97-469: The boomslang ( / ˈ b oʊ m s l ɑː ŋ / or / ˈ b ɔː m s l ə ŋ / ; Dispholidus typus ) is a highly venomous snake in the family Colubridae . The species is native to Sub-Saharan Africa . Its common name means "tree snake" in Dutch and Afrikaans – boom meaning "tree", and slang meaning "snake". In Afrikaans , the name is pronounced [ˈbuəmslaŋ] . The boomslang
194-581: A Bantu cultural core. DNA evidence has found that the Swahili people were of mixed African and Asian (particularly Persian) ancestry. The Kilwa Sultanate was a medieval sultanate centred at Kilwa , in modern-day Tanzania. At its height, its authority stretched over the entire length of the Swahili Coast , including Kenya. Since the 10th century, rulers of Kilwa would go on to build elaborate coral mosques and introduce copper coinage. Swahili,
291-566: A Bantu language with Arabic , Persian , and other Middle-Eastern and South Asian loanwords , later developed as a lingua franca for trade between the different peoples. Since the turn of the 20th century, Swahili has adopted numerous loanwords and calques from English, many of them originating during English colonial rule. The Swahili built Mombasa into a major port city and established trade links with other nearby city-states, as well as commercial centres in Persia, Arabia, and even India. By
388-421: A bite, sting, or similar action. The toxin is delivered through a specially evolved venom apparatus , such as fangs or a stinger , in a process called envenomation . Venom is often distinguished from poison , which is a toxin that is passively delivered by being ingested, inhaled, or absorbed through the skin, and toxungen , which is actively transferred to the external surface of another animal via
485-428: A chemical arms race. Predator/prey pairs are expected to coevolve over long periods of time. As the predator capitalizes on susceptible individuals, the surviving individuals are limited to those able to evade predation. Resistance typically increases over time as the predator becomes increasingly unable to subdue resistant prey. The cost of developing venom resistance is high for both predator and prey. The payoff for
582-596: A colonist's perspective is found in the memoir Out of Africa by Danish author Baroness Karen von Blixen-Finecke, published in 1937. By the 1930s, approximately 30,000 white settlers lived in the area and gained a political voice because of their contribution to the market economy. The central highlands were already home to over a million members of the Kikuyu people, most of whom had no land claims in European terms and lived as itinerant farmers. To protect their interests,
679-516: A fatal dose. He made notes on the symptoms he experienced almost to the end. D. S. Chapman reported eight serious envenomations by boomslangs between 1919 and 1962, two of which were lethal. Boomslang monovalent antivenom was developed during the 1940s. The South African Vaccine Producers manufactures a monovalent antivenom for use in boomslang envenomations. Treatment of bites may also require complete blood transfusions , especially if over 24-48 hours have passed without antivenom. The boomslang
776-555: A few hours without any noticeable effects, victims of boomslang bites may falsely believe that their attack was simply a dry or bluff strike. The pathophysiological mechanisms of the venom are different with every snake, resulting in different clinical manifestations with every patient. An adult boomslang has 1.6 to 8 mg of venom. Its median lethal dose (LD 50 ) in mice is 0.1 mg/kg (intravenously). 0.071 mg/kg (IV) has also been reported. 12.5 mg/kg ( subcutaneously ) and 1.3–1.8 mg/kg ( intraperitoneal ). Based on
873-560: A genus other than Dispholidus . The average adult boomslang is 100–160 centimetres (3.3–5.2 ft) in total length (including tail). Some exceed 183 centimetres (6.00 ft). The eyes are exceptionally large, and the head has a characteristic egg-like shape. Colouration is highly variable. Males are light green with black or blue scale edges, but adult females may be brown demonstrating sexual dimorphism . Weight varies from 175 to 510 g (0.386 to 1.124 lb), with an average weight of 299.4 g (0.660 lb). In this species,
970-541: A great amount of land, in the plains where people did not put up much resistance. The Nandi peoples managed to oppose the Masai, while the Taveta peoples fled to the forests on the eastern edge of Mount Kilimanjaro , although they later were forced to leave the land due to the threat of smallpox . An outbreak of either rinderpest or pleuropneumonia greatly affected the Masai's cattle, while an epidemic of smallpox affected
1067-625: A physical delivery mechanism. Venom has evolved in terrestrial and marine environments and in a wide variety of animals: both predators and prey, and both vertebrates and invertebrates . Venoms kill through the action of at least four major classes of toxin, namely necrotoxins and cytotoxins , which kill cells; neurotoxins , which affect nervous systems; myotoxins , which damage muscles; and haemotoxins , which disrupt blood clotting . Venomous animals cause tens of thousands of human deaths per year. Venoms are often complex mixtures of toxins of differing types. Toxins from venom are used to treat
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#17327933027291164-541: A protective mucus that acts as a chemical camouflage or macromolecular mimicry preventing "not self" recognition by the sea anemone and nematocyst discharge. Clownfish may acclimate their mucus to resemble that of a specific species of sea anemone. Kenya Kenya , officially the Republic of Kenya ( Swahili : Jamhuri ya Kenya ), is a country in East Africa . With a population of more than 47.6 million in
