91-419: This is an accepted version of this page A bee is a flying insect. Bee , BEE or The Bee may also refer to: Bee Apiformes (from Latin 'apis' ) Bees are winged insects closely related to wasps and ants , known for their roles in pollination and, in the case of the best-known bee species, the western honey bee , for producing honey . Bees are a monophyletic lineage within
182-401: A division of labour into reproductive and non-reproductive adults, plus overlapping generations. This division of labour creates specialized groups within eusocial societies which are called castes . In some species, groups of cohabiting females may be sisters, and if there is a division of labour within the group, they are considered semisocial . The group is called eusocial if, in addition,
273-663: A bee, be it a solitary or social species, involves the laying of an egg, the development through several moults of a legless larva , a pupation stage during which the insect undergoes complete metamorphosis , followed by the emergence of a winged adult. The number of eggs laid by a female during her lifetime can vary from eight or less in some solitary bees, to more than a million in highly social species. Most solitary bees and bumble bees in temperate climates overwinter as adults or pupae and emerge in spring when increasing numbers of flowering plants come into bloom. The males usually emerge first and search for females with which to mate. Like
364-429: A compartment (a "cell") with an egg and some provisions for the resulting larva, then seals it off. A nest may consist of numerous cells. When the nest is in wood, usually the last (those closer to the entrance) contain eggs that will become males. The adult does not provide care for the brood once the egg is laid, and usually dies after making one or more nests. The males typically emerge first and are ready for mating when
455-466: A distinct set of genera which are commonly known by their nesting behavior or preferences, namely: carpenter bees, sweat bees , mason bees, plasterer bees , squash bees , dwarf carpenter bees , leafcutter bees, alkali bees and digger bees . Most solitary bees are fossorial , digging nests in the ground in a variety of soil textures and conditions, while others create nests in hollow reeds or twigs, or holes in wood . The female typically creates
546-698: A fleet to reach Peru by sea, and sought the military support of various rebels from the Viceroyalty of Peru. In 1822 the two men met at the Guayaquil Conference in Ecuador, where they failed to agree on governance strategies for the liberated nations. Two years later Bolívar's forces beat the Spanish at the Battle of Ayacucho , securing the independence of Peru and the rest of South America. In
637-525: A larva's food is supplied gradually as it develops, as is the case in honey bees and some bumblebees. Most other bees, including familiar insects such as carpenter bees , leafcutter bees and mason bees are solitary in the sense that every female is fertile, and typically inhabits a nest she constructs herself. There is no division of labor so these nests lack queens and worker bees for these species. Solitary bees typically produce neither honey nor beeswax . Bees collect pollen to feed their young, and have
728-440: A length of 39 millimetres (1.54 in). Bees feed on nectar and pollen , the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for their larvae . Vertebrate predators of bees include primates and birds such as bee-eaters ; insect predators include beewolves and dragonflies . Bee pollination is important both ecologically and commercially , and
819-459: A long history of colonialism , the overwhelming majority of South Americans speak Spanish or Portuguese , and societies and states are rich in Western traditions . Relative to Africa, Asia, and Europe, post-1900 South America has been a peaceful continent with few wars, although high rates of violent crime remain a concern in some countries. South America occupies the southern portion of
910-729: A male is haploid (has only one copy of each gene), his daughters (which are diploid , with two copies of each gene) share 100% of his genes and 50% of their mother's. Therefore, they share 75% of their genes with each other. This mechanism of sex determination gives rise to what W. D. Hamilton termed "supersisters", more closely related to their sisters than they would be to their own offspring. Workers often do not reproduce, but they can pass on more of their genes by helping to raise their sisters (as queens) than they would by having their own offspring (each of which would only have 50% of their genes), assuming they would produce similar numbers. This unusual situation has been proposed as an explanation of
1001-682: A part of Oceania, also administered by Chile), the Galápagos Islands (administered by Ecuador , sometimes considered part of Oceania), and Tierra del Fuego (split between Argentina and Chile). In the Atlantic Ocean, Brazil administers Fernando de Noronha , Trindade and Martim Vaz , and the Saint Peter and Saint Paul Archipelago , while the Falkland Islands (Spanish: Islas Malvinas ) and South Georgia and
