142-532: Brachyopterus is a genus of prehistoric eurypterid of the family Rhenopteridae . It is one of the earliest known eurypterids, having been recovered from Middle Ordovician deposits in Montgomeryshire , Wales . Though other species have been assigned to it in the past, Brachyopterus is today recognized as containing one valid species, B. stubblefieldi . Brachyopterus is distinguished by its small size, compound eyes with axes converging anteriorly on
284-578: A cosmopolitan distribution . Though the eurypterids continued to be abundant and diversify during the Early Devonian (for instance leading to the evolution of the pterygotid Jaekelopterus , the largest of all arthropods), the group was one of many heavily affected by the Late Devonian extinction . The extinction event, only known to affect marine life (particularly trilobites, brachiopods and reef -building organisms) effectively crippled
426-509: A cuticle composed of proteins and chitin . As in other chelicerates , the body was divided into two tagmata (sections); the frontal prosoma (head) and posterior opisthosoma (abdomen). The prosoma was covered by a carapace (sometimes called the "prosomal shield") on which both compound eyes and the ocelli (simple eye-like sensory organs) were located. The prosoma also bore six pairs of appendages which are usually referred to as appendage pairs I to VI. The first pair of appendages,
568-408: A lung , plastron or a pseudotrachea . Plastrons are organs that some arthropods evolved secondarily to breathe air underwater. This is considered an unlikely explanation since eurypterids had evolved in water from the start and they would not have organs evolved from air-breathing organs present. In addition, plastrons are generally exposed on outer parts of the body while the eurypterid gill tract
710-561: A writing system , built huge pyramids and temples , had a complex calendar , and developed the concept of zero around 400 CE. The first recorded European references to North America are in Norse sagas where it is referred to as Vinland . The earliest verifiable instance of pre-Columbian trans-oceanic contact by any European culture with the North America mainland has been dated to around 1000 CE. The site , situated at
852-461: A dual respiratory system was present, which would have allowed for short periods of time in terrestrial environments. The name Eurypterida comes from the Ancient Greek words εὐρύς ( eurús ), meaning 'broad' or 'wide', and πτερόν ( pterón ), meaning 'wing', referring to the pair of wide swimming appendages present in many members of the group. The eurypterid order includes
994-742: A gait like that of most modern insects. The weight of its long abdomen would have been balanced by two heavy and specialized frontal appendages, and the center of gravity might have been adjustable by raising and positioning the tail. Preserved fossilized eurypterid trackways tend to be large and heteropodous and often have an associated telson drag mark along the mid-line (as with the Scottish Hibbertopterus track). Such trackways have been discovered on every continent except for South America. In some places where eurypterid fossil remains are otherwise rare, such as in South Africa and
1136-416: A manner similar to modern horseshoe crabs, by grabbing and shredding food with their appendages before pushing it into their mouth using their chelicerae. Fossils preserving digestive tracts have been reported from fossils of various eurypterids, among them Carcinosoma , Acutiramus and Eurypterus . Though a potential anal opening has been reported from the telson of a specimen of Buffalopterus , it
1278-425: A meter (1.64 ft) even if the extended chelicerae are not included. Two other eurypterids have also been estimated to have reached lengths of 2.5 metres; Erettopterus grandis (closely related to Jaekelopterus ) and Hibbertopterus wittebergensis , but E. grandis is very fragmentary and the H. wittenbergensis size estimate is based on trackway evidence, not fossil remains. The family of Jaekelopterus ,
1420-597: A part of Denmark) and Mexico classified as Latin American). It is unknown with certainty how and when first human populations first reached North America. People were known to live in the Americas at least 20,000 years ago, but various evidence points to possibly earlier dates. The Paleo-Indian period in North America followed the Last Glacial Period, and lasted until about 10,000 years ago when
1562-664: A period of mountain building called the Laramide orogeny , between 80 and 55 mya. The formation of the Isthmus of Panama that connected the continent to South America arguably occurred approximately 12 to 15 mya, and the Great Lakes (as well as many other northern freshwater lakes and rivers) were carved by receding glaciers about 10,000 years ago. North America is the source of much of what humanity knows about geologic time periods. The geographic area that would later become
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#17327979741381704-590: A powerful earthquake killed seven people in 2009. Volcanic eruptions are common in the region. In 1968, the Arenal Volcano , in Costa Rica, erupted and killed 87 people. Fertile soils from weathered volcanic lavas have made it possible to sustain dense populations in agriculturally productive highland areas. Central America has many mountain ranges ; the longest are the Sierra Madre de Chiapas ,
1846-471: A rowing type of propulsion similar to that of crabs and water beetles . Larger individuals may have been capable of underwater flying (or subaqueous flight ) in which the motion and shape of the paddles are enough to generate lift , similar to the swimming of sea turtles and sea lions . This type of movement has a relatively slower acceleration rate than the rowing type, especially since adults have proportionally smaller paddles than juveniles. However, since
1988-408: A subtrapezoid to subpentagonal prosoma (head). All of its legs are walking legs; the first three pairs are short with spines, except when modified into clasping organs; the last two pairs are moderately long, keeled and tapering in width to terminal claws. The last leg falls short of the penultimate abdominal segment. The abdomen is narrow and ends in a short styliform telson . Brachyopterus date from
2130-540: A variety of reactions, including curiosity, trading, cooperation, resignation, and resistance. The indigenous population declined substantially following European arrival, primarily due to the introduction of Eurasian diseases, such as smallpox , to which the indigenous peoples lacked immunity, and because of violent conflicts with Europeans. Indigenous culture changed significantly and their affiliation with political and cultural groups also changed. Several linguistic groups died out , and others changed quite quickly. On
2272-553: Is a genital appendage. This appendage, an elongated rod with an internal duct, is found in two distinct morphs, generally referred to as "type A" and "type B". These genital appendages are often preserved prominently in fossils and have been the subject of various interpretations of eurypterid reproduction and sexual dimorphism. Type A appendages are generally longer than those of type B. In some genera they are divided into different numbers of sections, such as in Eurypterus where
2414-495: Is a lightweight build. Factors such as locomotion, energy costs in molting and respiration, as well as the actual physical properties of the exoskeleton , limit the size that arthropods can reach. A lightweight construction significantly decreases the influence of these factors. Pterygotids were particularly lightweight, with most fossilized large body segments preserving as thin and unmineralized. Lightweight adaptations are present in other giant paleozoic arthropods as well, such as
2556-665: Is a very large continent that extends from north of the Arctic Circle to south of the Tropic of Cancer . Greenland, along with the Canadian Shield , is tundra with average temperatures ranging from 10 to 20 °C (50 to 68 °F), but central Greenland is composed of a very large ice sheet. This tundra radiates throughout Canada, but its border ends near the Rocky Mountains (but still contains Alaska) and at
