89-445: See text Angophora is a genus of nine species of trees and shrubs in the myrtle family, Myrtaceae . Endemic to eastern Australia, they differ from other eucalypts in having juvenile and adult leaves arranged in opposite pairs, sepals reduced to projections on the edge of the floral cup , four or five overlapping, more or less round petals , and a papery or thin, woody, often strongly ribbed capsule . Species are found between
178-603: A supercontinent . The remnants of Gondwana make up around two-thirds of today's continental area, including South America , Africa , Antarctica , Australia , Zealandia , Arabia , and the Indian Subcontinent . Gondwana was formed by the accretion of several cratons (large stable blocks of the Earth's crust), beginning c. 800 to 650 Ma with the East African Orogeny ,
267-642: A complex series of orogenic events assembled the eastern parts of Gondwana (eastern Africa, Arabian-Nubian Shield, Seychelles, Madagascar, India, Sri Lanka, East Antarctica, and Australia) c. 750 to 530 Ma . First, the Arabian-Nubian Shield collided with eastern Africa (in the Kenya-Tanzania region) in the East African Orogeny c. 750 to 620 Ma . Then Australia and East Antarctica were merged with
356-599: A continuous arc chain, the direction of subduction was different between the Australian-Tasmanian and New Zealand-Antarctica arc segments. Many terranes were accreted to Eurasia during Gondwana's existence, but the Cambrian or Precambrian origin of many of these terranes remains uncertain. For example, some Palaeozoic terranes and microcontinents that now make up Central Asia, often called the "Kazakh" and "Mongolian terranes", were progressively amalgamated into
445-746: A diversified assemblage of true insects. In Gondwana, in contrast, ice and, in Australia, volcanism decimated the Devonian flora to a low-diversity seed fern flora – the pteridophytes were increasingly replaced by the gymnosperms which were to dominate until the Mid-Cretaceous. Australia, however, was still located near the Equator during the Early Carboniferous, and during this period, temnospondyl and lepospondyl amphibians and
534-472: A few million years, reached its peak at c. 200 Ma , and coincided with the Triassic–Jurassic extinction event . The reformed Gondwanan continent was not precisely the same as that which had existed before Pangaea formed; for example, most of Florida and southern Georgia and Alabama is underlain by rocks that were originally part of Gondwana, but this region stayed attached to North America when
623-594: A gradual decline during the Triassic while ferns, though never dominant, managed to diversify. The brief period of icehouse conditions during the Triassic–Jurassic extinction event had a dramatic impact on dinosaurs but left plants largely unaffected. The Jurassic was mostly one of hot-house conditions and, while vertebrates managed to diversify in this environment, plants have left little evidence of such development, apart from Cheiroleidiacean conifers and Caytoniales and other groups of seed ferns. In terms of biomass,
712-480: A much older supercontinent, Rodinia , to amalgamate. One of those orogenic belts, the Mozambique Belt , formed 800 to 650 Ma and was originally interpreted as the suture between East (India, Madagascar, Antarctica, and Australia) and West Gondwana (Africa and South America). Three orogenies were recognised during the 1990s as a result of data sets compiled on behalf of oil and mining companies:
801-518: A north-east motion about 90 million years ago. While subduction direction changed, it remained oblique (and not perpendicular) to the coast of South America, and the direction change affected several subduction zone -parallel faults including Atacama , Domeyko and Liquiñe-Ofqui . Insular India began to collide with Asia circa 70 Ma , forming the Indian subcontinent , since which more than 1,400 km (870 mi) of crust has been absorbed by
890-639: A number of species related to those of the laurissilva of Valdivia, through the connection of the Antarctic flora . These include gymnosperms and the deciduous species of Nothofagus , as well as the New Zealand laurel, Corynocarpus laevigatus , and Laurelia novae-zelandiae . New Caledonia and New Zealand became separated from Australia by continental drift 85 million years ago. The islands still retain plants that originated in Gondwana and spread to
979-544: A pattern that reflects the Jurassic break-up of Pangaea. The Cretaceous saw the arrival of the angiosperms , or flowering plants, a group that probably evolved in western Gondwana (South America–Africa). From there the angiosperms diversified in two stages: the monocots and magnoliids evolved in the Early Cretaceous, followed by the hammamelid dicots . By the Mid-Cretaceous, angiosperms constituted half of
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#17327910003661068-521: A peak in biodiversity – the end-Permian extinction was enormous and so was the radiation that followed. Two families of conifers, Podocarpaceae and Araucariaceae , dominated Gondwana in the Early Triassic, but Dicroidium , an extinct genus of fork-leaved seed ferns, dominated woodlands and forests of Gondwana during most of the Triassic. Conifers evolved and radiated during the period, with six of eight extant families already present before