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#17327873430051092-533: A single season colony cycle, even in the tropics, and only mated females hibernate. A few species have long active seasons and attain colony sizes in the hundreds, such as Halictus hesperus . Some species are eusocial in parts of their range and solitary in others, or have a mix of eusocial and solitary nests in the same population. The orchid bees (Apidae) include some primitively eusocial species with similar biology. Some allodapine bees (Apidae) form primitively eusocial colonies, with progressive provisioning :
1183-481: A site called Chavín de Huantar in modern Peru at an elevation of 3,177 meters (10,423 ft). Chavín civilization spanned 900 BC to 300 BC. In the central coast of Peru, around the beginning of the 1st millennium AD, Moche (100 BC – 700 AD, at the northern coast of Peru), Paracas and Nazca (400 BC – 800 AD, Peru) cultures flourished with centralized states with permanent militia improving agriculture through irrigation and new styles of ceramic art. At
1274-689: A subtropical climate of the dinaric type. At the highest points of the Andean region, climates are colder than the ones occurring at the highest point of the Norwegian fjords. In the Andean plateaus, the warm climate prevails, although it is tempered by the altitude, while in the coastal strip, there is an equatorial climate of the Guinean type. From this point until the north of the Chilean coast appear, successively, Mediterranean oceanic climate , temperate of
1365-512: Is Cretotrigona prisca , a corbiculate bee of Late Cretaceous age (~70 mya) found in New Jersey amber . A fossil from the early Cretaceous (~100 mya), Melittosphex burmensis , was initially considered "an extinct lineage of pollen-collecting Apoidea sister to the modern bees", but subsequent research has rejected the claim that Melittosphex is a bee, or even a member of the superfamily Apoidea to which bees belong, instead treating
1456-1075: Is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere , with a considerably smaller portion in the Northern Hemisphere . It can also be described as the southern subregion of the Americas . South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean ; North America and the Caribbean Sea lie to the northwest. The continent includes twelve sovereign states : Argentina , Bolivia , Brazil , Chile , Colombia , Ecuador , Guyana , Paraguay , Peru , Suriname , Uruguay , and Venezuela ; two dependent territories :
1547-526: Is because the most extensive part of the continent is in the equatorial zone (the region has more areas of equatorial plains than any other region), therefore giving the Southern Cone more oceanic influence, which moderates year round temperatures. The average annual temperatures in the Amazon basin oscillate around 27 °C (81 °F), with low thermal amplitudes and high rainfall indices. Between
1638-476: Is believed that mites eat fungi that attack pollen, so the relationship in this case may be mutualistic . Molecular phylogeny was used by Debevic et al , 2012, to demonstrate that the bees ( Anthophila ) arose from deep within the Crabronidae sensu lato , which was thus rendered paraphyletic . In their study, the placement of the monogeneric Heterogynaidae was uncertain. The small family Mellinidae
1729-460: Is by far the most populous South American country, with almost half of the continent's population, followed by Colombia, Argentina, Venezuela, and Peru. In recent decades, Brazil has also generated half of the continent's GDP and has become the continent's first regional power. Most of the population lives near the continent's western or eastern coasts while the interior and the far south are sparsely populated. The geography of western South America
1820-605: Is dominated by the Andes mountains; in contrast, the eastern part contains both highland regions and vast lowlands where rivers such as the Amazon , Orinoco and Paraná flow. Most of the continent lies in the tropics, except for a large part of the Southern Cone located in the middle latitudes . The continent's cultural and ethnic outlook has its origin with the interaction of Indigenous peoples with European conquerors and immigrants and, more locally, with African slaves . Given
1911-614: Is now Colombia. They established the Muisca Confederation of many clans, or cacicazgos , that had a free trade network among themselves. Many were goldsmiths and farmers. Other important Pre-Columbian cultures include: the Cañaris (in south central Ecuador), Chimú Empire (1300–1470, Peruvian northern coast), Chachapoyas , and the Aymaran kingdoms (1000–1450, Western Bolivia and southern Peru). Holding their capital at
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#17327873430052002-480: Is one of the most biodiverse continents on Earth. It is home to many unique species of animals including the llama , anaconda , piranha , jaguar , vicuña , and tapir , and to one of the largest known insects in the world, the Titan beetle . The Amazon rainforests possess high biodiversity , with Brazil estimated to contain 10% of Earth's species . 83% of South America's large mammals ( megafauna ) became extinct at
2093-444: Is the ancestral state for all eusocial species so far investigated, so it is likely that haplodiploidy contributed to the evolution of eusociality in bees. Bees may be solitary or may live in various types of communities. Eusociality appears to have originated from at least three independent origins in halictid bees. The most advanced of these are species with eusocial colonies; these are characterised by cooperative brood care and
