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98-505: Tulips are spring-blooming perennial herbaceous bulbiferous geophytes in the Tulipa genus . Their flowers are usually large, showy, and brightly coloured, generally red, orange, pink, yellow, or white. They often have a different coloured blotch at the base of the tepals , internally. Because of a degree of variability within the populations and a long history of cultivation, classification has been complex and controversial. The tulip

196-443: A century plant can live for 80 years and grow 30 meters tall before flowering and dying. However, most perennials are polycarpic (or iteroparous ), flowering over many seasons in their lifetime. Perennials invest more resources than annuals into roots, crowns, and other structures that allow them to live from one year to the next. They often have a competitive advantage because they can commence their growth and leaf out earlier in

294-940: A photolyase and the other, a protein involved in nucleotide excision repair ). Perennials that are cultivated include: woody plants like fruit trees grown for their edible fruits; shrubs and trees grown as landscaping ornamentals ; herbaceous food crops like asparagus , rhubarb , strawberries ; and subtropical plants not hardy in colder areas such as tomatoes , eggplant , and coleus (which are treated as annuals in colder areas). Perennials also include plants grown for their flowering and other ornamental value including bulbs (like tulips, narcissus, and gladiolus); lawn grass, and other groundcovers , (such as periwinkle and Dichondra ). Each type of plant must be separated differently; for example, plants with fibrous root systems like daylilies, Siberian iris, or grasses can be pried apart with two garden forks inserted back to back, or cut by knives. However, plants such as bearded irises have

392-456: A band stretching from Southern Europe to Central Asia, but since the seventeenth century have become widely naturalised and cultivated ( see map ). In their natural state, they are adapted to steppes and mountainous areas with temperate climates . Flowering in the spring, they become dormant in the summer once the flowers and leaves die back, emerging above ground as a shoot from the underground bulb in early spring. Growing wild over much of

490-464: A common element of steppe and winter-rain Mediterranean vegetation . They thrive in climates with long, cool springs and dry summers. Tulips are most commonly found in meadows , steppes and chaparral , but also introduced in fields, orchards, roadsides and abandoned gardens. Botrytis tulipae is a major fungal disease affecting tulips, causing cell death and eventually the rotting of

588-559: A narcissus. Carolus Clusius is largely responsible for the spread of tulip bulbs in the final years of the 16th century; he planted tulips at the Vienna Imperial Botanical Gardens in 1573. He finished the first major work on tulips in 1592 and made note of the colour variations. After he was appointed the director of the Leiden University 's newly established Hortus Botanicus , he planted both

686-413: A penis. A female spotted hyena's apparent penis is in fact an enlarged clitoris, which contains an external birth canal. It can be difficult to determine the sex of spotted hyenas until sexual maturity , when they may become pregnant. When a female spotted hyena gives birth, she passes the cub through the cervix internally, but then passes it out through the elongated clitoris. The term hermaphrodite

784-524: A private gardener by clipping the stem and flower of an individual specimen. Commercial growers usually harvest the tulip bulbs in late summer and grade them into sizes; bulbs large enough to flower are sorted and sold, while smaller bulbs are sorted into sizes and replanted for sale in the future. Because tulip bulbs don't reliably come back every year, tulip varieties that fall out of favour with present aesthetic values have traditionally gone extinct. Unlike other flowers that do not suffer this same limitation,

882-470: A root system of rhizomes; these root systems should be planted with the top of the rhizome just above ground level, with leaves from the following year showing. The point of dividing perennials is to increase the amount of a single breed of plant in your garden. In the United States more than 900 million dollars worth of potted herbaceous perennial plants were sold in 2019. Although most of humanity

980-472: A sequential hermaphrodite is also stipulated to only change sex once. ) This contrasts with simultaneous hermaphrodites, in which an individual possesses fully functional male and female genitalia. Sequential hermaphroditism is common in fish (particularly teleost fish) and many gastropods (such as the common slipper shell ). Sequential hermaphroditism can best be understood in terms of behavioral ecology and evolutionary life history theory , as described in

1078-835: A single terminal flower, or when pluriflor as two to three (e.g. Tulipa turkestanica ), but up to four, flowers on the end of a floriferous stem ( scape ), which is single arising from amongst the basal leaf rosette. In structure, the flower is generally cup or star-shaped. As with other members of Liliaceae the perianth is undifferentiated ( perigonium ) and biseriate (two whorled ), formed from six free (i.e. apotepalous ) caducous tepals arranged into two separate whorls of three parts ( trimerous ) each. The two whorls represent three petals and three sepals but are termed tepals because they are nearly identical. The tepals are usually petaloid (petal-like), being brightly coloured, but each whorl may be different, or have different coloured blotches at their bases, forming darker colouration on