1157-853: A series of events severally restricted the Proto-ACC: change to shallow marine conditions along the North Scotia Ridge; closure of the Fuegan Seaway, the deep sea that existed in Tierra del Fuego; and uplift of the Patagonian Cordillera. This, together with the reactivated Iceland plume , contributed to global warming. During the Miocene, the Drake Passage began to widen, and as water flow between South America and
1246-694: A single jaw from Australia. The closure of the Rheic Ocean and the formation of Pangaea in the Carboniferous resulted in the rerouting of ocean currents that initiated an Ice House period. As Gondwana began to rotate clockwise, Australia shifted south to more temperate latitudes. An ice cap initially covered most of southern Africa and South America but spread to eventually cover most of the supercontinent, save for northernmost Africa-South America and eastern Australia. Giant lycopod and horsetail forests continued to evolve in tropical Laurasia together with
1335-495: A toothed margin). The flowers have a base number of five petals, though in several genera, the petals are minute or absent. The stamens are usually very conspicuous, brightly coloured, and numerous. Scientists hypothesize that the family Myrtaceae arose between 60 and 56 million years ago (Mya) during the Paleocene era. Pollen fossils have been sourced to the ancient supercontinent Gondwana . The breakup of Gondwana during
1424-911: Is a list species of accepted by the Australian Plant Census as at March 2020: Angophoras are found in coastal Queensland, New South Wales and Victoria from the Atherton Tableland to eastern Victoria. Myrtaceae About 130; see list Myrtaceae ( / m ə r ˈ t eɪ s i ˌ aɪ , - s iː ˌ iː / ), the myrtle family , is a family of dicotyledonous plants placed within the order Myrtales . Myrtle , pōhutukawa , bay rum tree , clove , guava , acca (feijoa) , allspice , and eucalyptus are some notable members of this group. All species are woody, contain essential oils , and have flower parts in multiples of four or five. The leaves are evergreen , alternate to mostly opposite, simple, and usually entire (i.e., without
1513-442: Is a papery or thin, slightly woody, hairy capsule with longitudinal ribs. The genus Angophora was first formally described in 1797 by Antonio José Cavanilles in his book Icones et Descriptiones Plantarum . Angophora is closely related to Corymbia and Eucalyptus , and all three genera are often referred to as "eucalypts". Collectively the eucalypts dominate many Australian ecosystems . Taxonomists have long recognised
1602-818: Is now the Transantarctic Mountains ): the Antarctic Peninsula , Marie Byrd Land , Zealandia , and Thurston Island ; the Falkland Islands and Ellsworth–Whitmore Mountains (in Antarctica) were rotated 90° in opposite directions; and South America south of the Gastre Fault (often referred to as Patagonia ) was pushed westward. The history of the Africa-Antarctica break-up can be studied in great detail in
1691-576: Is now the southern Weddell Sea where initial break-up occurred during the Jurassic c. 180 to 160 Ma . Gondwana began to break up in the early Jurassic following the extensive and fast emplacement of the Karoo-Ferrar flood basalts c. 184 Ma . Before the Karoo plume initiated rifting between Africa and Antarctica , it separated a series of smaller continental blocks from Gondwana's southern, Proto-Pacific margin (along what
1780-541: Is presumed to have been linked to the subduction of cold oceanic lithosphere . During the mid to Late Cretaceous ( c. 90 million years ago ), the Andean orogeny changed significantly in character. Warmer and younger oceanic lithosphere is believed to have started to be subducted beneath South America around this time. Such kind of subduction is held responsible not only for the intense contractional deformation that different lithologies were subject to, but also
1869-787: The Antarctic Peninsula increased, the renewed ACC resulted in cooler global climate. Since the Eocene, the northward movement of the Australian Plate has resulted in an arc-continent collision with the Philippine and Caroline plates and the uplift of the New Guinea Highlands . From the Oligocene to the late Miocene, the climate in Australia, dominated by warm and humid rainforests before this collision, began to alternate between open forest and rainforest before
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#17327910003661958-886: The Antongil Block in far eastern Madagascar, the Seychelles , and the Napier and Rayner Complexes in East Antarctica ). The Azania continent (much of central Madagascar , the Horn of Africa and parts of Yemen and Arabia) was an island in the Mozambique Ocean. The continent Australia/ Mawson was still separated from India, eastern Africa, and Kalahari by c. 600 Ma , when most of western Gondwana had already been amalgamated. By c. 550 Ma, India had reached its Gondwanan position, which initiated