2698-529: Is also possible and the structure may represent the unfused tips of the appendages. Located between the dorsal and ventral surfaces of the Blattfüsse associated with the type A appendages is a set of organs traditionally described as either "tubular organs" or "horn organs". These organs are most often interpreted as spermathecae (organs for storing sperm ), though this function is yet to be proven conclusively. In arthropods, spermathecae are used to store
2840-524: Is an ancient craton which forms the geologic core of North America; it formed between 1.5 and 1.0 billion years ago during the Proterozoic eon. The Canadian Shield is the largest exposure of this craton. From the Late Paleozoic to Early Mesozoic eras, North America was joined with the other modern-day continents as part of the supercontinent Pangaea , with Eurasia to its east. One of
2982-546: Is in present-day Panama at the Darien Gap on the Colombia -Panama border, placing almost all of Panama within North America. Alternatively, some geologists physiographically locate its southern limit at the Isthmus of Tehuantepec , Mexico, with Central America extending southeastward to South America from this point. The Caribbean islands, or West Indies, are considered part of North America. The continental coastline
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#17327979741383124-733: Is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian , from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though
3266-489: Is located behind the Blattfüssen . Instead, among arthropod respiratory organs, the eurypterid gill tracts most closely resemble the pseudotracheae found in modern isopods . These organs, called pseudotracheae, because of some resemblance to the tracheae (windpipes) of air-breathing organisms, are lung-like and present within the pleopods (back legs) of isopods. The structure of the pseudotracheae has been compared to
3408-648: Is long and irregular. The Gulf of Mexico is the largest body of water indenting the continent, followed by Hudson Bay . Others include the Gulf of Saint Lawrence and the Gulf of California . Before the Central American isthmus formed, the region had been underwater. The islands of the West Indies delineate a submerged former land bridge , which had connected North and South America via what are now Florida and Venezuela . There are several islands off
3550-487: Is made up of the first six exoskeleton segments fused together into a larger structure. The seventh segment (thus the first opisthosomal segment) is referred to as the metastoma and the eighth segment (distinctly plate-like) is called the operculum and contains the genital aperature. The underside of this segment is occupied by the genital operculum, a structure originally evolved from ancestral seventh and eighth pair of appendages. In its center, as in modern horseshoe crabs,
3692-521: Is more likely that the anus was opened through the thin cuticle between the last segment before the telson and the telson itself, as in modern horseshoe crabs. Eurypterid coprolites discovered in deposits of Ordovician age in Ohio containing fragments of a trilobite and eurypterid Megalograptus ohioensis in association with full specimens of the same eurypterid species have been suggested to represent evidence of cannibalism . Similar coprolites referred to
3834-544: Is much more of a marine influence in many of the sections yielding Adelophthalmus than has previously been acknowledged." Similarly, a study of the eurypterid Hibbertopterus from the Carboniferous of New Mexico concluded that the habitat of some eurypterids "may need to be re-evaluated". The sole surviving eurypterine family, Adelophthalmidae, was represented by only a single genus, Adelophthalmus . The hibbertopterids, mycteroptids and Adelophthalmus survived into
3976-561: Is part of North America geographically. In a geologic sense, Bermuda is not part of the Americas, but an oceanic island that was formed on the fissure of the Mid-Atlantic Ridge over 100 million years ago (mya). The nearest landmass to it is Cape Hatteras , North Carolina . However, Bermuda is often thought of as part of North America, especially given its historical, political and cultural ties to Virginia and other parts of
4118-638: Is possible that many eurypterid species thought to be distinct actually represent juvenile specimens of other species, with paleontologists rarely considering the influence of ontogeny when describing new species. Studies on a well-preserved fossil assemblage of eurypterids from the Pragian -aged Beartooth Butte Formation in Cottonwood Canyon , Wyoming , composed of multiple specimens of various developmental stages of eurypterids Jaekelopterus and Strobilopterus , revealed that eurypterid ontogeny
4260-707: Is the Morrison Formation of the western U.S. Canada is geologically one of the oldest regions in the world, with more than half of the region consisting of Precambrian rocks that have been above sea level since the beginning of the Palaeozoic era. Canada's mineral resources are diverse and extensive. Across the Canadian Shield and in the north there are large iron, nickel, zinc , copper, gold, lead, molybdenum , and uranium reserves. Large diamond concentrations have been recently developed in
4402-527: Is the humid subtropical climate . This area has the wettest cities in the contiguous U.S. , with annual precipitation reaching 67 in (1,700 mm) in Mobile, Alabama . Stretching from the borders of the humid continental and subtropical climates, and going west to the Sierra Nevada , south to the southern tip of Durango , north to the border with tundra climate, the steppe / desert climates are
Brachyopterus - Misplaced Pages Continue
4544-439: Is the first record of land locomotion by a eurypterid. The trackway provides evidence that some eurypterids could survive in terrestrial environments, at least for short periods of time, and reveals information about the stylonurine gait. In Hibbertopterus , as in most eurypterids, the pairs of appendages are different in size (referred to as a heteropodous limb condition). These differently sized pairs would have moved in phase, and
4686-816: Is the metastoma becoming proportionally less wide. This ontogenetic change has been observed in members of several superfamilies, such as the Eurypteroidea, the Pterygotioidea and the Moselopteroidea . No fossil gut contents from eurypterids are known, so direct evidence of their diet is lacking. The eurypterid biology is particularly suggestive of a carnivorous lifestyle. Not only were many large (in general, most predators tend to be larger than their prey), but they had stereoscopic vision (the ability to perceive depth). The legs of many eurypterids were covered in thin spines, used both for locomotion and
4828-418: Is unlikely the "gill tract" contained functional gills when comparing the organ to gills in other invertebrates and even fish. Previous interpretations often identified the eurypterid "gills" as homologous with those of other groups (hence the terminology), with gas exchange occurring within the spongy tract and a pattern of branchio-cardiac and dendritic veins (as in related groups) carrying oxygenated blood into
4970-454: The Age of Discovery and the early modern period . Present-day cultural and ethnic patterns reflect interactions between European colonists, indigenous peoples , enslaved Africans , immigrants from Europe, Asia, and descendants of these respective groups. Europe's colonization in North America led to most North Americans speaking European languages, such as English , Spanish , and French , and
5112-519: The Archaic period began. The classic stage followed the Archaic period, and lasted from approximately the 6th to 13th centuries. Beginning in 1000 AD, the Norse were the first Europeans to begin exploring and ultimately colonizing areas of North America. In 1492, the exploratory voyages of Christopher Columbus led to a transatlantic exchange , including migrations of European settlers during
