136-504: A hand is a body part. Hand or HAND may also refer to: Hand A hand is a prehensile , multi- fingered appendage located at the end of the forearm or forelimb of primates such as humans , chimpanzees , monkeys , and lemurs . A few other vertebrates such as the koala (which has two opposable thumbs on each "hand" and fingerprints extremely similar to human fingerprints ) are often described as having "hands" instead of paws on their front limbs. The raccoon
272-624: A gazetteer compiled by Chang Qu ( 常璩 ) during the Western Jin Dynasty (265–316), reported the discovery of dragon bones at Wucheng in Sichuan Province. Villagers in central China have long unearthed fossilized "dragon bones" for use in traditional medicines . In Europe , dinosaur fossils were generally believed to be the remains of giants and other biblical creatures. Scholarly descriptions of what would now be recognized as dinosaur bones first appeared in
408-523: A sacrum composed of three or more fused vertebrae (three are found in some other archosaurs, but only two are found in Herrerasaurus ); and a perforate acetabulum , or hip socket, with a hole at the center of its inside surface (closed in Saturnalia tupiniquim , for example). Another difficulty of determining distinctly dinosaurian features is that early dinosaurs and other archosaurs from
544-421: A "pillar-erect" configuration of the hip joint, where instead of having a projection from the femur insert on a socket on the hip, the upper pelvic bone was rotated to form an overhanging shelf. Dinosaur fossils have been known for millennia, although their true nature was not recognized. The Chinese considered them to be dragon bones and documented them as such. For example, Huayang Guo Zhi ( 華陽國志 ),
680-427: A compact fist, presumably for fighting purposes. The fist is compact and thus effective as a weapon. It also provides protection for the fingers. However, this is not widely accepted to be one of the primary selective pressures acting on hand morphology throughout human evolution, with tool use and production being thought to be far more influential. Dinosaur Dinosaurs are a diverse group of reptiles of
816-488: A controversial taxon that was recently confirmed to exist after archived photos were uncovered. Bruhathkayosaurus was a titanosaur and would have most likely weighed more than even Marrapunisaurus . Recent size estimates in 2023 have placed this sauropod reaching lengths of up to 44 m (144 ft) long and a colossal weight range of around 110 000 – 170 000 kg ( 240 000 – 370 000 lb), if these upper estimates up true, Bruhathkayosaurus would have rivaled
952-448: A corresponding inwardly facing distinct head on the femur. Their erect posture enabled early dinosaurs to breathe easily while moving, which likely permitted stamina and activity levels that surpassed those of "sprawling" reptiles . Erect limbs probably also helped support the evolution of large size by reducing bending stresses on limbs. Some non-dinosaurian archosaurs, including rauisuchians , also had erect limbs but achieved this by
1088-652: A dinosaur. In it he described and named a sauropod tooth , " Rutellum impicatum ", that had been found in Caswell, near Witney , Oxfordshire. Between 1815 and 1824, the Rev William Buckland , the first Reader of Geology at the University of Oxford, collected more fossilized bones of Megalosaurus and became the first person to describe a non-avian dinosaur in a scientific journal . The second non-avian dinosaur genus to be identified, Iguanodon ,
1224-537: A femur of Tyrannosaurus preserved soft, flexible tissue within, including blood vessels , bone matrix , and connective tissue (bone fibers) that had retained their microscopic structure. This discovery suggested that original soft tissues could be preserved over geological time, with multiple mechanisms having been proposed. Later, in 2009, Schweitzer and colleagues reported that a Brachylophosaurus femur preserved similar microstructures, and immunohistochemical techniques (based on antibody binding) demonstrated
1360-422: A group known as archosaurs, which also includes modern crocodilians. Within the archosaur group, dinosaurs are differentiated most noticeably by their gait. Dinosaur legs extend directly beneath the body, whereas the legs of lizards and crocodilians sprawl out to either side. Collectively, dinosaurs as a clade are divided into two primary branches, Saurischia and Ornithischia. Saurischia includes those taxa sharing
1496-399: A higher degree, the hands of other primates are anatomically similar and the dexterity of the human hand can not be explained solely on anatomical factors. The neural machinery underlying hand movements is a major contributing factor; primates have evolved direct connections between neurons in cortical motor areas and spinal motoneurons , giving the cerebral cortex monosynaptic control over
SECTION 10
#17327810388551632-486: A more recent common ancestor with birds than with Ornithischia, while Ornithischia includes all taxa sharing a more recent common ancestor with Triceratops than with Saurischia. Anatomically, these two groups can be distinguished most noticeably by their pelvic structure. Early saurischians—"lizard-hipped", from the Greek sauros ( σαῦρος ) meaning "lizard" and ischion ( ἰσχίον ) meaning "hip joint"—retained
1768-583: A now lost partial vertebral neural arch described in 1878. Extrapolating from the illustration of this bone, the animal may have been 58 meters (190 ft) long and weighed 122 400 kg ( 269 800 lb). However, recent research have placed Amphicoelias from the long, gracile diplodocid to the shorter but much stockier rebbachisaurid. Now renamed as Maraapunisaurus , this sauropod now stands as much as 40 meters (130 ft) long and weigh as much as 120 000 kg ( 260 000 lb). Another contender of this title includes Bruhathkayosaurus ,
1904-523: A pencil—reflects individual brain functioning. Among humans, the hands play an important function in body language and sign language . Likewise, the ten digits of two hands and the twelve phalanges of four fingers (touchable by the thumb) have given rise to number systems and calculation techniques. Many mammals and other animals have grasping appendages similar in form to a hand such as paws , claws , and talons, but these are not scientifically considered to be grasping hands. The scientific use of
2040-433: A period of time, with estimates ranging from 5–10 million years to 21 million years. When dinosaurs appeared, they were not the dominant terrestrial animals. The terrestrial habitats were occupied by various types of archosauromorphs and therapsids , like cynodonts and rhynchosaurs . Their main competitors were the pseudosuchians , such as aetosaurs , ornithosuchids and rauisuchians, which were more successful than
2176-663: A potential food source, radiated in the Late Triassic. Early sauropodomorphs did not have sophisticated mechanisms for processing food in the mouth, and so must have employed other means of breaking down food farther along the digestive tract. The general homogeneity of dinosaurian faunas continued into the Middle and Late Jurassic, where most localities had predators consisting of ceratosaurians , megalosauroids , and allosauroids , and herbivores consisting of stegosaurian ornithischians and large sauropods. Examples of this include
2312-681: A presentation about fossil reptiles to the British Association for the Advancement of Science in 1841, but reports of the time show that Owen did not mention the word "dinosaur", nor recognize dinosaurs as a distinct group of reptiles in his address. He introduced the Dinosauria only in the revised text version of his talk published in April 1842. With the backing of Prince Albert , the husband of Queen Victoria , Owen established