1261-464: A sting. In bees and wasps , the stinger is a modified ovipositor (egg-laying device). In Polistes fuscatus , the female continuously releases a venom that contains a sex pheromone that induces copulatory behavior in males. In wasps such as Polistes exclamans , venom is used as an alarm pheromone, coordinating a response from the nest and attracting nearby wasps to attack the predator. In some species, such as Parischnogaster striatula , venom
1358-470: A wide range of diseases, explored in over 5,000 scientific papers. In medicine, snake venom proteins are used to treat conditions including thrombosis , arthritis , and some cancers . Gila monster venom contains exenatide , used to treat type 2 diabetes . Solenopsins extracted from fire ant venom has demonstrated biomedical applications, ranging from cancer treatment to psoriasis . A branch of science, venomics , has been established to study
1455-422: A wide range of medical conditions including thrombosis , arthritis , and some cancers . Studies in venomics are investigating the potential use of venom toxins for many other conditions. The use of venom across a wide variety of taxa is an example of convergent evolution . It is difficult to conclude exactly how this trait came to be so intensely widespread and diversified. The multigene families that encode
1552-414: Is a colubrid snake within the subfamily Colubrinae . It belongs to the genus Dispholidus , which contains two other species, D. pembae and D. punctatus . The boomslang is thought to be closely related to members of the genera Thelotornis , Thrasops , Rhamnophis , and Xyelodontophis , with which it forms the taxonomic tribe Dispholidini . Close relationships can be shown in
1649-400: Is a timid snake, and bites generally occur only when people attempt to handle, catch, pursue or kill the animal. When confronted and cornered, it inflates its neck and assumes an S-shaped striking pose, a key indicator of any snake species feeling threatened. Venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of
1746-539: Is applied all over the body as an antimicrobial protection. Many caterpillars have defensive venom glands associated with specialized bristles on the body called urticating hairs . These are usually merely irritating, but those of the Lonomia moth can be fatal to humans. Bees synthesize and employ an acidic venom ( apitoxin ) to defend their hives and food stores, whereas wasps use a chemically different venom to paralyse prey, so their prey remains alive to provision
1843-534: Is found in a few other reptiles such as the Mexican beaded lizard , the gila monster , and some monitor lizards, including the Komodo dragon . Mass spectrometry showed that the mixture of proteins present in their venom is as complex as the mixture of proteins found in snake venom. Some lizards possess a venom gland; they form a hypothetical clade, Toxicofera , containing the suborders Serpentes and Iguania and
1940-459: Is higher. Rattlesnakes have responded locally by increasing the effectiveness of their venom. The kingsnakes of the Americas are constrictors that prey on many venomous snakes. They have evolved resistance which does not vary with age or exposure. They are immune to the venom of snakes in their immediate environment, like copperheads, cottonmouths, and North American rattlesnakes, but not to
2037-638: Is not restricted to trees and can often be found on the ground hunting, feeding, or taking shelter. It will occasionally hide underground when the weather is harsh. The boomslang is oviparous , and an adult female can produce up to 30 eggs, which are deposited in a hollow tree trunk or rotting log. The eggs have a relatively long (3 months on average) incubation period . Male hatchlings are grey with blue speckles, and female hatchlings are pale brown. They attain their adult colouration after several years. Hatchlings are about 20 cm (7.9 in) in length and pose no threat to humans, but are dangerously venomous by
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#17327933027292134-834: Is now Kenya and southwestern Somalia, from 1889 to 1907. Other important cities include Kisumu and Nakuru . Kenya is bordered by South Sudan to the northwest, Ethiopia to the north, Somalia to the east, Uganda to the west, Tanzania to the south, and the Indian Ocean to the southeast. Kenya's geography , climate and population vary widely, ranging from cold snow-capped mountaintops (Batian, Nelion and Point Lenana on Mount Kenya ) with vast surrounding forests, wildlife and fertile agricultural regions to temperate climates in western and rift valley counties and further on to dry less fertile arid and semi-arid areas and absolute deserts ( Chalbi Desert and Nyiri Desert ). Kenya's earliest inhabitants were hunter-gatherers , like
2231-460: Is proven that reptile and platypus venom have independently evolved, it is thought that there are certain protein structures that are favored to evolve into toxic molecules. This provides more evidence of why venom has become a homoplastic trait and why very different animals have convergently evolved. Envenomation resulted in 57,000 human deaths in 2013, down from 76,000 deaths in 1990. Venoms, found in over 173,000 species, have potential to treat
