162-480: Araripesuchus is a genus of extinct crocodyliform that existed during the Cretaceous period of the late Mesozoic era some 125 to 66 million years ago. Araripesuchus is generally considered to be a notosuchian (belonging to the clade Mesoeucrocodylia ), characterized by the varied teeth types and distinct skull elements. Seven species have been referred to Araripesuchus , though it has been argued that
324-585: A Zanzibari traveler, refers to the island as "Malagaskar" when recounting his journeys, including as part of the Emin Pasha Relief Expedition . In 1905, Charles Basset wrote in his doctoral thesis that Malagasikara was the way the island is referred to by its natives, who emphasized that they were Malagasy , and not Madagasy . Traditionally, archaeologists have estimated that the earliest settlers arrived in successive waves in outrigger canoes from South Borneo , possibly throughout
486-438: A food chain who lose their prey. "Species coextinction is a manifestation of one of the interconnectednesses of organisms in complex ecosystems ... While coextinction may not be the most important cause of species extinctions, it is certainly an insidious one." Coextinction is especially common when a keystone species goes extinct. Models suggest that coextinction is the most common form of biodiversity loss . There may be
648-669: A nautilus to the Royal Society that was more than two feet in diameter, and morphologically distinct from any known living species. Hooke theorized that this was simply because the species lived in the deep ocean and no one had discovered them yet. While he contended that it was possible a species could be "lost", he thought this highly unlikely. Similarly, in 1695, Sir Thomas Molyneux published an account of enormous antlers found in Ireland that did not belong to any extant taxa in that area. Molyneux reasoned that they came from
810-760: A protracted transition towards its fourth and current republic, with constitutional governance being restored in January 2014. Madagascar is a member of the United Nations (UN), the African Union (AU), the Southern African Development Community (SADC), and the Organisation Internationale de la Francophonie . Malagasy and French are both official languages of the state. Christianity
972-415: A species or a population is the variety of genetic information in its living members. A large gene pool (extensive genetic diversity ) is associated with robust populations that can survive bouts of intense selection . Meanwhile, low genetic diversity (see inbreeding and population bottlenecks ) reduces the range of adaptions possible. Replacing native with alien genes narrows genetic diversity within
1134-492: A strongman . Mid-2021 marked the beginning of the 2021–2022 Madagascar famine which, due to a severe drought, caused hundreds of thousands of people to face food insecurity and over one million people were on the verge of a famine . In November 2023, Rajoelina was re-elected to another term with 58.95% of the vote in the first round of the election amidst an opposition boycott and a controversy about his acquisition of French citizenship and subsequent eligibility. Turnout
1296-436: A viable population for species preservation and possible future reintroduction to the wild, through use of carefully planned breeding programs . The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction". This is especially common with extinction of keystone species . A 2018 study indicated that the sixth mass extinction started in
1458-479: A Lazarus species when extant individuals were described in 2019. Attenborough's long-beaked echidna ( Zaglossus attenboroughi ) is an example of a Lazarus species from Papua New Guinea that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023. Some species currently thought to be extinct have had continued speculation that they may still exist, and in
1620-503: A cascade of coextinction across the trophic levels . Such effects are most severe in mutualistic and parasitic relationships. An example of coextinction is the Haast's eagle and the moa : the Haast's eagle was a predator that became extinct because its food source became extinct. The moa were several species of flightless birds that were a food source for the Haast's eagle. Extinction as
1782-453: A century of warring and famine, Imerina was reunited in 1793 by King Andrianampoinimerina (1787–1810). From his initial capital Ambohimanga , and later from the Rova of Antananarivo , this Merina king rapidly expanded his rule over neighbouring principalities. His ambition to bring the entire island under his control was largely achieved by his son and successor, King Radama I (1810–28), who
SECTION 10
#17327917584251944-429: A common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups. A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at
2106-399: A fact that was accepted by most scientists. The primary debate focused on whether this turnover caused by extinction was gradual or abrupt in nature. Cuvier understood extinction to be the result of cataclysmic events that wipe out huge numbers of species, as opposed to the gradual decline of a species over time. His catastrophic view of the nature of extinction garnered him many opponents in
2268-564: A few, fragmentary jaw elements, reside at the Museum National d'Histoire Naturelle in Paris . The holotype's fragmentary nature meant that its placement in the genus was disputed until more remains were found in 2009 by Sereno and Larsson; these, along with the specimens of A. tsangatsangana, confirmed its place. Araripesuchus patagonicus was described from a patagonian specimen (MUC-PV 269) in 2000. Another species to be assigned to
2430-708: A higher risk of extinction and die out faster than less sexually dimorphic species, the least sexually dimorphic species surviving for millions of years while the most sexually dimorphic species die out within mere thousands of years. Earlier studies based on counting the number of currently living species in modern taxa have shown a higher number of species in more sexually dimorphic taxa which have been interpreted as higher survival in taxa with more sexual selection, but such studies of modern species only measure indirect effects of extinction and are subject to error sources such as dying and doomed taxa speciating more due to splitting of habitat ranges into more small isolated groups during
2592-405: A large range, a lack of individuals of both sexes (in sexually reproducing species), or other reasons. Pinpointing the extinction (or pseudoextinction ) of a species requires a clear definition of that species . If it is to be declared extinct, the species in question must be uniquely distinguishable from any ancestor or daughter species, and from any other closely related species. Extinction of
2754-607: A later point. The coelacanth , a fish related to lungfish and tetrapods , is an example of a Lazarus taxon that was known only from the fossil record and was considered to have been extinct since the end of the Cretaceous Period . In 1938, however, a living specimen was found off the Chalumna River (now Tyolomnqa) on the east coast of South Africa. Calliostoma bullatum , a species of deepwater sea snail originally described from fossils in 1844 proved to be
2916-583: A lower jaw. A more complete specimen, AMNH 24450 is held by the American Museum of Natural History . A second species, A. wegeneri was described in 1981. This species was discovered from Early Cretaceous deposits of Niger on the African continent, as opposed to the South American paleodistribution of the other species in the genus. The type specimen for the species, GDF-700 consisting of
3078-594: A mathematical model that falls in all positions. By contrast, conservation biology uses the extinction vortex model to classify extinctions by cause. When concerns about human extinction have been raised, for example in Sir Martin Rees ' 2003 book Our Final Hour , those concerns lie with the effects of climate change or technological disaster. Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago. Currently, environmental groups and some governments are concerned with
3240-527: A multi-party democracy and a separation of powers that placed significant control in the hands of the National Assembly. The new constitution also emphasized human rights, social and political freedoms, and free trade. Zafy's term, however, was marred by economic decline, allegations of corruption, and his introduction of legislation to give himself greater powers. He was consequently impeached in 1996, and an interim president, Norbert Ratsirahonana ,
3402-456: A natural part of the evolutionary process. Only recently have extinctions been recorded and scientists have become alarmed at the current high rate of extinctions . Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during
SECTION 20