2448-399: A primitive trait, while the palms of other extant higher primates are elongated to the extent that some of the thumb's original function has been lost (most notably in highly arboreal primates such as the spider monkey ). In humans, the big toe is thus more derived than the thumb. There is a hypothesis suggesting the form of the modern human hand is especially conducive to the formation of
2584-599: A review paper by Paul Sereno in 1998, were accompanied by increases in the number of published phylogenetic trees for dinosaurs. Dinosaur fossils are not limited to bones, but also include imprints or mineralized remains of skin coverings, organs, and other tissues. Of these, skin coverings based on keratin proteins are most easily preserved because of their cross-linked , hydrophobic molecular structure. Fossils of keratin-based skin coverings or bony skin coverings are known from most major groups of dinosaurs. Dinosaur fossils with scaly skin impressions have been found since
2720-549: A series of studies that likewise argued for active lifestyles in dinosaurs based on anatomical and ecological evidence (see § Physiology ), which were subsequently summarized in his 1986 book The Dinosaur Heresies . New revelations were supported by an increase in dinosaur discoveries. Major new dinosaur discoveries have been made by paleontologists working in previously unexplored regions, including India, South America, Madagascar, Antarctica, and most significantly China. Across theropods, sauropodomorphs, and ornithischians,
2856-603: A sister group to Dinosauria, including a large anterior trochanter, metatarsals II and IV of subequal length, reduced contact between ischium and pubis, the presence of a cnemial crest on the tibia and of an ascending process on the astragalus, and many others. A variety of other skeletal features are shared by dinosaurs. However, because they either are common to other groups of archosaurs or were not present in all early dinosaurs, these features are not considered to be synapomorphies. For example, as diapsids , dinosaurs ancestrally had two pairs of Infratemporal fenestrae (openings in
SECTION 20
#17327810388552992-427: A transitional element to the fifth metacarpal. Together with the thumb, the four fingers form four oblique arches, of which the arch of the index finger functionally is the most important, especially for precision grip, while the arch of the little finger contribute an important locking mechanism for power grip. The thumb is undoubtedly the "master digit" of the hand, giving value to all the other fingers. Together with
3128-490: A universally agreed-upon list of their distinguishing features, nearly all dinosaurs discovered so far share certain modifications to the ancestral archosaurian skeleton, or are clearly descendants of older dinosaurs showing these modifications. Although some later groups of dinosaurs featured further modified versions of these traits, they are considered typical for Dinosauria; the earliest dinosaurs had them and passed them on to their descendants. Such modifications, originating in
3264-468: A variety of waterbirds , diversified rapidly at the beginning of the Paleogene period, entering ecological niches left vacant by the extinction of Mesozoic dinosaur groups such as the arboreal enantiornithines , aquatic hesperornithines , and even the larger terrestrial theropods (in the form of Gastornis , eogruiids , bathornithids , ratites, geranoidids , mihirungs , and " terror birds "). It
3400-430: Is 39.7 meters (130 ft) long. There were larger dinosaurs, but knowledge of them is based entirely on a small number of fragmentary fossils. Most of the largest herbivorous specimens on record were discovered in the 1970s or later, and include the massive Argentinosaurus , which may have weighed 80 000 to 100 000 kilograms (88 to 110 short tons) and reached lengths of 30 to 40 meters (98 to 131 ft); some of
3536-710: Is a synapsid ). None of them had the erect hind limb posture characteristic of true dinosaurs. Dinosaurs were the dominant terrestrial vertebrates of the Mesozoic Era , especially the Jurassic and Cretaceous periods. Other groups of animals were restricted in size and niches; mammals , for example, rarely exceeded the size of a domestic cat and were generally rodent-sized carnivores of small prey. Dinosaurs have always been recognized as an extremely varied group: over 900 non-avian dinosaur genera have been confidently identified (2018) with 1124 species (2016). Estimates put
3672-470: Is an inexact art, and reconstructing the muscles and other organs of the living animal is, at best, a process of educated guesswork. The tallest and heaviest dinosaur known from good skeletons is Giraffatitan brancai (previously classified as a species of Brachiosaurus ). Its remains were discovered in Tanzania between 1907 and 1912. Bones from several similar-sized individuals were incorporated into
3808-755: Is based in part on preservation bias , as large, sturdy bones are more likely to last until they are fossilized. Many dinosaurs were quite small, some measuring about 50 centimeters (20 inches) in length. The first dinosaur fossils were recognized in the early 19th century, with the name "dinosaur" (meaning "terrible lizard") being coined by Sir Richard Owen in 1842 to refer to these "great fossil lizards". Since then, mounted fossil dinosaur skeletons have been major attractions at museums worldwide, and dinosaurs have become an enduring part of popular culture . The large sizes of some dinosaurs, as well as their seemingly monstrous and fantastic nature, have ensured their regular appearance in best-selling books and films, such as
3944-466: Is between 1 and 10 metric tons (1.1 and 11.0 short tons). This contrasts sharply with the average size of Cenozoic mammals, estimated by the National Museum of Natural History as about 2 to 5 kg (4.4 to 11.0 lb). The sauropods were the largest and heaviest dinosaurs. For much of the dinosaur era, the smallest sauropods were larger than anything else in their habitat, and the largest
4080-621: Is derived from a variety of fossil and non-fossil records, including fossilized bones, feces , trackways , gastroliths , feathers , impressions of skin, internal organs and other soft tissues . Many fields of study contribute to our understanding of dinosaurs, including physics (especially biomechanics ), chemistry , biology , and the Earth sciences (of which paleontology is a sub-discipline). Two topics of particular interest and study have been dinosaur size and behavior. Current evidence suggests that dinosaur average size varied through
4216-474: Is located on one of the sides, parallel to the arm. A reliable way of identifying human hands is from the presence of opposable thumbs. Opposable thumbs are identified by the ability to be brought opposite to the fingers, a muscle action known as opposition. The skeleton of the human hand consists of 27 bones: the eight short carpal bones of the wrist are organized into a proximal row ( scaphoid , lunate , triquetral and pisiform ) which articulates with
Hand (disambiguation) - Misplaced Pages Continue
4352-591: Is now at the American Museum of Natural History in New York City, while Marsh's is at the Peabody Museum of Natural History at Yale University . World War II caused a pause in palaeontological research; after the war, research attention was also diverted increasingly to fossil mammals rather than dinosaurs, which were seen as sluggish and cold-blooded. At the end of the 1960s, however,
4488-654: Is often stated that mammals out-competed the neornithines for dominance of most terrestrial niches but many of these groups co-existed with rich mammalian faunas for most of the Cenozoic Era. Terror birds and bathornithids occupied carnivorous guilds alongside predatory mammals, and ratites are still fairly successful as midsized herbivores; eogruiids similarly lasted from the Eocene to Pliocene , becoming extinct only very recently after over 20 million years of co-existence with many mammal groups. Dinosaurs belong to