2328-533: Is slightly compressed. The dorsal scales , which are arranged in 19 or 21 rows, are very narrow, oblique, strongly keeled, with apical pits . The tail is long, and the subcaudal scales are paired. Ventral scales are 164–201; the anal plate is divided; and the subcaudals are 91–131. The boomslang is endemic to Sub-Saharan Africa , from The Gambia , Guinea , Senegal and most of Western Africa (including Benin , Cameroon , Ghana , Nigeria , Togo ) through Central and Eastern Africa ( Democratic Republic of
2425-427: Is slow-acting, symptoms may not become apparent until many hours after a bite. Although the absence of symptoms provides sufficient time for procuring antivenom , it can also provide victims with false reassurances, leading to underestimation of the seriousness of the bite. Snakes of any species may sometimes fail to inject venom when they bite (a so-called "dry bite" or "bluff strike", enacted in-defense), wherein, after
2522-538: Is the second largest in eastern and central Africa, after Ethiopia, with Nairobi serving as a major regional commercial hub. Agriculture is the largest sector; tea and coffee are traditional cash crops, while fresh flowers are a fast-growing export. The service industry is also a major economic driver, particularly tourism. Kenya is a member of the East African Community trade bloc, though some international trade organisations categorise it as part of
2619-1006: The Cnidaria , sea urchins among the Echinodermata , and cone snails and cephalopods , including octopuses , among the Molluscs . Venom is found in some 200 cartilaginous fishes, including stingrays , sharks , and chimaeras ; the catfishes (about 1000 venomous species); and 11 clades of spiny-rayed fishes ( Acanthomorpha ), containing the scorpionfishes (over 300 species), stonefishes (over 80 species), gurnard perches , blennies , rabbitfishes , surgeonfishes , some velvetfishes , some toadfishes , coral crouchers , red velvetfishes , scats , rockfishes , deepwater scorpionfishes , waspfishes , weevers , and stargazers . Some salamanders can extrude sharp venom-tipped ribs. Two frog species in Brazil have tiny spines around
2716-540: The Greater Horn of Africa . Africa is Kenya's largest export market, followed by the European Union. The Republic of Kenya is named after Mount Kenya . The earliest recorded version of the modern name was written by German explorer Johann Ludwig Krapf in the 19th century. While travelling with a Kamba caravan led by the long-distance trader Chief Kivoi , Krapf spotted the mountain peak and asked what it
2813-638: The Kalenjin , Samburu , Luo , Turkana , and Maasai . By the first millennium AD, Bantu -speaking farmers had moved into the region, initially along the coast. The Bantus originated in West Africa along the Benue River in what is now eastern Nigeria and western Cameroon . The Bantu migration brought new developments in agriculture and ironworking to the region. Bantu groups in Kenya include
2910-606: The Kikuyu , Luhya , Kamba , Kisii , Meru , Kuria , Aembu , Ambeere , Wadawida -Watuweta, Wapokomo, and Mijikenda , among others. Notable prehistoric sites in the interior of Kenya include the (possibly archaeoastronomical) site Namoratunga on the west side of Lake Turkana and the walled settlement of Thimlich Ohinga in Migori County . The Kenyan coast had served as host to communities of ironworkers and Bantu subsistence farmers, hunters, and fishers who supported
3007-639: The Ming Dynasty , visited the East African coast on one of his last ' treasure voyages '. Malindi authorities also welcomed the Portuguese explorer Vasco da Gama in 1498. During the 18th and 19th century C.E, the Masai people moved into central and southern Rift Valley plains of Kenya, from a region north of Lake Rudolf (now Lake Turkana ). Although there were not many, they managed to conquer
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3104-524: The Pleistocene epoch. During excavations at Lake Turkana in 1984, palaeoanthropologist Richard Leakey , assisted by Kamoya Kimeu , discovered the Turkana Boy , a 1.6-million-year-old Homo erectus fossil. East Africa, including Kenya, is one of the earliest regions where modern humans ( Homo sapiens ) are believed to have lived. Evidence was found in 2018, dating to about 320,000 years ago, of
3201-707: The Somali Republic to the north. A ceasefire was eventually reached with the signing of the Arusha Memorandum in October 1967, but relative insecurity prevailed through 1969. To discourage further invasions, Kenya signed a defence pact with Ethiopia in 1969, which is still in effect. On 12 December 1964, the Republic of Kenya was proclaimed, and Jomo Kenyatta became Kenya's first president. Under Kenyatta, corruption became widespread throughout
3298-424: The cladogram below: Oligodon Thrasops flavigularis Thrasops jacksonii Thrasops capensis Dispholidus typus , Boomslang Thelotornis kirtlandii Hapsidophrys Philothamnus Two subspecies are recognised, including the nominotypical subspecies . The trinomial authority in parentheses for D. t. typus indicates that the subspecies was originally described in
3395-439: The phosphodiester bonds of DNA ; and neurotoxins, which disrupt signalling in the nervous system. Snake venom causes symptoms including pain, swelling, tissue necrosis, low blood pressure, convulsions, haemorrhage (varying by species of snake), respiratory paralysis, kidney failure, coma, and death. Snake venom may have originated with duplication of genes that had been expressed in the salivary glands of ancestors. Venom