#17327917584253564-441: A new mega-predator or by transporting animals and plants from one part of the world to another. Such introductions have been occurring for thousands of years, sometimes intentionally (e.g. livestock released by sailors on islands as a future source of food) and sometimes accidentally (e.g. rats escaping from boats). In most cases, the introductions are unsuccessful, but when an invasive alien species does become established,
3726-610: A population a higher chance in the short term of surviving an adverse change in conditions. Effects that cause or reward a loss in genetic diversity can increase the chances of extinction of a species. Population bottlenecks can dramatically reduce genetic diversity by severely limiting the number of reproducing individuals and make inbreeding more frequent. Extinction sometimes results for species evolved to specific ecologies that are subjected to genetic pollution —i.e., uncontrolled hybridization , introgression and genetic swamping that lead to homogenization or out-competition from
3888-407: A race of animals to become extinct. A series of fossils were discovered in the late 17th century that appeared unlike any living species. As a result, the scientific community embarked on a voyage of creative rationalization, seeking to understand what had happened to these species within a framework that did not account for total extinction. In October 1686, Robert Hooke presented an impression of
4050-502: A reduction in agricultural productivity. Furthermore, increased erosion contributes to poorer water quality by elevating the levels of sediment and pollutants in rivers and streams. Habitat degradation through toxicity can kill off a species very rapidly, by killing all living members through contamination or sterilizing them. It can also occur over longer periods at lower toxicity levels by affecting life span, reproductive capacity, or competitiveness. Habitat degradation can also take
4212-499: A result of climate change has been confirmed by fossil studies. Particularly, the extinction of amphibians during the Carboniferous Rainforest Collapse , 305 million years ago. A 2003 review across 14 biodiversity research centers predicted that, because of climate change, 15–37% of land species would be "committed to extinction" by 2050. The ecologically rich areas that would potentially suffer
4374-606: A result of the tangena ordeal, constituting roughly 20 percent of the population. The combination of regular warfare, disease, difficult forced labor, and harsh measures of justice resulted in a high mortality rate among soldiers and civilians alike during her 33-year reign; the population of Madagascar is estimated to have declined from around 5 million to 2.5 million between 1833 and 1839. Among those who continued to reside in Imerina were Jean Laborde , an entrepreneur who developed munitions and other industries on behalf of
4536-530: A sister taxon of Anatosuchus , questioning the monophyly of the genus. Lumping all species into one genus would lead to Uruguaysuchus taking priority, rendering Araripesuchus a junior synonym of Uruguaysuchus . Araripesuchus remains have been recovered from the continents of South America and Africa suggesting a Gondwanan origin for the evolution of the genus. At around the time of Araripesuchus' existence, South America and Africa were physically adjacent to each other. The various species evolved from
4698-479: A species (or replacement by a daughter species) plays a key role in the punctuated equilibrium hypothesis of Stephen Jay Gould and Niles Eldredge . In ecology , extinction is sometimes used informally to refer to local extinction , in which a species ceases to exist in the chosen area of study, despite still existing elsewhere. Local extinctions may be made good by the reintroduction of individuals of that species taken from other locations; wolf reintroduction
4860-439: A species may come suddenly when an otherwise healthy species is wiped out completely, as when toxic pollution renders its entire habitat unliveable; or may occur gradually over thousands or millions of years, such as when a species gradually loses out in competition for food to better adapted competitors. Extinction may occur a long time after the events that set it in motion, a phenomenon known as extinction debt . Assessing
5022-404: A species or group of species. "Just as each species is unique", write Beverly and Stephen C. Stearns , "so is each extinction ... the causes for each are varied—some subtle and complex, others obvious and simple". Most simply, any species that cannot survive and reproduce in its environment and cannot move to a new environment where it can do so, dies out and becomes extinct. Extinction of
Araripesuchus - Misplaced Pages Continue
5184-518: A subsequent report, IPBES listed unsustainable fishing, hunting and logging as being some of the primary drivers of the global extinction crisis. In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750, but likely many more. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming. A dagger symbol (†) placed next to
5346-576: A variety of afflictions. The drugs vinblastine and vincristine are vinca alkaloids , used to treat Hodgkin lymphoma , leukemia , and other cancers, were derived from the Madagascar periwinkle . The traveler's palm , known locally as ravinala and endemic to the eastern rain forests, is highly iconic of Madagascar and is featured in the national emblem as well as the Air Madagascar logo. Like its flora, Madagascar's fauna
5508-442: Is a biodiversity hotspot and one of the world's 17 megadiverse countries , with over 90% of wildlife being endemic. The island has a subtropical to tropical maritime climate. Madagascar was first permanently settled during or before the mid-first millennium AD by Austronesian peoples , presumably arriving on outrigger canoes from present-day Indonesia . These were joined around the ninth century AD by Bantu groups crossing
5670-465: Is also evidence to suggest that this event was preceded by another mass extinction, known as Olson's Extinction . The Cretaceous–Paleogene extinction event (K–Pg) occurred 66 million years ago, at the end of the Cretaceous period; it is best known for having wiped out non-avian dinosaurs , among many other species. According to a 1998 survey of 400 biologists conducted by New York 's American Museum of Natural History , nearly 70% believed that
5832-527: Is also historically attested. A British state paper in 1699 records the arrival of eighty to ninety passengers from "Malagaskar" to what would eventually become New York City . An 1882 edition of the British newspaper The Graphic referred to "Malagascar" as the name of the island, stating that it is etymologically a word of Malay origin, and may be related to the name of Malacca . In 1891, Saleh bin Osman,
5994-444: Is an example of this. Species that are not globally extinct are termed extant . Those species that are extant, yet are threatened with extinction, are referred to as threatened or endangered species . Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the conservation status "extinct in the wild" (EW) . Species listed under this status by
6156-585: Is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct Hyracotherium , which was an early horse that shares a common ancestor with the modern horse , is pseudoextinct, rather than extinct, because there are several extant species of Equus , including zebra and donkey ; however, as fossil species typically leave no genetic material behind, one cannot say whether Hyracotherium evolved into more modern horse species or merely evolved from
6318-621: Is diverse and exhibits a high rate of endemism. Lemurs have been characterized as "Madagascar's flagship mammal species" by Conservation International. In the absence of monkeys and other competitors, these primates have adapted to a wide range of habitats and diversified into numerous species. As of 2012 , there were officially 103 species and subspecies of lemur , 39 of which were described by zoologists between 2000 and 2008. They are almost all classified as rare, vulnerable, or endangered. At least 17 species of lemur have become extinct since humans arrived on Madagascar, all of which were larger than
6480-417: Is estimated as 100 to 1,000 times "background" rates (the average extinction rates in the evolutionary time scale of planet Earth), faster than at any other time in human history, while future rates are likely 10,000 times higher. However, some groups are going extinct much faster. Biologists Paul R. Ehrlich and Stuart Pimm , among others, contend that human population growth and overconsumption are
6642-414: Is estimated that there are currently around 8.7 million species of eukaryotes globally, and possibly many times more if microorganisms , like bacteria , are included. Notable extinct animal species include non-avian dinosaurs , saber-toothed cats , dodos , mammoths , ground sloths , thylacines , trilobites , golden toads , and passenger pigeons . Through evolution , species arise through
