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95-496: Historically, it was an important salmon fishery, but it has now been infected with the salmon parasite Gyrodactylus salaris . This Nordland location article is a stub . You can help Misplaced Pages by expanding it . This article related to a river in Norway is a stub . You can help Misplaced Pages by expanding it . Salmon all other members of Salmoninae Salmon ( / ˈ s æ m ən / ; pl. : salmon)

190-506: A common ancestor . The Eocene salmon's fossil from British Columbia provides evidence that the divergence between Pacific and Atlantic salmon had not yet occurred 40 million years ago. Both the fossil record and analysis of mitochondrial DNA suggest the divergence occurred 10 to 20 million years ago during the Miocene . This independent evidence from DNA analysis and the fossil record indicate that salmon divergence occurred long before

285-826: A Gaulish substrate, although there is some debate. One of these is considered certain, because this fact is clearly attested in the Gaulish-language epigraphy on the pottery found at la Graufesenque ( A.D. 1st century). There, the Greek word paropsid-es (written in Latin) appears as paraxsid-i . The consonant clusters /ps/ and /pt/ shifted to /xs/ and /xt/, e.g. Lat capsa > *kaxsa > caisse ( ≠ Italian cassa ) or captīvus > *kaxtivus > OF chaitif (mod. chétif ; cf. Irish cacht 'servant'; ≠ Italian cattiv-ità , Portuguese cativo , Spanish cautivo ). This phonetic evolution

380-421: A definitive influence on the development of Old French, which partly explains why the earliest attested Old French documents are older than the earliest attestations in other Romance languages (e.g. Strasbourg Oaths , Sequence of Saint Eulalia ). It is the result of an earlier gap created between Classical Latin and its evolved forms, which slowly reduced and eventually severed the mutual intelligibility between

475-620: A feudal elite and commerce. The area of Old French in contemporary terms corresponded to the northern parts of the Kingdom of France (including Anjou and Normandy , which in the 12th century were ruled by the Plantagenet kings of England ), Upper Burgundy and the Duchy of Lorraine . The Norman dialect was also spread to England and Ireland , and during the Crusades , Old French

570-442: A few species of salmon remain in fresh water throughout their life cycle, the majority are anadromous and migrate to the ocean for maturation: in these species, smolts spend a portion of their out-migration time in brackish water, where their body chemistry becomes accustomed to osmoregulation in the ocean. This body chemistry change is hormone-driven, causing physiological adjustments in the function of osmoregulatory organs such as

665-477: A fish returns after just one year's sea feeding, it is called a grilse in Canada, Britain, and Ireland. Grilse may be present at spawning, and go unnoticed by large males, releasing their own sperm on the eggs. Prior to spawning, depending on the species , salmon undergo changes. They may grow a hump, develop canine-like teeth, or develop a kype (a pronounced curvature of the jaws in male salmon). All change from

760-420: A fraindre, Fors Sarragoce qu'est en une montaigne; Li reis Marsilies la tient, ki Deu nen aimet, Mahomet sert ed Apolin reclaimet: Ne·s poet guarder que mals ne l'i ataignet! ˈt͡ʃarləs li ˈre͜is, ˈnɔstr‿empəˈræðrə ˈmaɲəs ˈsɛt ˈant͡s ˈtot͡s ˈple͜ins ˈað esˈtæθ en esˈpaɲə ˈtræs k‿en la ˈmɛr konˈkist la ˈtɛr alˈta͜iɲə t͡ʃasˈtɛl ni ˈaθ ki dəˈvant ˈly͜i rəˈma͜iɲəθ ˈmyrs nə t͡siˈtæθ n‿i ˈɛst rəˈmæs

855-736: A future in aquaculture that eliminates the need for land, freshwater, or fertilizer to raise fish. Salmon population levels are of concern in the Atlantic and in some parts of the Pacific. The population of wild salmon declined markedly in recent decades, especially North Atlantic populations, which spawn in the waters of western Europe and eastern Canada, and wild salmon in the Snake and Columbia River systems in northwestern United States. Old French language Old French ( franceis , françois , romanz ; French : ancien français )

950-569: A high demand for wild forage fish . As a predator, salmon require large nutritional intakes of protein , and farmed salmon consume more fish than they generate as a final product. On a dry weight basis, 2–4 kg of wild-caught fish are needed to produce one kilogram of salmon. As the salmon farming industry expands, it requires more forage fish for feed, at a time when 75% of the world's monitored fisheries are already near to or have exceeded their maximum sustainable yield . The industrial-scale extraction of wild forage fish for salmon farming affects

1045-548: A lot of work on Henneguya salminicola was done by scientists at the Pacific Biological Station in Nanaimo in the mid-1980s, in particular, an overview report which states, "the fish that have the longest fresh water residence time as juveniles have the most noticeable infections. Hence in order of prevalence , coho are most infected followed by sockeye, chinook, chum and pink. As well, the report says, at