2047-538: The Atherton Tableland in Queensland and south through New South Wales to eastern Victoria, Australia. Plants in the genus Angophora are trees, occasionally shrubs, with rough bark except for A. costata . The juvenile leaves differ from adult leaves in being hairy with raised oil glands . Both juvenile and adult leaves are arranged in opposite pairs, the adult leaves usually glabrous and paler on
2136-1075: The Australian Plate are now separated by the Capricorn Plate and its diffuse boundaries. During the opening of the Indian Ocean, the Kerguelen hotspot first formed the Kerguelen Plateau on the Antarctic Plate c. 118 to 95 Ma and then the Ninety East Ridge on the Indian Plate at c. 100 Ma . The Kerguelen Plateau and the Broken Ridge , the southern end of the Ninety East Ridge, are now separated by
2225-844: The Campbell Plateau , Chatham Rise , Lord Howe Rise , Norfolk Ridge , and New Caledonia , from West Antarctica c. 84 Ma . The opening of the South Atlantic Ocean divided West Gondwana (South America and Africa), but there is considerable debate over the exact timing of this break-up. Rifting propagated from south to north along Triassic–Early Jurassic lineaments, but intra-continental rifts also began to develop within both continents in Jurassic–Cretaceous sedimentary basins, subdividing each continent into three sub-plates. Rifting began c. 190 Ma at Falkland latitudes, forcing Patagonia to move relative to
2314-550: The Central Atlantic opened . Antarctica, the centre of the supercontinent, shared boundaries with all other Gondwana continents and the fragmentation of Gondwana propagated clockwise around it. The break-up was the result of the eruption of the Karoo-Ferrar igneous province , one of the Earth's most extensive large igneous provinces (LIP) c. 200 to 170 Ma , but the oldest magnetic anomalies between South America, Africa, and Antarctica are found in what
2403-590: The Cretaceous period (145 to 66 Mya) geographically isolated disjunct taxa and allowed for rapid speciation; in particular, genera once considered members of the now-defunct Leptospermoideae alliance are now isolated within Oceania . Generally, experts agree that vicariance is responsible for the differentiation of Myrtaceae taxa, except in the cases of Leptospermum species now located on New Zealand and New Caledonia , islands which may have been submerged at
2492-834: The Drake Passage and the deepening of the Tasman Gateway. The oldest oceanic crust in the Drake Passage, however, is 34 to 29 Ma -old which indicates that the spreading between the Antarctic and South American plates began near the Eocene/Oligocene boundary. Deep sea environments in Tierra del Fuego and the North Scotia Ridge during the Eocene and Oligocene indicate a "Proto-ACC" opened during this period. Later, 26 to 14 Ma ,
2581-527: The East African Orogeny ( 650 to 800 Ma ) and Kuunga orogeny (including the Malagasy Orogeny in southern Madagascar) ( 550 Ma ), the collision between East Gondwana and East Africa in two steps, and the Brasiliano orogeny ( 660 to 530 Ma ), the successive collision between South American and African cratons . The last stages of Gondwanan assembly overlapped with
2670-757: The Great American Interchange . The break-up of Gondwana can be said to continue in eastern Africa at the Afar Triple Junction , which separates the Arabian , Nubian , and Somali plates, resulting in rifting in the Red Sea and East African Rift . In the Early Cenozoic , Australia was still connected to Antarctica c. 35–40° south of its current location and both continents were largely unglaciated. A rift between
2759-803: The Himalayan - Tibetan orogen. During the Cenozoic, the orogen resulted in the construction of the Tibetan Plateau between the Tethyan Himalayas in the south and the Kunlun and Qilian mountains in the north. Later, South America was connected to North America via the Isthmus of Panama , cutting off a circulation of warm water and thereby making the Arctic colder, as well as allowing
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2848-845: The Ordovician . This is the Cuyania or Precordillera terrane of the Famatinian orogeny in northwest Argentina which may have continued the line of the Appalachians southwards. Chilenia terrane accreted later against Cuyania. The collision of the Patagonian terrane with the southwestern Gondwanan occurred in the late Paleozoic. Subduction-related igneous rocks from beneath the North Patagonian Massif have been dated at 320–330 million years old, indicating that
2937-565: The Paleogene (from around 66 to 23 million years ago (Ma)). Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica , Laurentia , and Siberia were separated from it. To differentiate it from the Indian region of the same name (see § Name ), it is also commonly called Gondwanaland . Regions that were part of Gondwana shared floral and zoological elements that persist to