4624-408: Is supplemented by the precision grip between the thumb and the distal finger pads made possible by the opposable thumbs. Hominidae (great apes including humans) acquired an erect bipedal posture about 3.6 million years ago , which freed the hands from the task of locomotion and paved the way for the precision and range of motion in human hands. Functional analyses of the features unique to
4760-497: Is to the neck of a metacarpal. One can also have a broken finger . The prehensile hands and feet of primates evolved from the mobile hands of semi- arboreal tree shrews that lived about 60 million years ago . This development has been accompanied by important changes in the brain and the relocation of the eyes to the front of the face, together allowing the muscle control and stereoscopic vision necessary for controlled grasping. This grasping, also known as power grip,
4896-505: Is usually described as having "hands" though opposable thumbs are lacking. Some evolutionary anatomists use the term hand to refer to the appendage of digits on the forelimb more generally—for example, in the context of whether the three digits of the bird hand involved the same homologous loss of two digits as in the dinosaur hand. The human hand usually has five digits: four fingers plus one thumb ; these are often referred to collectively as five fingers , however, whereby
5032-535: The Jurassic Park franchise. Persistent public enthusiasm for the animals has resulted in significant funding for dinosaur science, and new discoveries are regularly covered by the media. Under phylogenetic nomenclature , dinosaurs are usually defined as the group consisting of the most recent common ancestor (MRCA) of Triceratops and modern birds (Neornithes), and all its descendants. It has also been suggested that Dinosauria be defined with respect to
5168-437: The blue whale and Perucetus colossus as one of the largest animals to have ever existed. The largest carnivorous dinosaur was Spinosaurus , reaching a length of 12.6 to 18 meters (41 to 59 ft) and weighing 7 to 20.9 metric tons (7.7 to 23.0 short tons). Other large carnivorous theropods included Giganotosaurus , Carcharodontosaurus , and Tyrannosaurus . Therizinosaurus and Deinocheirus were among
5304-695: The Anisian epoch of the Triassic, approximately 243 million years ago, which is the age of Nyasasaurus from the Manda Formation of Tanzania. However, its known fossils are too fragmentary to identify it as a dinosaur or only a close relative. The referral of the Manda Formation to the Anisian is also uncertain. Regardless, dinosaurs existed alongside non-dinosaurian ornithodirans for
5440-462: The Carnian pluvial event . Dinosaur evolution after the Triassic followed changes in vegetation and the location of continents. In the Late Triassic and Early Jurassic, the continents were connected as the single landmass Pangaea , and there was a worldwide dinosaur fauna mostly composed of coelophysoid carnivores and early sauropodomorph herbivores. Gymnosperm plants (particularly conifers ),
5576-637: The Ischigualasto and Santa Maria Formations of Argentina and Brazil, and the Pebbly Arkose Formation of Zimbabwe. The Ischigualasto Formation ( radiometrically dated at 231–230 million years old ) has produced the early saurischian Eoraptor , originally considered a member of the Herrerasauridae but now considered to be an early sauropodomorph, along with the herrerasaurids Herrerasaurus and Sanjuansaurus , and
Hand (disambiguation) - Misplaced Pages Continue
5712-562: The Late Jurassic epoch , and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs —birds—and the extinct non-avian dinosaurs , which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic , morphological and ecological standpoints. Birds, at over 11,000 living species , are among
5848-491: The Late Triassic epoch are often poorly known and were similar in many ways; these animals have sometimes been misidentified in the literature. Dinosaurs stand with their hind limbs erect in a manner similar to most modern mammals , but distinct from most other reptiles, whose limbs sprawl out to either side. This posture is due to the development of a laterally facing recess in the pelvis (usually an open socket) and
5984-657: The Morrison Formation of North America and Tendaguru Beds of Tanzania. Dinosaurs in China show some differences, with specialized metriacanthosaurid theropods and unusual, long-necked sauropods like Mamenchisaurus . Ankylosaurians and ornithopods were also becoming more common, but primitive sauropodomorphs had become extinct. Conifers and pteridophytes were the most common plants. Sauropods, like earlier sauropodomorphs, were not oral processors, but ornithischians were evolving various means of dealing with food in
6120-497: The Natural History Museum, London , to display the national collection of dinosaur fossils and other biological and geological exhibits. In 1858, William Parker Foulke discovered the first known American dinosaur, in marl pits in the small town of Haddonfield, New Jersey . (Although fossils had been found before, their nature had not been correctly discerned.) The creature was named Hadrosaurus foulkii . It
6256-621: The Pietraroja Plattenkalk of Italy. It preserves portions of the intestines, colon, liver, muscles, and windpipe. Concurrently, a line of work led by Mary Higby Schweitzer , Jack Horner , and colleagues reported various occurrences of preserved soft tissues and proteins within dinosaur bone fossils. Various mineralized structures that likely represented red blood cells and collagen fibres had been found by Schweitzer and others in tyrannosaurid bones as early as 1991. However, in 2005, Schweitzer and colleagues reported that
6392-708: The United States and displayed in Pittsburgh 's Carnegie Museum of Natural History in 1907. The longest dinosaur known from good fossil material is Patagotitan : the skeleton mount in the American Museum of Natural History in New York is 37 meters (121 ft) long. The Museo Municipal Carmen Funes in Plaza Huincul , Argentina, has an Argentinosaurus reconstructed skeleton mount that
6528-473: The carpal tunnel and contribute to the deep and superficial palmar arches . Several muscle tendons attaching to the TCL and the distal carpals also contribute to maintaining the carpal arch. Compared to the carpal arches, the arch formed by the distal ends of the metacarpal bones is flexible due to the mobility of the peripheral metacarpals (thumb and little finger). As these two metacarpals approach each other,
6664-600: The chimpanzee–human last common ancestor (CHLCA) and absent in modern humans are still present in the hands of Australopithecus , Paranthropus , and Homo floresiensis . This suggests that the derived changes in modern humans and Neanderthals did not evolve until 2.5 to 1.5 million years ago or after the appearance of the earliest Acheulian stone tools, and that these changes are associated with tool-related tasks beyond those observed in other hominins. The thumbs of Ardipithecus ramidus , an early hominin, are almost as robust as in humans, so this may be
6800-713: The clade Dinosauria . They first appeared during the Triassic period , between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is a subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya and their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs , having evolved from earlier theropods during
6936-419: The dermis of palmoplantar skin inhibit melanin production and thus the ability to tan , and promote the thickening of the stratum lucidum and stratum corneum layers of the epidermis . All parts of the skin involved in grasping are covered by papillary ridges ( fingerprints ) acting as friction pads. In contrast, the hairy skin on the dorsal side is thin, soft, and pliable, so that the skin can recoil when
SECTION 50
