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Hoarding or caching in animal behavior is the storage of food in locations hidden from the sight of both conspecifics (animals of the same or closely related species) and members of other species. Most commonly, the function of hoarding or caching is to store food in times of surplus for times when food is less plentiful. However, there is evidence that a certain amount of caching or hoarding is actually undertaken with the aim of ripening the food so stored, and this practice is thus referred to as ‘ripening caching’. The term hoarding is most typically used for rodents , whereas caching is more commonly used in reference to birds , but the behaviors in both animal groups are quite similar.

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86-538: The Brazil nut ( Bertholletia excelsa ) is a South American tree in the family Lecythidaceae , and it is also the name of the tree's commercially harvested edible seeds . It is one of the largest and longest-lived trees in the Amazon rainforest . The fruit and its nutshell – containing the edible Brazil nut – are relatively large and weigh as much as 2 kg (4.4 lb) in total. As food, Brazil nuts are notable for diverse content of micronutrients , especially

172-736: A tropical forest without destroying it. The nuts are most often gathered by migrant workers known as castañeros (in Spanish ) or castanheiros (in Portuguese ). Logging is a significant threat to the sustainability of the Brazil nut-harvesting industry. Analysis of tree ages in areas that are harvested shows that moderate and intense gathering takes so many seeds that not enough are left to replace older trees as they die. Sites with light gathering activities had many young trees, while sites with intense gathering practices had nearly none. In 2003,

258-469: A Brazil nut tree requires previous authorization from the Brazilian Institute of Environment and Renewable Natural Resources . Brazil nut oil is used as a lubricant in clocks and in the manufacturing of paint and cosmetics , such as soap and perfume. Because of its hardness, the Brazil nutshell is often pulverized and used as an abrasive to polish materials, such as metals and ceramics, in

344-527: A clear connection between hamsters and hoarding: Polish chomikować , from chomik – hamster; Hebrew hamster ; oger (אוגר) comes from to hoard ; le'egor (לאגור). A disadvantage of larder hoarding is that if a cache is raided, this is far more problematic for the animal than if it were a scatter hoarder. While the hoard is much easier to remember the location of, these larger hoards must also be more staunchly defended. Most species are particularly wary of onlooking individuals during caching and ensure that

430-698: A fleet to reach Peru by sea, and sought the military support of various rebels from the Viceroyalty of Peru. In 1822 the two men met at the Guayaquil Conference in Ecuador, where they failed to agree on governance strategies for the liberated nations. Two years later Bolívar's forces beat the Spanish at the Battle of Ayacucho , securing the independence of Peru and the rest of South America. In

516-530: A fungus which is the main food for the colony. Contrary to popular belief, there is no scientific evidence that crocodilians such as the American Alligator cache large prey underwater to consume later. Scatter hoarding is the formation of a large number of small hoards. This behavior is present in both birds (especially the Canada jay ) and small mammals, mainly squirrels and other rodents, such as

602-400: A hard, woody shell 8–12 mm ( 3 ⁄ 8 – 1 ⁄ 2  in) thick, which contains eight to 24 wedge-shaped seeds 4–5 cm ( 1 + 5 ⁄ 8 –2 in) long (the "Brazil nuts") packed like the segments of an orange, but not limited to one whorl of segments. Up to three whorls can be stacked onto each other, with the polar ends of the segments of the middle whorl nestling into

688-877: A high amount of selenium . The wood of the Brazil nut tree is prized for its quality in carpentry , flooring , and heavy construction. In Portuguese-speaking countries, like Brazil, they are variously called " castanha-do-brasil " (meaning "chestnut from Brazil " in Portuguese), " castanha-do-pará " (meaning "chestnut from Pará " in Portuguese), with other names: castanha-da-amazônia, castanha-do-acre, " noz amazônica " (meaning "Amazonian nut" in Portuguese), noz boliviana, tocari ("probably of Carib origin"), and tururi (from Tupi turu'ri ) also used. In various Spanish-speaking countries of South America , Brazil nuts are called castañas de Brasil , nuez de Brasil , or castañas de Pará (or Para) . In North America, as early as 1896, Brazil nuts were sometimes known by

774-498: A jaguar hangs partially eaten prey from a tree to be eaten within a few days, or long term, where the food is hidden and retrieved many months later. Caching is a common adaptation to seasonal changes in food availability. In regions where winters are harsh, food availability typically becomes low, and caching food during the times of high food availability in the warmer months provides a significant survival advantage. For species that hoard perishable food weather can significantly affect