3492-410: The 15th century, Portuguese voyager Duarte Barbosa claimed that "Mombasa is a place of great traffic and has a good harbour in which there are always moored small craft of many kinds and also great ships, both of which are bound from Sofala and others which come from Cambay and Melinde and others which sail to the island of Zanzibar." In the 17th century, the Swahili coast was conquered and came under
3589-470: The 1963 independence constitution with a new one. As a result, the elections of 2007 took place following the procedure set by the old constitution. Kibaki was reelected in highly contested elections marred by political and ethnic violence . The main opposition leader, Raila Odinga, claimed the election was rigged and that he was the rightfully elected president. In the ensuing violence, 1,500 people were killed and another 600,000 internally displaced, making it
3686-415: The 2002 elections. Moi's plan to be replaced by Uhuru Kenyatta failed, and Mwai Kibaki , running for the opposition coalition "National Rainbow Coalition" ( NARC ), was elected president. David Anderson (2003) reports the elections were judged free and fair by local and international observers, and seemed to mark a turning point in Kenya's democratic evolution. In 2005, Kenyans rejected a plan to replace
3783-550: The 2019 census, Kenya is the 28th-most-populous country in the world and 7th most populous in Africa. Kenya's capital and largest city is Nairobi , while its oldest and second-largest city, is the major port city of Mombasa , situated on Mombasa Island in the Indian Ocean and the surrounding mainland. Mombasa was the capital of the British East Africa Protectorate , which included most of what
3880-739: The Air Force. It was quickly suppressed by forces commanded by Chief of General Staff Mahamoud Mohamed , a veteran Somali military official. They included the General Service Unit (GSU)—a paramilitary wing of the police—and later the regular police. On the heels of the Garissa Massacre of 1980, Kenyan troops committed the Wagalla massacre in 1984 against thousands of civilians in Wajir County . An official probe into
3977-623: The Congo , western Ethiopia , Kenya , South Sudan , Tanzania , Uganda ). It is found across much of Southern Africa , in a wide array of habitats, with some of the species' densest populations being in Angola , Botswana , Eswatini , Malawi , Mozambique , Namibia , South Africa , Zambia and Zimbabwe . The boomslang is an excellent climber and is highly arboreal, living mainly in forested areas. D. typus lives in karoo shrubs, savannahs , lowland forests, and in grasslands. The boomslang
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4074-513: The Embu call it "Kirinyaa". All three names have the same meaning. Ludwig Krapf recorded the name as both Kenia and Kegnia . Some have said that this was a precise notation of the African pronunciation / ˈ k ɛ n j ə / . An 1882 map drawn by Joseph Thompsons, a Scottish geologist and naturalist, indicated Mt. Kenya as Mt. Kenia. The mountain's name was accepted, pars pro toto , as
4171-522: The Masai themselves. After the death of the Masai Mbatian , the chief laibon (medicine man), the Masai split into warring factions. The Masai caused much strife in the areas they conquered; however, cooperation between such groups as the Luo people , Luhya people , and Gusii people is evidenced by shared vocabulary for modern implements and similar economic regimes. Although Arab traders remained in
4268-825: The Protectorate of Kenya so that all of Kenya would become one sovereign state. In this way, Kenya became an independent country under the Kenya Independence Act 1963 of the United Kingdom. On 12 December 1964, Kenya became a republic under the name "Republic of Kenya". Concurrently, the Kenyan army fought the Shifta War against ethnic Somali rebels inhabiting the Northern Frontier District who wanted to join their kin in
4365-561: The UK and the colony led to the Mau Mau revolution , which began in 1952, and the declaration of independence in 1963. After independence, Kenya remained a member of the Commonwealth of Nations . The current constitution was adopted in 2010 and replaced the 1963 independence constitution . Kenya is a presidential representative democratic republic, in which elected officials represent
4462-473: The United Kingdom. Kenya itself was the site of fighting between Allied forces and Italian troops in 1940–41, when Italian forces invaded. Wajir and Malindi were bombed as well. From October 1952 to December 1959, Kenya was in a state of emergency arising from the Mau Mau rebellion against British rule. The Mau Mau, also known as the Kenya Land and Freedom Army, were primarily Kikuyu people. During
4559-466: The area, trade routes were disrupted by the hostile Masai, though there was trade in ivory between these factions. The first foreigners to successfully get past the Masai were Johann Ludwig Krapf and Johannes Rebmann , two German missionaries who established a mission in Rabai , not too far from Mombasa . The pair were the first Europeans to sight Mount Kenya . The colonial history of Kenya dates from
4656-598: The article by stating the cost of the stability in terms of human rights abuses. The opposition party started by Oginga Odinga — Kenya People's Union (KPU)—was banned in 1969 after the Kisumu Massacre and KPU leaders were still in detention without trial in gross violation of the U.N. Declaration of Human Rights . The Kenya Students Union, Jehovah Witnesses and all opposition parties were outlawed. Kenyatta ruled until his death on 22 August 1978. After Kenyatta died, Daniel arap Moi became president. He retained