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1176-769: A teaching garden and his private garden with tulips in late 1593. Thus, 1594 is considered the date of the tulip's first flowering in the Netherlands, despite reports of the cultivation of tulips in private gardens in Antwerp and Amsterdam two or three decades earlier. These tulips at Leiden would eventually lead to both the tulip mania and the tulip industry in the Netherlands. Over two raids, in 1596 and in 1598, more than one hundred bulbs were stolen from his garden. Tulips spread rapidly across Europe, and more opulent varieties such as double tulips were already known in Europe by

1274-479: A very few (e.g. Zostera ) occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including trees and shrubs. Perennial plants are usually better long-term competitors, especially under stable, resource-poor conditions. This

1372-609: A wide variety of colours, black tulips have historically been difficult to achieve. The Queen of the Night tulip is as close to black as a flower gets, though it is, in fact, a dark and glossy maroonish purple. The first truly black tulip was bred in 1986 by a Dutch flower grower in Bovenkarspel, Netherlands. The specimen was created by cross-breeding two deep purple tulips, the Queen of the Night and Wienerwald tulips. The tulip's fruit

1470-417: A year or more of growth before plants are large enough to flower. Tulips grown from seeds often need five to eight years before plants are of flowering size. To prevent cross-pollination, increase the growth rate of bulbs and increase the vigour and size of offsets, the flower and stems of a field of commercial tulips are usually topped using large tractor-mounted mowing heads. The same goals can be achieved by

1568-414: Is also found in some fish species , but is rare in other vertebrate groups. Most hermaphroditic species exhibit some degree of self-fertilization. The distribution of self-fertilization rates among animals is similar to that of plants, suggesting that similar pressures are operating to direct the evolution of selfing in animals and plants. A rough estimate of the number of hermaphroditic animal species

1666-644: Is cauline (born on a stem), strap-shaped, with a waxy coating, and the leaves are alternate (alternately arranged on the stem), diminishing in size the further up the stem. These fleshy blades are often bluish-green in colour. The bulbs are truncated basally and elongated towards the apex. They are covered by a protective tunic (tunicate) which can be glabrous or hairy inside. The tulip's flowers are usually large and are actinomorphic (radially symmetric) and hermaphrodite (contain both male ( androecium ) and female ( gynoecium ) characteristics), generally erect, or more rarely pendulous , and are arranged more usually as

1764-444: Is 65,000, about 5% of all animal species, or 33% excluding insects. Insects are almost exclusively gonochoric, and no definitive cases of hermaphroditism have been demonstrated in this group. There are no known hermaphroditic species among mammals or birds . About 94% of flowering plant species are either hermaphroditic (all flowers produce both male and female gametes) or monoecious , where both male and female flowers occur on

1862-526: Is a genus of the lily family, Liliaceae , once one of the largest families of monocots , but which molecular phylogenetics has reduced to a monophyletic grouping with only 15 genera. Within Liliaceae, Tulipa is placed within Lilioideae , one of three subfamilies, with two tribes . Tribe Lilieae includes seven other genera in addition to Tulipa . The genus, which includes about 75 species ,

1960-552: Is a globose or ellipsoid capsule with a leathery covering and an ellipsoid to globe shape. Each capsule contains numerous flat, disc-shaped seeds in two rows per chamber. These light to dark brown seeds have very thin seed coats and endosperm that do not normally fill the entire seed. Tulipanin is an anthocyanin found in tulips. It is the 3-rutinoside of delphinidin . The chemical compounds named tuliposides and tulipalins can also be found in tulips and are responsible for allergies. Tulipalin A, or α-methylene-γ-butyrolactone,

2058-456: Is a common allergen , generated by hydrolysis of the glucoside tuliposide A. It induces a dermatitis that is mostly occupational and affects tulip bulb sorters and florists who cut the stems and leaves. Tulipanin A and B are toxic to horses, cats and dogs. The colour of a tulip is formed from two pigments working in concert; a base colour that is always yellow or white, and a second laid-on anthocyanin colour. The mix of these two hues determines

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2156-436: Is a member of the lily family , Liliaceae , along with 14 other genera, where it is most closely related to Amana , Erythronium , and Gagea in the tribe Lilieae . There are about 75 species , and these are divided among four subgenera . The name "tulip" is thought to be derived from a Persian word for turban , which it may have been thought to resemble by those who discovered it. Tulips were originally found in

2254-810: Is both male and female. And an hermaphrodite (which is also called Androgynus ) shall be heire, either as male or female, according to that kind of sexe which doth prevaile." During the Victorian era , medical authors attempted to ascertain whether or not humans could be hermaphrodites, adopting a precise biological definition to the term. From that period until the early 21st century, individuals with ovotesticular syndrome were termed true hermaphrodites if their gonadal tissue contained both testicular and ovarian tissue, and pseudohermaphrodites if their external appearance ( phenotype ) differed from sex expected from internal gonads. This language has fallen out of favor due to misconceptions and stigma associated with