#17327810388557072-427: The largest and smallest dinosaurs to have ever existed. This is because only a tiny percentage of animals were ever fossilized and most of these remain buried in the earth. Few non-avian dinosaur specimens that are recovered are complete skeletons, and impressions of skin and other soft tissues are rare. Rebuilding a complete skeleton by comparing the size and morphology of bones to those of similar, better-known species
7208-523: The ulnar nerve may result in a condition in which some of the fingers cannot be flexed. A common fracture of the hand is a scaphoid fracture —a fracture of the scaphoid bone , one of the carpal bones. This is the commonest carpal bone fracture and can be slow to heal due to a limited blood flow to the bone. There are various types of fracture to the base of the thumb; these are known as Rolando fractures , Bennet's fracture , and Gamekeeper's thumb . Another common fracture, known as Boxer's fracture ,
7344-534: The "lizard-hipped" dinosaurs—birds evolved from earlier dinosaurs with "lizard hips". The following is a simplified classification of dinosaur groups based on their evolutionary relationships, and those of the main dinosaur groups Theropoda, Sauropodomorpha and Ornithischia, compiled by Justin Tweet. Further details and other hypotheses of classification may be found on individual articles. Timeline of major dinosaur groups per Holtz (2007) . Knowledge about dinosaurs
7480-677: The 1970s , however, has indicated that dinosaurs were active animals with elevated metabolisms and numerous adaptations for social interaction. Some were herbivorous , others carnivorous . Evidence suggests that all dinosaurs were egg-laying , and that nest -building was a trait shared by many dinosaurs, both avian and non-avian. While dinosaurs were ancestrally bipedal , many extinct groups included quadrupedal species, and some were able to shift between these stances. Elaborate display structures such as horns or crests are common to all dinosaur groups, and some extinct groups developed skeletal modifications such as bony armor and spines . While
7616-409: The 19th century. Samuel Beckles discovered a sauropod forelimb with preserved skin in 1852 that was incorrectly attributed to a crocodile; it was correctly attributed by Marsh in 1888 and subject to further study by Reginald Hooley in 1917. Among ornithischians, in 1884 Jacob Wortman found skin impressions on the first known specimen of Edmontosaurus annectens , which were largely destroyed during
7752-511: The English paleontologist Sir Richard Owen coined the term "dinosaur", using it to refer to the "distinct tribe or sub-order of Saurian Reptiles" that were then being recognized in England and around the world. The term is derived from Ancient Greek δεινός (deinos) 'terrible, potent or fearfully great' and σαῦρος (sauros) 'lizard or reptile'. Though
7888-581: The Greek ornitheios (ὀρνίθειος) meaning "of a bird" and ischion (ἰσχίον) meaning "hip joint"—had a pelvis that superficially resembled a bird's pelvis: the pubic bone was oriented caudally (rear-pointing). Unlike birds, the ornithischian pubis also usually had an additional forward-pointing process. Ornithischia includes a variety of species that were primarily herbivores. Despite the terms "bird hip" (Ornithischia) and "lizard hip" (Saurischia), birds are not part of Ornithischia. Birds instead belong to Saurischia,
8024-508: The Late Cretaceous, was the evolution of flowering plants . At the same time, several groups of dinosaurian herbivores evolved more sophisticated ways to orally process food. Ceratopsians developed a method of slicing with teeth stacked on each other in batteries, and iguanodontians refined a method of grinding with dental batteries , taken to its extreme in hadrosaurids. Some sauropods also evolved tooth batteries, best exemplified by
8160-785: The MRCA of Megalosaurus and Iguanodon , because these were two of the three genera cited by Richard Owen when he recognized the Dinosauria. Both definitions cover the same known genera: Dinosauria = Ornithischia + Saurischia . This includes major groups such as ankylosaurians (armored herbivorous quadrupeds), stegosaurians (plated herbivorous quadrupeds), ceratopsians (bipedal or quadrupedal herbivores with neck frills ), pachycephalosaurians (bipedal herbivores with thick skulls), ornithopods (bipedal or quadrupedal herbivores including " duck-bills "), theropods (mostly bipedal carnivores and birds), and sauropodomorphs (mostly large herbivorous quadrupeds with long necks and tails). Birds are
8296-401: The Middle to Late Triassic epochs, roughly 20 million years after the devastating Permian–Triassic extinction event wiped out an estimated 96% of all marine species and 70% of terrestrial vertebrate species approximately 252 million years ago. The oldest dinosaur fossils known from substantial remains date to the Carnian epoch of the Triassic period and have been found primarily in
SECTION 60
#17327810388558432-524: The Triassic, Early Jurassic, Late Jurassic and Cretaceous. Predatory theropod dinosaurs, which occupied most terrestrial carnivore niches during the Mesozoic, most often fall into the 100-to-1,000 kg (220-to-2,200 lb) category when sorted by estimated weight into categories based on order of magnitude , whereas recent predatory carnivoran mammals peak in the 10-to-100 kg (22-to-220 lb) category. The mode of Mesozoic dinosaur body masses
8568-595: The United States, known as dinosaur mania. Dinosaur mania was exemplified by the fierce rivalry between Edward Drinker Cope and Othniel Charles Marsh , both of whom raced to be the first to find new dinosaurs in what came to be known as the Bone Wars . This fight between the two scientists lasted for over 30 years, ending in 1897 when Cope died after spending his entire fortune on the dinosaur hunt. Many valuable dinosaur specimens were damaged or destroyed due to
8704-402: The back of the hand), the deep palmar arch , and the superficial palmar arch . Together these three arches and their anastomoses provide oxygenated blood to the palm, the fingers, and the thumb. The hand is drained by the dorsal venous network of the hand with deoxygenated blood leaving the hand via the cephalic vein and the basilic vein . The glabrous (hairless) skin on the front of
8840-673: The best-known genera are remarkable for their large size, many Mesozoic dinosaurs were human-sized or smaller, and modern birds are generally small in size. Dinosaurs today inhabit every continent, and fossils show that they had achieved global distribution by the Early Jurassic epoch at latest. Modern birds inhabit most available habitats, from terrestrial to marine, and there is evidence that some non-avian dinosaurs (such as Microraptor ) could fly or at least glide, and others, such as spinosaurids , had semiaquatic habits. While recent discoveries have made it more difficult to present
8976-455: The body . Other prehistoric animals, including pterosaurs , mosasaurs , ichthyosaurs , plesiosaurs , and Dimetrodon , while often popularly conceived of as dinosaurs, are not taxonomically classified as dinosaurs. Pterosaurs are distantly related to dinosaurs, being members of the clade Ornithodira . The other groups mentioned are, like dinosaurs and pterosaurs, members of Sauropsida (the reptile and bird clade), except Dimetrodon (which
9112-429: The bone as the lower extremity of the femur of a large animal, and recognized that it was too large to belong to any known species. He therefore concluded it to be the femur of a huge human, perhaps a Titan or another type of giant featured in legends. Edward Lhuyd , a friend of Sir Isaac Newton , published Lithophylacii Britannici ichnographia (1699), the first scientific treatment of what would now be recognized as
9248-418: The bones of the forearm, and a distal row ( trapezium , trapezoid , capitate and hamate ), which articulates with the bases of the five metacarpal bones of the hand. The heads of the metacarpals will each in turn articulate with the bases of the proximal phalanx of the fingers and thumb. These articulations with the fingers are the metacarpophalangeal joints known as the knuckles. At the palmar aspect of