4753-453: The atrocities was later ordered in 2011. The election held in 1988 saw the advent of the mlolongo (queuing) system, where voters were supposed to line up behind their favoured candidates instead of casting a secret ballot. This was seen as the climax of a very undemocratic regime and led to widespread agitation for constitutional reform. Several contentious clauses, including the one that allowed for only one political party, were changed in
4850-475: The beginning of the end of Moi's leadership and the rule of KANU. Moi retained the presidency and George Saitoti became vice president. Although it held on to power, KANU won 100 seats and lost 88 seats to the six opposition parties. The 1992 elections marked the beginning of multiparty politics after more than 25 years of KANU rule. Following skirmishes in the aftermath of the elections, 5,000 people were killed and another 75,000 displaced from their homes. In
4947-541: The boomslang brumates for short periods, often curling up inside the enclosed nest of a weaverbird . Many venomous species of the Colubridae are harmless to humans—unless one has a known rare allergy—largely because of small venom glands and inefficient teeth that are situated at the back of the mouth. These species, including the boomslang, are collectively known as "rear-fanged" (or opisthoglyphous ) snakes, as their venom-injecting teeth are situated farther back in
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#17327933027295044-561: The boomslang is primarily a hemotoxin ; it works via a process in which many small clots form in the blood, causing the victim’s circulatory system to improperly coagulate, resulting in excessive bleeding and death. The venom has been observed to cause haemorrhaging in tissues such as muscle and the brain (among other organs), while, at the same time, clogging capillaries with tiny blood clots. Other signs and symptoms include headache , nausea , sleepiness, and confusion, leading to cardiac arrest and unconsciousness. Because boomslang venom
5141-573: The bulk of the skilled labour required for construction. They and most of their descendants later remained in Kenya and formed the core of several distinct Indian communities, such as the Ismaili Muslim and Sikh communities. While building the railway through Tsavo , a number of the Indian railway workers and local African labourers were attacked by two lions known as the Tsavo maneaters . At
5238-546: The centuries, the Kenyan coast has played host to many merchants and explorers. Among the cities that line the Kenyan coast is Malindi. It has remained an important Swahili settlement since the 14th century and once rivalled Mombasa for dominance in the African Great Lakes region. Malindi has traditionally been a friendly port city for foreign powers. In 1414, the Chinese trader and explorer Zheng He , representing
5335-585: The coast and the interior between 250 BC and 500 AD. European contact began in 1500 AD with the Portuguese Empire , and effective colonisation of Kenya began in the 19th century during the European exploration of Africa . Modern-day Kenya emerged from a protectorate established by the British Empire in 1895 and the subsequent Kenya Colony , which began in 1920. Numerous disputes between
5432-614: The coastal hotel business, with Kenyatta personally owning the Leonard Beach Hotel. Kenyatta's mixed legacy was highlighted at the 10-year anniversary of Kenya's independence. A December 1973 article in The New York Times praised Kenyatta's leadership and Kenya for emerging as a model of pragmatism and conservatism. Kenya's GDP had increased at an annual rate of 6.6%, higher than the population growth rate of more than 3%. But Amnesty International responded to
5529-606: The colonial administration's crackdown, over 11,000 freedom fighters had been killed, along with 100 British troops and 2,000 Kenyan loyalist soldiers. War crimes were committed on both sides of the conflict, including the publicised Lari massacre and the Hola massacre . The governor requested and obtained British and African troops, including the King's African Rifles . The British began counter-insurgency operations. In May 1953, General Sir George Erskine took charge as commander-in-chief of
5626-847: The colony's armed forces, with the personal backing of Winston Churchill . The capture of Waruhiu Itote (nom de guerre "General China" ) on 15 January 1954 and the subsequent interrogation led to a better understanding of the Mau Mau command structure for the British. Operation Anvil opened on 24 April 1954, after weeks of planning by the army with the approval of the War Council. The operation effectively placed Nairobi under military siege. Nairobi's occupants were screened and suspected Mau Mau supporters moved to detention camps. More than 80,000 Kikuyu were held in detention camps without trial, often subject to brutal treatment. The Home Guard formed
5723-430: The constitution to allow Moi to remain president for another term. Subsequently, Moi stood for reelection and won a 5th term in 1997. His win was strongly criticised by his major opponents, Kibaki and Odinga , as fraudulent. Following this win, Moi was constitutionally barred from another presidential term. Beginning in 1998, he attempted to influence the country's succession politics to have Uhuru Kenyatta elected in
5820-524: The core of the government's strategy as it was composed of loyalist Africans, not foreign forces such as the British Army and King's African Rifles. The capture of Dedan Kimathi on 21 October 1956 in Nyeri signified the ultimate defeat of the Mau Mau and essentially ended the military offensive. During this period, substantial governmental changes to land tenure occurred. The most important of these