5254-567: The Bering Strait during the Early-Middle Holocene . Prior to the arrival of European explorers and colonists in North America, the natives of North America were divided into many different polities, ranging from small bands of a few families to large empires. They lived in several culture areas , which roughly correspond to geographic and biological zones that defined the representative cultures and lifestyles of
5396-600: The British Columbia Coast , Western Canada , and Northern Canada . In the U.S., they include New England , the Mid-Atlantic , South Atlantic states , East North Central states , West North Central states , East South Central states , West South Central states , Mountain states , and Pacific states . The Great Lakes region and the Pacific Northwest include areas in both Canada and
5538-913: The Caribbean Plate , whereas the Juan de Fuca Plate and Cocos Plate border the North American Plate on its western frontier. The continent can be divided into four great regions (each of which contains many subregions): the Great Plains stretching from the Gulf of Mexico to the Canadian Arctic ; the geologically young, mountainous west, including the Rocky Mountains , the Great Basin , California , and Alaska ;
5680-568: The Cordillera Isabelia , and the Cordillera de Talamanca . Between the mountain ranges lie fertile valleys that are suitable for the people; in fact, most of the population of Honduras, Costa Rica, and Guatemala live in valleys. Valleys are also suitable for the production of coffee, beans, and other crops. The indigenous peoples of the Americas have many creation myths , based on which they assert that they have been present on
5822-465: The East Coast to eastern North Dakota , and stretching down to Kansas , is the humid continental climate featuring intense seasons, with a large amount of annual precipitation, with places like New York City averaging 50 in (1,300 mm). Starting at the southern border of the humid continental climate and stretching to the Gulf of Mexico (whilst encompassing the eastern half of Texas)
Brachyopterus - Misplaced Pages Continue
5964-620: The Inuit of the high Arctic arrived in North America much later than other native groups, evidenced by the disappearance of Dorset culture artifacts from the archaeological record and their replacement by the Thule people . During the thousands of years of native habitation on the continent, cultures changed and shifted. One of the oldest yet discovered is the Clovis culture (c. 9550–9050 BCE) in modern New Mexico . Later groups include
6106-719: The Mississippian culture and related Mound building cultures, found in the Mississippi River valley and the Pueblo culture of what is now the Four Corners . The more southern cultural groups of North America were responsible for the domestication of many common crops now used around the world, such as tomatoes, squash , and maize . As a result of the development of agriculture in the south, many other cultural advances were made there. The Mayans developed
6248-532: The Stylonuroidea , Kokomopteroidea and Mycteropoidea as well as eurypterine groups such as the Pterygotioidea, Eurypteroidea and Waeringopteroidea . The most successful eurypterid by far was the Middle to Late Silurian Eurypterus , a generalist , equally likely to have engaged in predation or scavenging . Thought to have hunted mainly small and soft-bodied invertebrates, such as worms , species of
6390-516: The United States . North America covers an area of about 24,709,000 square kilometers (9,540,000 square miles), representing approximately 16.5% of the Earth 's land area and 4.8% of its total surface area. It is the third-largest continent by size after Asia and Africa , and the fourth-largest continent by population after Asia, Africa, and Europe . As of 2021 , North America's population
6532-536: The coxae (limb segments) used for feeding. These appendages were generally walking legs that were cylindrical in shape and were covered in spines in some species. In most lineages, the limbs tended to get larger the farther back they were. In the Eurypterina suborder , the larger of the two eurypterid suborders, the sixth pair of appendages was also modified into a swimming paddle to aid in traversing aquatic environments. The opisthosoma comprised 12 segments and
6674-488: The first official name given to Mexico. North America includes several regions and subregions, each of which have their own respective cultural, economic, and geographic regions. Economic regions include several regions formalized in 20th- and 21st-century trade agreements, including NAFTA between Canada , Mexico , and the United States , and CAFTA between Central America , the Dominican Republic , and
6816-442: The rhizodonts , were the new apex predators in marine environments. However, various recent findings raise doubts about this, and suggest that these eurypterids were euryhaline forms that lived in marginal marine environments, such as estuaries, deltas, lagoons, and coastal ponds. One argument is paleobiogeographical; pterygotoid distribution seems to require oceanic dispersal. A recent review of Adelophthalmoidea admitted that "There
6958-405: The spermatophore received from males. This would imply that the type A appendage is the female morph and the type B appendage is the male. Further evidence for the type A appendages representing the female morph of genital appendages comes in their more complex construction (a general trend for female arthropod genitalia). It is possible that the greater length of the type A appendage means that it
7100-678: The telson , the posteriormost division of the body, which in most species took the form of a blade-like shape. In some lineages, notably the Pterygotioidea , the Hibbertopteridae and the Mycteroptidae , the telson was flattened and may have been used as a rudder while swimming. Some genera within the superfamily Carcinosomatoidea , notably Eusarcana , had a telson similar to that of modern scorpions and may have been capable of using it to inject venom . The coxae of
7242-458: The 1490s, building cities, putting the resident indigenous populations to work, raising crops for Spanish settlers and panning gold to enrich the Spaniards. Much of the indigenous population died due to disease and overwork, spurring the Spaniards on to claim new lands and peoples. An expedition under the command of Spanish settler, Hernán Cortés , sailed westward in 1519 to what turned out to be
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#17327979741387384-630: The Arctic, making Canada one of the world's largest producers. Throughout the Shield, there are many mining towns extracting these minerals. The largest, and best known, is Sudbury , Ontario. Sudbury is an exception to the normal process of forming minerals in the Shield since there is significant evidence that the Sudbury Basin is an ancient meteorite impact crater . The nearby, but less-known Temagami Magnetic Anomaly has striking similarities to
7526-651: The Caribbean islands. France took the western half of Hispaniola and developed Saint-Domingue as a cane sugar producing colony worked by black slave labor. Britain took Barbados and Jamaica , and the Dutch and Danes took islands previously claimed by Spain. Britain did not begin settling on the North American mainland until a hundred years after the first Spanish settlements, since it sought first to control nearby Ireland . The first permanent English settlement
7668-635: The Cod fish", in reference to the abundance of cod fish on the East Coast . Waldseemüller used the Latin version of Vespucci's name, Americus Vespucius, in its feminine form of "America", following the examples of "Europa", "Asia", and "Africa". Americus originated from Medieval Latin Emericus (see Saint Emeric of Hungary ), coming from the Old High German name Emmerich . Map makers later extended