9384-475: The common theropods, and ankylosaurids and early ceratopsians like Psittacosaurus became important herbivores. Meanwhile, Australia was home to a fauna of basal ankylosaurians, hypsilophodonts , and iguanodontians. The stegosaurians appear to have gone extinct at some point in the late Early Cretaceous or early Late Cretaceous . A major change in the Early Cretaceous, which would be amplified in
9520-437: The densest areas of nerve endings in the body, and are the richest source of tactile feedback. They also have the greatest positioning capability of the body; thus, the sense of touch is intimately associated with hands. Like other paired organs (eyes, feet, legs) each hand is dominantly controlled by the opposing brain hemisphere , so that handedness —the preferred hand choice for single-handed activities such as writing with
9656-731: The dinosaur subgroup Maniraptora , which are coelurosaurs , which are theropods, which are saurischians. Research by Matthew G. Baron, David B. Norman , and Paul M. Barrett in 2017 suggested a radical revision of dinosaurian systematics. Phylogenetic analysis by Baron et al. recovered the Ornithischia as being closer to the Theropoda than the Sauropodomorpha, as opposed to the traditional union of theropods with sauropodomorphs. This would cause sauropods and kin to fall outside traditional dinosaurs, so they re-defined Dinosauria as
9792-427: The dinosaurs' modern-day surviving avian lineage (birds) are generally small due to the constraints of flight, many prehistoric dinosaurs (non-avian and avian) were large-bodied—the largest sauropod dinosaurs are estimated to have reached lengths of 39.7 meters (130 feet) and heights of 18 m (59 ft) and were the largest land animals of all time. The misconception that non-avian dinosaurs were uniformly gigantic
9928-686: The dinosaurs. Most of these other animals became extinct in the Triassic, in one of two events. First, at about 215 million years ago, a variety of basal archosauromorphs, including the protorosaurs , became extinct. This was followed by the Triassic–Jurassic extinction event (about 201 million years ago), that saw the end of most of the other groups of early archosaurs, like aetosaurs, ornithosuchids, phytosaurs , and rauisuchians. Rhynchosaurs and dicynodonts survived (at least in some areas) at least as late as early –mid Norian and late Norian or earliest Rhaetian stages , respectively, and
10064-519: The disorders that can cause this is Catel-Manzke syndrome . The fingers may be fused in a disorder known as syndactyly . Or there may be an absence of one or more central fingers—a condition known as ectrodactyly . Additionally, some people are born without one or both hands ( amelia ). Hereditary multiple exostoses of the forearm—also known as hereditary multiple osteochondromas—is another cause of hand and forearm deformity in children and adults. There are several cutaneous conditions that can affect
10200-585: The early saurischian Alwalkeria , are known from the Upper Maleri and Lower Maleri Formations of India. The Carnian-aged Chañares Formation of Argentina preserves primitive, dinosaur-like ornithodirans such as Lagosuchus and Lagerpeton in Argentina , making it another important site for understanding dinosaur evolution. These ornithodirans support the model of early dinosaurs as small, bipedal predators. Dinosaurs may have appeared as early as
10336-462: The end of the Cretaceous, caused the extinction of all dinosaur groups except for the neornithine birds. Some other diapsid groups, including crocodilians , dyrosaurs , sebecosuchians , turtles, lizards , snakes , sphenodontians , and choristoderans , also survived the event. The surviving lineages of neornithine birds, including the ancestors of modern ratites , ducks and chickens , and
10472-456: The exact date of their extinction is uncertain. These losses left behind a land fauna of crocodylomorphs , dinosaurs, mammals, pterosaurians, and turtles . The first few lines of early dinosaurs diversified through the Carnian and Norian stages of the Triassic, possibly by occupying the niches of the groups that became extinct. Also notably, there was a heightened rate of extinction during
10608-447: The exact number varies between people: whereas a pair of sesamoid bones are found at virtually all thumb metacarpophalangeal joints, sesamoid bones are also common at the interphalangeal joint of the thumb (72.9%) and at the metacarpophalangeal joints of the little finger (82.5%) and the index finger (48%). In rare cases, sesamoid bones have been found in all the metacarpophalangeal joints and all distal interphalangeal joints except that of
10744-433: The field of dinosaur research experienced a surge in activity that remains ongoing. Several seminal studies led to this activity. First, John Ostrom discovered the bird-like dromaeosaurid theropod Deinonychus and described it in 1969. Its anatomy indicated that it was an active predator that was likely warm-blooded, in marked contrast to the then-prevailing image of dinosaurs. Concurrently, Robert T. Bakker published
10880-426: The fingers and toes". The ratio of the length of the index finger to the length of the ring finger in adults is affected by the level of exposure to male sex hormones of the embryo in utero . This digit ratio is below 1 for both sexes but it is lower in males than in females on average. A number of genetic disorders affect the hand. Polydactyly is the presence of more than the usual number of fingers. One of
11016-434: The fingers are stretched. On the dorsal side, the skin can be moved across the hand up to 3 cm (1.2 in); an important input the cutaneous mechanoreceptors . The web of the hand is a "fold of skin which connects the digits". These webs, located between each set of digits, are known as skin folds (interdigital folds or plica interdigitalis). They are defined as "one of the folds of skin, or rudimentary web, between
11152-417: The first metacarpophalangeal joints are small, almost spherical bones called the sesamoid bones. The fourteen phalanges make up the fingers and thumb, and are numbered I-V (thumb to little finger) when the hand is viewed from an anatomical position (palm up). The four fingers each consist of three phalanx bones: proximal, middle, and distal. The thumb only consists of a proximal and distal phalanx. Together with
11288-418: The groove on the dorsum of inferior side of ulna. The hand is innervated by the radial , median , and ulnar nerves . The radial nerve supplies the finger extensors and the thumb abductor , thus the muscles that extends at the wrist and metacarpophalangeal joints (knuckles); and that abducts and extends the thumb. The median nerve supplies the flexors of the wrist and digits, the abductors and opponens of
11424-456: The hand including the nails . The autoimmune disease rheumatoid arthritis can affect the hand, particularly the joints of the fingers. Some conditions can be treated by hand surgery . These include carpal tunnel syndrome , a painful condition of the hand and fingers caused by compression of the median nerve , and Dupuytren's contracture , a condition in which fingers bend towards the palm and cannot be straightened. Similarly, injury to
11560-457: The hand of modern humans have shown that they are consistent with the stresses and requirements associated with the effective use of paleolithic stone tools. It is possible that the refinement of the bipedal posture in the earliest hominids evolved to facilitate the use of the trunk as leverage in accelerating the hand. While the human hand has unique anatomical features, including a longer thumb and fingers that can be controlled individually to