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1140-1242: A new orthography for the latter; among the earliest examples are parts of the Oaths of Strasbourg and the Sequence of Saint Eulalia . Some Gaulish words influenced Vulgar Latin and, through this, other Romance languages. For example, classical Latin equus was uniformly replaced in Vulgar Latin by caballus 'nag, work horse', derived from Gaulish caballos (cf. Welsh ceffyl , Breton kefel ), yielding ModF cheval , Occitan caval ( chaval ), Catalan cavall , Spanish caballo , Portuguese cavalo , Italian cavallo , Romanian cal , and, by extension, English cavalry and chivalry (both via different forms of [Old] French: Old Norman and Francien ). An estimated 200 words of Gaulish etymology survive in Modern French, for example chêne , 'oak tree', and charrue , 'plough'. Within historical phonology and studies of language contact , various phonological changes have been posited as caused by

1235-655: A radical change had the effect of rendering Latin sermons completely unintelligible to the general Romance-speaking public, which prompted officials a few years later, at the Third Council of Tours , to instruct priests to read sermons aloud in the old way, in rusticam romanam linguam or 'plain Roman[ce] speech'. As there was now no unambiguous way to indicate whether a given text was to be read aloud as Latin or Romance, various attempts were made in France to devise

1330-412: A technically demanding kind of sport fishing , not necessarily intuitive for beginning fishermen. A conflict exists between commercial fishermen and recreational fishermen for the right to salmon stock resources . Commercial fishing in estuaries and coastal areas is often restricted so enough salmon can return to their natal rivers where they can spawn and be available for sport fishing. On parts of

1425-557: A very distinctive identity compared to the other future Romance languages. The first noticeable influence is the substitution of the Latin melodic accent with a Germanic stress and its result was diphthongization , differentiation between long and short vowels, the fall of the unaccented syllable and of the final vowels: Additionally, two phonemes that had long since died out in Vulgar Latin were reintroduced: [h] and [w] (> OF g(u)- , ONF w- cf. Picard w- ): In contrast,

1520-505: A ˈfra͜indrə ˈfɔrs saraˈgot͡sə k‿ˈɛst en ˈynə monˈtaɲə li ˈre͜is marˈsiʎəs la ˈti͜ɛnt, ki ˈdɛ͜u nən ˈa͜iməθ mahoˈmɛt ˈsɛrt eð apoˈlin rəˈkla͜iməθ nə‿s ˈpu͜ɛt gwarˈdær kə ˈmals nə l‿i aˈta͜iɲəθ Charles the king, our great emperor, Has been in Spain for seven full years: He has conquered the lofty land up to the sea. No castle remains standing before him; No wall or city is left to destroy Other than Saragossa, which lies atop

1615-858: Is a major contributor to the world production of farmed finfish, representing about US$ 10 billion annually. Other commonly cultured fish species include tilapia , catfish , sea bass , carp and bream . Salmon farming is significant in Chile , Norway , Scotland , Canada and the Faroe Islands ; it is the source for most salmon consumed in the United States and Europe. Atlantic salmon are also, in very small volumes, farmed in Russia and Tasmania , Australia. Salmon are carnivorous , and need to be fed meals produced from catching other wild forage fish and other marine organisms. Salmon farming leads to

1710-490: Is called Vulgar Latin , the common spoken language of the Western Roman Empire . Vulgar Latin differed from Classical Latin in phonology and morphology as well as exhibiting lexical differences; however, they were mutually intelligible until the 7th century when Classical Latin 'died' as a daily spoken language, and had to be learned as a second language (though it was long thought of as the formal version of

1805-609: Is common in its later stages with the shift of the Latin cluster /kt/ in Old French ( Lat factum > fait , ≠ Italian fatto , Portuguese feito , Spanish hecho ; or lactem * > lait , ≠ Italian latte , Portuguese leite , Spanish leche ). This means that both /pt/ and /kt/ must have first merged into /kt/ in the history of Old French, after which this /kt/ shifted to /xt/. In parallel, /ps/ and /ks/ merged into /ks/ before shifting to /xs/, apparently under Gaulish influence. The Celtic Gaulish language

1900-680: Is derived from Middle English : samoun , samon and saumon , which in turn are from Anglo-Norman : saumon , from Old French : saumon , and from Latin : salmō (which in turn might have originated from salire , meaning "to leap". ). The unpronounced "l" absent from Middle English was later added as a Latinisation to make the word closer to its Latin root. The term salmon has mostly displaced its now dialectal synonym lax , in turn from Middle English : lax , from Old English : leax , from Proto-Germanic : * lahsaz from Proto-Indo-European : *lakso- . The seven commercially important species of salmon occur in two genera of

1995-441: Is quite high (perhaps as high as 40 to 50%). To lay her roe , the female salmon uses her tail (caudal fin), to create a low-pressure zone, lifting gravel to be swept downstream, excavating a shallow depression, called a redd. The redd may sometimes contain 5,000 eggs covering 2.8 m (30 sq ft). The eggs usually range from orange to red. One or more males approach the female in her redd, depositing sperm, or milt, over