2184-582: Is the largest country in South America, covering a little less than half of the continent's land area and encompassing around half of the continent's population. The remaining countries and territories are divided among four subregions : the Andean states , Caribbean South America , The Guianas , and the Southern Cone . Physiographically, South America also includes some of the nearby islands. The Dutch ABC islands ( Aruba , Bonaire , and Curaçao ),
2275-555: Is thought to have occurred due to rapid climatic cooling around the Eocene-Oligocene boundary , leading to the extinction of some bee lineages such as the tribe Melikertini . Over the Paleogene and Neogene , different bee lineages continued to spread all over the world, and the shifting habitats and connectedness of continents led to the isolation and evolution of many new bee tribes. The oldest non-compression bee fossil
2366-519: The Altiplano , Tiahuanaco or Tiwanaku (100 BC – 1200 AD, Bolivia) managed a large commercial network based on religion. Around the 7th century, both Tiahuanaco and Wari or Huari Empire (600–1200, Central and northern Peru) expanded its influence to all the Andean region, imposing the Huari urbanism and Tiahuanaco religious iconography. The Muisca were the main indigenous civilization in what
2457-586: The Archipelago of San Andrés, Providencia and Santa Catalina ( San Andrés Island , Providencia Island , and Santa Catalina Island etc., which are administered by Colombia ) are politically parts of South American countries but physiographically parts of North America. Other islands often associated with geopolitical South America are the Chiloé Archipelago and Robinson Crusoe Island (both administered by Chile), Easter Island (culturally
2548-820: The Atacama Desert ; the wettest place on earth, López de Micay in Colombia; the largest rainforest, the Amazon rainforest ; the highest capital city, La Paz , Bolivia ; the highest commercially navigable lake in the world, Lake Titicaca ; and, excluding research stations in Antarctica , the world's southernmost permanently inhabited community, Puerto Toro , Chile. South America's major mineral resources are gold, silver, copper, iron ore, tin , and petroleum. These resources have brought high income to its countries, especially in times of war or of rapid economic growth by industrialized countries elsewhere. However,
2639-717: The Bering Land Bridge (now the Bering Strait ) at least 15,000 years ago from the territory that is present-day Russia. They migrated south through North America, and eventually reached South America through the Isthmus of Panama . Amongst the oldest evidence for human presence in South America is the Monte Verde II site in Chile, suggested to date to around 14,500 years ago. From around 13,000 years ago,
2730-634: The Caribbean colonies and Brazil, as European nations built up economically slave-dependent colonies in the New World . Nearly 40% of all African slaves trafficked to the Americas went to Brazil. An estimated 4.9 million slaves from Africa came to Brazil during the period from 1501 to 1866. In contrast to other European colonies in the Americas which mainly used the labor of African slaves, Spanish colonists mainly enslaved indigenous Americans. In 1750,
2821-541: The Early Cretaceous (about 124 million years ago) on the supercontinent of West Gondwana , just prior to its breakup into South America and Africa . The supercontinent is thought to have been a largely xeric environment at this time; modern bee diversity hotspots are also in xeric and seasonal temperate environments, suggesting strong niche conservatism among bees ever since their origins. Genomic analysis indicates that despite only appearing much later in
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2912-1030: The Falkland Islands and South Georgia and the South Sandwich Islands ; and one internal territory : French Guiana . The ABC islands of the Dutch Caribbean (north of Venezuela in the Leeward Antilles ), Ascension Island (a part of Saint Helena, Ascension and Tristan da Cunha ), Bouvet Island (a dependency of Norway ), Panama , and the Caribbean island country of Trinidad and Tobago may also be considered parts of South America. South America has an area of 17,840,000 square kilometers (6,890,000 sq mi). Its population as of 2021 has been estimated at more than 434 million. South America ranks fourth in area (after Asia, Africa, and North America) and fifth in population (after Asia, Africa, Europe, and North America). Brazil
3003-470: The Fishtail projectile point style became widespread across South America, with its disappearance around 11,000 years ago coincident with the disappearance of South America's megafauna. Maize was present in northern South America by around 6,000 years ago. By 2000 BC, many agrarian communities had been settled throughout the Andes and the surrounding regions. Fishing became a widespread practice along
3094-472: The Fortaleza area, contemporaneous with the pyramids of Ancient Egypt , one of the oldest-known civilizations in the Americas and one of the six sites where civilization separately originated in the ancient world. Caral–Supe governing class established a trade network and developed agriculture then followed by Chavín by 900 BC, according to some estimates and archaeological finds. Artifacts were found at