3026-481: The Pangaea supercontinent during the Carboniferous. Pangaea began to break up in the Mid-Jurassic when the Central Atlantic opened . In the western end of Pangaea, the collision between Gondwana and Laurasia closed the Rheic and Palaeo-Tethys oceans. The obliquity of this closure resulted in the docking of some northern terranes in the Marathon , Ouachita , Alleghanian , and Variscan orogenies, respectively. Southern terranes, such as Chortis and Oaxaca , on
3115-499: The Southeast Indian Ridge . Separation between Australia and East Antarctica began c. 132 Ma with seafloor spreading occurring c. 96 Ma . A shallow seaway developed over the South Tasman Rise during the Early Cenozoic and as oceanic crust started to separate the continents during the Eocene c. 35.5 Ma global ocean temperature dropped significantly. A dramatic shift from arc- to rift magmatism c. 100 Ma separated Zealandia , including New Zealand ,
3204-442: The Uralian orogeny and Laurasia . Pangaea was finally amalgamated in the Late Carboniferous-Early Permian, but the oblique forces continued until Pangaea began to rift in the Triassic. In the eastern end, collisions occurred slightly later. The North China , South China , and Indochina blocks rifted from Gondwana during the middle Paleozoic and opened the Proto-Tethys Ocean . North China docked with Mongolia and Siberia during
3293-422: The Variscan orogeny close to the Carboniferous–Permian boundary. South-east Asia was made of Gondwanan and Cathaysian continental fragments that were assembled during the Mid-Palaeozoic and Cenozoic. This process can be divided into three phases of rifting along Gondwana's northern margin: first, in the Devonian, North and South China , together with Tarim and Quidam (north-western China) rifted, opening
3382-403: The uplift and erosion known to have occurred from the Late Cretaceous onward. Plate tectonic reorganisation since the mid-Cretaceous might also have been linked to the opening of the South Atlantic Ocean . Another change related to mid-Cretaceous plate tectonic rearrangement was the change of subduction direction of the oceanic lithosphere that went from having south-east motion to having
3471-415: The Americas, apart from Metrosideros in Chile and Argentina . Genera with fleshy fruits have their greatest concentrations in eastern Australia and Malesia (the Australasian realm ) and the Neotropics . Eucalyptus is a dominant, nearly ubiquitous genus in the more mesic parts of Australia and extends north sporadically to the Philippines . Eucalyptus regnans is the tallest flowering plant in
3560-502: The Carboniferous–Permian, followed by South China. The Cimmerian blocks then rifted from Gondwana to form the Palaeo-Thethys and Neo-Tethys oceans in the Late Carboniferous, and docked with Asia during the Triassic and Jurassic. Western Pangaea began to rift while the eastern end was still being assembled. The formation of Pangaea and its mountains had a tremendous impact on global climate and sea levels, which resulted in glaciations and continent-wide sedimentation. In North America,
3649-405: The Early Cretaceous, and West Burma and Woyla during the Late Cretaceous. Gondwana's long, northern margin remained a mostly passive margin throughout the Palaeozoic. The Early Permian opening of the Neo-Tethys Ocean along this margin produced a long series of terranes, many of which were and still are being deformed in the Himalaya Orogeny . These terranes are, from Turkey to north-eastern India:
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3738-403: The Earth's surface. It fused with Euramerica during the Carboniferous to form Pangea . It began to separate from northern Pangea ( Laurasia ) during the Triassic , and started to fragment during the Early Jurassic (around 180 million years ago). The final stages of break-up, involving the separation of Antarctica from South America (forming the Drake Passage ) and Australia, occurred during
3827-448: The Equator during this period and remained a lifeless and barren landscape. West Gondwana drifted north during the Devonian , bringing Gondwana and Laurasia close together. Global cooling contributed to the Late Devonian extinction (19% of marine families and 50% of genera went extinct) and glaciation occurred in South America. Before Pangaea had formed, terrestrial plants, such as pteridophytes , began to diversify rapidly resulting in
3916-400: The Equator on landmasses then limited to Laurasia and, in Gondwana, to Australia. In the late Silurian, two distinctive lineages, zosterophylls and rhyniophytes , had colonised the tropics. The former evolved into the lycopods that were to dominate the Gondwanan vegetation over a long period, whilst the latter evolved into horsetails and gymnosperms . Most of Gondwana was located far from
4005-446: The Jurassic flora was dominated by conifer families and other gymnosperms that had evolved during the Triassic. The Pteridophytes that had dominated during the Palaeozoic were now marginalised, except for ferns. In contrast to Laurentia, very few insect fossils have been found in Gondwana, to a considerable extent because of widespread deserts and volcanism. While plants had a cosmopolitan distribution, dinosaurs evolved and diversified in