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2090-966: Is strictly a fish parasite that cannot live in or affect warm blooded animals, including man". According to Klaus Schallie, Molluscan Shellfish Program Specialist with the Canadian Food Inspection Agency , " Henneguya salminicola is found in southern B.C. also and in all species of salmon. I have previously examined smoked chum salmon sides that were riddled with cysts and some sockeye runs in Barkley Sound (southern B.C., west coast of Vancouver Island ) are noted for their high incidence of infestation." Sea lice , particularly Lepeophtheirus salmonis and various Caligus species, including C. clemensi and C. rogercresseyi , can cause deadly infestations of both farm-grown and wild salmon. Sea lice are ectoparasites which feed on mucus, blood, and skin, and migrate and latch onto

2185-488: Is the common name for several commercially important species of euryhaline ray-finned fish from the genera Salmo and Oncorhynchus of the family Salmonidae , native to tributaries of the North Atlantic ( Salmo ) and North Pacific ( Oncorhynchus ) basins. Other closely related fish in the same family include trout , char , grayling , whitefish , lenok and taimen , all coldwater fish of

2280-542: Is thought to have survived into the 6th century in France, despite considerable cultural Romanization. Coexisting with Latin, Gaulish helped shape the Vulgar Latin dialects that developed into French, with effects including loanwords and calques (including oui , the word for "yes"), sound changes shaped by Gaulish influence, and influences in conjugation and word order. A computational study from 2003 suggests that early gender shifts may have been motivated by

2375-480: Is under development in Alaska . There, the young salmon are released into the ocean far from any wild salmon streams. When it is time for them to spawn, they return to where they were released, where fishermen can catch them. An alternative method to hatcheries is to use spawning channels. These are artificial streams , usually parallel to an existing stream, with concrete or rip-rap sides and gravel bottoms. Water from

2470-594: The langue d'oïl as early as the 9th century and is attested as a distinct Gallo-Romance variety by the 12th century. Dialects or variants of Old French include: Some modern languages are derived from Old French dialects other than Classical French, which is based on the Île-de-France dialect. They include Angevin , Berrichon , Bourguignon-Morvandiau , Champenois , Franc-Comtois , Gallo, Lorrain, Norman , Picard, Poitevin , Saintongeais , and Walloon. Beginning with Plautus ' time (254–184 b.c. ), one can see phonological changes between Classical Latin and what

2565-531: The Bibliothèque bleue – that a standardized Classical French spread throughout France alongside the regional dialects. The material and cultural conditions in France and associated territories around the year 1100 triggered what Charles Homer Haskins termed the " Renaissance of the 12th century ", resulting in a profusion of creative works in a variety of genres. Old French gave way to Middle French in

2660-509: The Roman de Fauvel in 1310 and 1314, a satire on abuses in the medieval church, filled with medieval motets , lais , rondeaux and other new secular forms of poetry and music (mostly anonymous, but with several pieces by Philippe de Vitry , who would coin the expression ars nova to distinguish the new musical practice from the music of the immediately preceding age). The best-known poet and composer of ars nova secular music and chansons of

2755-624: The Cook Inlet Aquaculture Association , as a way to increase salmon populations in situations where they have declined due to overharvesting , construction of dams and habitat destruction or fragmentation . Negative consequences to this sort of population manipulation include genetic "dilution" of the wild stocks. Many jurisdictions are now beginning to discourage supplemental fish planting in favour of harvest controls, and habitat improvement and protection. A variant method of fish stocking , called ocean ranching,

2850-489: The Japanese word for trout. On the other hand, the steelhead and sea trout , the anadromous forms of rainbow trout and brown trout respectively, are from the same genera as salmon and live identical migratory lives, but neither is termed "salmon" . The extinct Eosalmo driftwoodensis , the oldest known Salmoninae fish in the fossil record , helps scientists figure how the different species of salmon diverged from

2945-526: The Pacific Northwest and Alaska , salmon are keystone species . The migration of salmon represent a massive retrograde nutrient transfer, rich in nitrogen , sulfur , carbon and phosphorus , from the ocean to the inland freshwater ecosystems . Predation by piscivorous land animals (such as ospreys , bears and otters ) along the journey serve to transfer the nutrients from the water to land, and decomposition of salmon carcass benefits

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3040-461: The Pacific coast of Canada , the louse-induced mortality of pink salmon in some regions is commonly over 80%. The risk of injury caused by underwater pile driving has been studied by Dr. Halvorsen and her co-workers. The study concluded that the fish are at risk of injury if the cumulative sound exposure level exceeds 210 dB relative to 1 μPa s. As can be seen from the production chart at

3135-514: The Quaternary glaciation began the cycle of glacial advance and retreat. There are several other species of fish which are colloquially called "salmon" but are not true salmon. Of those listed below, the Danube salmon or huchen is a large freshwater salmonid closely related (from the same subfamily) to the seven species of salmon above, but others are fishes of unrelated orders , given