5917-473: The cost of physiological resistance is an increased chance of survival for prey, but it allows predators to expand into underutilised trophic niches. The California ground squirrel has varying degrees of resistance to the venom of the Northern Pacific rattlesnake . The resistance involves toxin scavenging and depends on the population. Where rattlesnake populations are denser, squirrel resistance
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#17327933027296014-479: The country. The building of the railway was resisted by some ethnic groups—notably the Nandi , led by Orkoiyot Koitalel Arap Samoei from 1890 to 1900—but the British eventually built it. The Nandi were the first ethnic group to be put in a native reserve to stop them from disrupting the building of the railway. During the railway construction era, there was a significant influx of Indian workers, who provided
6111-406: The crown of their skulls which, on impact, deliver venom into their targets. Some 450 species of snake are venomous. Snake venom is produced by glands below the eye (the mandibular glands ) and delivered to the target through tubular or channeled fangs. Snake venoms contain a variety of peptide toxins, including proteases , which hydrolyze protein peptide bonds; nucleases , which hydrolyze
6208-588: The direct rule of the Omani Arabs , who expanded the slave trade to meet the demands of plantations in Oman and Zanzibar . Initially, these traders came mainly from Oman, but later many came from Zanzibar (such as Tippu Tip ). In addition, the Portuguese started buying slaves from the Omani and Zanzibari traders in response to the interruption of the transatlantic slave trade by British abolitionists. Throughout
6305-441: The early emergence of modern behaviours , including long-distance trade networks (involving goods such as obsidian), the use of pigments, and the possible making of projectile points. The authors of three 2018 studies on the site suggest that complex and modern behaviours had already begun in Africa around the time of the emergence of Homo sapiens . The first inhabitants of present-day Kenya were hunter-gatherer groups, akin to
6402-411: The economy with agriculture, fishing, metal production, and trade with foreign countries. These communities formed the earliest city-states in the region, which were collectively known as Azania . By the 1st century CE, many of the city-states such as Mombasa , Malindi , and Zanzibar began to establish trading relations with Arabs . This led to the increased economic growth of the Swahili states,
6499-710: The establishment of a German Empire protectorate over the Sultan of Zanzibar 's coastal possessions in 1885, followed by the arrival of the Imperial British East Africa Company in 1888. Imperial rivalry was prevented by the Heligoland–Zanzibar Treaty , Germany handed its East African coastal holdings to Britain in 1890. The transfer by Germany to Britain was followed by the building of the Uganda Railway passing through
6596-483: The families Varanidae , Anguidae , and Helodermatidae . Euchambersia , an extinct genus of therocephalians , is hypothesized to have had venom glands attached to its canine teeth. A few species of living mammals are venomous, including solenodons , shrews , the European mole , vampire bats , male platypuses , and slow lorises . Shrews have venomous saliva and most likely evolved their trait similarly to snakes. The presence of tarsal spurs akin to those of
6693-431: The following years. In 1991, Kenya transitioned to a multiparty political system after 26 years of single-party rule. On 28 October 1992, Moi dissolved parliament, five months before the end of his term. As a result, preparations began for all elective seats in parliament as well as the president. The election was scheduled to take place on 7 December 1992, but delays led to its postponement to 29 December. Apart from KANU,
6790-521: The food chambers of their young. The use of venom is much more widespread than just these examples; many other insects, such as true bugs and many ants , also produce venom. The ant species Polyrhachis dives uses venom topically for the sterilisation of pathogens. There are venomous invertebrates in several phyla , including jellyfish such as the dangerous box jellyfish , the Portuguese man-of-war (a siphonophore) and sea anemones among
6887-515: The government, civil service, and business community. Kenyatta and his family were tied up with this corruption as they enriched themselves through the mass purchase of property after 1963. Their acquisitions in the Central, Rift Valley, and Coast Provinces aroused great anger among landless Kenyans. His family used his presidential position to circumvent legal or administrative obstacles to acquiring property. The Kenyatta family also heavily invested in
6984-431: The head is distinct from the neck and the canthus rostralis is distinct. The pupil of the very large eye is round. The boomslang has excellent eyesight and often moves its head from side to side to get a better view of objects directly in front. The maxillary teeth are small anteriorly, seven or eight in number, followed by three very large, grooved fangs situated below each eye. The mandibular teeth are subequal. The body