2352-434: Is common in bryophytes and some vascular plants. Historically, the term hermaphrodite was used in law to refer to people whose sex was in doubt. The 12th-century Decretum Gratiani states that "Whether an hermaphrodite may witness a testament, depends on which sex prevails" ("Hermafroditus an ad testamentum adhiberi possit, qualitas sexus incalescentis ostendit."). Alexander ab Alexandro (1461–1523) stated, using

2450-517: Is divided into four subgenera . The word tulip , first mentioned in western Europe in or around 1554 and seemingly derived from the "Turkish Letters" of diplomat Ogier Ghiselin de Busbecq , first appeared in English as tulipa or tulipant , entering the language by way of French: tulipe and its obsolete form tulipan or by way of Modern Latin tulipa , from Ottoman Turkish tülbend (" muslin " or " gauze "), and may be ultimately derived from

2548-783: Is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring. Annual plants have an advantage in disturbed environments because of their faster growth and reproduction rates. Each section contains a short list of species related to that topic, these are an example as the true lists would fill several books. Perennials grown for their decorative flowers include very many species and types. Some examples include: The majority of fruit bearing plants are perennial even in temperate climates. Examples include: Many herbs are perennial, including these examples: Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Examples of some of

2646-483: Is fed by the re-sowing of the seeds of annual grain crops, (either naturally or by the manual efforts of humans), perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion , capture dissolved nitrogen before it can contaminate ground and surface water, and out-compete weeds (reducing the need for herbicides ). These potential benefits of perennials have resulted in new attempts to increase

2744-899: Is not mentioned by any writer from antiquity, therefore it seems probable that tulips were introduced into Anatolia only with the advance of the Seljuks . In the Ottoman Empire , numerous types of tulips were cultivated and bred, and today, 14 species can still be found in Turkey. Tulips are mentioned by Omar Kayam and Jalāl ad-Dīn Rûmi . Species of tulips in Turkey typically come in red, less commonly in white or yellow. The Ottoman Turks had discovered that these wild tulips were great changelings, freely hybridizing (though it takes 7 years to show colour) but also subject to mutations that produced spontaneous changes in form and colour. A paper by Arthur Baker reports that in 1574, Sultan Selim II ordered

2842-422: Is not to be confused with ovotesticular syndrome in mammals, which is a separate and unrelated phenomenon. While people with the condition were previously called " true hermaphrodites" in medical literature, this usage is now considered to be outdated as of 2006 and misleading, as people with ovotesticular syndrome do not have functional sets of both male and female organs. The term hermaphrodite derives from

2940-409: Is the opposite of hermaphroditic. The individuals of many taxonomic groups of animals , primarily invertebrates, are hermaphrodites, capable of producing viable gametes of both sexes . In the great majority of tunicates , mollusks , and earthworms , hermaphroditism is a normal condition, enabling a form of sexual reproduction in which either partner can act as the female or male. Hermaphroditism

3038-429: Is too cold or dry. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season. Some perennial plants are protected from wildfires because they have underground roots that produce adventitious shoots, bulbs, crowns , or stems ; other perennials like trees and shrubs may have thick cork layers that protect

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3136-699: Is unknown who first brought the tulip to Northwestern Europe, the most widely accepted story is that it was Oghier Ghislain de Busbecq , an ambassador for Emperor Ferdinand I to Suleyman the Magnificent . According to a letter, he saw "an abundance of flowers everywhere; Narcissus , hyacinths and those in Turkish called Lale, much to our astonishment because it was almost midwinter, a season unfriendly to flowers." However, in 1559, an account by Conrad Gessner describes tulips flowering in Augsburg , Swabia in

3234-544: Is used in botany to describe, for example, a perfect flower that has both staminate (male, pollen-producing) and carpellate (female, ovule-producing) parts. The overwhelming majority of flowering plant species are hermaphroditic. Flowering plant species with separate, imperfect, male and female flowers on the same individual are called monoecious . Monoecy only occurs in about 7% of flowering plant species. Monoecious plants are often referred to as hermaphroditic because they produce both male and female gametes. However,

3332-514: The Latin : hermaphroditus , from Ancient Greek : ἑρμαφρόδιτος , romanized :  hermaphroditos , which derives from Hermaphroditus (Ἑρμαφρόδιτος), the son of Hermes and Aphrodite in Greek mythology . According to Ovid , he fused with the nymph Salmacis resulting in one individual possessing physical traits of male and female sexes. According to the earlier Diodorus Siculus , he

3430-590: The Near East and Central Asia, tulips had probably been cultivated in Persia from the 10th century. By the 15th century, tulips were among the most prized flowers; becoming the symbol of the later Ottomans. Tulips were cultivated in Byzantine Constantinople as early as 1055 but they did not come to the attention of Northern Europeans until the sixteenth century, when Northern European diplomats to

3528-615: The Ottoman court observed and reported on them. They were rapidly introduced into Northern Europe and became a much-sought-after commodity during tulip mania . Tulips were frequently depicted in Dutch Golden Age paintings , and have become associated with the Netherlands , the major producer for world markets, ever since. In the seventeenth-century Netherlands, during the time of the tulip mania, an infection of tulip bulbs by