3185-715: The Maracaibo Lake and the mouth of the Orinoco, predominates an equatorial climate of the type Congolese, that also includes parts of the Brazilian territory. The east-central Brazilian plateau has a humid and warm tropical climate. The northern and eastern parts of the Argentine pampas have a humid subtropical climate with dry winters and humid summers of the Chinese type, while the western and eastern ranges have
3276-599: The Orinoco Llanos and in the Guianas Plateau , the precipitation levels go from moderate to high. The Pacific coast of Colombia and northern Ecuador are rainy regions, with Chocó in Colombia being the rainiest place in the world along with the northern slopes of Indian Himalayas. The Atacama Desert, along this stretch of coast, is one of the driest regions in the world. The central and southern parts of Chile are subject to extratropical cyclones , and most of
3367-547: The Portuguese Crown abolished the enslavement of indigenous peoples in colonial Brazil , under the belief that they were unfit for labor and less effective than enslaved Africans. Enslaved Africans were brought to the Americas on slave ships , under inhuman conditions and ill-treatment, and those who survived were sold in slave markets . After independence, all South American countries maintained slavery for some time. The first South American country to abolish slavery
3458-738: The Portuguese Kingdom of Brazil , Dom Pedro I (also Pedro IV of Portugal), son of the Portuguese King Dom João VI , proclaimed the independent Kingdom of Brazil in 1822, which later became the Empire of Brazil . Despite the Portuguese loyalties of garrisons in Bahia , Cisplatina and Pará , independence was diplomatically accepted by Portugal in 1825, on condition of a high compensation paid by Brazil mediated by
3549-665: The South American Plate . South America is home to several superlatives, including the world's highest uninterrupted waterfall , Angel Falls in Venezuela; the highest single-drop waterfall Kaieteur Falls in Guyana; the largest river by volume, the Amazon River ; the longest mountain range, the Andes (whose highest mountain is Aconcagua at 6,962 m or 22,841 ft); the driest non-polar place on earth,
3640-613: The Spanish American wars of independence between the patriots, who promoted such autonomy, and the royalists , who supported Spanish authority over the Americas. The Juntas, in both Spain and the Americas, promoted the ideas of the Enlightenment . Five years after the beginning of the war, Ferdinand VII returned to the throne and began the Absolutist Restoration , as the royalists got the upper hand in
3731-707: The Stenotritidae ), and by the end of the Cretaceous, South American bees had also colonized North America. The North American fossil taxon Cretotrigona belongs to a group that is no longer found in North America, suggesting that many bee lineages went extinct during the Cretaceous-Paleogene extinction event . Following the K-Pg extinction, surviving bee lineages continued to spread into
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3822-712: The native populations had no immune resistance – caused large-scale depopulation of the native population under Spanish control. Systems of forced labor, such as the haciendas and mining industry's mit'a also contributed to the depopulation. After this, enslaved Africans , who had developed immunities to these diseases, were quickly brought in to replace them. The Spaniards were committed to converting their native subjects to Christianity and were quick to purge any native cultural practices that hindered this end; however, many initial attempts at this were only partially successful, as native groups simply blended Catholicism with their established beliefs and practices. Furthermore,
3913-692: The Americas. The continent is generally delimited on the northwest by the Darién watershed along the Colombia–Panama border , although some may consider the border instead to be the Panama Canal . Geopolitically and geographically, all of Panama – including the segment east of the Panama Canal in the isthmus – is typically included in North America alone and among the countries of Central America . Almost all of mainland South America sits on
4004-692: The Argentine Patagonia is desert . In the Pampas of Argentina, Uruguay and South of Brazil the rainfall is moderate, with rains well distributed during the year. The moderately dry conditions of the Chaco oppose the intense rainfall of the eastern region of Paraguay. In the semiarid coast of the Brazilian Northeast the rains are linked to a monsoon regime. Important factors in the determination of climates are sea currents, such as
4095-748: The Breton type and, already in Tierra del Fuego , cold climate of the Siberian type. The distribution of rainfall is related to the regime of winds and air masses. In most of the tropical region east of the Andes, winds blowing from the northeast, east and southeast carry moisture from the Atlantic, causing abundant rainfall. However, due to a consistently strong wind shear and a weak Intertropical Convergence Zone , South Atlantic tropical cyclones are rare. In
4186-588: The Bumblebee Specialist Group to review the threat status of all bumblebee species worldwide using the IUCN Red List criteria. There are many more species of primitively eusocial than highly eusocial bees, but they have been studied less often. Most are in the family Halictidae , or "sweat bees". Colonies are typically small, with a dozen or fewer workers, on average. Queens and workers differ only in size, if at all. Most species have