Araripesuchus - Misplaced Pages Continue
6804-468: Is the case, it is unknown when the name was adopted by the inhabitants of the island. No single Malagasy-language name predating Madagasikara appears to have been used by the local population to refer to the island, although some communities had their name for part or all of the lands they inhabited. One hypothesis relates Madagascar to the word Malay , referring to the Austronesian origin of
6966-502: Is the country's predominant religion, with a significant minority still practising traditional faiths . Madagascar is classified as a least developed country by the UN. Ecotourism and agriculture, paired with greater investments in education, health and private enterprise, are key elements of its development strategy. Despite substantial economic growth since the early 2000s, income disparities have widened, and quality of life remains low for
7128-592: Is the destruction of ocean floors by bottom trawling . Diminished resources or introduction of new competitor species also often accompany habitat degradation. Global warming has allowed some species to expand their range, bringing competition to other species that previously occupied that area. Sometimes these new competitors are predators and directly affect prey species, while at other times they may merely outcompete vulnerable species for limited resources. Vital resources including water and food can also be limited during habitat degradation, leading to extinction. In
7290-525: Is the most important determinant of genus extinction at background rates but becomes increasingly irrelevant as mass extinction arises. Limited geographic range is a cause both of small population size and of greater vulnerability to local environmental catastrophes. Extinction rates can be affected not just by population size, but by any factor that affects evolvability , including balancing selection , cryptic genetic variation , phenotypic plasticity , and robustness . A diverse or deep gene pool gives
7452-495: The 1973 oil crisis , resulted in the rapid collapse of Madagascar's economy and a sharp decline in living standards, and the country had become completely bankrupt by 1979. The Ratsiraka administration accepted the conditions of transparency, anti-corruption measures and free market policies imposed by the International Monetary Fund, World Bank and various bilateral donors in exchange for their bailout of
7614-486: The High Transitional Authority , an interim governing body responsible for moving the country toward presidential elections. In 2010, a new constitution was adopted by referendum , establishing a Fourth Republic, which sustained the democratic, multi-party structure established in the previous constitution. Hery Rajaonarimampianina was declared the winner of the 2013 presidential election , which
7776-483: The International Union for Conservation of Nature (IUCN) are not known to have any living specimens in the wild and are maintained only in zoos or other artificial environments. Some of these species are functionally extinct, as they are no longer part of their natural habitat and it is unlikely the species will ever be restored to the wild. When possible, modern zoological institutions try to maintain
7938-583: The Jurassic period, around 185 million years ago. The Indo-Madagascar landmass separated from Antarctica and Australia around 125 million years ago and Madagascar separated from the Indian landmass about 84–92 million years ago during the Late Cretaceous . This long history of separation from other continents has allowed plants and animals on the island to evolve in relative isolation. Along
8100-485: The Late Pleistocene could take up to 5 to 7 million years to restore mammal diversity to what it was before the human era. Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed ( anagenesis ) into a successor, or split into more than one ( cladogenesis ). Pseudoextinction
8262-767: The Madagascar Plan that had identified the island as a potential site for the deportation of Europe's Jews. During the Second World War , the island was the site of the Battle of Madagascar between the Vichy French and an Allied expeditionary force. The occupation of France during the Second World War tarnished the prestige of the colonial administration in Madagascar and galvanized
SECTION 50
#17327917584258424-562: The Malagasy language using the Roman alphabet , translated the Bible, and introduced a variety of new technologies to the island. Radama's successor, Queen Ranavalona I (1828–61), responded to increasing political and cultural encroachment on the part of Britain and France by issuing a royal edict prohibiting the practice of Christianity in Madagascar and pressuring most foreigners to leave
8586-584: The Malagasy people in modern-day Indonesia . In a map by Muhammad al-Idrisi dating from the year 1154, the island is named Gesira Malai , or "Malay island" in Arabic . The inversion of this name to Malai Gesira , as it was known by the Greeks , is thought to be the precursor of the modern name of the island. The name "Malay island" was later rendered in Latin as Malichu , an abbreviated form of Malai Insula , in
8748-564: The Mozambique Channel from East Africa. Other groups continued to settle on Madagascar over time, each one making lasting contributions to Malagasy cultural life. Consequently, there are 18 or more classified peoples of Madagascar , the most numerous being the Merina of the central highlands. Until the late 18th century, the island of Madagascar was ruled by a fragmented assortment of shifting sociopolitical alliances. Beginning in
8910-525: The Portuguese India Armadas . Matatana was the first Portuguese settlement on the south coast, 10 km west of Fort Dauphin . In 1508, settlers there built a tower, a small village, and a stone column. This settlement was established in 1513 at the behest of the viceroy of Portuguese India , Jeronimo de Azevedo . Contacts continued from the 1550s. Several colonization and conversion missions were ordered by King João III and by
9072-781: The Tsaratanana Massif is the island's highest point, followed by Boby Peak 2,658 m (8,720 ft) in the Andringitra Massif , and Tsiafajavona 2,643 m (8,671 ft) in the Ankaratra Massif. To the east, the Canal des Pangalanes is a chain of human-made and natural lakes connected by canals built by the French just inland from the east coast and running parallel to it for some 600 km (370 mi). The western and southern sides, which lie in
9234-629: The Vazimba . Probably the descendants of an earlier and less technologically advanced Austronesian settlement wave, the Vazimba were assimilated or expelled from the highlands by the Merina kings Andriamanelo , Ralambo , and Andrianjaka in the 16th and early 17th centuries. Today, the spirits of the Vazimba are revered as tompontany (ancestral masters of the land) by many traditional Malagasy communities. The written history of Madagascar began with
9396-422: The rain shadow of the central highlands, are home to dry deciduous forests , spiny forests , and deserts and xeric shrublands . Due to their lower population densities, Madagascar's dry deciduous forests have been better preserved than the eastern rain forests or the original woodlands of the central plateau. The western coast features many protected harbors, but silting is a major problem caused by sediment from
9558-410: The slender-billed curlew ( Numenius tenuirostris ), not seen since 2007. As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years, although this varies widely between taxa. A variety of causes can contribute directly or indirectly to the extinction of
9720-488: The strata of the Paris basin. They saw alternating saltwater and freshwater deposits, as well as patterns of the appearance and disappearance of fossils throughout the record. From these patterns, Cuvier inferred historic cycles of catastrophic flooding, extinction, and repopulation of the earth with new species. Cuvier's fossil evidence showed that very different life forms existed in the past than those that exist today,
9882-433: The trans-Atlantic slave trade . The small island of Nosy Boroha off the northeastern coast of Madagascar has been proposed by some historians as the site of the legendary pirate utopia of Libertalia . Many European sailors were shipwrecked on the coasts of the island, among them Robert Drury , whose journal is one of the few written depictions of life in southern Madagascar during the 18th century. European accounts until
SECTION 60
#173279175842510044-544: The "eighth continent", and the island has been classified by Conservation International as a biodiversity hotspot. Madagascar is classed as one of 17 megadiverse countries . The country is home to seven terrestrial ecoregions: Madagascar lowland forests , Madagascar subhumid forests , Madagascar dry deciduous forests , Madagascar ericoid thickets , Madagascar spiny forests , Madagascar succulent woodlands , and Madagascar mangroves . More than 80 percent of Madagascar's 14,883 plant species are found nowhere else in