3626-526: The Persian : دلبند delband (" Turban "), this name being applied because of a perceived resemblance of the shape of a tulip flower to that of a turban. This may have been due to a translation error in early times when it was fashionable in the Ottoman Empire to wear tulips on turbans. The translator possibly confused the flower for the turban. Ogier Ghiselin de Busbecq stated that the "Turks" used

3724-733: The Topkapı Sarayı in Istanbul . It is also reported that shortly after arriving in Constantinople in 1554, Ogier Ghislain de Busbecq, ambassador of the Austrian Habsburgs to the court of Suleyman the Magnificent, claimed to have introduced the tulip to Europe by sending a consignment of bulbs west. The fact that the tulip's first official trip west took it from one court to the other could have contributed to its ascendency. Sultan Ahmet III maintained famous tulip gardens in

3822-475: The astronomical symbol for the planet Jupiter . Perennial plants can be short-lived (only a few years) or long-lived. They include a wide assortment of plant groups from non-flowering plants like ferns and liverworts to highly diverse flowering plants like orchids , grasses , and woody plants . Plants that flower and fruit only once and then die are termed monocarpic or semelparous ; these species may live for many years before they flower. For example,

3920-550: The ceramic tulipiere was devised for the display of cut flowers stem by stem. Vases and bouquets, usually including tulips, often appeared in Dutch still-life painting . To this day, tulips are associated with the Netherlands, and the cultivated forms of the tulip are often called "Dutch tulips". The Netherlands has the world's largest permanent display of tulips at the Keukenhof . The majority of tulip cultivars are classified in

4018-419: The ovaries are superior, with three chambers. The "Semper Augustus" was the most expensive tulip during the 17th-century tulip mania . After seeing the tulip in the garden of Dr. Adriaen Pauw, a director of the new East India Company , Nicolas van Wassenaer wrote in 1624 that "The colour is white, with Carmine on a blue base, and with an unbroken flame right to the top". With limited specimens in existence at

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4116-581: The sexual differentiation stage; however, this is now known to be incorrect, as humans are simply undifferentiated before this stage and possess a paramesonephric duct , a mesonephric duct , and a genital tubercle . The evolution of anisogamy may have contributed to the evolution of simultaneous hermaphroditism and sequential hermaphroditism, it remains unclear if the evolution of anisogamy first led to hermaphroditism or gonochorism . A 2023 study argued that hermaphroditism can evolve directly from mating types under certain circumstances, such as if

4214-482: The tulip breaking virus created variegated patterns in the tulip flowers that were much admired and valued. While truly broken tulips are not cultivated anymore, the closest available specimens today are part of the group known as the Rembrandts – so named because Rembrandt painted some of the most admired breaks of his time. Breeding programmes have produced thousands of hybrid and cultivars in addition to

4312-539: The 'Didieri Group' with unicolorous tepals. Tulip bulbs are typically planted around late summer and fall, in well-drained soils. Tulips should be planted 10 to 15 cm (4 to 6 inches) apart from each other. The recommended hole depth is 10 to 20 cm (4 to 8 inches) deep and is measured from the top of the bulb to the surface. Therefore, larger tulip bulbs would require deeper holes. Species of tulips are normally planted deeper. Perennial Perennials (especially small flowering plants ) that grow and bloom over

4410-531: The American East, white-tailed deer eat tulips, with no apparent ill effects. However, tulips are poisonous to domestic animals e.g. horses, cats, and dogs. Cultivation of the tulip began in Iran ( Persia ), probably in the 10th century. Early cultivars must have emerged from hybridisation in gardens from wild collected plants, which were then favoured, possibly due to flower size or growth vigour. The tulip

4508-648: The Balkans are native, subsequent identification of Tulipa sylvestris subsp. australis as a native of the Iberian peninsula and adjacent North Africa shows that this may be a simplification. In addition to these regions in the west tulips have been identified in Greece, Cyprus and the Balkans . In the south, Iran marks its furthest extent, while the northern limit is Ukraine. Although tulips are also throughout most of

4606-540: The Dutch tulipomania gained their delicately feathered patterns from an infection with the tulip breaking virus , a mosaic virus that was carried by the green peach aphid , Myzus persicae . While the virus produces fantastically streaked flowers, it also weakens plants and reduces the number of offsets produced. Dutch growers would go to extraordinary lengths during tulipomania to make tulips break, borrowing alchemists’ techniques and resorting to sprinkling paint powders of

4704-469: The Dutch. Between 1634 and 1637, the enthusiasm for the new flowers in Holland triggered a speculative frenzy now known as the tulip mania that eventually led to the collapse of the market three years later. Tulip bulbs had become so expensive that they were treated as a form of currency, or rather, as futures, forcing the Dutch government to introduce trading restrictions on the bulbs. Around this time,