7081-518: The introduction of Islam , Arabic influences on the Swahili Bantu language , cultural diffusion , as well as the Swahili city-states becoming members of a larger trade network. Many historians had long believed that the city-states were established by Arab or Persian traders, but archaeological evidence has led scholars to recognise the city-states as an indigenous development which, though subjected to foreign influence due to trade, retained
7178-606: The land, capturing British supplies, and remaining undefeated. He eventually surrendered in Northern Rhodesia (today Zambia) 14 days after the Armistice was signed in 1918. To chase von Lettow, the British deployed the British Indian Army troops from India but needed large numbers of porters to overcome the formidable logistics of transporting supplies far into the interior on foot. The Carrier Corps
7275-637: The many toxins that they contain; some venoms are complex mixtures of toxins of differing types. Major classes of toxin in venoms include: Venom is widely distributed taxonomically, being found in both invertebrates and vertebrates, in aquatic and terrestrial animals, and among both predators and prey. The major groups of venomous animals are described below. Venomous arthropods include spiders , which use fangs on their chelicerae to inject venom , and centipedes , which use forcipules — modified legs — to deliver venom, while scorpions and stinging insects inject venom with
7372-636: The modern Khoisan speakers. These people were later largely replaced by agropastoralist Cushitic (ancestral to Kenya's Cushitic speakers) from the Horn of Africa . During the early Holocene , the regional climate shifted from dry to wetter conditions, providing an opportunity for the development of cultural traditions such as agriculture and herding , in a more favourable environment. Around 500 BC, Nilotic -speaking pastoralists (ancestral to Kenya's Nilotic speakers) started migrating from present-day southern Sudan into Kenya. Nilotic groups in Kenya include
7469-414: The mouth than elapids or vipers , and thus require the snake to bite, hold-on, and "chew" the venom into its victim. However, many opisthoglyphous snakes, including the boomslang, actually possess a highly potent and deadly venom, regardless of their teeth being placed further back in the mouth; the boomslang is able to open its jaws up to 170° when biting, enabling sufficient envenomation . The venom of
7566-571: The name of the country. It did not come into widespread official use during the early colonial period, when the country was referred to as the East African Protectorate . The official name was changed to the Colony of Kenya in 1920. Hominids such as Homo habilis (1.8 to 2.5 million years ago) and Homo erectus (1.9 million to 350,000 years ago) are possible direct ancestors of modern Homo sapiens , and lived in Kenya in
7663-468: The next five years, many political alliances were formed in preparation for the next elections. In 1994, Jaramogi Oginga Odinga died and several coalitions joined his FORD Kenya party to form a new party, United National Democratic Alliance. This party was plagued with disagreements. In 1995, Richard Leakey formed the Safina party, but it was denied registration until November 1997. In 1996, KANU revised
7760-608: The outbreak of World War I in August 1914, the governors of British East Africa (as the protectorate was generally known) and German East Africa initially agreed on a truce in an attempt to keep the young colonies out of direct hostilities. But Lieutenant Colonel Paul von Lettow-Vorbeck , the German military commander, determined to tie down as many British resources as possible. Completely cut off from Germany, Lettow-Vorbeck conducted an effective guerrilla warfare campaign , living off
7857-617: The people and the president is the head of state and government. Kenya is a member of the United Nations , the Commonwealth , World Bank , International Monetary Fund , World Trade Organization , COMESA , International Criminal Court , as well as other international organisations. It is also a major non-NATO ally of the United States . With a GNI of 1,840, Kenya is a lower-middle-income economy. Kenya's economy
7954-443: The platypus in many non- therian Mammaliaformes groups suggests that venom was an ancestral characteristic among mammals. Extensive research on platypuses shows that their toxin was initially formed from gene duplication, but data provides evidence that the further evolution of platypus venom does not rely as much on gene duplication as was once thought. Modified sweat glands are what evolved into platypus venom glands. Although it
8051-521: The present-day Hadza people . According to archaeological dating of associated artifacts and skeletal material, Cushitic speakers first settled in Kenya's lowlands between 3,200 and 1,300 BC, a phase known as the Lowland Savanna Pastoral Neolithic . Nilotic -speaking pastoralists (ancestral to Kenya's Nilotic speakers) began migrating from present-day South Sudan into Kenya around 500 BC. Bantu people settled at
8148-416: The presidency, running unopposed in elections held in 1979, 1983 ( snap elections ), and 1988, all of which were held under the single-party constitution. The 1983 elections were held a year early, and were a direct result of a failed military coup on 2 August 1982. The 1982 coup was masterminded by a low-ranking Air Force serviceman, Senior Private Hezekiah Ochuka , and was staged mainly by enlisted men of
8245-414: The proteins associated with venom and how individual components of venom can be used for pharmaceutical means. Venom is used as a trophic weapon by many predator species. The coevolution between predators and prey is the driving force of venom resistance, which has evolved multiple times throughout the animal kingdom. The coevolution between venomous predators and venom-resistant prey has been described as