4094-406: The Kuunga orogeny (also known as the Pinjarra orogeny). Meanwhile, on the other side of the newly forming Africa, Kalahari collided with Congo and Rio de la Plata which closed the Adamastor Ocean . c. 540–530 Ma, the closure of the Mozambique Ocean brought India next to Australia–East Antarctica, and both North and South China were in proximity to Australia. As the rest of Gondwana formed,
4183-422: The Late Permian, many known from South Africa and Australia. Beetles and cockroaches remained minor elements in this fauna. Tetrapod fossils from the Early Permian have only been found in Laurasia but they became common in Gondwana later during the Permian. The arrival of the therapsids resulted in the first plant-vertebrate-insect ecosystem. During the Mid- to Late Triassic, hot-house conditions coincided with
4272-405: The Miocene, a warm and humid climate developed with pockets of rainforests in central Australia, but before the end of the period, colder and drier climate severely reduced this rainforest. A brief period of increased rainfall in the Pliocene was followed by drier climate which favoured grassland. Since then, the fluctuation between wet interglacial periods and dry glacial periods has developed into
4361-443: The Neoproterozoic to Palaeozoic phase of the Terra Australis Orogen , a series of terranes were rafted from the proto-Andean margin when the Iapteus Ocean opened, to be added back to Gondwana during the closure of that ocean. During the Paleozoic, some blocks which helped to form parts of the Southern Cone of South America, include a piece transferred from Laurentia when the west edge of Gondwana scraped against southeast Laurentia in
4450-441: The Palaeo-Tethys behind them. These terranes accreted to Asia during Late Devonian and Permian. Second, in the Late Carboniferous to Early Permian, Cimmerian terranes opened Meso-Tethys Ocean; Sibumasu and Qiangtang were added to south-east Asia during Late Permian and Early Jurassic. Third, in the Late Triassic to Late Jurassic, Lhasa , West Burma , Woyla terranes opened the Neo-Tethys Ocean; Lhasa collided with Asia during
4539-456: The South Atlantic (Brazil and Cameroon ) dating to around 120 million years ago , suggesting that some form of land connection still existed between Africa and South America as recently as the early Aptian . The first phases of Andean orogeny in the Jurassic and Early Cretaceous were characterised by extensional tectonics , rifting , the development of back-arc basins and the emplacement of large batholiths . This development
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#17327910003664628-465: The Southern Hemisphere, has a "Gondwanan distribution" and is often described as an archaic, or relict , lineage. The distributions in the Proteaceae is, nevertheless, the result of both Gondwanan rafting and later oceanic dispersal. During the Silurian, Gondwana extended from the Equator (Australia) to the South Pole (North Africa and South America) whilst Laurasia was located on the Equator opposite to Australia. A short-lived Late Ordovician glaciation
4717-408: The Taurides in southern Turkey; the Lesser Caucasus Terrane in Georgia; the Sanand, Alborz, and Lut terranes in Iran; the Mangysglak or Kopetdag Terrane in the Caspian Sea; the Afghan Terrane; the Karakorum Terrane in northern Pakistan; and the Lhasa and Qiangtang terranes in Tibet. The Permian–Triassic widening of the Neo-Tethys pushed all these terranes across the Equator and over to Eurasia. During
4806-575: The baccate (fleshy) fruits evolved twice from capsular fruits and, as such, the two-subfamily classification does not accurately portray the phylogenetic history of the family. Thus, many workers are now using a recent analysis by Wilson et al. (2001) as a starting point for further analyses of the family. This study pronounced both Leptospermoideae and Myrtoideae invalid, but retained several smaller suballiances shown to be monophyletic through matK analysis. The genera Heteropyxis and Psiloxylon have been separated as separate families by many authors in
4895-550: The base of the Absaroka sequence coincides with the Alleghanian and Ouachita orogenies and are indicative of a large-scale change in the mode of deposition far away from the Pangaean orogenies. Ultimately, these changes contributed to the Permian–Triassic extinction event and left large deposits of hydrocarbons, coal, evaporite, and metals. The breakup of Pangaea began with the Central Atlantic magmatic province (CAMP) between South America, Africa, North America, and Europe. CAMP covered more than seven million square kilometres over
4984-467: The collision of India and Madagascar with East Africa, and culminating in c. 600 to 530 Ma with the overlapping Brasiliano and Kuunga orogenies, the collision of South America with Africa, and the addition of Australia and Antarctica, respectively. Eventually, Gondwana became the largest piece of continental crust of the Palaeozoic Era, covering an area of some 100,000,000 km (39,000,000 sq mi), about one-fifth of