4802-621: The Kadi of A‘azāz in Syria to send him 50,000 tulip bulbs. However, John Harvey points out several problems with this source, and there is also the possibility that tulips and hyacinth ( sümbüll ), originally Indian spikenard ( Nardostachys jatamansi ) have been confused. Sultan Selim also imported 300,000 bulbs of Kefe Lale (also known as Cafe-Lale, from the medieval name Kaffa, probably Tulipa schrenkii ) from Kefe in Crimea , for his gardens in

4900-529: The Mediterranean and Europe, these regions do not form part of the natural distribution. Tulips were brought to Europe by travellers and merchants from Anatolia and Central Asia for cultivation, from where they escaped and naturalised ( see map ). For instance, less than half of those species found in Turkey are actually native. These have been referred to as neo-tulipae. Tulips are indigenous to mountainous areas with temperate climates , where they are

4998-460: The Netherlands. The gardening book Revnak'ı Bostan (Beauty of the Garden) by Sahibül Reis ülhaç Ibrahim Ibn ülhaç Mehmet, written in 1660 does not mention the tulip at all, but contains advice on growing hyacinths and lilies . However, there is considerable confusion of terminology, and tulips may have been subsumed under hyacinth, a mistake several European botanists were to perpetuate. In 1515,

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5096-521: The Tulip's historical forms do not survive alongside their modern incarnations. In horticulture, tulips are divided into fifteen groups (Divisions) mostly based on flower morphology and plant size. They may also be classified by their flowering season: A number of names are based on naturalised garden tulips and are usually referred to as neo-tulipae. These are often difficult to trace back to their original cultivar, and in some cases have been occurring in

5194-574: The adults take several years to change from female to male, the broodstock are extremely valuable individuals. Simultaneous hermaphrodites (or homogamous hermaphrodites) are individuals in which both male and female sexual organs are present and functional at the same time. Self-fertilization often occurs. When spotted hyenas were first scientifically observed by explorers, they were thought to be hermaphrodites. Early observations of wild spotted hyenas led researchers to believe that all spotted hyenas, male or female, were born with what looked to be

5292-446: The advantage of generating larger seedlings that can better compete with other plants. Perennials also produce seeds over many years. An important aspect of cold acclimation is overexpression of DNA repair genes. In Thinopyrum intermedium a perennial relative of common wheat Triticum aestivum , conditions of freezing stress were shown to be associated with large increases in expression of two DNA repair genes (one gene product

5390-607: The colours. Tulip growth is also dependent on temperature conditions. Slightly germinated plants show greater growth if subjected to a period of cool dormancy, known as vernalisation . Furthermore, although flower development is induced at warmer temperatures (20–25 °C or 68–77 °F), elongation of the flower stalk and proper flowering is dependent on an extended period of low temperature (< 10 °C or 50 °F). Tulip bulbs imported to warm-winter areas are often planted in autumn to be treated as annuals. The colour of tulip flowers also varies with growing conditions. In

5488-425: The desired hue or pigeon droppings onto flower roots. Tulips affected by the mosaic virus are called "broken"; while such plants can occasionally revert to a plain or solid colouring, they will remain infected and have to be destroyed. Today the virus is almost eradicated from tulip growers' fields. The multicoloured patterns of modern varieties result from breeding; they normally have solid, un-feathered borders between

5586-510: The dormancy period is over and new growth begins. In climates that are warm all year long, perennials may grow continuously. Annuals which complete their life cycle in one growing season, in contrast with perennials, produce seeds as the next generation and die; the seeds may survive cold or dry periods or germinate soon after dispersal depending on the climate. Some perennials retain their foliage year-round; these are evergreen perennials. Deciduous perennials shed all their leaves part of

5684-675: The early 17th century to c. 1960, is held by Polly Nicholson at Blackland House , near Calne in Wiltshire. It is believed the first tulips in the United States were grown near Spring Pond at the Fay Estate in Lynn and Salem , Massachusetts. From 1847 to 1865, Richard Sullivan Fay, Esq., one of Lynn's wealthiest men, settled on 500 acres (2 km; 202 ha) located partly in present-day Lynn and partly in present-day Salem. Mr. Fay imported many different trees and plants from all parts of

5782-518: The early 17th century, entire properties were exchanged as payment for a single tulip bulb. The value of the flower gave it an aura of mystique, and numerous publications describing varieties in lavish garden manuals were published, cashing in on the value of the flower. An export business was built up in France, supplying Dutch, Flemish, German and English buyers. The trade drifted slowly from the French to

5880-551: The early 17th century. These curiosities fitted well in an age when natural oddities were cherished especially in the Netherlands, France, Germany and England, where the spice trade with the East Indies had made many people wealthy. Nouveaux riches seeking wealthy displays embraced the exotic plant market, especially in the Low Countries where gardens had become fashionable. A craze for bulbs soon grew in France, where in