7810-539: The Devonian, large two meter (6.5+ ft) pterygotids such as Acutiramus were already present during the Late Silurian. Their ecology ranged from generalized predatory behavior to ambush predation and some, such as Pterygotus itself, were active apex predators in Late Silurian marine ecosystems. The pterygotids were also evidently capable of crossing oceans, becoming one of only two eurypterid groups to achieve
7952-496: The Middle Ordovician . Kjellesvig-Waering (1966) concluded that "there is no other genus that warrants comparison, or, indeed at our present state of knowledge, reveals any close affinities with this very unusual genus", though the genus was able to be classified as a stylonurine . In later years, further similarities have been noted between Brachyopterus and the rhenopterids . Lamsdell et al. (2010) classified it as
8094-825: The Middle Ordovician suggests that eurypterids either originated during the Early Ordovician and experienced a rapid and explosive radiation and diversification soon after the first forms evolved, or that the group originated much earlier, perhaps during the Cambrian period. As such, the exact eurypterid time of origin remains unknown. Though fossils referred to as "primitive eurypterids" have occasionally been described from deposits of Cambrian or even Precambrian age, they are not recognized as eurypterids, and sometimes not even as related forms, today. Some animals previously seen as primitive eurypterids, such as
8236-485: The Middle Ordovician, 467.3 million years ago . There are also reports of even earlier fossil eurypterids in the Fezouata Biota of Late Tremadocian (Early Ordovician) age in Morocco , but these have yet to be thoroughly studied, and are likely to be peytoiid appendages. Pentecopterus was a relatively derived eurypterid, part of the megalograptid family within the carcinosomatoid superfamily. Its derived position suggests that most eurypterid clades, at least within
8378-483: The Middle Silurian and the Early Devonian, with an absolute peak in diversity during the Pridoli epoch , 423 to 419.2 million years ago, of the very latest Silurian. This peak in diversity has been recognized since the early twentieth century; of the approximately 150 species of eurypterids known in 1916, more than half were from the Silurian and a third were from the Late Silurian alone. Though stylonurine eurypterids generally remained rare and low in number, as had been
8520-419: The North America's southeastern coast, Spanish explorer Juan Ponce de León , who had accompanied Columbus's second voyage, visited and named in 1513 La Florida . As the colonial period unfolded, Spain, England, and France appropriated and claimed extensive territories in North America eastern and southern coastlines. Spain established permanent settlements on the Caribbean islands of Hispaniola and Cuba in
8662-525: The Permian. North America North America is a continent in the Northern and Western Hemispheres . North America is bordered to the north by the Arctic Ocean , to the east by the Atlantic Ocean , to the southeast by South America and the Caribbean Sea , and to the west and south by the Pacific Ocean . The region includes the Bahamas , Bermuda , Canada , the Caribbean , Central America , Clipperton Island , Greenland , Mexico , Saint Pierre and Miquelon , Turks and Caicos Islands , and
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#17327979741388804-494: The Pterygotidae, is noted for several unusually large species. Both Acutiramus , whose largest member A. bohemicus measured 2.1 meters (6.9 ft), and Pterygotus , whose largest species P. grandidentatus measured 1.75 meters (5.7 ft), were gigantic. Several different contributing factors to the large size of the pterygotids have been suggested, including courtship behaviour, predation and competition over environmental resources. Giant eurypterids were not limited to
8946-478: The Stylonurina, this appendage takes the form of a long and slender walking leg, while in the Eurypterina, the leg is modified and broadened into a swimming paddle. Other than the swimming paddle, the legs of many eurypterines were far too small to do much more than allow them to crawl across the sea floor . In contrast, a number of stylonurines had elongated and powerful legs that might have allowed them to walk on land (similar to modern crabs ). A fossil trackway
9088-416: The Sudbury Basin. Its magnetic anomalies are very similar to the Sudbury Basin, and so it could be a second metal-rich impact crater. The Shield is also covered by vast boreal forests that support an important logging industry. The United States can be divided into twelve main geological provinces: Each province has its own geologic history and unique features. The geology of Alaska is typical of that of
9230-410: The U.S. North America occupies the northern portion of the landmass generally referred to as the New World, the Western Hemisphere , the Americas, or simply America, which, in many countries, is considered a single continent with North America a subcontinent . North America is the third-largest continent by area after Asia and Africa . North America's only land connection to South America
9372-429: The United States has been the source of more varieties of dinosaurs than any other modern country. According to paleontologist Peter Dodson, this is primarily due to stratigraphy, climate and geography, human resources, and history. Much of the Mesozoic Era is represented by exposed outcrops in the many arid regions of the continent. The most significant Late Jurassic dinosaur-bearing fossil deposit in North America
9514-550: The United States, and Mexico. France , Italy , Portugal , Spain , Romania , Greece , and the countries of Latin America use a six-continent model , with the Americas viewed as a single continent and North America designating a subcontinent comprising Canada , the United States , Mexico, and Saint Pierre and Miquelon (politically part of France), and often including Greenland and Bermuda . North America has historically been known by other names, including Spanish North America, New Spain , and América Septentrional,
9656-527: The United States. North America is divided linguistically and culturally into two primary regions, Anglo-America and Latin America . Anglo-America includes most of North America, Belize , and Caribbean islands with English -speaking populations. There are also regions, including Louisiana and Quebec , with large Francophone populations; in Quebec , French is the official language. . The southern portion of North America includes Central America and non-English speaking Caribbean nations. The north of
9798-432: The abundance and diversity previously seen within the eurypterids. A major decline in diversity had already begun during the Early Devonian and eurypterids were rare in marine environments by the Late Devonian. During the Frasnian stage four families went extinct, and the later Famennian saw an additional five families going extinct. As marine groups were the most affected, the eurypterids were primarily impacted within
9940-472: The ancient continent of Laurentia , and demersal (living on the seafloor ) and basal animals from the continents Avalonia and Gondwana. The Laurentian predators, classified in the family Megalograptidae (comprising the genera Echinognathus , Megalograptus and Pentecopterus ), are likely to represent the first truly successful eurypterid group, experiencing a small radiation during the Late Ordovician. Eurypterids were most diverse and abundant between
10082-442: The animal in question could possibly have measured just short of 2 meters (6.6 ft) in length. More robust than the pterygotids, this giant Hibbertopterus would possibly have rivalled the largest pterygotids in weight, if not surpassed them, and as such be among the heaviest arthropods. The two eurypterid suborders, Eurypterina and Stylonurina , are distinguished primarily by the morphology of their final pair of appendages. In