8342-404: The ruling party, other parties represented in the elections included FORD Kenya and FORD Asili. This election was marked by large-scale intimidation of opponents and harassment of election officials. It resulted in an economic crisis propagated by ethnic violence as the president was accused of rigging electoral results to retain power. This election was a turning point for Kenya as it signified
8439-403: The settlers banned the growing of coffee and introduced a hut tax, and the landless were granted less and less land in exchange for their labour. A massive exodus to the cities ensued as their ability to make a living from the land dwindled. By the 1950s, there were 80,000 white settlers living in Kenya. Throughout World War II , Kenya was an important source of manpower and agriculture for
8536-527: The tentacles of venomous sea anemones (an obligatory symbiosis for the fish), and are resistant to their venom. Only 10 known species of anemones are hosts to clownfish and only certain pairs of anemones and clownfish are compatible. All sea anemones produce venoms delivered through discharging nematocysts and mucous secretions. The toxins are composed of peptides and proteins. They are used to acquire prey and to deter predators by causing pain, loss of muscular coordination, and tissue damage. Clownfish have
8633-576: The time they reach a length around 45 cm (18 in) and a girth as thick as an adult's smallest finger. D. typus is diurnal and almost exclusively arboreal . It is reclusive, and moves from branch to branch when pursued by anything too large to eat. Its diet includes chameleons and other arboreal lizards , frogs , and occasionally small mammals , birds , and eggs from nesting birds and reptiles, all of which it swallows whole. The boomslang will also feed on other snakes, including cannibalising members of its own species. During cool weather,
8730-403: The toxins of venomous animals are actively selected , creating more diverse toxins with specific functions. Venoms adapt to their environment and victims, evolving to become maximally efficient on a predator 's particular prey (particularly the precise ion channels within the prey). Consequently, venoms become specialized to an animal's standard diet. Venoms cause their biological effects via
8827-433: The venom of, for example, king cobras or black mambas. Among marine animals, eels are resistant to sea snake venoms, which contain complex mixtures of neurotoxins, myotoxins, and nephrotoxins, varying according to species. Eels are especially resistant to the venom of sea snakes that specialise in feeding on them, implying coevolution; non-prey fishes have little resistance to sea snake venom. Clownfish always live among
8924-423: The very low venom quantities produced by D. typus , and the very serious effects found in a good part of the reported cases in humans, it has been suggested that the venom's LD 50 is lower in humans than in mice, with only 2 to 3 mg being enough to potentially kill a healthy adult. In 1957, herpetologist Karl Schmidt died after being bitten by a juvenile boomslang, which he had doubted could produce
9021-594: The willingness of Somalis in Kenya to join Somalia . The result of the referendum showed that 86% of Somalis in Kenya wanted to join Somalia , yet the British colonial administration rejected the result and the Somalis remained in Kenya. The first direct elections for native Kenyans to the Legislative Council took place in 1957. Despite British hopes of handing power to "moderate" local rivals, it
9118-439: Was called. Kivoi told him " Kĩ-Nyaa " or " Kĩlĩma- Kĩinyaa ", probably because the pattern of black rock and white snow on its peaks reminded him of the feathers of the male ostrich. In archaic Kikuyu , the word 'nyaga' or more commonly 'manyaganyaga' is used to describe an extremely bright object. The Agikuyu, who inhabit the slopes of Mt. Kenya, call it Kĩrĩma Kĩrĩnyaga (literally 'the mountain with brightness') in Kikuyu , while
9215-512: Was formed and ultimately mobilised over 400,000 Africans, contributing to their long-term politicisation. In 1920, the East Africa Protectorate was turned into a colony and renamed Kenya after its highest mountain. During the early part of the 20th century, the interior central highlands were settled by British and other European farmers, who became wealthy farming coffee and tea. One depiction of this period of change from
9312-536: Was the Kenya African National Union (KANU) of Jomo Kenyatta that formed a government. The Colony of Kenya and the Protectorate of Kenya each came to an end on 12 December 1963, with independence conferred on all of Kenya. The U.K. ceded sovereignty over the Colony of Kenya. The Sultan of Zanzibar agreed that simultaneous with independence for the colony, he would cease to have sovereignty over
9409-549: Was the Swynnerton Plan , which was used to both reward loyalists and punish Mau Mau. This left roughly 1/3rd of Kikuyu bereft of any tenancy land arrangement and thus propertyless at the time of independence. Before Kenya got its independence, Somali ethnic people in present-day Kenya in the areas of Northern Frontier Districts petitioned Her Majesty's Government not to be included in Kenya. The colonial government decided to hold Kenya's first referendum in 1962 to check
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