10206-639: The 20 biodiversity goals laid out by the Aichi Biodiversity Targets in 2010, only 6 were "partially achieved" by the deadline of 2020. The report warned that biodiversity will continue to decline if the status quo is not changed, in particular the "currently unsustainable patterns of production and consumption, population growth and technological developments". In a 2021 report published in the journal Frontiers in Conservation Science , some top scientists asserted that even if
10368-524: The 46th largest country in the world. Its capital and largest city is Antananarivo . Following the prehistoric breakup of the supercontinent Gondwana , Madagascar split from Africa during the Early Jurassic , around 180 million years ago, and split from the Indian subcontinent around 90 million years ago, allowing native plants and animals to evolve in relative isolation; consequently, it
10530-587: The African island of Madagascar . Analysis of this specimen solidifies the position of A. wegneri as a member of the genus. A. tsangatsangana is the geologically youngest known of this genus. The sixth species, A. rattoides , was found in the Kem Kem Beds of the Sahara in a similar location to the specimens of A. wegeneri found by Sereno and Larsson, and is known only from parts of dentary bones, up to
10692-416: The African species support the inclusion of all five first described species into the same genus. The genus was originally assigned by Price to the family Uruguaysuchidae in the original 1959 description. This classification was followed by Buffetaut in 1981 with the description of A. wegeneri also within the same family. However, in their 2000 description of A. patagonicus , Ortega et al. avoided placing
10854-451: The Aichi Biodiversity Targets set for 2020 had been achieved, it would not have resulted in a significant mitigation of biodiversity loss. They added that failure of the global community to reach these targets is hardly surprising given that biodiversity loss is "nowhere close to the top of any country's priorities, trailing far behind other concerns such as employment, healthcare, economic growth, or currency stability." For much of history,
11016-638: The Arabs, who established trading posts along the northwest coast by at least the 10th century and introduced Islam , the Arabic script (used to transcribe the Malagasy language in a form of writing known as sorabe ), Arab astrology, and other cultural elements. European contact began in 1500, when the Portuguese sea captain Diogo Dias sighted the island, while participating in the 2nd Armada of
11178-618: The Earth is currently in the early stages of a human-caused mass extinction, known as the Holocene extinction . In that survey, the same proportion of respondents agreed with the prediction that up to 20% of all living populations could become extinct within 30 years (by 2028). A 2014 special edition of Science declared there is widespread consensus on the issue of human-driven mass species extinctions. A 2020 study published in PNAS stated that
11340-581: The Merina monarchy and sending the royal family into exile on Réunion Island and to Algeria. The conquest was followed by ten years of civil war, due to the Menalamba insurrection. The "pacification" carried out by the French administration lasted more than fifteen years, in response to the rural guerrillas scattered throughout the country. In total, the repression of this resistance to colonial conquest caused several tens of thousands of Malagasy victims. Under colonial rule, plantations were established for
11502-584: The North American moose and that the animal had once been common on the British Isles . Rather than suggest that this indicated the possibility of species going extinct, he argued that although organisms could become locally extinct, they could never be entirely lost and would continue to exist in some unknown region of the globe. The antlers were later confirmed to be from the extinct deer Megaloceros . Hooke and Molyneux's line of thinking
11664-522: The Paris basin, could be formed by a slow rise and fall of sea levels . The concept of extinction was integral to Charles Darwin 's On the Origin of Species , with less fit lineages disappearing over time. For Darwin, extinction was a constant side effect of competition . Because of the wide reach of On the Origin of Species , it was widely accepted that extinction occurred gradually and evenly (a concept now referred to as background extinction ). It
11826-493: The Rova palace compound was turned into a museum. Additional schools were built, particularly in rural and coastal areas where the schools of the Merina had not reached. Education became mandatory between the ages of 6 and 13 and focused primarily on the French language and practical skills. Huge mining and forestry concessions were granted to large companies. Native chiefs loyal to the French administration were also granted part of
11988-548: The United States government, to force the removal of Native Americans , many of whom relied on the bison for food. Madagascar Madagascar , officially the Republic of Madagascar , is an island country comprising the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa , it is the world's fourth largest island , the second-largest island country and
12150-548: The Viceroy of India, including one in 1553 by Baltazar Lobo de Sousa. In that mission, according to detailed descriptions by chroniclers Diogo do Couto and João de Barros, emissaries reached the inland via rivers and bays, exchanging goods and even converting one of the local kings. The French established trading posts along the east coast in the late 17th century. From about 1774 to 1824, Madagascar gained prominence among pirates and European traders, particularly those involved in
12312-597: The absolute power of the monarch. Following the coup, the courtiers offered Radama's queen, Rasoherina , the opportunity to rule, if she would accept a power-sharing arrangement with the Prime Minister: a new social contract that would be sealed by a political marriage between them. Queen Rasoherina accepted, first marrying Rainivoninahitriniony, then later deposing him and marrying his brother, Prime Minister Rainilaiarivony , who would go on to marry Queen Ranavalona II and Queen Ranavalona III in succession. Over
12474-429: The accumulation of slightly deleterious mutations , then a population will go extinct. Smaller populations have fewer beneficial mutations entering the population each generation, slowing adaptation. It is also easier for slightly deleterious mutations to fix in small populations; the resulting positive feedback loop between small population size and low fitness can cause mutational meltdown . Limited geographic range
12636-557: The adoption of a constitution in 1959 and full independence on 26 June 1960. Since regaining independence, Madagascar has transitioned through four republics with corresponding revisions to its constitution. The First Republic (1960–72), under the leadership of French-appointed President Philibert Tsiranana , was characterized by a continuation of strong economic and political ties to France. Many high-level technical positions were filled by French expatriates, and French teachers, textbooks and curricula continued to be used in schools around
12798-431: The back and paired dorsal ones along the tail. Each side of the tail also had a single row of osteoderms, and there were paired ventral osteoderms across most of the belly and underside as well. The osteoderms were not strongly keeled, which, along with the long limb bones and shoulder, hip and ankle joints that suggest upright posture, indicate that Araripesuchus was probably more active on land than on water. The name of
12960-481: The basis that the Lambert Charter had not been respected, France invaded Madagascar in 1883 in what became known as the first Franco-Hova War . At the end of the war, Madagascar ceded the northern port town of Antsiranana (Diego Suarez) to France and paid 560,000 francs to Lambert's heirs. In 1890, the British accepted the full formal imposition of a French protectorate on the island, but French authority
13122-404: The case. A. rattoides also had the distinctive feature of a highly enlarged and forward-pointing first dentary tooth referred to as an incisiform, resembling the elongated incisors found in rodents (hence the specific epithet). All species of Araripesuchus had relatively large orbits and hence eyes . They also had thin osteoderms that covered the entire body, multiple rows of them across
13284-429: The central highlands with its capital at the royal palace of Antananarivo, emerged at around the same time under the leadership of King Andriamanelo. Upon its emergence in the early 17th century, the highland kingdom of Imerina was initially a minor power relative to the larger coastal kingdoms and grew even weaker in the early 18th century when King Andriamasinavalona divided it among his four sons. Following almost