5073-414: The colonisation of Gondwana. The Baragwanathia Flora, found only in the Yea Beds of Victoria, Australia, occurs in two strata separated by 1,700 m (5,600 ft) or 30 Ma; the upper assemblage is more diverse and includes Baragwanathia, the first primitive herbaceous lycopod to evolve from the zosterophylls. During the Devonian, giant club mosses replaced the Baragwanathia Flora, introducing
5162-492: The continent Kazakhstania in the late Silurian . Whether these blocks originated on the shores of Gondwana is not known. In the Early Palaeozoic, the Armorican terrane , which today form large parts of France, was part of either Peri-Gondwana or core Gondwana; the Rheic Ocean closed in front of it and the Palaeo-Tethys Ocean opened behind it. Precambrian rocks from the Iberian Peninsula suggest that it, too, formed part of core Gondwana before its detachment as an orocline in
5251-424: The continent became the arid or semiarid landscape it is today. The adjective "Gondwanan" is in common use in biogeography when referring to patterns of distribution of living organisms, typically when the organisms are restricted to two or more of the now-discontinuous regions that were once part of Gondwana, including the Antarctic flora . For example, the plant family Proteaceae , known from all continents in
5340-559: The development of rifts systems on both continents, including the Central African Rift System and the Central African Shear Zone which lasted until c. 85 Ma . At Brazilian latitudes spreading is more difficult to assess because of the lack of palaeo-magnetic data, but rifting occurred in Nigeria at the Benue Trough c. 118 Ma . North of the Equator the rifting began after 120.4 Ma and continued until c. 100 to 96 Ma . Dinosaur footprints representing identical species assemblages are known from opposite sides of
5429-427: The eastern areas of Australia , but could occur in other neighbouring countries where some Corymbia species are native. Weevils in the tribe Cryptoplini mostly use Myrtaceae as hosts. Their larvae can develop in flower and fruit buds, or in galls (often galls already formed by other insects). Gondwana Gondwana ( / ɡ ɒ n d ˈ w ɑː n ə / ) was a large landmass, sometimes referred to as
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#17327910003665518-437: The end of it. Bennettitales and Pentoxylales , two now extinct orders of gymnospermous plants, evolved in the Late Triassic and became important in the Jurassic and Cretaceous. It is possible that gymnosperm biodiversity surpassed later angiosperm biodiversity and that the evolution of angiosperms began during the Triassic but, if so, in Laurasia rather than in Gondwana. Two Gondwanan classes, lycophytes and sphenophytes , saw
5607-650: The eruption of the Deccan basalts , whose eruption site may survive as the Réunion hotspot . The Seychelles and the Maldives are now separated by the Central Indian Ridge . During the initial break-up in the Early Jurassic a marine transgression swept over the Horn of Africa covering Triassic planation surfaces with sandstone , limestone , shale , marls and evaporites . East Gondwana, comprising Antarctica, Madagascar, India, and Australia, began to separate from Africa. East Gondwana then began to break up c. 132.5 to 96 Ma when India moved northwest from Australia-Antarctica. The Indian Plate and
5696-439: The evolution of Voltziales , one of the few plant orders to survive the end-Permian extinction (57% of marine families and 83% of genera went extinct) and which came to dominate in the Late Permian and from whom true conifers evolved. Tall lycopods and horsetails dominated the wetlands of Gondwana in the Early Permian. Insects co-evolved with glossopterids across Gondwana and diversified with more than 200 species in 21 orders by
5785-452: The first amniote reptilians evolved, all closely related to the Laurasian fauna, but spreading ice eventually drove these animals away from Gondwana entirely. The Gondwana ice sheet melted, and sea levels dropped during the Permian and Triassic global warming. During this period, the extinct glossopterids colonised Gondwana and reached peak diversity in the Late Permian when coal-forming forests covered much of Gondwana. The period also saw
5874-402: The first trees, and by the Late Devonian this first forest was accompanied by the progymnosperms , including the first large trees Archaeopteris . The Late Devonian extinction probably also resulted in osteolepiform fishes evolving into the amphibian tetrapods , the earliest land vertebrates, in Greenland and Russia. The only traces of this evolution in Gondwana are amphibian footprints and
5963-459: The flora in northeastern Australia. There is, however, no obvious connection between this spectacular angiosperm radiation and any known extinction event nor with vertebrate/insect evolution. Insect orders associated with pollination, such as beetles , flies , butterflies and moths , and wasps, bees, and ants , radiated continuously from the Permian-Triassic, long before the arrival of the angiosperms. Well-preserved insect fossils have been found in