4277-597: The Eocene-Oligocene boundary, around 34 Mya, of the Florissant shale. The Halictidae first appear in the Early Eocene with species found in amber. The Stenotritidae are known from fossil brood cells of Pleistocene age. The earliest animal-pollinated flowers were shallow, cup-shaped blooms pollinated by insects such as beetles , so the syndrome of insect pollination was well established before
4368-413: The Northern Hemisphere, colonizing Europe from Africa by the Paleocene , and then spreading east to Asia . This was facilitated by the warming climate around the same time, allowing bees to move to higher latitudes following the spread of tropical and subtropical habitats. By the Eocene (~45 mya) there was already considerable diversity among eusocial bee lineages. A second extinction event among bees
4459-447: The South Sandwich Islands ( biogeographically and hydrologically associated with Antarctica) have been administered as two British Overseas Territories under the Crown , whose sovereignty over the islands is disputed by Argentina . An isolated volcanic island on the South American Plate , Ascension Island is geologically a part of South America. Administered as a dependency of Saint Helena, Ascension and Tristan da Cunha ,
4550-830: The Spaniards brought their language to the degree they did with their religion, although the Roman Catholic Church 's evangelization in Quechua , Aymara , and Guaraní actually contributed to the continuous use of these native languages albeit only in the oral form. Eventually, the natives and the Spaniards interbred, forming a mestizo class. At the beginning, many mestizos of the Andean region were offspring of Amerindian mothers and Spanish fathers. After independence, most mestizos had native fathers and European or mestizo mothers. Many native artworks were considered pagan idols and destroyed by Spanish explorers; this included many gold and silver sculptures and other artifacts found in South America, which were melted down before their transport to Spain or Portugal. Spaniards and Portuguese brought
4641-483: The Spanish and Portuguese colonies. First, Napoleon invaded Portugal, but the House of Braganza avoided capture by escaping to Brazil . Napoleon captured King Ferdinand VII of Spain, and appointed his own brother instead. This appointment provoked popular resistance, which created Juntas to rule in the name of the captured king. Many cities in the Spanish colonies, however, considered themselves equally authorized to appoint local Juntas like those of Spain. This began
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#17327873430054732-423: The United Kingdom. The newly independent nations began a process of fragmentation, with several civil and international wars. However, it was not as strong as in Central America. Some countries created from provinces of larger countries stayed as such up to modern times (such as Paraguay or Uruguay), while others were reconquered and reincorporated into their former countries (such as the Republic of Entre Ríos and
4823-488: The appearance of being social. Large groups of solitary bee nests are called aggregations , to distinguish them from colonies . In some species, multiple females share a common nest, but each makes and provisions her own cells independently. This type of group is called "communal" and is not uncommon. The primary advantage appears to be that a nest entrance is easier to defend from predators and parasites when multiple females use that same entrance regularly. The life cycle of
4914-789: The bee families is based on Hedtke et al., 2013, which places the former families Dasypodaidae and Meganomiidae as subfamilies inside the Melittidae. English names, where available, are given in parentheses. Melittidae (inc. Dasypodainae , Meganomiinae ) at least 50 Mya [REDACTED] Apidae (inc. honeybees, cuckoo bees, carpenter bees) ≈87 Mya [REDACTED] Megachilidae (mason, leafcutter bees) ≈50 Mya [REDACTED] Andrenidae (mining bees) ≈34 Mya [REDACTED] Halictidae (sweat bees) ≈50 Mya [REDACTED] Colletidae (plasterer bees) ≈25 Mya [REDACTED] Stenotritidae (large Australian bees) ≈2 Mya [REDACTED] Bees differ from closely related groups such as wasps by having branched or plume-like setae (hairs), combs on
5005-617: The best known insects. Their colonies are established by swarms , consisting of a queen and several thousand workers. There are 29 subspecies of one of these species, Apis mellifera , native to Europe, the Middle East, and Africa. Africanized bees are a hybrid strain of A. mellifera that escaped from experiments involving crossing European and African subspecies; they are extremely defensive. Stingless bees are also highly eusocial . They practise mass provisioning , with complex nest architecture and perennial colonies also established via swarming. Many bumblebees are eusocial, similar to
5096-437: The coast, helping establish fish as a primary source of food. Irrigation systems were also developed at this time, which aided in the rise of an agrarian society . South American cultures began domesticating llamas , vicuñas , guanacos , and alpacas in the highlands of the Andes circa 3500 BC. Besides their use as sources of meat and wool, these animals were used for transportation of goods. The rise of plant growing and