13446-597: The city in September 1895, the column bombarded the royal palace with heavy artillery, causing heavy casualties and leading Queen Ranavalona III to surrender. Popular resistance to the French capture of Antananarivo —known as the Menalamba rebellion —broke out in December 1895, and was not suppressed until the end of 1897. France annexed Madagascar in 1896 and declared the island a colony the following year, dissolving
13608-475: The coastal rainforests for cultivation. The first settlers encountered Madagascar's abundance of megafauna , including 17 species of giant lemurs , the large flightless elephant birds (including possibly the largest bird to ever exist, Aepyornis maximus ), the giant fossa , and several species of Malagasy hippopotamus , which have since become extinct because of hunting and habitat destruction. By 600 AD, groups of these early settlers had begun clearing
13770-517: The coasts of Madagascar, conquering the island and signing a treaty with its inhabitants. They remained for eight months and erected at different points of the island eight pillars on which they engraved "Magadoxo", a name which later, by corruption became Madagascar. Jan Huyghen van Linschoten , a Dutch traveler who copied Portuguese works and maps, confirmed this event by saying "Madagascar has its name from 'makdishu' (Mogadishu)" whose "shayk" invaded it. The name Malagasikara , or Malagascar ,
13932-600: The consequences can be catastrophic. Invasive alien species can affect native species directly by eating them, competing with them, and introducing pathogens or parasites that sicken or kill them; or indirectly by destroying or degrading their habitat. Human populations may themselves act as invasive predators. According to the "overkill hypothesis", the swift extinction of the megafauna in areas such as Australia (40,000 years before present), North and South America (12,000 years before present), Madagascar , Hawaii (AD 300–1000), and New Zealand (AD 1300–1500), resulted from
14094-418: The contemporary extinction crisis "may be the most serious environmental threat to the persistence of civilization, because it is irreversible." Biologist E. O. Wilson estimated in 2002 that if current rates of human destruction of the biosphere continue, one-half of all plant and animal species of life on earth will be extinct in 100 years. More significantly, the current rate of global species extinctions
14256-419: The country of Mauritius to the east, as well as the state of Comoros and the French territory of Mayotte to the north west. The nearest mainland state is Mozambique , located to the west. The prehistoric breakup of the supercontinent Gondwana resulted in the separation of East Gondwana (comprising Madagascar, Antarctica, Australia and the Indian subcontinent) and West Gondwana (Africa–South America) during
14418-427: The country. Popular resentment over Tsiranana's tolerance for this "neo-colonial" arrangement inspired a series of farmer and student protests that overturned his administration in 1972. Gabriel Ramanantsoa , a major general in the army, was appointed interim president and prime minister that same year, but low public approval forced him to step down in 1975. Colonel Richard Ratsimandrava , appointed to succeed him,
14580-404: The course of Rainilaiarivony's 31-year tenure as prime minister, numerous policies were adopted to modernize and consolidate the power of the central government. Schools were constructed throughout the island and attendance was made mandatory. Army organization was improved and British consultants were employed to train and professionalize soldiers. Polygamy was outlawed and Christianity declared
14742-541: The death of its last member if it loses the capacity to reproduce and recover. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa , where a species presumed extinct abruptly "reappears" (typically in the fossil record ) after a period of apparent absence. More than 99% of all species that ever lived on Earth , amounting to over five billion species, are estimated to have died out. It
14904-476: The deliberate destruction of some species, such as dangerous viruses , and the total destruction of other problematic species has been suggested. Other species were deliberately driven to extinction, or nearly so, due to poaching or because they were "undesirable", or to push for other human agendas. One example was the near extinction of the American bison , which was nearly wiped out by mass hunts sanctioned by
15066-515: The early 19th century, most of it was united and ruled as the Kingdom of Madagascar by a series of Merina nobles . The monarchy was ended in 1897 by the annexation by France , from which Madagascar gained independence in 1960. The country has since undergone four major constitutional periods, termed republics, and has been governed as a constitutional democracy since 1992. Following a political crisis and military coup in 2009 , Madagascar underwent
15228-526: The early 20th century identified Malagasy people as being of Jewish origin . The wealth generated by maritime trade spurred the rise of organized kingdoms on the island, some of which had grown quite powerful by the 17th century. Among these were the Betsimisaraka alliance of the eastern coast and the Sakalava chiefdoms of Menabe and Boina on the west coast. The Kingdom of Imerina, located in
15390-403: The endangered wild water buffalo is most threatened with extinction by genetic pollution from the abundant domestic water buffalo ). Such extinctions are not always apparent from morphological (non-genetic) observations. Some degree of gene flow is a normal evolutionary process; nevertheless, hybridization (with or without introgression) threatens rare species' existence. The gene pool of
15552-591: The event of rediscovery would be considered Lazarus species. Examples include the thylacine , or Tasmanian tiger ( Thylacinus cynocephalus ), the last known example of which died in Hobart Zoo in Tasmania in 1936; the Japanese wolf ( Canis lupus hodophilax ), last sighted over 100 years ago; the American ivory-billed woodpecker ( Campephilus principalis ), with the last universally accepted sighting in 1944; and
15714-443: The extinction crisis. According to the International Union for Conservation of Nature (IUCN), 784 extinctions have been recorded since the year 1500, the arbitrary date selected to define "recent" extinctions, up to the year 2004; with many more likely to have gone unnoticed. Several species have also been listed as extinct since 2004. If adaptation increasing population fitness is slower than environmental degradation plus
15876-475: The extinction of species caused by humanity, and they try to prevent further extinctions through a variety of conservation programs. Humans can cause extinction of a species through overharvesting , pollution , habitat destruction , introduction of invasive species (such as new predators and food competitors ), overhunting, and other influences. Explosive, unsustainable human population growth and increasing per capita consumption are essential drivers of
16038-517: The family Uruguaysuchidae. This family was found to be the most basal group of Notosuchia. Below is a cladogram from the analysis: Uruguaysuchus A. wegeneri A. buitreraensis A. gomesii A. patagonicus Libycosuchus Simosuchus Malawisuchus Candidodon Chimaerasuchus Sphagesaurus Baurusuchus Bretesuchus Iberosuchus Notosuchus Comahuesuchus Mariliasuchus However, recent phylogenetic analyses placed A. wegeneri as
16200-540: The field of zoology , and biology in general, and has also become an area of concern outside the scientific community. A number of organizations, such as the Worldwide Fund for Nature , have been created with the goal of preserving species from extinction. Governments have attempted, through enacting laws, to avoid habitat destruction, agricultural over-harvesting, and pollution . While many human-caused extinctions have been accidental, humans have also engaged in
16362-401: The first African species discovered, A. wegeneri , was questioned for a while by various authors. Ortega et al. argued for the assignment of the errant species to another genus based on phylogenetic analysis Further analysis, combined with the discovery of the second African species A. tsangatsangana has shed more light on the placement of A. wegeneri within the genus. When analyzed together,
16524-412: The first documents written that might explain why Marco Polo called it Madagascar is in a 1609 book on Madagascar by Jerome Megiser. Jerome Megiser describes an event in which the kings of Mogadishu and Adal traveled to Madagascar with a fleet of around twenty-five thousand men in order to invade the wealthy islands of Taprobane and Sumatra. However, a tempest threw them off course and they landed on