10224-454: The appendage via tracts, but these supposed tracts remain unpreserved in available fossil material. Type B appendages, assumed male, would have produced, stored and perhaps shaped spermatophore in a heart-shaped structure on the dorsal surface of the appendage. A broad genital opening would have allowed large amounts of spermatophore to be released at once. The long furca associated with type B appendages, perhaps capable of being lowered like
10366-547: The area had shifted to the Aztec Empire , whose capital city Tenochtitlan was located further north in the Valley of Mexico . The Aztecs were conquered in 1521 by Hernán Cortés . During the so-called Age of Discovery , Europeans explored overseas and staked claims to various parts of North America, much of which was already settled by indigenous peoples. Upon Europeans' arrival in the " New World ", indigenous peoples had
10508-414: The body. The primary analogy used in previous studies has been horseshoe crabs, though their gill structure and that of eurypterids are remarkably different. In horseshoe crabs, the gills are more complex and composed of many lamellae (plates) which give a larger surface area used for gas exchange. In addition, the gill tract of eurypterids is proportionally much too small to support them if it is analogous to
10650-523: The case during the preceding Ordovician, eurypterine eurypterids experienced a rapid rise in diversity and number. In most Silurian fossil beds, eurypterine eurypterids account for 90% of all eurypterids present. Though some were likely already present by the Late Ordovician (simply missing from the fossil record so far), a vast majority of eurypterid groups are first recorded in strata of Silurian age. These include both stylonurine groups such as
10792-420: The coastlines and shallow inland seas of Euramerica. During the Late Silurian the pterygotid eurypterids, large and specialized forms with several new adaptations, such as large and flattened telsons capable of being used as rudders, and large and specialized chelicerae with enlarged pincers for handling (and potentially in some cases killing) prey appeared. Though the largest members of the family appeared in
10934-432: The continent maintains recognized regions as well. In contrast to the common definition of North America, which encompasses the whole North American continent, the term "North America" is sometimes used more narrowly to refer only to four nations, Canada, Greenland, Mexico, and the U.S. The U.S. Census Bureau includes Saint Pierre and Miquelon, but excludes Mexico from its definition. The term Northern America refers to
11076-981: The continent's coasts; principally, the Arctic Archipelago, the Bahamas , Turks and Caicos , the Greater and Lesser Antilles , the Aleutian Islands (some of which are in the Eastern Hemisphere proper), the Alexander Archipelago , the many thousand islands of the British Columbia Coast, and Newfoundland. Greenland , a self-governing Danish island, and the world's largest , is on the same tectonic plate (the North American Plate ) and
11218-599: The continent. The vast majority of North America is on the North American Plate. Parts of western Mexico, including Baja California, and of California , including the cities of San Diego , Los Angeles , and Santa Cruz , lie on the eastern edge of the Pacific Plate , with the two plates meeting along the San Andreas Fault . The southernmost portion of the continent and much of the West Indies lie on
11360-659: The cordillera, while the major islands of Hawaii consist of Neogene volcanics erupted over a hot spot . Central America is geologically active with volcanic eruptions and earthquakes occurring from time to time. In 1976 Guatemala was hit by a major earthquake , killing 23,000 people; Managua, the capital of Nicaragua, was devastated by earthquakes in 1931 and 1972, the last one killing about 5,000 people; three earthquakes devastated El Salvador, one in 1986 and two in 2001; one earthquake devastated northern and central Costa Rica in 2009, killing at least 34 people; in Honduras
11502-748: The cultures of the region commonly reflect Western traditions . However, relatively small parts of North America in Canada, the United States, Mexico, and Central America have indigenous populations that continue adhering to their respective pre-European colonial cultural and linguistic traditions. The Americas were named after the Italian explorer Amerigo Vespucci by German cartographers Martin Waldseemüller and Matthias Ringmann . Vespucci explored South America between 1497 and 1502, and
11644-695: The cuticle) after which they underwent rapid and immediate growth. Some arthropods, such as insects and many crustaceans, undergo extreme changes over the course of maturing. Chelicerates, including eurypterids, are in general considered to be direct developers, undergoing no extreme changes after hatching (though extra body segments and extra limbs may be gained over the course of ontogeny in some lineages, such as xiphosurans and sea spiders ). Whether eurypterids were true direct developers (with hatchlings more or less being identical to adults) or hemianamorphic direct developers (with extra segments and limbs potentially being added during ontogeny) has been controversial in
11786-571: The driest in the U.S. Highland climates cut from north to south of the continent, where subtropical or temperate climates occur just below the tropics, as in central Mexico and Guatemala . Tropical climates appear in the island regions and in the subcontinent's bottleneck, found in countries and states bathed by the Caribbean Sea or to the south of the Gulf of Mexico and the Pacific Ocean . Precipitation patterns vary across
11928-470: The end of the Canadian Shield, near the Great Lakes . Climate west of the Cascade Range is described as being temperate weather with average precipitation 20 inches (510 millimeters). Climate in coastal California is described to be Mediterranean , with average temperatures in cities like San Francisco ranging from 57 to 70 °F (14 to 21 °C) over the course of the year. Stretching from
12070-409: The eurypterine suborder, had already been established at this point during the Middle Ordovician. The earliest known stylonurine eurypterid, Brachyopterus , is also Middle Ordovician in age. The presence of members of both suborders indicates that primitive stem-eurypterids would have preceded them, though these are so far unknown in the fossil record. The presence of several eurypterid clades during
12212-474: The eurypterine suborder. Only one group of stylonurines (the family Parastylonuridae ) went extinct in the Early Devonian. Only two families of eurypterines survived into the Late Devonian at all ( Adelophthalmidae and Waeringopteridae). The eurypterines experienced their most major declines in the Early Devonian, during which over 50% of their diversity was lost in just 10 million years. Stylonurines, on
12354-515: The eurypterine swimming paddles varied from group to group. In the Eurypteroidea , the paddles were similar in shape to oars. The condition of the joints in their appendages ensured their paddles could only be moved in near-horizontal planes, not upwards or downwards. Some other groups, such as the Pterygotioidea, would not have possessed this condition and were probably able to swim faster. Most eurypterines are generally agreed to have utilized
12496-454: The family Pterygotidae. An isolated 12.7 centimeters (5.0 in) long fossil metastoma of the carcinosomatoid eurypterid Carcinosoma punctatum indicates the animal would have reached a length of 2.2 meters (7.2 ft) in life, rivalling the pterygotids in size. Another giant was Pentecopterus decorahensis , a primitive carcinosomatoid, which is estimated to have reached lengths of 1.7 meters (5.6 ft). Typical of large eurypterids
12638-562: The first American inhabitants sailed from Beringia some 13,000 years ago, with widespread habitation of the Americas during the end of the Last Glacial Period , in what is known as the Late Glacial Maximum , around 12,500 years ago. The oldest petroglyphs in North America date from 15,000 to 10,000 years before present. Genetic research and anthropology indicate additional waves of migration from Asia via
