are found in standardized literary forms in literature , while others are attested only in inscriptions. There are also several historical forms. Homeric Greek is a literary form of Archaic Greek (derived primarily from Ionic and Aeolic) used in
13338-495: Was likely more of a fore-flipper swimmer, but could probably swim with either pair. One species, Enaliarctos emlongi , exhibited notable sexual dimorphism , suggesting that this physical characteristic may have been an important driver of pinniped evolution. A closer relative of extant pinnipeds was Pteronarctos , which lived in Oregon 19–15 mya. As in modern seals, the maxilla or upper jaw bone of Pteroarctos intersects with
13455-404: Was not true of all extinct walruses, the most prominent feature of the living species is its long tusks. These are elongated canines , which are present in both male and female walruses and can reach a length of 1 m (3 ft 3 in) and weigh up to 5.4 kg (12 lb). Tusks are slightly longer and thicker among males, which use them for fighting, dominance and display; the strongest males with
13572-527: Was the first to recognize the pinnipeds as a distinct taxonomic unit; in 1811 he gave the name Pinnipedia to both a family and an order . American zoologist Joel Asaph Allen reviewed the world's pinnipeds in an 1880 monograph , History of North American pinnipeds, a monograph of the walruses, sea-lions, sea-bears and seals of North America . In this publication, he traced the history of names, gave keys to families and genera, described North American species and provided synopses of species in other parts of
13689-517: Was wiped out after Norse settlement around 1213–1330 AD. An isolated population is restricted, year-round, to the central and western regions of the Laptev Sea , from the eastern Kara Sea to the westernmost regions of the East Siberian Sea . The current population of these Laptev walruses has been estimated at between 5-10,000. Even though walruses can dive to depths beyond 500 meters, they spend most of their time in shallow waters (and
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