5978-619: The eastern shore of the Caspian Sea through Turkmenistan , Bukhara , Samarkand and Tashkent (Uzbekistan), to the eastern end of the range in the Pamir-Alai and Tien-Shan mountains in Central Asia, which form the centre of diversity . Further to the east, Tulipa is found in the western Himalayas , southern Siberia , Inner Mongolia, and as far as the northwest of China. While authorities have stated that no tulips west of

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6076-439: The fertilization is well organized and the average size of groups is small. Simultaneous hermaphroditism that exclusively reproduces through self-fertilization has evolved many times in plants and animals, but it might not last long evolutionarily. Joan Roughgarden and Priya Iyer argued that the last common ancestor for animals was hermaphroditic and that transitions from hermaphroditism to gonochorism were more numerous than

6174-476: The garden of Councillor Heinrich Herwart. In Central and Northern Europe, tulip bulbs are generally removed from the ground in June and must be replanted by September for the winter. It is doubtful that Busbecq could have had the tulip bulbs harvested, shipped to Germany and replanted between March 1558 and Gessner's description the following year. Pietro Andrea Mattioli illustrated a tulip in 1565 but identified it as

6272-541: The growing season, and can grow taller than annuals. In doing so they can better compete for space and collect more light. Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of vegetative reproduction rather than seeding. These structures include bulbs , tubers , woody crowns, rhizomes , turions , woody stems, or crowns which allows them to survive periods of dormancy over cold or dry seasons; these structures typically store carbohydrates which are used once

6370-431: The growth of ovarian tissues ). Thus, testicular and ovarian tissues will both be present in the same individual. Of all total recorded cases of ovotesticular DSD, in only 8% percent of all cases was SRY present, leaving the rest of cases that could be explained to other or less common causes, with the vast majority simply being currently unexplainable. Fetuses were previously thought to be phenotypically female before

6468-620: The hormones produced due to environmental situations (i.e., seasons), reproduction, and stage of development to begin and halt the ability to grow or flower. There is also a distinction between the ability to grow and the actual task of growth. For example, most trees regain the ability to grow during winter but do not initiate physical growth until the spring and summer months. The start of dormancy can be seen in perennial plants through withering flowers, loss of leaves on trees, and halting of reproduction in both flowering and budding plants. Perennial species may produce relatively large seeds that have

6566-553: The individual flowers are not hermaphroditic if they only produce gametes of one sex. 65% of gymnosperm species are dioecious, but conifers are almost all monoecious. Some plants can change their sex throughout their lifetime, a phenomenon called sequential hermaphroditism . In andromonoecious species, the plants produce perfect (hermaphrodite) flowers and separate fertile male flowers that are sterile as female. Andromonoecy occurs in about 4000 species of flowering plants (2% of flowering plants). In gynomonoecious species,

6664-434: The interior surface. The inner petals have a small, delicate cleft at the top, while the sturdier outer ones form uninterrupted ovals. The flowers have six distinct, basifixed introrse stamens arranged in two whorls of three, which vary in length and may be glabrous or hairy. The filaments are shorter than the tepals and dilated towards their base. The style is short or absent and each stigma has three distinct lobes, and

6762-420: The more completely perennial vegetables are: Many aquatic plants are perennial even though many do not have woody tissue. Examples include: Hermaphrodite#Plants A hermaphrodite ( / h ər ˈ m æ f r ə ˌ d aɪ t / ) is a sexually reproducing organism that produces both male and female gametes . Animal species in which individuals are either male or female are gonochoric , which

6860-560: The original species (known in horticulture as botanical tulips). They are popular throughout the world, both as ornamental garden plants and as cut flowers . Tulips are perennial herbaceous bulbiferous geophytes that bloom in spring and die back after flowering to an underground storage bulb. Depending on the species, tulip plants can be between 10 and 70 cm (4 and 28 inches) high. Tulip stems have few leaves. Larger species tend to have multiple leaves. Plants typically have two to six leaves, some species up to 12. The tulip's leaf

6958-427: The plains of India, as he did with other plants like melons and grapes. The tulip represents the official symbol of Turkey. In Moorish Andalus , a "Makedonian bulb" ( basal al-maqdunis ) or "bucket- Narcissus " ( naryis qadusi ) was cultivated as an ornamental plant in gardens. It was supposed to have come from Alexandria and may have been Tulipa sylvestris , but the identification is not wholly secure. Although it

7056-423: The plant. Other pathogens include anthracnose , bacterial soft rot , blight caused by Sclerotium rolfsii , bulb nematodes , other rots including blue molds , black molds and mushy rot. The fungus Trichoderma viride can infect tulips, producing dried leaf tips and reduced growth, although symptoms are usually mild and only present on bulbs growing in glasshouses . Variegated tulips admired during

7154-519: The plants produce hermaphrodite flowers and separate male-sterile pistillate flowers. One example is the meadow saxifrage, Saxifraga granulata . Charles Darwin gave several other examples in his 1877 book "The Different Forms of Flowers on Plants of the Same Species". About 57% of moss species and 68% of liverworts are unisexual , meaning that their gametophytes produce either male or female gametes, but not both. Sequential hermaphroditism