6052-433: The fracture zones and magnetic anomalies flanking the Southwest Indian Ridge . The Madagascar block and the Mascarene Plateau , stretching from the Seychelles to Réunion , were broken off India, causing Madagascar and Insular India to be separate landmasses : elements of this break-up nearly coincide with the Cretaceous–Paleogene extinction event . The India–Madagascar–Seychelles separations appear to coincide with
6141-571: The lake deposits of the Santana Formation in Brazil, the Koonwarra Lake fauna in Australia, and the Orapa diamond mine in Botswana. Dinosaurs continued to prosper but, as the angiosperm diversified, conifers, bennettitaleans and pentoxylaleans disappeared from Gondwana c. 115 Ma together with the specialised herbivorous ornithischians , whilst generalist browsers, such as several families of sauropodomorph Saurischia , prevailed. The Cretaceous–Paleogene extinction event killed off all dinosaurs except birds, but plant evolution in Gondwana
6230-413: The lower surface. The flower buds are arranged in groups of three or seven. The flower has four or five small sepals, reduced to small projections on the rim of the floral cup. There are four or five more or less round, keeled, overlapping petals and whorls of many creamy white stamens . Unlike in Eucalyptus and Corymbia , the petals and sepals are not fused to form a cap-like operculum . The fruit
6319-422: The mature seed pods of Corymbia torelliana , resulting in mellitochory as the seeds get stuck onto the corbiculae of the bees and sometimes are successfully disposed of by colony members that remove them. But usually, they get stuck in the hives or near hive entrances instead, hence also making it a minor nuisance for some keepers as they can take up a lot of space. Fortunately, this is only known to occur in
6408-701: The opening of the Iapetus Ocean between Laurentia and western Gondwana. During this interval, the Cambrian explosion occurred. Laurentia was docked against the western shores of a united Gondwana for a brief period near the Precambrian/Cambrian boundary, forming the short-lived and still disputed supercontinent Pannotia . The Mozambique Ocean separated the Congo – Tanzania – Bangweulu Block of central Africa from Neoproterozoic India (India,
6497-600: The other hand, remained largely unaffected by the collision along the southern shores of Laurentia. Some Peri-Gondwanan terranes, such as Yucatán and Florida , were buffered from collisions by major promontories. Other terranes, such as Carolina and Meguma , were directly involved in the collision. The final collision resulted in the Variscan- Appalachian Mountains , stretching from present-day Mexico to southern Europe. Meanwhile, Baltica collided with Siberia and Kazakhstania which resulted in
6586-486: The past as Heteropyxidaceae and Psiloxylaceae. However, Wilson et al. included them in Myrtaceae. These two genera are presently believed to be the earliest arising and surviving lineages of Myrtaceae. The most recent classification recognizes 17 tribes and two subfamilies, Myrtoideae and Psiloxyloideae, based on a phylogenetic analysis of plastid DNA. Many new species are being described annually from throughout
6675-561: The present arid regime. Australia has thus experienced various climate changes over a 15-million-year period with a gradual decrease in precipitation. The Tasman Gateway between Australia and Antarctica began to open c. 40 to 30 Ma . Palaeontological evidence indicates the Antarctic Circumpolar Current (ACC) was established in the Late Oligocene c. 23 Ma with the full opening of
6764-530: The present day. The continent of Gondwana was named by the Austrian scientist Eduard Suess , after the region in central India of the same name , which is derived from Sanskrit for "forest of the Gonds ". The name had been previously used in a geological context, first by H. B. Medlicott in 1872, from which the Gondwana sedimentary sequences ( Permian - Triassic ) are also described. Some scientists prefer
6853-461: The range of Myrtaceae. Likewise, new genera are being described nearly yearly. Following Wilson (2011) Subfamily Psiloxyloideae Subfamily Myrtoideae 127 genera are currently accepted: Myrtaceae is foraged by many stingless bees, especially by species such as Melipona bicolor which gather pollen from this plant family. Some Australian species such as Tetragonula hockingsi and T. carbonaria are also known to collect resin from
6942-576: The relationships between the eucalypt taxa, but have not agreed upon a classification scheme. Some have proposed merging Angophora and Corymbia into genus Eucalyptus as subgenera , a plan which was immediately rejected by others. Some authors maintain Angophora as a genus, while others continue to debate the issue. Among the eucalypts, Angophora species were nicknamed "apples" by European settlers, who thought they resembled apple trees . Many are still known commonly as apples today. The following