5187-461: The concentration in producing one, or few, major export commodities has often hindered the development and diversification of its economies. The fluctuation in the price of commodities in international markets has led historically to major highs and lows, booms and busts, in the economies of South American states, often causing political instability. This has led for calls to diversify production and increase trade within South America itself. Brazil
5278-424: The conflict. The independence of South America was secured by Simón Bolívar (Venezuela) and José de San Martín (Argentina), the two most important Libertadores . Bolívar led a great uprising in the north, then led his army south towards Lima , the capital of the Viceroyalty of Peru . Meanwhile, San Martín led an army across the Andes Mountains, along with Chilean expatriates, and liberated Chile. He organized
5369-405: The current Humboldt and Falklands . The equatorial current of the South Atlantic strikes the coast of the Northeast and there is divided into two others: the current of Brazil and a coastal current that flows to the northwest towards the Antilles , where there it moves towards northeast course thus forming the most Important and famous ocean current in the world, the Gulf Stream . South America
5460-401: The decline in wild bees has increased the value of pollination by commercially managed hives of honey bees. The analysis of 353 wild bee and hoverfly species across Britain from 1980 to 2013 found the insects have been lost from a quarter of the places they inhabited in 1980. Human beekeeping or apiculture ( meliponiculture for stingless bees) has been practised for millennia, since at least
5551-427: The end of the Pleistocene around 11,000 years ago as part of the Quaternary extinction event , among the highest of any continent, with the casualties including saber-toothed cats , ground sloths , glyptodonts , gomphotheres , the equines Hippidion and Equus neogeus , and all remaining South American native ungulates . South America is thought to have been first inhabited by humans when people were crossing
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#17327873430055642-417: The eusocial Vespidae such as hornets in that the queen initiates a nest on her own rather than by swarming. Bumblebee colonies typically have from 50 to 200 bees at peak population, which occurs in mid to late summer. Nest architecture is simple, limited by the size of the pre-existing nest cavity, and colonies rarely last more than a year. In 2011, the International Union for Conservation of Nature set up
5733-480: The family Crabronidae , which were predators of other insects. The switch from insect prey to pollen may have resulted from the consumption of prey insects which were flower visitors and were partially covered with pollen when they were fed to the wasp larvae. This same evolutionary scenario may have occurred within the vespoid wasps, where the pollen wasps evolved from predatory ancestors. Based on phylogenetic analysis, bees are thought to have originated during
5824-422: The females emerge. Solitary bees are very unlikely to sting (only in self-defense, if ever), and some (esp. in the family Andrenidae ) are stingless. While solitary, females each make individual nests. Some species, such as the European mason bee Hoplitis anthocopoides , and the Dawson's Burrowing bee , Amegilla dawsoni, are gregarious, preferring to make nests near others of the same species, and giving
5915-605: The first appearance of bees. The novelty is that bees are specialized as pollination agents, with behavioral and physical modifications that specifically enhance pollination, and are the most efficient pollinating insects. In a process of coevolution , flowers developed floral rewards such as nectar and longer tubes, and bees developed longer tongues to extract the nectar. Bees also developed structures known as scopal hairs and pollen baskets to collect and carry pollen. The location and type differ among and between groups of bees. Most species have scopal hairs on their hind legs or on
6006-470: The forelimbs for cleaning their antennae, small anatomical differences in limb structure, and the venation of the hind wings; and in females, by having the seventh dorsal abdominal plate divided into two half-plates. Bees have the following characteristics: The largest species of bee is thought to be Wallace's giant bee Megachile pluto , whose females can attain a length of 39 millimetres (1.54 in). The smallest species may be dwarf stingless bees in
6097-407: The fossil record, all modern bee families had already diverged from one another by the end of the Cretaceous. The Melittidae , Apidae , and Megachilidae had already evolved on the supercontinent prior to its fragmentation. Further divergences were facilitated by West Gondwana's breakup around 100 million years ago, leading to a deep Africa-South America split within both the Apidae and Megachilidae,
6188-502: The great city of Cusco , the Inca civilization dominated the Andes region from 1438 to 1533. Known as Tawantin suyu , and "the land of the four regions", in Quechua , the Inca Empire was highly distinct and developed. Inca rule extended to nearly a hundred linguistic or ethnic communities, some nine to fourteen million people connected by a 25,000 kilometer road system . Cities were built with precise, unmatched stonework, constructed over many levels of mountain terrain. Terrace farming