3230-672: The Salmo and Oncorhynchus genera also contain a number of trout species informally referred to as salmon. Within Salmo , the Adriatic salmon ( Salmo obtusirostris ) and Black Sea salmon ( Salmo labrax ) have both been named as salmon in English , although they fall outside the generally recognized seven salmon species. The masu salmon ( Oncorhynchus masou ) is actually considered a trout ("cherry trout") in Japan , with masu actually being

3325-525: The chansons de geste is The Song of Roland (earliest version composed in the late 11th century). Bertrand de Bar-sur-Aube in his Girart de Vienne set out a grouping of the chansons de geste into three cycles : the Geste du roi centering on Charlemagne, the Geste de Garin de Monglane (whose central character was William of Orange ), and the Geste de Doon de Mayence or the "rebel vassal cycle",

3420-497: The forest ecosystem . In the case of Pacific salmon , most (if not all) of the salmon that survive to reach the headwater spawning grounds will die after laying eggs and their dead bodies sink to cover the gravel beds, with the nutrients released from the biodegradation of their corpses providing a significant boost to these otherwise biomass -poor shallow streams. Grizzly bears function as ecosystem engineers , capturing salmon and carrying them into adjacent dry land to eat

3515-524: The myrtle zone . These dams can be overtopped at high tide and hold water at low tide. This provides refuges for juvenile salmon so they do not have to swim into large channels where they are subject to predation by larger fish. It has been discovered that rivers which have seen a decline or disappearance of anadromous lampreys , loss of the lampreys also affects the salmon in a negative way. Like salmon, anadromous lampreys stop feeding and die after spawning, and their decomposing bodies release nutrients into

3610-590: The otolith (annuli), analogous to the growth rings visible in a tree trunk. Freshwater growth shows as densely crowded rings, sea growth as widely spaced rings; spawning is marked by significant erosion as body mass is converted into eggs and milt. Freshwater streams and estuaries provide important habitat for many salmon species. They feed on terrestrial and aquatic insects , amphipods , and other crustaceans while young, and primarily on other fish when older. Eggs are laid in deeper water with larger gravel and need cool water and good water flow (to supply oxygen) to

3705-616: The subarctic and cooler temperate regions with some sporadic endorheic populations in Central Asia . Salmon are typically anadromous : they hatch in the shallow gravel beds of freshwater headstreams and spend their juvenile years in rivers , lakes and freshwater wetlands , migrate to the ocean as adults and live like sea fish , then return to their freshwater birthplace to reproduce . However, populations of several species are restricted to fresh waters (i.e. landlocked) throughout their lives. Folklore has it that

3800-849: The subfamily Salmoninae . The genus Salmo contains the Atlantic salmon , found in both sides of the North Atlantic , as well as more than 40 other species commonly named as trout . The genus Oncorhynchus contains 12 recognised species which occur naturally only in the North Pacific, six of which are known as Pacific salmon while the remainder are considered trout. Outside their native habitats, Chinook salmon have been successfully introduced in New Zealand and Patagonia , while coho , sockeye and Atlantic salmon have been established in Patagonia, as well.     Both

3895-596: The Gallo-Romance that prefigures French – after the Reichenau and Kassel glosses (8th and 9th centuries) – are the Oaths of Strasbourg (treaties and charters into which King Charles the Bald entered in 842): Pro Deo amur et pro Christian poblo et nostro commun salvament, d'ist di en avant, in quant Deus savir et podir me dunat, si salvarai eo cist meon fradre Karlo, et in aiudha et in cadhuna cosa ... (For

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3990-747: The Italian, Portuguese and Spanish words of Germanic origin borrowed from French or directly from Germanic retain /gw/ ~ /g/ , e.g. Italian, Spanish guerra 'war', alongside /g/ in French guerre ). These examples show a clear consequence of bilingualism, that sometimes even changed the first syllable of the Latin words. One example of a Latin word influencing an OLF loan is framboise 'raspberry', from OF frambeise , from OLF *brāmbesi 'blackberry' (cf. Dutch braambes , braambezie ; akin to German Brombeere , English dial. bramberry ) blended with LL fraga or OF fraie 'strawberry', which explains

4085-479: The North American West Coast salmon sport fishing has completely replaced inshore commercial salmon fishing. In most cases, the commercial value of a salmon sold as seafood can be several times less than the value attributed to the same fish caught by a sport fisherman. This is "a powerful economic argument for allocating stock resources preferentially to sport fishing". Salmon aquaculture

4180-565: The Old French dialects diverged into a number of distinct langues d'oïl , among which Middle French proper was the dialect of the Île-de-France region. During the Early Modern period , French was established as the official language of the Kingdom of France throughout the realm, including the langue d'oc -speaking territories in the south. It was only in the 17th to 18th centuries – with the development especially of popular literature of

4275-570: The Pacific Ocean, the second host releases a stage infective to salmon. The parasite is then carried in the salmon until the next spawning cycle. The myxosporean parasite that causes whirling disease in trout has a similar life cycle. However, as opposed to whirling disease, the Henneguya infestation does not appear to cause disease in the host salmon—even heavily infected fish tend to return to spawn successfully. According to Dr. Kieser,