12780-476: The found tracks each being about 7.6 centimeters (3.0 in) in diameter. Other eurypterid ichnogenera include Merostomichnites (though it is likely that many specimens actually represent trackways of crustaceans) and Arcuites (which preserves grooves made by the swimming appendages). In eurypterids, the respiratory organs were located on the ventral body wall (the underside of the opisthosoma). Blattfüsse , evolved from opisthosomal appendages, covered
12922-582: The gathering of food. In some groups, these spiny appendages became heavily specialized. In some eurypterids in the Carcinosomatoidea, forward-facing appendages were large and possessed enormously elongated spines (as in Mixopterus and Megalograptus ). In derived members of the Pterygotioidea, the appendages were completely without spines, but had specialized claws instead. Other eurypterids, lacking these specialized appendages, likely fed in
13064-656: The genus Strabops from the Cambrian of Missouri , are now classified as aglaspidids or strabopids . The aglaspidids, once seen as primitive chelicerates, are now seen as a group more closely related to trilobites. The fossil record of Ordovician eurypterids is quite poor. The majority of eurypterids once reportedly known from the Ordovician have since proven to be misidentifications or pseudofossils . Today only 11 species can be confidently identified as representing Ordovician eurypterids. These taxa fall into two distinct ecological categories; large and active predators from
13206-525: The genus (of which the most common is the type species, E. remipes ) account for more than 90% (perhaps as many as 95%) of all known fossil eurypterid specimens. Despite their vast number, Eurypterus are only known from a relatively short temporal range, first appearing during the Late Llandovery epoch (around 432 million years ago) and being extinct by the end of the Pridoli epoch. Eurypterus
13348-847: The geographic center of either the 50 states, the conterminous United States, or the North American continent." Nonetheless, there is a 4.6-meter (15 ft) field stone obelisk in Rugby claiming to mark the center. The North American continental pole of inaccessibility is located 1,650 km (1,030 mi) from the nearest coastline, between Allen and Kyle, South Dakota at 43°22′N 101°58′W / 43.36°N 101.97°W / 43.36; -101.97 ( Pole of Inaccessibility North America ) . Canada can be divided into roughly seven physiographic divisions: The lower 48 U.S. states can be divided into roughly eight physiographic divisions: Mexico can be divided into roughly fifteen physiographic divisions: North America
13490-429: The giant millipede Arthropleura , and are possibly vital for the evolution of giant size in arthropods. In addition to the lightweight giant eurypterids, some deep-bodied forms in the family Hibbertopteridae were also very large. A carapace from the Carboniferous of Scotland referred to the species Hibbertoperus scouleri measures 65 cm (26 in) wide. As Hibbertopterus was very wide compared to its length,
13632-412: The gills of other groups. To be functional gills, they would have to have been highly efficient and would have required a highly efficient circulatory system. It is considered unlikely, however, that these factors would be enough to explain the large discrepancy between gill tract size and body size. It has been suggested instead that the "gill tract" was an organ for breathing air, perhaps actually being
13774-587: The group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event . They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event (or sometime shortly before) 251.9 million years ago. Although popularly called "sea scorpions", only the earliest eurypterids were marine ; many later forms lived in brackish or fresh water , and they were not true scorpions . Some studies suggest that
13916-432: The group lived primarily in the waters around and within the ancient supercontinent of Euramerica . Only a handful of eurypterid groups spread beyond the confines of Euramerica and a few genera, such as Adelophthalmus and Pterygotus , achieved a cosmopolitan distribution with fossils being found worldwide. Like all other arthropods , eurypterids possessed segmented bodies and jointed appendages (limbs) covered in
14058-419: The indigenous people who lived there, including the bison hunters of the Great Plains and the farmers of Mesoamerica . Native groups also are classified by their language families , which included Athapascan and Uto-Aztecan languages. Indigenous peoples with similar languages did not always share the same material culture , however, and were not necessarily always allies. Anthropologists speculate that
14200-580: The invaginations leading to asphyxiation . Furthermore, most eurypterids would have been aquatic their entire lives. No matter how much time was spent on land, organs for respiration in underwater environments must have been present. True gills, expected to have been located within the branchial chamber within the Blattfüssen , remain unknown in eurypterids. Like all arthropods, eurypterids matured and grew through static developmental stages referred to as instars . These instars were punctuated by periods during which eurypterids went through ecdysis (molting of
14342-468: The land since its creation, but there is no evidence that humans evolved there. The specifics of the initial settlement of the Americas by ancient Asians are subject to ongoing research and discussion. The traditional theory has been that hunters entered the Bering Land Bridge between eastern Siberia and present-day Alaska from 27,000 to 14,000 years ago. A growing viewpoint is that
14484-491: The larger sizes of adults mean a higher drag coefficient , using this type of propulsion is more energy-efficient. Some eurypterines, such as Mixopterus (as inferred from attributed fossil trackways), were not necessarily good swimmers. It likely kept mostly to the bottom, using its swimming paddles for occasional bursts of movements vertically, with the fourth and fifth pairs of appendages positioned backwards to produce minor movement forwards. While walking, it probably used
14626-505: The largest known arthropod ever to have lived, is Jaekelopterus rhenaniae . A chelicera from the Emsian Klerf Formation of Willwerath, Germany measured 36.4 centimeters (14.3 in) in length, but is missing a quarter of its length, suggesting that the full chelicera would have been 45.5 centimeters (17.9 in) long. If the proportions between body length and chelicerae match those of its closest relatives, where
14768-523: The largest known arthropods ever to have lived. The largest, Jaekelopterus , reached 2.5 meters (8.2 ft) in length. Eurypterids were not uniformly large and most species were less than 20 centimeters (8 in) long; the smallest eurypterid, Alkenopterus , was only 2.03 centimeters (0.80 in) long. Eurypterid fossils have been recovered from every continent. A majority of fossils are from fossil sites in North America and Europe because
14910-497: The last ever radiation within the eurypterids, which gave rise to several new forms capable of "sweep-feeding" (raking through the substrate in search of prey). Only three eurypterid families—Adelophthalmidae, Hibbertopteridae and Mycteroptidae—survived the extinction event in its entirety. It was assumed that these were all freshwater animals, which would have rendered the eurypterids extinct in marine environments, and with marine eurypterid predators gone, sarcopterygians , such as
15052-657: The mainland in Mexico. With local indigenous allies, the Spanish conquered the Aztec empire in central Mexico in 1521. Spain then established permanent cities in Mexico, Central America, and Spanish South America in the sixteenth century. Once Spaniards conquered the high civilization of the Aztecs and Incas, the Caribbean was a backwater of the Spanish empire. Other European powers began to intrude on areas claimed by Spain, including