7252-725: The pollen to move several feet to another plant but only within their blossoms." Tulips can be propagated through bulb offsets , seeds or micropropagation . Offsets and tissue culture methods are means of asexual propagation for producing genetic clones of the parent plant, which maintains cultivar genetic integrity. Seeds are most often used to propagate species and subspecies or to create new hybrids . Many tulip species can cross-pollinate with each other, and when wild tulip populations overlap geographically with other tulip species or subspecies, they often hybridise and create mixed populations. Most commercial tulip cultivars are complex hybrids, and often sterile . Offsets require

7350-474: The result of an atypical set of sex chromosomes. One common cause of variations in sex characteristics traits is the crossing over of the testis-determining factor (SRY) from the Y chromosome to the X chromosome during meiosis . The SRY is then activated in only certain areas, causing development of testes in some areas by beginning a series of events starting with the upregulation of the transcription factor (SOX9), and in other areas not being active (causing

7448-487: The reverse. Other scientists have criticized this argument; saying it’s based on paraphyletic Spiralia , assignments of sexual modes for the phylum level than the species level, and methods exclusively based on maximum parsimony . Hermaphroditism is polyphyletic in invertebrates where it evolved from gonochorism and gonochorism is also ancestral to hermaphroditic fishes. According to Nelson Çabej simultaneous hermaphroditism in animals most likely evolved due to

7546-442: The same plant. There are also mixed breeding systems , in both plants and animals, where hermaphrodite individuals coexist with males (called androdioecy ) or with females (called gynodioecy ), or all three exist in the same species (called trioecy ). Sometimes, both male and hermaphrodite flowers occur on the same plant ( andromonoecy ) or both female and hermaphrodite flowers occur on the same plant ( gynomonoecy ). Hermaphrodism

7644-622: The scholar Qasim from Herat in contrast had identified both wild and garden tulips (lale) as anemones ( shaqayq al-nu'man ) but described the crown imperial as laleh kakli . In a Turkic text written before 1495, the Chagatay Husayn Bayqarah mentions tulips ( lale ). Babur , the founder of the Mughal Empire , also names tulips in the Baburnama . He may actually have introduced them from Afghanistan to

7742-465: The seed yield of perennial species, which could result in the creation of new perennial grain crops. Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass . A perennial rice developed in 2018, was reported in 2023, to have provided a similar yield to replanted annual rice when evaluated over eight consecutive harvests. Perennial plants dominate many natural ecosystems on land and in fresh water, with only

7840-568: The size-advantage mode first proposed by Michael T. Ghiselin which states that if an individual of a certain sex could significantly increase its reproductive success after reaching a certain size, it would be to their advantage to switch to that sex. Sequential hermaphrodites can be divided into three broad categories: Dichogamy can have both conservation-related implications for humans, as mentioned above, as well as economic implications. For instance, groupers are favoured fish for eating in many Asian countries and are often aquacultured . Since

7938-668: The spring and summer, die back every autumn and winter, and then return in the spring from their rootstock or other overwintering structure, are known as herbaceous perennials . However, depending on the rigours of the local climate (temperature, moisture, organic content in the soil, microorganisms), a plant that is a perennial in its native habitat, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings, or from divisions. Tomato vines, for example, live several years in their natural tropical/ subtropical habitat but are grown as annuals in temperate regions because their above-ground biomass does not survive

8036-431: The stems. Herbaceous perennials from temperate and alpine regions of the world can tolerate the cold during winter. Perennial plants may remain dormant for long periods and then recommence growth and reproduction when the environment is more suitable, while most annual plants complete their life cycle during one growing period, and biennials have two growing periods. The meristem of perennial plants communicates with

8134-458: The summer highland pastures ( Yayla ) at Spil Dağı above the town of Manisa . They seem to have consisted of wild tulips. However, of the 14 tulip species known from Turkey, only four are considered to be of local origin, so wild tulips from Iran and Central Asia may have been brought into Turkey during the Seljuk and especially Ottoman periods. Also, Sultan Ahmet imported domestic tulip bulbs from

8232-471: The taxon Tulipa ×gesneriana . They have usually several species in their direct background, but most have been derived from Tulipa suaveolens (today often regarded as a synonym with Tulipa schrenkii ). Tulipa ×gesneriana is in itself an early hybrid of complex origin and is probably not the same taxon as was described by Conrad Gessner in the 16th century. The UK's National Collection of English florists' tulips and Dutch historic tulips, dating from

8330-648: The term hermaphrodite, that the people who bore the sexes of both man and woman were regarded by the Athenians and the Romans as monsters, and thrown into the sea at Athens and into the Tiber at Rome. Similarly, the 17th-century English jurist and judge Edward Coke (Lord Coke), wrote in his Institutes of the Lawes of England on laws of succession stating, "Every heire is either a male, a female, or an hermaphrodite, that