16686-610: The first round. In January 2019 the High Constitutional Court declared Rajoelina as the winner of the elections and the new president. In June 2019 parliamentary elections the Rajoelina's won absolute majority of the seats of the National Assembly. It received 84 seats and the supporters of former president Ravalomana got only 16 seats of 151 seats of the National Assembly. 51 seats of deputies were independent or represented small parties. Rajoelina could rule as
16848-512: The forests of the central highlands. Arab traders first reached the island between the 7th and 9th centuries. A wave of Bantu -speaking migrants from southeastern Africa arrived around the year 1000. Around this time, zebu from South India were first brought, intermingling with sanga found in East Africa. Irrigated paddy fields were developed in the central highland Betsileo Kingdom and were extended with terraced paddies throughout
17010-435: The form of a physical destruction of niche habitats. The widespread destruction of tropical rainforests and replacement with open pastureland is widely cited as an example of this; elimination of the dense forest eliminated the infrastructure needed by many species to survive. For example, a fern that depends on dense shade for protection from direct sunlight can no longer survive without forest to shelter it. Another example
17172-474: The fourteenth alveolus. It was described in the same paper as Kaprosuchus , Laganosuchus and Anatosuchus ; the four were therefore popularized by the authors as 'RatCroc', 'BoarCroc', 'PancakeCroc' and 'DuckCroc' respectively. As of 2024, seven species were recognized within the genus Araripesuchus : A. gomesii ( type species ), A. buitreraensis , A. manzanensis , A. patagonicus , A. rattoides , A. tsangatsangana and A. wegeneri . The placement of
17334-648: The genus was coined in 1959 with the description of the type species Araripesuchus gomesii , a notosuchian crocodylian from the famed Santana Group of the Araripe Basin in Brazil . The holotype used to describe the genus, 423-R is currently in the care of the Divisão de Mineralogia e Geologia do Departamento Nacional da Produção Mineral in Rio de Janeiro . 423-R consists of a single skull articulating with part of
17496-443: The genus, was Araripesuchus buitreraensis , described in 2005. This species was described from a single skull (MPCA-PV 235) retrieved from Late Cretaceous deposits in what is now Argentina . At 130 millimeters, the skull is the largest Araripesuchus specimen discovered to date. A fifth species, Araripesuchus tsangatsangana was described in 2006. This species' type specimen was discovered from latest Late Cretaceous deposits from
17658-602: The growing independence movement, leading to the Malagasy Uprising of 1947. This movement led the French to establish reformed institutions in 1956 under the Loi Cadre (Overseas Reform Act), and Madagascar moved peacefully towards independence. The Malagasy Republic was proclaimed on 14 October 1958, as an autonomous state within the French Community . A period of provisional government ended with
17820-580: The habitat retreat of taxa approaching extinction. Possible causes of the higher extinction risk in species with more sexual selection shown by the comprehensive fossil studies that rule out such error sources include expensive sexually selected ornaments having negative effects on the ability to survive natural selection , as well as sexual selection removing a diversity of genes that under current ecological conditions are neutral for natural selection but some of which may be important for surviving climate change. There have been at least five mass extinctions in
17982-676: The heaviest losses include the Cape Floristic Region and the Caribbean Basin . These areas might see a doubling of present carbon dioxide levels and rising temperatures that could eliminate 56,000 plant and 3,700 animal species. Climate change has also been found to be a factor in habitat loss and desertification . Studies of fossils following species from the time they evolved to their extinction show that species with high sexual dimorphism , especially characteristics in males that are used to compete for mating, are at
18144-654: The heavy precipitation supports the area's rainforest ecosystem. The central highlands are both drier and cooler while the west is drier still, and a semi-arid climate prevails in the southwest and southern interior of the island. Tropical cyclones cause damage to infrastructure and local economies as well as loss of life. In 2004, Cyclone Gafilo became the strongest cyclone ever recorded to hit Madagascar. The storm killed 172 people, left 214,260 homeless and caused more than US$ 250 million in damage. In February 2022, Cyclone Batsirai killed 121 people, weeks after Cyclone Ana killed 55 and displaced 130,000 people on
18306-471: The high levels of inland erosion carried by rivers crossing the broad western plains. The combination of southeastern trade winds and northwestern monsoons produces a hot rainy season (November–April) with frequently destructive cyclones , and a relatively cooler dry season (May–October). Rain clouds originating over the Indian Ocean discharge much of their moisture over the island's eastern coast;
18468-447: The history of life on earth, and four in the last 350 million years in which many species have disappeared in a relatively short period of geological time. A massive eruptive event that released large quantities of tephra particles into the atmosphere is considered to be one likely cause of the " Permian–Triassic extinction event " about 250 million years ago, which is estimated to have killed 90% of species then existing. There
18630-475: The human era since the Late Pleistocene would require 5 to 7 million years to recover. According to the 2019 Global Assessment Report on Biodiversity and Ecosystem Services by IPBES , the biomass of wild mammals has fallen by 82%, natural ecosystems have lost about half their area and a million species are at risk of extinction—all largely as a result of human actions. Twenty-five percent of plant and animal species are threatened with extinction. In
18792-523: The initiative of the communist activist François Vittori, who was imprisoned as a result. The 1930s saw the Malagasy anti-colonial movement gain further momentum. Malagasy trade unionism began to appear underground and the Communist Party of the Madagascar region was formed. But in 1939, all the organizations were dissolved by the administration of the colony, which opted for the Vichy regime. The MDRM
18954-481: The international community deemed fair and transparent. In 2018 the first round of the presidential election was held on 7 November and the second round was held on 10 December. Three former presidents and the most recent president were the main candidates of the elections. Rajoelina won the second round of the elections. Ravalomana lost the second round and he did not accept the results because of allegations of fraud. Rajaonarimampianina received very modest support in
19116-431: The introduced ( or hybrid ) species. Endemic populations can face such extinctions when new populations are imported or selectively bred by people, or when habitat modification brings previously isolated species into contact. Extinction is likeliest for rare species coming into contact with more abundant ones; interbreeding can swamp the rarer gene pool and create hybrids, depleting the purebred gene pool (for example,
19278-482: The island and further expand the Kingdom of Merina to encompass most of Madagascar. Residents of Madagascar could accuse one another of various crimes, including theft, Christianity and especially witchcraft, for which the ordeal of tangena was routinely obligatory. Between 1828 and 1861, the tangena ordeal caused about 3,000 deaths annually. In 1838, it was estimated that as many as 100,000 people in Imerina died as
19440-449: The island earlier than the mid-first millennium AD are not strongly supported. However, there is scattered evidence for much earlier human visits and presence. (See History of Madagascar ). Archaeological finds such as cut marks on bones found in the northwest and stone tools in the northeast indicate that Madagascar was visited by foragers around 2000 BCE. Upon arrival, early settlers practiced slash-and-burn agriculture to clear
19602-499: The island. A 2022 analysis found that the expected costs for Madagascar, to adapt to and avert the environmental consequences of climate change , are going to be high. As a result of the island's long isolation from neighbouring continents, Madagascar is home to various endemic plants and animals found nowhere else on Earth. Approximately 90% of all plant and animal species found in Madagascar are endemic . This distinctive ecology has led some ecologists to refer to Madagascar as