15194-927: The middle into the main range of the Rockies and the coast ranges in California , Oregon , Washington , and British Columbia , with the Great Basin—a lower area containing smaller ranges and low-lying deserts—in between. The highest peak is Denali in Alaska. The U.S. Geographical Survey (USGS) states that the geographic center of North America is "6 miles [10 km] west of Balta, Pierce County, North Dakota " at about 48°10′N 100°10′W / 48.167°N 100.167°W / 48.167; -100.167 , about 24 kilometers (15 mi) from Rugby, North Dakota . The USGS further states that "No marked or monumented point has been established by any government agency as
15336-526: The most basal of the rhenopterids and a sister taxon to more derived rhenopterids (such as Kiaeropterus and Rhenopterus ). Eurypterid Eurypterids , often informally called sea scorpions , are a group of extinct arthropods that form the order Eurypterida . The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago . The group
15478-454: The name America to North America. In 1538, Gerardus Mercator used the term America on his world map of the entire Western Hemisphere . On his subsequent 1569 map , Mercator called North America "America or New India" ( America sive India Nova ). The Spanish Empire called its territories in North and South America "Las Indias", and the name given to the state body that oversaw the region
15620-493: The northernmost countries and territories of North America: the U.S., Bermuda , Canada, Greenland, and St. Pierre and Miquelon. Although the term does not refer to a unified region, Middle America includes Mexico, Central America, and the Caribbean. North America's largest countries by land area are Canada and the U.S., both of which have well-defined and recognized subregions. In Canada, these include (from east to west) Atlantic Canada , Central Canada , Canadian Prairies ,
15762-440: The northernmost extent of the island named Newfoundland , has provided unmistakable evidence of Norse settlement. Norse explorer Leif Erikson (c. 970–1020 CE) is thought to have visited the area. Erikson was the first European to make landfall on the continent (excluding Greenland). The Mayan culture was still present in southern Mexico and Guatemala when the Spanish conquistadors arrived, but political dominance in
15904-507: The only pair placed before the mouth, is called the chelicerae ( homologous to the fangs of spiders). They were equipped with small pincers used to manipulate food fragments and push them into the mouth. In one lineage, the Pterygotidae , the chelicerae were large and long, with strong, well-developed teeth on specialised chelae (claws). The subsequent pairs of appendages, numbers II to VI, possessed gnathobases (or "tooth-plates") on
16046-812: The opisthosoma was covered in structures evolved from modified opisthosomal appendages. Throughout the opisthosoma, these structures formed plate-like structures termed Blattfüsse ( lit. ' leaf-feet ' in German). These created a branchial chamber (gill tract) between preceding Blattfüsse and the ventral surface of the opisthosoma itself, which contained the respiratory organs. The second to sixth opisthosomal segments also contained oval or triangular organs that have been interpreted as organs that aid in respiration. These organs, termed Kiemenplatten or "gill tracts", would potentially have aided eurypterids to breathe air above water, while Blattfüssen , similar to organs in modern horseshoe crabs , would cover
16188-424: The other hand, persisted through the period with more or less consistent diversity and abundance but were affected during the Late Devonian, when many of the older groups were replaced by new forms in the families Mycteroptidae and Hibbertopteridae. It is possible that the catastrophic extinction patterns seen in the eurypterine suborder were related to the emergence of more derived fish. Eurypterine decline began at
16330-552: The parts that serve for underwater respiration . The appendages of opisthosomal segments 1 and 2 (the seventh and eighth segments overall) were fused into a structure termed the genital operculum, occupying most of the underside of the opisthosomal segment 2. Near the anterior margin of this structure, the genital appendage (also called the Zipfel or the median abdominal appendage) protruded. This appendage, often preserved very prominently, has consistently been interpreted as part of
16472-433: The past. Hemianamorphic direct development has been observed in many arthropod groups, such as trilobites , megacheirans , basal crustaceans and basal myriapods . True direct development has on occasion been referred to as a trait unique to arachnids . There have been few studies on eurypterid ontogeny as there is a general lack of specimens in the fossil record that can confidently be stated to represent juveniles. It
16614-434: The point when jawless fish first became more developed and coincides with the emergence of placoderms (armored fish) in both North America and Europe. Stylonurines of the surviving hibbertopterid and mycteroptid families completely avoided competition with fish by evolving towards a new and distinct ecological niche. These families experienced a radiation and diversification through the Late Devonian and Early Carboniferous,
16756-605: The raised but relatively flat plateau of the Canadian Shield in the northeast; and the varied eastern region, which includes the Appalachian Mountains, the coastal plain along the Atlantic seaboard, and the Florida peninsula. Mexico, with its long plateaus and cordilleras , falls largely in the western region, although the eastern coastal plain does extend south along the Gulf. The western mountains are split in
16898-452: The ratio between claw size and body length is relatively consistent, the specimen of Jaekelopterus that possessed the chelicera in question would have measured between 233 and 259 centimeters (7.64 and 8.50 ft), an average 2.5 meters (8.2 ft), in length. With the chelicerae extended, another meter (3.28 ft) would be added to this length. This estimate exceeds the maximum body size of all other known giant arthropods by almost half
17040-500: The region, and as such rainforest , monsoon , and savanna types can be found, with rains and high temperatures throughout the year. Notable North American fauna include the bison , black bear , jaguar , cougar , prairie dog , turkey , pronghorn , raccoon , coyote , and monarch butterfly . Notable plants that were domesticated in North America include tobacco , maize , squash , tomato , sunflower , blueberry , avocado , cotton , chile pepper , and vanilla . Laurentia
17182-445: The reproduction and sexual dimorphism of eurypterids is difficult, as they are only known from fossilized shells and carapaces. In some cases, there might not be enough apparent differences to separate the sexes based on morphology alone. Sometimes two sexes of the same species have been interpreted as two different species, as was the case with two species of Drepanopterus ( D. bembycoides and D. lobatus ). The eurypterid prosoma
17324-458: The reproductive system and occurs in two recognized types, assumed to correspond to male and female. Eurypterids were highly variable in size, depending on factors such as lifestyle, living environment and taxonomic affinity . Sizes around 100 centimeters (3.3 ft) are common in most eurypterid groups. The smallest eurypterid, Alkenopterus burglahrensis , measured just 2.03 centimeters (0.80 in) in length. The largest eurypterid, and
17466-435: The rest of the former supercontinent Gondwana , the discoveries of trackways both predate and outnumber eurypterid body fossils. Eurypterid trackways have been referred to several ichnogenera, most notably Palmichnium (defined as a series of four tracks often with an associated drag mark in the mid-line), wherein the holotype of the ichnospecies P. kosinkiorum preserves the largest eurypterid footprints known to date with
17608-487: The results of the formation of Pangaea was the Appalachian Mountains , which formed some 480 mya, making it among the oldest mountain ranges in the world. When Pangaea began to rift around 200 mya, North America became part of Laurasia , before it separated from Eurasia as its own continent during the mid- Cretaceous period. The Rockies and other western mountain ranges began forming around this time from