11696-448: The hand, that of the thumb is the most mobile (and the least longitudinal). While the ray formed by the little finger and its associated metacarpal bone still offers some mobility, the remaining rays are firmly rigid. The phalangeal joints of the index finger, however, offer some independence to its finger, due to the arrangement of its flexor and extension tendons. The carpal bones form two transversal rows, each forming an arch concave on
11832-400: The hand, the palm, is relatively thick and can be bent along the hand's flexure lines where the skin is tightly bound to the underlying tissue and bones. Compared to the rest of the body's skin, the hands' palms (as well as the soles of the feet ) are usually lighter—and even much lighter in dark-skinned individuals, compared to the other side of the hand. Indeed, genes specifically expressed in
11968-486: The hip structure of their ancestors, with a pubis bone directed cranially , or forward. This basic form was modified by rotating the pubis backward to varying degrees in several groups ( Herrerasaurus , therizinosauroids, dromaeosaurids, and birds ). Saurischia includes the theropods (exclusively bipedal and with a wide variety of diets) and sauropodomorphs (long-necked herbivores which include advanced, quadrupedal groups). By contrast, ornithischians—"bird-hipped", from
12104-461: The human hand are plesiomorphic (shared by both ancestors and extant primate species); the elongated thumbs and short hands more closely resemble the hand proportions of Miocene apes than those of extant primates. Humans did not evolve from knuckle-walking apes, and chimpanzees and gorillas independently acquired elongated metacarpals as part of their adaptation to their modes of locomotion. Several primitive hand features most likely present in
12240-413: The human hand include: There are five digits attached to the hand, notably with a nail fixed to the end in place of the normal claw . The four fingers can be folded over the palm which allows the grasping of objects. Each finger, starting with the one closest to the thumb, has a colloquial name to distinguish it from the others: The thumb (connected to the first metacarpal bone and trapezium )
12376-413: The human hand, including pentadactyly (having five fingers), the hairless skin of the palm and fingers, and the os centrale found in human embryos, prosimians, and apes. Furthermore, the precursors of the intrinsic muscles of the hand are present in the earliest fishes, reflecting that the hand evolved from the pectoral fin and thus is much older than the arm in evolutionary terms. The proportions of
12512-450: The index and middle finger, it forms the dynamic tridactyl configuration responsible for most grips not requiring force. The ring and little fingers are more static, a reserve ready to interact with the palm when great force is needed. The muscles acting on the hand can be subdivided into two groups: the extrinsic and intrinsic muscle groups. The extrinsic muscle groups are the long flexors and extensors . They are called extrinsic because
12648-421: The index finger. For example, in some individuals, the ulnar nerve supplies the entire ring finger and the ulnar side of the middle finger, whilst, in others, the median nerve supplies the entire ring finger. The hand is supplied with blood from two arteries, the ulnar artery and the radial artery . These arteries form three arches over the dorsal and palmar aspects of the hand, the dorsal carpal arch (across
12784-480: The interlocking shapes of the carpal bones, and the wrist is therefore more stable in flexion than in extension. The distal carpal arch affects the function of the CMC joints and the hands, but not the function of the wrist or the proximal carpal arch. The ligaments that maintain the distal carpal arches are the transverse carpal ligament and the intercarpal ligaments (also oriented transversally). These ligaments also form
12920-432: The last common ancestor of Triceratops horridus , Passer domesticus and Diplodocus carnegii , and all of its descendants, to ensure that sauropods and kin remain included as dinosaurs. They also resurrected the clade Ornithoscelida to refer to the group containing Ornithischia and Theropoda. Using one of the above definitions, dinosaurs can be generally described as archosaurs with hind limbs held erect beneath
13056-572: The late 17th century in England. Part of a bone, now known to have been the femur of a Megalosaurus , was recovered from a limestone quarry at Cornwell near Chipping Norton , Oxfordshire, in 1676. The fragment was sent to Robert Plot , Professor of Chemistry at the University of Oxford and first curator of the Ashmolean Museum , who published a description in his The Natural History of Oxford-shire (1677). He correctly identified
13192-420: The long finger. The articulations are: The fixed and mobile parts of the hand adapt to various everyday tasks by forming bony arches: longitudinal arches (the rays formed by the finger bones and their associated metacarpal bones), transverse arches (formed by the carpal bones and distal ends of the metacarpal bones), and oblique arches (between the thumb and four fingers): Of the longitudinal arches or rays of
13328-478: The longest were the 33.5-meter (110 ft) long Diplodocus hallorum (formerly Seismosaurus ), the 33-to-34-meter (108 to 112 ft) long Supersaurus , and 37-meter (121 ft) long Patagotitan ; and the tallest, the 18-meter (59 ft) tall Sauroposeidon , which could have reached a sixth-floor window. There were a few dinosaurs that was considered either the heaviest and longest. The most famous one include Amphicoelias fragillimus , known only from
13464-479: The most complete theropod specimens, while North American localities have produced the most complete sauropodomorph specimens. Prior to the dinosaur renaissance, dinosaurs were mostly classified using the traditional rank-based system of Linnaean taxonomy . The renaissance was also accompanied by the increasingly widespread application of cladistics , a more objective method of classification based on ancestry and shared traits, which has proved tremendously useful in
13600-495: The most diverse groups of vertebrates. Using fossil evidence, paleontologists have identified over 900 distinct genera and more than 1,000 different species of non-avian dinosaurs. Dinosaurs are represented on every continent by both extant species (birds) and fossil remains. Through the first half of the 20th century, before birds were recognized as dinosaurs, most of the scientific community believed dinosaurs to have been sluggish and cold-blooded . Most research conducted since
13736-509: The most important advantage was dietary. Large animals are more efficient at digestion than small animals, because food spends more time in their digestive systems. This also permits them to subsist on food with lower nutritive value than smaller animals. Sauropod remains are mostly found in rock formations interpreted as dry or seasonally dry, and the ability to eat large quantities of low-nutrient browse would have been advantageous in such environments. Scientists will probably never be certain of
13872-478: The most recent common ancestor of a certain taxonomic group, are called the synapomorphies of such a group. A detailed assessment of archosaur interrelations by Sterling Nesbitt confirmed or found the following twelve unambiguous synapomorphies, some previously known: Nesbitt found a number of further potential synapomorphies and discounted a number of synapomorphies previously suggested. Some of these are also present in silesaurids , which Nesbitt recovered as
14008-456: The motoneurons of the hand muscles; placing the hands "closer" to the brain. The recent evolution of the human hand is thus a direct result of the development of the central nervous system , and the hand, therefore, is a direct tool of our consciousness—the main source of differentiated tactile sensations—and a precise working organ enabling gestures—the expressions of our personalities. There are nevertheless several primitive features left in