6279-464: The group consists of a mother (the queen ) and her daughters ( workers ). When the castes are purely behavioural alternatives, with no morphological differentiation other than size, the system is considered primitively eusocial, as in many paper wasps ; when the castes are morphologically discrete, the system is considered highly eusocial. True honey bees (genus Apis , of which eight species are currently recognized) are highly eusocial, and are among
6370-417: The island is geopolitically a part of Africa. All of the world's major climate zones are present in South America. The distribution of the average temperatures in the region presents a constant regularity from the 30° of latitude south , when the isotherms tend, more and more, to be confused with the degrees of latitude. In temperate latitudes, winters and summers are milder than in North America. This
6461-487: The islands of Trinidad and Tobago ( Trinidad Island and Tobago Island etc.), the State of Nueva Esparta , and the Federal Dependencies of Venezuela sit on the northern portion of the South American continental shelf and are sometimes considered parts of the continent. Geopolitically, all the island countries and territories in the Caribbean have generally been grouped as a subregion of North America instead. By contrast, Aves Island (administered by Venezuela ) and
6552-656: The isolation of the Melittidae in Africa, and the origins of the Colletidae , Andrenidae and Halictidae in South America. The rapid radiation of the South American bee families is thought to have followed the concurrent radiation of flowering plants in the same region. Later in the Cretaceous (80 million years ago), colletid bees colonized Australia from South America (with an offshoot lineage evolving into
6643-534: The land outside Europe should be an exclusive duopoly between the two countries. The treaty established an imaginary line along a north–south meridian 370 leagues west of the Cape Verde Islands , roughly 46° 37' W. In terms of the treaty, all land to the west of the line (known to comprise most of the South American soil) would belong to Spain, and all land to the east, to Portugal. As accurate measurements of longitude were impossible at that time,
6734-476: The line was not strictly enforced, resulting in a Portuguese expansion of Brazil across the meridian. Beginning in the 1530s, the people and natural resources of South America were repeatedly exploited by foreign conquistadors , first from Spain and later from Portugal. These competing colonial nations claimed the land and resources as their own and divided it into colonies. European infectious diseases ( smallpox , influenza , measles , and typhus ) – to which
6825-661: The lineage as incertae sedis within the Aculeata . The Allodapini (within the Apidae) appeared around 53 Mya. The Colletidae appear as fossils only from the late Oligocene (~25 Mya) to early Miocene . The Melittidae are known from Palaeomacropis eocenicus in the Early Eocene . The Megachilidae are known from trace fossils (characteristic leaf cuttings) from the Middle Eocene . The Andrenidae are known from
6916-458: The mid-17th century. It became a Dutch colony in 1667. The indigenous peoples of the Americas in various European colonies were forced to work in European plantations and mines; along with enslaved Africans who were also introduced in the proceeding centuries via the slave trade . European colonists were heavily dependent on indigenous labor during the initial phases of settlement to maintain
7007-501: The multiple (at least nine) evolutions of eusociality within Hymenoptera . Haplodiploidy is neither necessary nor sufficient for eusociality. Some eusocial species such as termites are not haplodiploid. Conversely, all bees are haplodiploid but not all are eusocial, and among eusocial species many queens mate with multiple males, creating half-sisters that share only 25% of each other's genes. But, monogamy (queens mating singly)
7098-565: The necessary adaptations to do this. However, certain wasp species such as pollen wasps have similar behaviours, and a few species of bee scavenge from carcases to feed their offspring. Solitary bees are important pollinators; they gather pollen to provision their nests with food for their brood. Often it is mixed with nectar to form a paste-like consistency. Some solitary bees have advanced types of pollen-carrying structures on their bodies. Very few species of solitary bee are being cultured for commercial pollination. Most of these species belong to
7189-443: The other members of Hymenoptera bees are haplodiploid ; the sex of a bee is determined by whether or not the egg is fertilised. After mating, a female stores the sperm, and determines which sex is required at the time each individual egg is laid, fertilised eggs producing female offspring and unfertilised eggs, males. Tropical bees may have several generations in a year and no diapause stage. South America South America
7280-477: The planet that contains insect-pollinated flowering plants . The most common bees in the Northern Hemisphere are the Halictidae , or sweat bees, but they are small and often mistaken for wasps or flies. Bees range in size from tiny stingless bee species, whose workers are less than 2 millimetres (0.08 in) long, to the leafcutter bee Megachile pluto , the largest species of bee, whose females can attain