17750-405: The same genera. The primary function of the long, assumed female, type A appendages was likely to take up spermatophore from the substrate into the reproductive tract rather than to serve as an ovipositor, as arthropod ovipositors are generally longer than eurypterid type A appendages. By rotating the sides of the operculum, it would have been possible to lower the appendage from the body. Due to
17892-413: The short stride length indicates that Hibbertopterus crawled with an exceptionally slow speed, at least on land. The large telson was dragged along the ground and left a large central groove behind the animal. Slopes in the tracks at random intervals suggest that the motion was jerky. The gait of smaller stylonurines, such as Parastylonurus , was probably faster and more precise. The functionality of
18034-429: The sixth pair of appendages were overlaid by a plate that is referred to as the metastoma, originally derived from a complete exoskeleton segment. The opisthosoma itself can be divided either into a " mesosoma " (comprising segments 1 to 6) and " metasoma " (comprising segments 7 to 12) or into a "preabdomen" (generally comprising segments 1 to 7) and "postabdomen" (generally comprising segments 8 to 12). The underside of
18176-441: The species Lanarkopterus dolichoschelus from the Ordovician of Ohio contain fragments of jawless fish and fragments of smaller specimens of Lanarkopterus itself. Though apex predatory roles would have been limited to the very largest eurypterids, smaller eurypterids were likely formidable predators in their own right just like their larger relatives. As in many other entirely extinct groups, understanding and researching
18318-427: The spongy structure of the eurypterid gill tracts. It is possible the two organs functioned in the same way. Some researchers have suggested that eurypterids may have been adapted to an amphibious lifestyle, using the full gill tract structure as gills and the invaginations within it as pseudotrachea. This mode of life may not have been physiologically possible, however, since water pressure would have forced water into
18460-498: The structure. Though the Kiemenplatte is referred to as a "gill tract", it may not necessarily have functioned as actual gills. In other animals, gills are used for oxygen uptake from water and are outgrowths of the body wall. Despite eurypterids clearly being primarily aquatic animals that almost certainly evolved underwater (some eurypterids, such as the pterygotids, would even have been physically unable to walk on land), it
18602-468: The type A appendage is divided into three but the type B appendage into only two. Such division of the genital appendage is common in eurypterids, but the number is not universal; for instance, the appendages of both types in the family Pterygotidae are undivided. The type A appendage is also armed with two curved spines called furca (lit. 'fork' in Latin). The presence of furca in the type B appendage
18744-470: The type A appendage, could have been used to detect whether a substrate was suitable for spermatophore deposition. Until 1882 no eurypterids were known from before the Silurian. Contemporary discoveries since the 1880s have expanded the knowledge of early eurypterids from the Ordovician period. The earliest eurypterids known today, the megalograptid Pentecopterus , date from the Darriwilian stage of
18886-400: The underside and created a gill chamber where the "gill tracts" were located. Depending on the species, the eurypterid gill tract was either triangular or oval in shape and was possibly raised into a cushion-like state. The surface of this gill tract bore several spinules (small spines), which resulted in an enlarged surface area. It was composed of spongy tissue due to many invaginations in
19028-429: The way different plates overlay at its location, the appendage would have been impossible to move without muscular contractions moving around the operculum. It would have been kept in place when not it use. The furca on the type A appendages may have aided in breaking open the spermatophore to release the free sperm inside for uptake. The "horn organs," possibly spermathecae, are thought to have been connected directly to
19170-400: Was also restricted to the continent Euramerica (composed of the equatorial continents Avalonia, Baltica and Laurentia), which had been completely colonized by the genus during its merging and was unable to cross the vast expanses of ocean separating this continent from other parts of the world, such as the southern supercontinent Gondwana. As such, Eurypterus was limited geographically to
19312-605: Was called the Council of the Indies . The United Nations and its statistics division recognize North America as including three regions: Northern America, Central America , and the Caribbean . "Northern America" is a distinct term from "North America", excluding Central America, which itself may or may not include Mexico . In the limited context of regional trade agreements, the term is used to reference three nations: Canada,
19454-505: Was discovered in Carboniferous-aged fossil deposits of Scotland in 2005. It was attributed to the stylonurine eurypterid Hibbertopterus due to a matching size (the trackmaker was estimated to have been about 1.6 meters (5.2 ft) long) and inferred leg anatomy. It is the largest terrestrial trackway—measuring 6 meters (20 ft) long and averaging 95 centimeters (3.12 ft) in width—made by an arthropod found thus far. It
19596-561: Was estimated as over 592 million people in 23 independent states , or about 7.5% of the world's population. In human geography , the terms "North America" and "North American" can refer to Canada, the United States, Mexico, and Greenland or, alternatively, Canada, Greenland and the US (Mexico being classified as part of Latin America ) or simply Canada and the US (Greenland being classified as either Arctic or European (due to its political status as
19738-817: Was in Jamestown, Virginia in 1607, followed by additional colonial establishments on the east coast from present-day Georgia in the south to Massachusetts in the north, forming the Thirteen Colonies of British America. The English did not establish settlements north or east of the St. Lawrence Valley in present-day Canada until after the conclusion of the American Revolutionary War . Britain's early settlements in present-day Canada included St. John's, Newfoundland in 1630 and Halifax, Nova Scotia in 1749. The first permanent French settlement
19880-452: Was more or less parallel and similar to that of extinct and extant xiphosurans, with the largest exception being that eurypterids hatched with a full set of appendages and opisthosomal segments. Eurypterids were thus not hemianamorphic direct developers, but true direct developers like modern arachnids. The most frequently observed change occurring through ontogeny (except for some genera, such as Eurypterus , which appear to have been static)
20022-473: Was the first European to suggest that the Americas represented a landmass not then known to the Europeans. In 1507, Waldseemüller published a world map, and placed the word "America" on the continent of present-day South America. The continent north of present-day Mexico was then referred to as Parias. On a 1553 world map published by Petrus Apianus , North America was called "Baccalearum", meaning "realm of
20164-518: Was used as an ovipositor (used to deposit eggs). The different types of genital appendages are not necessarily the only feature that distinguishes between the sexes of eurypterids. Depending on the genus and species in question, other features such as size, the amount of ornamentation and the proportional width of the body can be the result of sexual dimorphism. In general, eurypterids with type B appendages (males) appear to have been proportionally wider than eurypterids with type A appendages (females) of
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