19764-671: The land. Forced labor was introduced in favor of the French companies and peasants were encouraged, through taxation, to work for wages (especially in the colonial concessions) to the detriment of small individual farms. However, the colonial period was accompanied by movements fighting for independence: the Menalamba, the Vy Vato Sakelika, the Democratic Movement for Malagasy Renovation (MDRM). In 1927, major demonstrations were organized in Antananarivo, notably on
19926-438: The length of the eastern coast runs a narrow and steep escarpment containing much of the island's remaining tropical lowland forest . To the west of this ridge lies a plateau in the center of the island ranging in altitude from 750 to 1,500 m (2,460 to 4,920 ft) above sea level. These central highlands , traditionally the homeland of the Merina people and the location of their historic capital at Antananarivo , are
20088-440: The main drivers of the modern extinction crisis. In January 2020, the UN's Convention on Biological Diversity drafted a plan to mitigate the contemporary extinction crisis by establishing a deadline of 2030 to protect 30% of the Earth's land and oceans and reduce pollution by 50%, with the goal of allowing for the restoration of ecosystems by 2050. The 2020 United Nations ' Global Biodiversity Outlook report stated that of
20250-498: The majority of the population. As of 2021, 68.4 percent of the population is considered to be multidimensionally poor . In the Malagasy language , the island of Madagascar is called Madagasikara ( Malagasy pronunciation: [madaɡasʲˈkʲarə̥] ) and its people are referred to as Malagasy . The origin of the name is uncertain, and is likely foreign, having been propagated in the Middle Ages by Europeans. If this
20412-503: The medieval Hereford Mappa Mundi as the name of Madagascar. Another hypothesis is that Madagascar is a corrupted transliteration of Mogadishu , the capital of Somalia and an important medieval Indian Ocean port. This would have resulted from 13th-century Venetian explorer Marco Polo confusing the two locations in his memoirs, in which he mentions the land of Madageiscar to the south of Socotra . This name would then have been popularized on Renaissance maps by Europeans. One of
20574-421: The modern understanding of extinction as the end of a species was incompatible with the prevailing worldview. Prior to the 19th century, much of Western society adhered to the belief that the world was created by God and as such was complete and perfect. This concept reached its heyday in the 1700s with the peak popularity of a theological concept called the great chain of being , in which all life on earth, from
20736-438: The monarchy, and Joseph-François Lambert , a French adventurer and slave trader, with whom then-Prince Radama II signed a controversial trade agreement termed the Lambert Charter. Succeeding his mother, Radama II attempted to relax the queen's stringent policies but was overthrown two years later by Prime Minister Rainivoninahitriniony and an alliance of Andriana (noble) and Hova (commoner) courtiers, who sought to end
20898-480: The most densely populated part of the island and are characterized by terraced, rice-growing valleys lying between grassy hills and patches of the subhumid forests that formerly covered the highland region. To the west of the highlands, the increasingly arid terrain gradually slopes down to the Mozambique Channel and mangrove swamps along the coast. Madagascar's highest peaks rise from three prominent highland massifs : Maromokotro 2,876 m (9,436 ft) in
21060-490: The name of a species or other taxon normally indicates its status as extinct. Examples of species and subspecies that are extinct include: A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over
21222-515: The nation's broken economy. Ratsiraka's dwindling popularity in the late 1980s reached a critical point in 1991 when presidential guards opened fire on unarmed protesters during a rally. Within two months, a transitional government had been established under the leadership of Albert Zafy (1993–96), who went on to win the 1992 presidential elections and inaugurate the Third Republic (1992–2010). The new Madagascar constitution established
21384-469: The natural course of events, species become extinct for a number of reasons, including but not limited to: extinction of a necessary host, prey or pollinator, interspecific competition , inability to deal with evolving diseases and changing environmental conditions (particularly sudden changes) which can act to introduce novel predators, or to remove prey. Recently in geological time, humans have become an additional cause of extinction of some species, either as
21546-423: The neighboring Kingdom of Imerina a century later. The rising intensity of land cultivation and the ever-increasing demand for zebu pasturage had largely transformed the central highlands from a forest ecosystem to grassland by the 17th century. The oral histories of the Merina people, who arrived in the central highlands between 600 and 1,000 years ago, describe encountering an established population they called
21708-410: The newly emerging school of uniformitarianism . Jean-Baptiste Lamarck , a gradualist and colleague of Cuvier, saw the fossils of different life forms as evidence of the mutable character of species. While Lamarck did not deny the possibility of extinction, he believed that it was exceptional and rare and that most of the change in species over time was due to gradual change. Unlike Cuvier, Lamarck
21870-430: The official religion of the court in 1869, was adopted alongside traditional beliefs among a growing portion of the populace. Legal codes were reformed on the basis of British common law and three European-style courts were established in the capital city. In his joint role as Commander-in-Chief, Rainilaiarivony also successfully ensured the defense of Madagascar against several French colonial incursions. Primarily on
22032-399: The original population, thereby increasing the chance of extinction. Habitat degradation is currently the main anthropogenic cause of species extinctions. The main cause of habitat degradation worldwide is agriculture, with urban sprawl , logging, mining, and some fishing practices close behind. The degradation of a species' habitat may alter the fitness landscape to such an extent that
22194-457: The period between 350 BC and 550 AD, while others are cautious about dates earlier than AD 250. In either case, these dates make Madagascar one of the most recent major landmasses on Earth to be settled by humans, predating the settlement of Iceland and New Zealand . It is proposed that Ma'anyan people were brought as laborers and slaves by Javan and Sumatran - Malays in their trading fleets to Madagascar. Dates of settlement of
22356-511: The phylogenetic position of this genus is uncertain, and that taxonomic revision is required. A. wegeneri is estimated to reach 81 centimetres (32 in) long while A. rattoides is estimated to reach up to 1 metre (3.3 ft). Araripesuchus can be distinguished by their laterally bulged edges of the snout, with the bulge being the most prominent around the area of an enlarged maxillary tooth. The snout and premaxilla are also smoother than that of most crocodyliforms, without foramina or
22518-418: The political crisis was gradually overcome by Ravalomanana's progressive economic and political policies, which encouraged investments in education and ecotourism, facilitated foreign direct investment, and cultivated trading partnerships both regionally and internationally. National GDP grew at an average rate of 7 percent per year under his administration. In the latter half of his second term, Ravalomanana
22680-702: The process of speciation —where new varieties of organisms arise and thrive when they are able to find and exploit an ecological niche —and species become extinct when they are no longer able to survive in changing conditions or against superior competition . The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called living fossils , survive with little to no morphological change for hundreds of millions of years. Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are
22842-407: The production of a variety of export crops. Slavery was abolished in 1896 and approximately 500,000 slaves were freed; many remained in their former masters' homes as servants or as sharecroppers; in many parts of the island strong discriminatory views against slave descendants are still held today. Wide paved boulevards and gathering places were constructed in the capital city of Antananarivo and
23004-410: The relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on nature and nurture . The question of whether more extinctions in the fossil record have been caused by evolution or by competition or by predation or by disease or by catastrophe is a subject of discussion; Mark Newman, the author of Modeling Extinction , argues for