7031-760: The remaining Gondwana c. 570 to 530 Ma in the Kuunga Orogeny. The later Malagasy orogeny at about 550–515 Mya affected Madagascar, eastern East Africa and southern India. In it, Neoproterozoic India collided with the already combined Azania and Congo–Tanzania–Bangweulu Block, suturing along the Mozambique Belt. The 18,000 km-long (11,000 mi) Terra Australis Orogen developed along Gondwana's western, southern, and eastern margins. Proto-Gondwanan Cambrian arc belts from this margin have been found in eastern Australia, Tasmania, New Zealand, and Antarctica. Though these belts formed
7120-665: The still static remainder of South America and Africa, and this westward movement lasted until the Early Cretaceous 126.7 Ma . From there rifting propagated northward during the Late Jurassic c. 150 Ma or Early Cretaceous c. 140 Ma most likely forcing dextral movements between sub-plates on either side. South of the Walvis Ridge and Rio Grande Rise the Paraná and Etendeka magmatics resulted in further ocean-floor spreading c. 130 to 135 Ma and
7209-399: The subduction process initiated in the early Carboniferous. This was relatively short-lived (lasting about 20 million years), and initial contact of the two landmasses occurred in the mid-Carboniferous, with broader collision during the early Permian. In the Devonian, an island arc named Chaitenia accreted to Patagonia in what is now south-central Chile. Gondwana and Laurasia formed
7298-525: The term "Gondwanaland" for the supercontinent to make a clear distinction between the region and the supercontinent. The assembly of Gondwana was a protracted process during the Neoproterozoic and Paleozoic , which remains incompletely understood because of the lack of paleo-magnetic data. Several orogenies , collectively known as the Pan-African orogeny , caused the continental fragments of
7387-420: The time of late Eocene differentiation. Recent estimates suggest the Myrtaceae include about 5,950 species in about 132 genera. The family has a wide distribution in tropical and warm-temperate regions of the world, and is common in many of the world's biodiversity hotspots . Genera with capsular fruits such as Eucalyptus , Corymbia , Angophora , Leptospermum , and Melaleuca are absent from
7476-636: The two developed but remained an embayment until the Eocene-Oligocene boundary when the Circumpolar Current developed and the glaciation of Antarctica began. Australia was warm and wet during the Palaeocene and dominated by rainforest. The opening of the Tasman Gateway at the Eocene-Oligocene boundary ( 33 Ma ) resulted in abrupt cooling but the Oligocene became a period of high rainfall with swamps in southeast Australia. During
7565-400: The world. Other important Australian genera are Callistemon (bottlebrushes), Syzygium , and Melaleuca (paperbarks). Species of the genus Osbornia , native to Australasia, are mangroves . Eugenia , Myrcia , and Calyptranthes are among the larger genera in the neotropics. Historically, the Myrtaceae were divided into two subfamilies. Subfamily Myrtoideae (about 75 genera)
7654-483: Was followed by a Silurian Hot House period. The End-Ordovician extinction , which resulted in 27% of marine invertebrate families and 57% of genera going extinct, occurred during this shift from Ice House to Hot House. By the end of the Ordovician, Cooksonia , a slender, ground-covering plant, became the first known vascular plant to establish itself on land. This first colonisation occurred exclusively around
7743-540: Was hardly affected. Gondwanatheria is an extinct group of non- therian mammals with a Gondwanan distribution (South America, Africa, Madagascar, India, Zealandia and Antarctica) during the Late Cretaceous and Palaeogene. Xenarthra and Afrotheria , two placental clades, are of Gondwanan origin and probably began to evolve separately c. 105 Ma when Africa and South America separated. The laurel forests of Australia, New Caledonia, and New Zealand have
7832-743: Was recognized as having dry, dehiscent fruits (capsules) and leaves arranged spirally or alternate. The Leptospermoideae are found mostly in Australasia, with a centre of diversity in Australia. Many genera in Western Australia have greatly reduced leaves and flowers typical of more xeric habitats. The division of the Myrtaceae into Leptospermoideae and Myrtoideae was challenged by a number of authors, including Johnson and Briggs (1984), who identified 14 tribes or clades within Myrtaceae, and found Myrtoideae to be polyphyletic. Molecular studies by several groups of authors, as of 2008, have confirmed
7921-513: Was recognized as having fleshy fruits and opposite, entire leaves. Most genera in this subfamily have one of three easily recognized types of embryos. The genera of Myrtoideae can be very difficult to distinguish in the absence of mature fruits. Myrtoideae are found worldwide in subtropical and tropical regions, with centers of diversity in the Neotropics, northeastern Australia, and Malesia. In contrast, subfamily Leptospermoideae (about 80 genera)
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