4370-497: The Renaissance short story ( conte or nouvelle ). Among the earliest works of rhetoric and logic to appear in Old French were the translations of Rhetorica ad Herennium and Boethius ' De topicis differentiis by John of Antioch in 1282. In northern Italy, authors developed Franco-Italian , a mixed language of Old French and Venetian or Lombard used in literary works in the 13th and 14th centuries. Old French

4465-432: The adjacent stream is piped into the top of the channel, sometimes via a header pond, to settle out sediment. Spawning success is often much better in channels than in adjacent streams due to the control of floods, which in some years can wash out the natural redds. Because of the lack of floods, spawning channels must sometimes be cleaned out to remove accumulated sediment. The same floods that destroy natural redds also clean

4560-608: The common name "salmon" simply due to similar shapes, behaviors and niches occupied: Salmon eggs are laid in freshwater streams typically at high latitudes. The eggs hatch into alevin or sac fry. The fry quickly develop into parr with camouflaging vertical stripes. The parr stay for six months to three years in their natal stream before becoming smolts, which are distinguished by their bright, silvery colour with scales that are easily rubbed off. Only 10% of all salmon eggs are estimated to survive to this stage. The smolt body chemistry changes, allowing them to live in salt water. While

4655-430: The developing embryos. Mortality of salmon in the early life stages is usually high due to natural predation and human-induced changes in habitat, such as siltation, high water temperatures, low oxygen concentration, loss of stream cover, and reductions in river flow. Estuaries and their associated wetlands provide vital nursery areas for the salmon prior to their departure to the open ocean. Wetlands not only help buffer

4750-590: The development of northern French culture in and around Île-de-France , which slowly but firmly asserted its ascendency over the more southerly areas of Aquitaine and Tolosa ( Toulouse ); however, the Capetians ' langue d'oïl , the forerunner of modern standard French, did not begin to become the common speech of all of France until after the French Revolution . In the Late Middle Ages,

4845-472: The elimination of beavers from large parts of the river system, salmon runs plummeted, even in the absence of many of the factors usually associated with the demise of salmon runs. Salmon recruitment can be affected by beavers' dams because dams can: Beaver dams are able to nurture salmon juveniles in estuarine tidal marshes where the salinity is less than 10 ppm. Beavers build small dams of generally less than 60 cm (2 ft) high in channels in

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4940-821: The estuary from silt and pollutants, but also provide important feeding and hiding areas. Salmon not killed by other means show greatly accelerated deterioration ( phenoptosis , or "programmed aging") at the end of their lives. Their bodies rapidly deteriorate right after they spawn as a result of the release of massive amounts of corticosteroids . Salmon are mid- level carnivores whose diet change according to their life stage. Salmon fry predominantly feed upon zooplanktons until they reach fingerling sizes, when they start to consume more aquatic invertebrates such as insect larvae , micro crustaceans and worms . As juveniles (parrs), they become more predatory and actively prey upon aquatic insects , small crustaceans, tadpoles and small bait fishes . They are also known to breach

5035-754: The first such text. At the beginning of the 13th century, Jean Bodel , in his Chanson de Saisnes , divided medieval French narrative literature into three subject areas: the Matter of France or Matter of Charlemagne ; the Matter of Rome ( romances in an ancient setting); and the Matter of Britain ( Arthurian romances and Breton lais ). The first of these is the subject area of the chansons de geste ("songs of exploits" or "songs of (heroic) deeds"), epic poems typically composed in ten-syllable assonanced (occasionally rhymed ) laisses . More than one hundred chansons de geste have survived in around three hundred manuscripts. The oldest and most celebrated of

5130-432: The fish return to the exact stream where they themselves hatched to spawn , and tracking studies have shown this to be mostly true. A portion of a returning salmon run may stray and spawn in different freshwater systems; the percent of straying depends on the species of salmon. Homing behavior has been shown to depend on olfactory memory . Salmon are important food fish and are intensively farmed in many parts of

5225-443: The fish. There they deposit nutrient-rich urine and feces and partially eaten carcasses. Bears preparing for hibernation tend to preferentially consume the more nutrient- and energy -rich salmon roes and brain over the actual flesh , and are estimated to discard up to half the salmon they've harvested uneaten on the forest floor, in densities that can reach 4,000 kg (8,800 lb) per hectare, providing as much as 24% of

5320-757: The gender of the corresponding word in Gaulish. The pronunciation, vocabulary, and syntax of the Vulgar Latin spoken in Roman Gaul in late antiquity were modified by the Old Frankish language , spoken by the Franks who settled in Gaul from the 5th century and conquered the future Old French-speaking area by the 530s. The name français itself is derived from the name of the Franks. The Old Frankish language had