23166-416: The same stock in the general area, radiating outward from a yet-unidentified origin point. The presence of specimens from Madagascar further strengthens this evolutionary radiation model. [REDACTED] [REDACTED] [REDACTED] [REDACTED] Extinct Extinction is the termination of a taxon by the death of its last member . A taxon may become functionally extinct before
23328-460: The species is no longer able to survive and becomes extinct. This may occur by direct effects, such as the environment becoming toxic , or indirectly, by limiting a species' ability to compete effectively for diminished resources or against new competitor species. Habitat destruction, particularly the removal of vegetation that stabilizes soil, enhances erosion and diminishes nutrient availability in terrestrial ecosystems. This degradation can lead to
23490-514: The species within the family. Instead, it was simply noted that Uruguaysuchus was a possible close relative of the genus. Ortega et al. and several other studies place Araripesuchus outside Notosuchia. In some phylogenetic analyses, it is placed closer to the clade Neosuchia , which includes modern crocodilians. In most recent analyses, however, Araripesuchus is placed as a basal notosuchian. The phylogenetic analysis of Soto et al. (2011) joined Araripesuchus with Uruguaysuchus , reinstating
23652-430: The sudden introduction of human beings to environments full of animals that had never seen them before and were therefore completely unadapted to their predation techniques. Coextinction refers to the loss of a species due to the extinction of another; for example, the extinction of parasitic insects following the loss of their hosts. Coextinction can also occur when a species loses its pollinator , or to predators in
23814-525: The territory. William Ellis of the London Missionary Society described his visits made during her reign in his book Three Visits to Madagascar during the years 1853, 1854, and 1856 . The Queen made heavy use of the traditional practice of fanompoana (forced labor as tax payment) to complete public works projects and develop a standing army of between 20,000 and 30,000 Merina soldiers, whom she deployed to pacify outlying regions of
23976-412: The tiniest microorganism to God, is linked in a continuous chain. The extinction of a species was impossible under this model, as it would create gaps or missing links in the chain and destroy the natural order. Thomas Jefferson was a firm supporter of the great chain of being and an opponent of extinction, famously denying the extinction of the woolly mammoth on the grounds that nature never allows
24138-567: The total extinction of the dodo and the extirpation of indigenous horses to the British Isles. He similarly argued against mass extinctions , believing that any extinction must be a gradual process. Lyell also showed that Cuvier's original interpretation of the Parisian strata was incorrect. Instead of the catastrophic floods inferred by Cuvier, Lyell demonstrated that patterns of saltwater and freshwater deposits , like those seen in
24300-432: The typical rugose texture. There are seven valid species within this genus, all with slightly differing maxillary or dentary structure. A. gomesii, A. wegeneri and A. tsangatsangana all have a mild concavity of the external alveolar margin of the premaxilla as viewed from the ventral surface; A. rattoides may also have this feature, although this part of its skull is not known, as the dentary suggests that this would be
24462-530: The wider scientific community of his theory. Cuvier was a well-regarded geologist, lauded for his ability to reconstruct the anatomy of an unknown species from a few fragments of bone. His primary evidence for extinction came from mammoth skulls found in the Paris basin . Cuvier recognized them as distinct from any known living species of elephant, and argued that it was highly unlikely such an enormous animal would go undiscovered. In 1812, Cuvier, along with Alexandre Brongniart and Geoffroy Saint-Hilaire , mapped
24624-561: The world, including five plant families. The family Didiereaceae , composed of four genera and 11 species, is limited to the spiny forests of southwestern Madagascar. Four-fifths of the world's Pachypodium species are endemic to the island. Three-fourths of Madagascar's 860 orchid species are found here alone, as are six of the world's nine baobab species. The island is home to around 170 palm species, three times as many as on all of mainland Africa; 165 of them are endemic. Many native plant species are used as herbal remedies for
24786-419: Was 46.36%, the lowest in a presidential election in the country's history. At 592,800 square kilometres (228,900 sq mi), Madagascar is the world's 46th largest country , the second-largest island country and the fourth-largest island . The country lies mostly between latitudes 12°S and 26°S , and longitudes 43°E and 51°E . Neighboring islands include the French territory of Réunion and
24948-465: Was accused by the colonial regime of being at the origin of the 1947 insurrection and was pursued by violent repression. The Merina royal tradition of taxes paid in the form of labor was continued under the French and used to construct a railway and roads linking key coastal cities to Antananarivo. Malagasy troops fought for France in World War I . In the 1930s, Nazi political thinkers developed
25110-475: Was appointed for the three months prior to the next presidential election. Ratsiraka was then voted back into power on a platform of decentralization and economic reforms for a second term which lasted from 1996 to 2001. The contested 2001 presidential elections in which then-mayor of Antananarivo, Marc Ravalomanana, eventually emerged victorious, caused a seven-month standoff in 2002 between supporters of Ravalomanana and Ratsiraka. The negative economic impact of
25272-611: Was assassinated six days into his tenure. General Gilles Andriamahazo ruled after Ratsimandrava for four months before being replaced by another military appointee: Vice Admiral Didier Ratsiraka , who ushered in the Marxist–Leninist Second Republic that ran under his tenure from 1975 to 1993. This period saw a political alignment with the Eastern Bloc countries and a shift toward economic insularity. These policies, coupled with economic pressures stemming from
25434-532: Was criticised by domestic and international observers who accused him of increasing authoritarianism and corruption. Opposition leader and then-mayor of Antananarivo, Andry Rajoelina , led a movement in early 2009 in which Ravalomanana was pushed from power in an unconstitutional process widely condemned as a coup d'état . In March 2009, Rajoelina was declared by the Supreme Court as the President of
25596-491: Was difficult to disprove. When parts of the world had not been thoroughly examined and charted, scientists could not rule out that animals found only in the fossil record were not simply "hiding" in unexplored regions of the Earth. Georges Cuvier is credited with establishing the modern conception of extinction in a 1796 lecture to the French Institute , though he would spend most of his career trying to convince
25758-431: Was not acknowledged by the government of Madagascar. To force capitulation, the French bombarded and occupied the harbor of Toamasina on the east coast, and Mahajanga on the west coast, in December 1894 and January 1895 respectively. A French military flying column then marched toward Antananarivo, losing many men to malaria and other diseases. Reinforcements came from Algeria and Sub-Saharan Africa . Upon reaching
25920-430: Was not until 1982, when David Raup and Jack Sepkoski published their seminal paper on mass extinctions, that Cuvier was vindicated and catastrophic extinction was accepted as an important mechanism . The current understanding of extinction is a synthesis of the cataclysmic extinction events proposed by Cuvier, and the background extinction events proposed by Lyell and Darwin. Extinction is an important research topic in
26082-442: Was recognized by the British government as King of Madagascar. Radama concluded a treaty in 1817 with the British governor of Mauritius to abolish the lucrative slave trade in return for British military and financial assistance. Artisan missionary envoys from the London Missionary Society began arriving in 1818 and included such key figures as James Cameron , David Jones and David Griffiths , who established schools, transcribed
26244-571: Was skeptical that catastrophic events of a scale large enough to cause total extinction were possible. In his geological history of the earth titled Hydrogeologie, Lamarck instead argued that the surface of the earth was shaped by gradual erosion and deposition by water, and that species changed over time in response to the changing environment. Charles Lyell , a noted geologist and founder of uniformitarianism , believed that past processes should be understood using present day processes. Like Lamarck, Lyell acknowledged that extinction could occur, noting
#424575