14144-466: The mouth, including potential cheek -like organs to keep food in the mouth, and jaw motions to grind food. Another notable evolutionary event of the Jurassic was the appearance of true birds, descended from maniraptoran coelurosaurians. By the Early Cretaceous and the ongoing breakup of Pangaea, dinosaurs were becoming strongly differentiated by landmass. The earliest part of this time saw
14280-455: The muscle belly is located on the forearm. The intrinsic muscle groups are the thenar (thumb) and hypothenar (little finger) muscles; the interosseous muscles ( four dorsally and three volarly ) originating between the metacarpal bones; and the lumbrical muscles arising from the deep flexor (and are special because they have no bony origin) to insert on the dorsal extensor hood mechanism. The fingers have two long flexors, located on
14416-768: The now-splitting supercontinent Gondwana , abelisaurids were the common theropods, and titanosaurian sauropods the common herbivores. Finally, in Europe, dromaeosaurids, rhabdodontid iguanodontians, nodosaurid ankylosaurians, and titanosaurian sauropods were prevalent. Flowering plants were greatly radiating, with the first grasses appearing by the end of the Cretaceous. Grinding hadrosaurids and shearing ceratopsians became very diverse across North America and Asia. Theropods were also radiating as herbivores or omnivores , with therizinosaurians and ornithomimosaurians becoming common. The Cretaceous–Paleogene extinction event, which occurred approximately 66 million years ago at
14552-756: The number of named genera began to increase exponentially in the 1990s. As of 2008, over 30 new species of dinosaurs were named each year. At least sauropodomorphs experienced a further increase in the number of named species in the 2010s, with an average of 9.3 new species having been named each year between 2009 and 2020. As a consequence, more sauropodomorphs were named between 1990 and 2020 than in all previous years combined. These new localities also led to improvements in overall specimen quality, with new species being increasingly named not on scrappy fossils but on more complete skeletons, sometimes from multiple individuals. Better specimens also led to new species being invalidated less frequently. Asian localities have produced
14688-438: The pair's rough methods: for example, their diggers often used dynamite to unearth bones. Modern paleontologists would find such methods crude and unacceptable, since blasting easily destroys fossil and stratigraphic evidence. Despite their unrefined methods, the contributions of Cope and Marsh to paleontology were vast: Marsh unearthed 86 new species of dinosaur and Cope discovered 56, a total of 142 new species. Cope's collection
14824-403: The palmar gutter deepens. The central-most metacarpal (middle finger) is the most rigid. It and its two neighbors are tied to the carpus by the interlocking shapes of the metacarpal bones. The thumb metacarpal only articulates with the trapezium and is therefore completely independent, while the fifth metacarpal (little finger) is semi-independent with the fourth metacarpal (ring finger) which forms
14960-407: The palmar side of the thumb, index, middle, and half ring fingers. Dorsal branches innervates the distal phalanges of the index, middle, and half ring fingers. The ulnar nerve supplies the ulnar third of the hand, both at the palm and the back of the hand, and the little and half ring fingers. There is a considerable variation to this general pattern, except for the little finger and volar surface of
15096-410: The palmar side. Because the proximal arch simultaneously has to adapt to the articular surface of the radius and to the distal carpal row, it is by necessity flexible. In contrast, the capitate, the "keystone" of the distal arch, moves together with the metacarpal bones and the distal arch is therefore rigid. The stability of these arches is more dependent of the ligaments and capsules of the wrist than of
15232-415: The phalanges of the fingers and thumb these metacarpal bones form five rays or poly-articulated chains. Because supination and pronation (rotation about the axis of the forearm) are added to the two axes of movements of the wrist, the ulna and radius are sometimes considered part of the skeleton of the hand. There are numerous sesamoid bones in the hand, small ossified nodes embedded in tendons;
15368-408: The presence of proteins such as collagen, elastin , and laminin . Both specimens yielded collagen protein sequences that were viable for molecular phylogenetic analyses , which grouped them with birds as would be expected. The extraction of fragmentary DNA has also been reported for both of these fossils, along with a specimen of Hypacrosaurus . In 2015, Sergio Bertazzo and colleagues reported
15504-458: The preservation of collagen fibres and red blood cells in eight Cretaceous dinosaur specimens that did not show any signs of exceptional preservation, indicating that soft tissue may be preserved more commonly than previously thought. Suggestions that these structures represent bacterial biofilms have been rejected, but cross-contamination remains a possibility that is difficult to detect. Dinosaurs diverged from their archosaur ancestors during
15640-422: The rebbachisaurid Nigersaurus . There were three general dinosaur faunas in the Late Cretaceous. In the northern continents of North America and Asia, the major theropods were tyrannosaurids and various types of smaller maniraptoran theropods, with a predominantly ornithischian herbivore assemblage of hadrosaurids, ceratopsians, ankylosaurids, and pachycephalosaurians. In the southern continents that had made up
15776-1413: The rocks that produced the Jehol (Early Cretaceous) and Yanliao (Mid-to-Late Jurassic) biotas of northeastern China, from which hundreds of dinosaur specimens bearing impressions of feather-like structures (both closely related to birds and otherwise, see § Origin of birds ) have been described by Xing Xu and colleagues. In living reptiles and mammals, pigment-storing cellular structures known as melanosomes are partially responsible for producing colouration. Both chemical traces of melanin and characteristically shaped melanosomes have been reported from feathers and scales of Jehol and Yanliao dinosaurs, including both theropods and ornithischians. This has enabled multiple full-body reconstructions of dinosaur colouration , such as for Sinosauropteryx and Psittacosaurus by Jakob Vinther and colleagues, and similar techniques have also been extended to dinosaur fossils from other localities. (However, some researchers have also suggested that fossilized melanosomes represent bacterial remains. ) Stomach contents in some Jehol and Yanliao dinosaurs closely related to birds have also provided indirect indications of diet and digestive system anatomy (e.g., crops ). More concrete evidence of internal anatomy has been reported in Scipionyx from
15912-450: The sauropodomorphs Bagualosaurus , Buriolestes , Guaibasaurus , Macrocollum , Nhandumirim , Pampadromaeus , Saturnalia , and Unaysaurus . The Pebbly Arkose Formation, which is of uncertain age but was likely comparable to the other two, has produced the sauropodomorph Mbiresaurus , along with an unnamed herrerasaurid. Less well-preserved remains of the sauropodomorphs Jaklapallisaurus and Nambalia , along with
16048-416: The sauropodomorphs Chromogisaurus , Eodromaeus , and Panphagia . Eoraptor 's likely resemblance to the common ancestor of all dinosaurs suggests that the first dinosaurs would have been small, bipedal predators . The Santa Maria Formation (radiometrically dated to be older, at 233.23 million years old ) has produced the herrerasaurids Gnathovorax and Staurikosaurus , along with
16184-584: The skeleton now mounted and on display at the Museum für Naturkunde in Berlin ; this mount is 12 meters (39 ft) tall and 21.8 to 22.5 meters (72 to 74 ft) long, and would have belonged to an animal that weighed between 30 000 and 60 000 kilograms ( 70 000 and 130 000 lb). The longest complete dinosaur is the 27 meters (89 ft) long Diplodocus , which was discovered in Wyoming in