5415-566: The gills, which leads to large increases in their ability to secrete salt. Hormones involved in increasing salinity tolerance include insulin-like growth factor I , cortisol , and thyroid hormones , which permits the fish to endure the transition from a freshwater environment to the ocean. The salmon spend about one to five years (depending on the species) in the open ocean, where they gradually become sexually mature. The adult salmon then return primarily to their natal streams to spawn. Atlantic salmon spend between one and four years at sea. When

5510-539: The incipient Middle French period was Guillaume de Machaut . Discussions about the origins of non-religious theater ( théâtre profane )—both drama and farce—in the Middle Ages remain controversial, but the idea of a continuous popular tradition stemming from Latin comedy and tragedy to the 9th century seems unlikely. Most historians place the origin of medieval drama in the church's liturgical dialogues and "tropes". Mystery plays were eventually transferred from

5605-684: The left, the global capture reported by different countries to the FAO of commercial wild salmon has remained fairly steady since 1990 at about one million tonnes per year. This is in contrast to farmed salmon (below) which has increased in the same period from about 0.6 million tonnes to well over two million tonnes. Nearly all captured wild salmon are Pacific salmon . The capture of wild Atlantic salmon has always been relatively small, and has declined steadily since 1990. In 2011 only 2,500 tonnes were reported. In contrast, about half of all farmed salmon are Atlantic salmon. Recreational salmon fishing can be

5700-458: The longer the fish remain in fresh water, and they then deteriorate further after they spawn, when they are known as kelts. In all species of Pacific salmon, the mature individuals die within a few days or weeks of spawning, a trait known as semelparity . Between 2 and 4% of Atlantic salmon kelts survive to spawn again, all females. However, even in those species of salmon that may survive to spawn more than once ( iteroparity ), postspawning mortality

5795-639: The loss of an intervening consonant. Manuscripts generally do not distinguish hiatus from true diphthongs, but modern scholarly transcription indicates it with a diaeresis , as in Modern French: Presented below is the first laisse of The Song of Roland along with a broad transcription reflecting reconstructed pronunciation c.  1050 . Charles li reis, nostre emperedre magnes, Set anz toz pleins at estét en Espaigne. Tres qu'en la mer conquist la tere altaigne, Chastel n'i at ki devant lui remaignet. Murs ne citét n'i est remés

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5890-570: The love of God and for the Christian people, and our common salvation, from this day forward, as God will give me the knowledge and the power, I will defend my brother Karlo with my help in everything ...) The second-oldest document in Old French is the Eulalia sequence , which is important for linguistic reconstruction of Old French pronunciation due to its consistent spelling. The royal House of Capet , founded by Hugh Capet in 987, inaugurated

5985-406: The mid-14th century, paving the way for early French Renaissance literature of the 15th century. The earliest extant French literary texts date from the ninth century, but very few texts before the 11th century have survived. The first literary works written in Old French were saints' lives . The Canticle of Saint Eulalie , written in the second half of the 9th century, is generally accepted as

6080-528: The monastery church to the chapter house or refectory hall and finally to the open air, and the vernacular was substituted for Latin. In the 12th century one finds the earliest extant passages in French appearing as refrains inserted into liturgical dramas in Latin, such as a Saint Nicholas (patron saint of the student clercs) play and a Saint Stephen play. An early French dramatic play is Le Jeu d'Adam ( c.  1150 ) written in octosyllabic rhymed couplets with Latin stage directions (implying that it

6175-555: The most famous characters of which were Renaud de Montauban and Girart de Roussillon . A fourth grouping, not listed by Bertrand, is the Crusade cycle , dealing with the First Crusade and its immediate aftermath. Jean Bodel 's other two categories—the "Matter of Rome" and the "Matter of Britain"—concern the French romance or roman . Around a hundred verse romances survive from the period 1150–1220. From around 1200 on,

6270-446: The parasitic infection into a number of cysts that contain milky fluid. This fluid is an accumulation of a large number of parasites. Henneguya and other parasites in the myxosporean group have complex life cycles, where the salmon is one of two hosts. The fish releases the spores after spawning. In the Henneguya case, the spores enter a second host, most likely an invertebrate, in the spawning stream. When juvenile salmon migrate to

6365-526: The preferred prey of seals and sea lions, which can eat 30 lampreys to every salmon, allowing more adult salmon to enter the rivers to spawn without being eaten by the marine mammals. According to Canadian biologist Dorothy Kieser, the myxozoan parasite Henneguya salminicola is commonly found in the flesh of salmonids. It has been recorded in the field samples of salmon returning to the Haida Gwaii Islands . The fish responds by walling off

6460-408: The problems of rising costs for buying hatchery fish feed . Yet another attractive alternative is the increased use of seaweed . Seaweed provides essential minerals and vitamins for growing organisms. It offers the advantage of providing natural amounts of dietary fiber and having a lower glycemic load than grain-based fish meal . In the best-case scenario, widespread use of seaweed could yield

6555-466: The regular streams. Spawning channels preserve the natural selection of natural streams, as there is no benefit, as in hatcheries, to use prophylactic chemicals to control diseases. Farm-raised salmon are fed the carotenoids astaxanthin and canthaxanthin to match their flesh colour to wild salmon to improve their marketability. Wild salmon get these carotenoids , primarily astaxanthin, from eating shellfish and krill . One proposed alternative to