7371-421: The subsequent appearance of permanent human settlements allowed for the multiple and overlapping beginnings of civilizations in South America. One of the earliest known South American civilizations was Caral–Supe , on the central Peruvian coast. Though a pre-ceramic culture, the monumental architecture of Caral–Supe created one of the first cities of the world, generally dated to around 3500 BC, at Huaricanga in
7462-411: The subsistence economy, and natives were often captured by expeditions. The importation of African slaves began midway through the 16th century, but the enslavement of indigenous peoples continued well into the 17th and 18th centuries. The Atlantic slave trade brought enslaved Africans primarily to South American colonies, beginning with the Portuguese since 1502. The main destinations of this phase were
7553-545: The superfamily Apoidea . They are currently considered a clade , called Anthophila . There are over 20,000 known species of bees in seven recognized biological families . Some species – including honey bees , bumblebees , and stingless bees – live socially in colonies while most species (>90%) – including mason bees , carpenter bees , leafcutter bees , and sweat bees – are solitary. Bees are found on every continent except Antarctica , in every habitat on
7644-492: The times of Ancient Egypt and Ancient Greece . Bees have appeared in mythology and folklore, through all phases of art and literature from ancient times to the present day, although primarily focused in the Northern Hemisphere where beekeeping is far more common. In Mesoamerica , the Mayans have practiced large-scale intensive meliponiculture since pre-Columbian times. The immediate ancestors of bees were stinging wasps in
7735-604: The tribe Meliponini whose workers are less than 2 millimetres (0.08 in) in length. According to inclusive fitness theory, organisms can gain fitness not just through increasing their own reproductive output, but also that of close relatives. In evolutionary terms, individuals should help relatives when Cost < Relatedness * Benefit . The requirements for eusociality are more easily fulfilled by haplodiploid species such as bees because of their unusual relatedness structure. In haplodiploid species, females develop from fertilized eggs and males from unfertilized eggs. Because
7826-501: The underside of their abdomens. Some species in the family Apidae have pollen baskets on their hind legs, while very few lack these and instead collect pollen in their crops. The appearance of these structures drove the adaptive radiation of the angiosperms , and, in turn, bees themselves. Bees coevolved not only with flowers but it is believed that some species coevolved with mites. Some provide tufts of hairs called acarinaria that appear to provide lodgings for mites; in return, it
7917-573: The western European architectural style to the continent, and helped to improve infrastructures like bridges, roads, and the sewer system of the cities they discovered or conquered. They also significantly increased economic and trade relations, not just between the old and new world but between the different South American regions and peoples. Finally, with the expansion of the Portuguese and Spanish languages, many cultures that were previously separated became united through that of Latin American. Guyana
8008-472: Was Chile in 1823, Uruguay in 1830, Bolivia in 1831, Guyana in 1833, Colombia and Ecuador in 1851, Argentina in 1853, Peru and Venezuela in 1854, Suriname in 1863, Paraguay in 1869, and in 1888 Brazil was the last South American nation and the last country in western world to abolish slavery. The European Peninsular War (1807–14), a theater of the Napoleonic Wars , changed the political situation of
8099-633: Was a useful form of agriculture. The Mapuche in Central and Southern Chile resisted the European and Chilean settlers, waging the Arauco War for more than 300 years. In 1494, Portugal and Spain, the two great maritime European powers of that time, on the expectation of new lands being discovered in the west, signed the Treaty of Tordesillas , by which they agreed, with the support of the Pope, that all
8190-562: Was initially colonized by the Dutch before coming under British control , though there was a brief period during the Napoleonic Wars when it was occupied by the French . The region was initially partitioned between the Dutch, French and British before fully coming under the control of Britain. Suriname was first explored by the Spanish in the 16th century and then settled by the English in
8281-895: Was not included in this analysis. Further studies by Sann et al. , 2018, elevated the subfamilies (plus one tribe and one subtribe) of Crabronidae sensu lato to family status. They also recovered the placement of Heterogyna within Nyssonini and sunk Heterogynaidae . The newly erected family, Ammoplanidae , formerly a subtribe of Pemphredoninae , was recovered as the most sister family to bees. Ampulicidae (Cockroach wasps) [REDACTED] Astatidae [REDACTED] Bembicidae [REDACTED] Sphecidae ( sensu stricto ) [REDACTED] Crabronidae ( sensu stricto ) [REDACTED] Mellinidae [REDACTED] Pemphredonidae [REDACTED] Philanthidae [REDACTED] Psenidae [REDACTED] Ammoplanidae Anthophila (bees) [REDACTED] This cladogram of
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