16320-406: The skull behind the eyes), and as members of the diapsid group Archosauria, had additional openings in the snout and lower jaw. Additionally, several characteristics once thought to be synapomorphies are now known to have appeared before dinosaurs, or were absent in the earliest dinosaurs and independently evolved by different dinosaur groups. These include an elongated scapula , or shoulder blade;
16456-434: The sole surviving dinosaurs. In traditional taxonomy , birds were considered a separate class that had evolved from dinosaurs, a distinct superorder . However, most contemporary paleontologists reject the traditional style of classification based on anatomical similarity, in favor of phylogenetic taxonomy based on deduced ancestry, in which each group is defined as all descendants of a given founding genus. Birds belong to
16592-515: The specimen's excavation. Owen and Hooley subsequently described skin impressions of Hypsilophodon and Iguanodon in 1885 and 1917. Since then, scale impressions have been most frequently found among hadrosaurids, where the impressions are known from nearly the entire body across multiple specimens. Starting from the 1990s, major discoveries of exceptionally preserved fossils in deposits known as conservation Lagerstätten contributed to research on dinosaur soft tissues. Chiefly among these were
16728-538: The spread of ankylosaurians, iguanodontians , and brachiosaurids through Europe, North America, and northern Africa . These were later supplemented or replaced in Africa by large spinosaurid and carcharodontosaurid theropods, and rebbachisaurid and titanosaurian sauropods, also found in South America . In Asia , maniraptoran coelurosaurians like dromaeosaurids, troodontids , and oviraptorosaurians became
16864-468: The study of dinosaur systematics and evolution. Cladistic analysis, among other techniques, helps to compensate for an often incomplete and fragmentary fossil record. Reference books summarizing the state of dinosaur research, such as David B. Weishampel and colleagues' The Dinosauria , made knowledge more accessible and spurred further interest in dinosaur research. The release of the first and second editions of The Dinosauria in 1990 and 2004, and of
17000-459: The taxonomic name has often been interpreted as a reference to dinosaurs' teeth, claws, and other fearsome characteristics, Owen intended it also to evoke their size and majesty. Owen recognized that the remains that had been found so far, Iguanodon , Megalosaurus and Hylaeosaurus , shared distinctive features, and so decided to present them as a distinct taxonomic group. As clarified by British geologist and historian Hugh Torrens, Owen had given
17136-403: The tendons of these form the anatomical snuff box . Also, the index finger and the little finger have an extra extensor used, for instance, for pointing. The extensors are situated within 6 separate compartments. The first four compartments are located in the grooves present on the dorsum of inferior side of radius while the 5th compartment is in between radius and ulna. The 6th compartment is in
17272-410: The term hand in this sense to distinguish the terminations of the front paws from the hind ones is an example of anthropomorphism . The only true grasping hands appear in the mammalian order of primates . Hands must also have opposable thumbs , as described later in the text. The hand is located at the distal end of each arm. Apes and monkeys are sometimes described as having four hands, because
17408-405: The thumb in opposition, making grasping possible. The extensors are located on the back of the forearm and are connected in a more complex way than the flexors to the dorsum of the fingers. The tendons unite with the interosseous and lumbrical muscles to form the extensorhood mechanism. The primary function of the extensors is to straighten out the digits. The thumb has two extensors in the forearm;
17544-415: The thumb is included as one of the fingers . It has 27 bones, not including the sesamoid bone , the number of which varies among people, 14 of which are the phalanges ( proximal , intermediate and distal ) of the fingers and thumb. The metacarpal bones connect the fingers and the carpal bones of the wrist . Each human hand has five metacarpals and eight carpal bones. Fingers contain some of
17680-461: The thumb, the first and second lumbrical. The ulnar nerve supplies the remaining intrinsic muscles of the hand. All muscles of the hand are innervated by the brachial plexus (C5–T1) and can be classified by innervation: The radial nerve supplies the skin on the back of the hand from the thumb to the ring finger and the dorsal aspects of the index, middle, and half ring fingers as far as the proximal interphalangeal joints. The median nerve supplies
17816-410: The toes are long and the hallux is opposable and looks more like a thumb , thus enabling the feet to be used as hands. The word "hand" is sometimes used by evolutionary anatomists to refer to the appendage of digits on the forelimb such as when researching the homology between the three digits of the bird hand and the dinosaur hand. An adult human male's hand weighs about a pound. Areas of
17952-849: The total number of dinosaur genera preserved in the fossil record at 1850, nearly 75% still undiscovered, and the number that ever existed (in or out of the fossil record) at 3,400. A 2016 estimate put the number of dinosaur species living in the Mesozoic at 1,543–2,468, compared to the number of modern-day birds (avian dinosaurs) at 10,806 species. Extinct dinosaurs, as well as modern birds, include genera that are herbivorous and others carnivorous, including seed-eaters, fish-eaters, insectivores, and omnivores. While dinosaurs were ancestrally bipedal (as are all modern birds), some evolved into quadrupeds, and others, such as Anchisaurus and Iguanodon , could walk as easily on two or four legs. Cranial modifications like horns and crests are common dinosaurian traits, and some extinct species had bony armor. Although
18088-435: The underside of the forearm. They insert by tendons to the phalanges of the fingers. The deep flexor attaches to the distal phalanx, and the superficial flexor attaches to the middle phalanx. The flexors allow for the actual bending of the fingers. The thumb has one long flexor and a short flexor in the thenar muscle group. The human thumb also has other muscles in the thenar group ( opponens and abductor brevis muscle ), moving
18224-457: Was an extremely important find: Hadrosaurus was one of the first nearly complete dinosaur skeletons found ( the first was in 1834, in Maidstone, England ), and it was clearly a bipedal creature. This was a revolutionary discovery as, until that point, most scientists had believed dinosaurs walked on four feet, like other lizards. Foulke's discoveries sparked a wave of interests in dinosaurs in
18360-489: Was an order of magnitude more massive than anything else that has since walked the Earth. Giant prehistoric mammals such as Paraceratherium (the largest land mammal ever) were dwarfed by the giant sauropods, and only modern whales approach or surpass them in size. There are several proposed advantages for the large size of sauropods, including protection from predation, reduction of energy use, and longevity, but it may be that
18496-450: Was purportedly discovered in 1822 by Mary Ann Mantell , the wife of English geologist Gideon Mantell , though this is disputed and some historians say Gideon had acquired remains years earlier. Gideon Mantell recognized similarities between his fossils and the bones of modern iguanas and published his findings in 1825. The study of these "great fossil lizards" soon became of great interest to European and American scientists, and in 1842
#854145