6650-504: The replacement [b] > [f] and in turn the final -se of framboise added to OF fraie to make freise , modern fraise (≠ Wallon frève , Occitan fraga , Romanian fragă , Italian fragola , fravola 'strawberry'). Mildred Pope estimated that perhaps still 15% of the vocabulary of Modern French derives from Germanic sources. This proportion was larger in Old French, because Middle French borrowed heavily from Latin and Italian. The earliest documents said to be written in

6745-407: The roe. The female then covers the eggs by disturbing the gravel at the upstream edge of the depression before moving on to make another redd. The female may make as many as seven redds before her supply of eggs is exhausted. Each year, the fish experiences a period of rapid growth, often in summer, and one of slower growth, normally in winter. This results in ring formation around an earbone called

6840-509: The salmon population. This system is referred to as ranching . It was very common in countries such as Sweden, before the Norwegians developed salmon farming, but is seldom done by private companies. As anyone may catch the salmon when they return to spawn, a company is limited in benefiting financially from their investment. Because of this, the ranching method has mainly been used by various public authorities and non-profit groups, such as

6935-421: The silvery blue of a fresh-run fish from the sea to a darker colour. Salmon can make amazing journeys, sometimes moving hundreds of miles upstream against strong currents and rapids to reproduce. Chinook and sockeye salmon from central Idaho, for example, travel over 1,400 km (900 mi) and climb nearly 2,100 m (7,000 ft) from the Pacific Ocean as they return to spawn. Condition tends to deteriorate

7030-546: The skin of wild salmon during free-swimming, planktonic nauplii and copepodid larval stages, which can persist for several days. Large numbers of highly populated, open-net salmon farms can create exceptionally large concentrations of sea lice; when exposed in river estuaries containing large numbers of open-net farms, many young wild salmon are infected, and do not survive as a result. Adult salmon may survive otherwise critical numbers of sea lice, but small, thin-skinned juvenile salmon migrating to sea are highly vulnerable. On

7125-494: The south of France. The mid-14th century witnessed the emergence of Middle French , the language of the French Renaissance in the Île-de-France region; this dialect was a predecessor to Modern French . Other dialects of Old French evolved themselves into modern forms ( Poitevin-Saintongeais , Gallo , Norman , Picard , Walloon , etc.), each with its linguistic features and history. The region where Old French

7220-547: The spoken language). Vulgar Latin was the ancestor of the Romance languages , including Old French. By the late 8th century, when the Carolingian Renaissance began, native speakers of Romance idioms continued to use Romance orthoepy rules while speaking and reading Latin. When the most prominent scholar of Western Europe at the time, English deacon Alcuin , was tasked by Charlemagne with improving

7315-404: The standards of Latin writing in France, not being a native Romance speaker himself, he prescribed a pronunciation based on a fairly literal interpretation of Latin spelling. For example, in a radical break from the traditional system, a word such as ⟨viridiarium⟩ ' orchard ' now had to be read aloud precisely as it was spelled rather than */verdʒjær/ (later spelled as OF 'vergier' ). Such

7410-458: The stream. Also, along with species like rainbow trout and Sacramento sucker , lampreys clean the gravel in the rivers during spawning. Their larvae, called ammocoetes, are filter feeders which contribute to the health of the waters. They are also a food source for the young salmon, and being fattier and oilier, it is assumed predators prefer them over salmon offspring, taking off some of the predation pressure on smolts. Adult lampreys are also

7505-534: The survivability of other wild predatory fish which rely on them for food. Research is ongoing into sustainable and plant-based salmon feeds. Intensive salmon farming uses open-net cages, which have low production costs. It has the drawback of allowing disease and sea lice to spread to local wild salmon stocks. Another form of salmon production, which is safer but less controllable, is to raise salmon in hatcheries until they are old enough to become independent. They are released into rivers in an attempt to increase

7600-514: The tendency was increasingly to write the romances in prose (many of the earlier verse romances were adapted into prose versions), although new verse romances continued to be written to the end of the 14th century. The most important romance of the 13th century is the Romance of the Rose , which breaks considerably from the conventions of the chivalric adventure story. Medieval French lyric poetry

7695-496: The time the studies were conducted, stocks from the middle and upper reaches of large river systems in British Columbia such as Fraser , Skeena , Nass and from mainland coastal streams in the southern half of B.C., "are more likely to have a low prevalence of infection." The report also states, "It should be stressed that Henneguya , economically deleterious though it is, is harmless from the view of public health . It

7790-467: The total nitrogen available to the riparian woodlands . The foliage of spruce trees up to 500 m (1,600 ft) from a stream where grizzlies fish salmon have been found to contain nitrogen originating from the fished salmon. Beavers also function as ecosystem engineers; in the process of tree-cutting and damming , beavers alter the local ecosystems extensively. Beaver ponds can provide critical habitat for juvenile salmon . An example of this