8428-505: The terminology " disorders of sex development " (also known as variations in sex characteristics .) This is particularly significant because of the relationship between medical terminology and medical intervention. Intersex civil society organizations , and many human rights institutions, have criticized medical interventions designed to make bodies more typically male or female. In some cases, variations in sex characteristics are caused by unusual levels of sex hormones, which may be

8526-399: The terms, and also a shift to nomenclature based on genetics. The term "intersex" described a wide variety of combinations of what are ambiguous biological characteristics. Intersex biology may include, for example, ambiguous-looking external genitalia, karyotypes that include mixed XX and XY chromosome pairs (46XX/46XY, 46XX/47XXY or 45X/XY mosaic ). Clinically, medicine currently uses

8624-433: The time and most owned by Pauw, his refusal to sell any flowers, despite wildly escalating offers, is believed by some to have sparked the mania. Tulip flowers come in a wide variety of colours, except pure blue (several tulips with "blue" in the name have a faint violet hue), and have absent nectaries . Tulip flowers are generally bereft of scent and are the coolest of floral characters. While tulips can be bred to display

8722-591: The tulip. Tulips are mainly distributed along a band corresponding to latitude 40° north, from southeast of Europe ( Greece , Albania , North Macedonia , Kosovo , Southern Serbia , Bulgaria , most part of Romania , Ukraine , Russia) and Turkey in the west, through the Levant (Syria, Israel, Palestinian Territories , Lebanon and Jordan) and the Sinai Peninsula . From there it extends eastwards through Jerevan (Armenia), and Baku ( Azerbaijan ) and on

8820-456: The visible unitary colour. The breaking of flowers occurs when a virus suppresses anthocyanin and the base colour is exposed as a streak. The great majority of tulips, both species and cultivars, have no discernable scent, but a few of both are scented to a degree, and Anna Pavord describes T. hungarica as "strongly scented", and among cultivars, some such as "Monte Carlo" and "Brown Sugar" are "scented", and "Creme Upstar" "fragrant". Tulipa

8918-399: The wild for many centuries. The history of naturalisation is unknown, but populations are usually associated with agricultural practices and are possibly linked to saffron cultivation. Some neo-tulipae have been brought into cultivation, and are often offered as botanical tulips. These cultivated plants can be classified into two Cultivar Groups: 'Grengiolensis Group', with picotee tepals, and

9016-405: The winter. There is also a class of evergreen perennials which lack woody stems, such as Bergenia which retain a mantle of leaves throughout the year. An intermediate class of plants is known as subshrubs , which retain a vestigial woody structure in winter, e.g. Penstemon . The symbol for a perennial plant, based on Species Plantarum by Linnaeus , is [REDACTED] , which is also

9114-432: The word tulipan to describe the flower. Extensive speculation has tried to understand why he would state this, given that the Turkish word for tulip is lale . It is from this speculation that tulipan being a translation error referring to turbans is derived. This etymology has been challenged and makes no assumptions about possible errors. At no point does Busbecq state this was the word used in Turkey, he simply states it

9212-540: The word used by his "Turk/Hungarian" guides. The Hungarian word tulipan may be adopted from an Indo-Aryan reference to the tulip as a symbol of resurrection, tala meaning "bottom or underworld" and pAna meaning "defence". Prior to arriving in Europe the Hungarians, and other Finno-Ugrians, embraced the Indo-Iranian cult of the dead, Yima/Yama, and would have been familiar with all of its symbols including

9310-450: The world and planted them among the meadows of the Fay Estate. The Netherlands is the world's main producer of commercial tulip plants, producing as many as 3 billion bulbs annually, the majority for export. "Unlike many flower species, tulips do not produce nectar to entice insect pollination. Instead, tulips rely on wind and land animals to move their pollen between reproductive organs. Because they are self-pollinating, they do not need

9408-439: The year. Deciduous perennials include herbaceous and woody plants; herbaceous plants have stems that lack hard, fibrous growth, while woody plants have stems with buds that survive above ground during dormancy. Some perennials are semi-deciduous, meaning they lose some of their leaves in either winter or summer. Deciduous perennials shed their leaves when growing conditions are no longer suitable for photosynthesis, such as when it

9506-513: Was born with a physical body combining male and female sexes. The word hermaphrodite entered the English lexicon as early as the late fourteenth century. Sequential hermaphrodites ( dichogamy ) occur in species in which the individual first develops as one sex, but can later change into the opposite sex. (Definitions differ on whether sequential hermaphroditism encompasses serial hermaphroditism; for authors who exclude serial hermaphroditism,

9604-494: Was used by the "Turks". On his way to Constantinople Busbecq states he travelled through Hungary and used Hungarian guides. Until recent times "Turk" was a common term when referring to Hungarians. The word tulipan is in fact the Hungarian word for tulip. As long as one recognizes "Turk" as a reference to Hungarians, no amount of speculation is required to reconcile the word's origin or form. Busbecq may have been simply repeating

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