7885-434: The two. The Old Low Franconian influence is also believed to be responsible for the differences between the langue d'oïl and the langue d'oc (Occitan), being that various parts of Northern France remained bilingual between Latin and Germanic for some time, and these areas correspond precisely to where the first documents in Old French were written. This Germanic language shaped the popular Latin spoken here and gave it

7980-597: The use of wild-caught fish as feed for the salmon, is the use of soy -based products. This should be better for the local environment of the fish farm, but producing soy beans has a high environmental cost for the producing region. The fish omega-3 fatty acid content would be reduced compared to fish-fed salmon. Another possible alternative is a yeast-based coproduct of bioethanol production, proteinaceous fermentation biomass. Substituting such products for engineered feed can result in equal (sometimes enhanced) growth in fish. With its increasing availability, this would address

8075-412: The verb trobar "to find, to invent"). By the late 13th century, the poetic tradition in France had begun to develop in ways that differed significantly from the troubadour poets, both in content and in the use of certain fixed forms. The new poetic (as well as musical: some of the earliest medieval music has lyrics composed in Old French by the earliest composers known by name) tendencies are apparent in

8170-417: The water to attack terrestrial insects such as grasshoppers and dragonflies , as well as consuming fish eggs (even those of other salmon). As adults, salmon behave like other mid-sized pelagic fish , eating a variety of sea creatures including smaller forage fish such as lanternfish , herrings , sand lances , mackerels and barracudina . They also eat krill , squid and polychaete worms . In

8265-581: The world, with Norway being the world's largest producer of farmed salmon, followed by Chile . They are also highly prized game fish for recreational fishing , by both freshwater and saltwater anglers . Many species of salmon have since been introduced and naturalized into non-native environments such as the Great Lakes of North America , Patagonia in South America and South Island of New Zealand . The Modern English term salmon

8360-772: Was also spoken in the Kingdom of Sicily , and in the Principality of Antioch and the Kingdom of Jerusalem in the Levant . As part of the emerging Gallo-Romance dialect continuum, the langues d'oïl were contrasted with the langues d'oc , at the time also called "Provençal", adjacent to the Old French area in the southwest, and with the Gallo-Italic group to the southeast. The Franco-Provençal group developed in Upper Burgundy, sharing features with both French and Provençal; it may have begun to diverge from

8455-414: Was constantly changing and evolving; however, the form in the late 12th century, as attested in a great deal of mostly poetic writings, can be considered standard. The writing system at this time was more phonetic than that used in most subsequent centuries. In particular, all written consonants (including final ones) were pronounced, except for s preceding non- stop consonants and t in et , and final e

8550-680: Was indebted to the poetic and cultural traditions in Southern France and Provence —including Toulouse and the Aquitaine region—where langue d'oc was spoken ( Occitan language ); in their turn, the Provençal poets were greatly influenced by poetic traditions from the Hispano-Arab world . Lyric poets in Old French are called trouvères – etymologically the same word as the troubadours of Provençal or langue d'oc (from

8645-590: Was pronounced [ ə ] . The phonological system can be summarised as follows: Notes: In Old French, the nasal vowels were not separate phonemes but only allophones of the oral vowels before a nasal consonant. The nasal consonant was fully pronounced; bon was pronounced [bõn] ( ModF [bɔ̃] ). Nasal vowels were present even in open syllables before nasals where Modern French has oral vowels, as in bone [bõnə] ( ModF bonne [bɔn] ). Notes: Notes: In addition to diphthongs, Old French had many instances of hiatus between adjacent vowels because of

8740-644: Was seen in the years following 1818 in the Columbia River Basin. In 1818, the British government made an agreement with the U.S. government to allow U.S. citizens access to the Columbia catchment (see Treaty of 1818 ). At the time, the Hudson's Bay Company sent word to trappers to extirpate all furbearers from the area in an effort to make the area less attractive to U.S. fur traders. In response to

8835-518: Was spoken natively roughly extended to the northern half of the Kingdom of France and its vassals (including parts of the Angevin Empire ), and the duchies of Upper and Lower Lorraine to the east (corresponding to modern north-eastern France and Belgian Wallonia ), but the influence of Old French was much wider, as it was carried to England and the Crusader states as the language of

8930-423: Was the language spoken in most of the northern half of France approximately between the late 8th and the mid-14th century. Rather than a unified language , Old French was a group of Romance dialects , mutually intelligible yet diverse . These dialects came to be collectively known as the langues d'oïl , contrasting with the langues d'oc , the emerging Occitano-Romance languages of Occitania , now

9025-617: Was written by Latin-speaking clerics for a lay public). A large body of fables survive in Old French; these include (mostly anonymous) literature dealing with the recurring trickster character of Reynard the Fox . Marie de France was also active in this genre, producing the Ysopet (Little Aesop ) series of fables in verse. Related to the fable was the more bawdy fabliau , which covered topics such as cuckolding and corrupt clergy. These fabliaux would be an important source for Chaucer and for

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