860-459: A long history of colonialism , the overwhelming majority of South Americans speak Spanish or Portuguese , and societies and states are rich in Western traditions . Relative to Africa, Asia, and Europe, post-1900 South America has been a peaceful continent with few wars, although high rates of violent crime remain a concern in some countries. South America occupies the southern portion of

946-682: A part of Oceania, also administered by Chile), the Galápagos Islands (administered by Ecuador , sometimes considered part of Oceania), and Tierra del Fuego (split between Argentina and Chile). In the Atlantic Ocean, Brazil administers Fernando de Noronha , Trindade and Martim Vaz , and the Saint Peter and Saint Paul Archipelago , while the Falkland Islands (Spanish: Islas Malvinas ) and South Georgia and

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1032-632: A particularly rich source of selenium , with just 28 g (1 oz) supplying 544 micrograms of selenium or 10 times the DV of 55 micrograms. However, the amount of selenium within batches of nuts may vary considerably. The high selenium content is used as a biomarker in studies of selenium intake and deficiency. Consumption of just one Brazil nut per day over 8 weeks was sufficient to restore selenium blood levels and increase HDL cholesterol in obese women. Brazil nut oil contains 48% unsaturated fatty acids composed mainly of oleic and linoleic acids ,

1118-481: A site called Chavín de Huantar in modern Peru at an elevation of 3,177 meters (10,423 ft). Chavín civilization spanned 900 BC to 300 BC. In the central coast of Peru, around the beginning of the 1st millennium AD, Moche (100 BC – 700 AD, at the northern coast of Peru), Paracas and Nazca (400 BC – 800 AD, Peru) cultures flourished with centralized states with permanent militia improving agriculture through irrigation and new styles of ceramic art. At

1204-496: A state of dormancy, for a tree to fall and sunlight to reach it, when it starts growing again. The Brazil nut family, the Lecythidaceae, is in the order Ericales , as are other well-known plants such as blueberries, cranberries, sapote , gutta-percha , tea , phlox , and persimmons . The tree is the only species in the monotypic genus Bertholletia , named after French chemist Claude Louis Berthollet . The Brazil nut

1290-427: A stone as an anvil . In 2020, global production of Brazil nuts (in shells) was 69,658 tonnes , most of which derive from wild harvests in tropical forests, especially the Amazon regions of Brazil and Bolivia which produced 92% of the world total (table). Since most of the production for international trade is harvested in the wild, the business arrangement has been advanced as a model for generating income from

1376-689: A subtropical climate of the dinaric type. At the highest points of the Andean region, climates are colder than the ones occurring at the highest point of the Norwegian fjords. In the Andean plateaus, the warm climate prevails, although it is tempered by the altitude, while in the coastal strip, there is an equatorial climate of the Guinean type. From this point until the north of the Chilean coast appear, successively, Mediterranean oceanic climate , temperate of

1462-435: A two-parted, deciduous calyx , six unequal cream-colored petals, and numerous stamens united into a broad, hood-shaped mass. Brazil nut trees produce fruit almost exclusively in pristine forests, as disturbed forests lack the large-bodied bees of the genera Bombus , Centris , Epicharis , Eulaema , and Xylocopa , which are the only ones capable of pollinating the tree's flowers, with different bee genera being

1548-526: Is because the most extensive part of the continent is in the equatorial zone (the region has more areas of equatorial plains than any other region), therefore giving the Southern Cone more oceanic influence, which moderates year round temperatures. The average annual temperatures in the Amazon basin oscillate around 27 °C (81 °F), with low thermal amplitudes and high rainfall indices. Between

1634-460: Is by far the most populous South American country, with almost half of the continent's population, followed by Colombia, Argentina, Venezuela, and Peru. In recent decades, Brazil has also generated half of the continent's GDP and has become the continent's first regional power. Most of the population lives near the continent's western or eastern coasts while the interior and the far south are sparsely populated. The geography of western South America

1720-553: Is dominated by the Andes mountains; in contrast, the eastern part contains both highland regions and vast lowlands where rivers such as the Amazon , Orinoco and Paraná flow. Most of the continent lies in the tropics, except for a large part of the Southern Cone located in the middle latitudes . The continent's cultural and ethnic outlook has its origin with the interaction of Indigenous peoples with European conquerors and immigrants and, more locally, with African slaves . Given

1806-636: Is native to the Guianas , Venezuela , Brazil , eastern Colombia , eastern Peru , and eastern Bolivia . It occurs as scattered trees in large forests on the banks of the Amazon River , Rio Negro , Tapajós , and the Orinoco . The fruit is heavy and rigid; when the fruits fall, they pose a serious threat to vehicles and potential for traumatic brain injury of people passing under the tree. Capuchin monkeys have been reported to open Brazil nuts using

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1892-475: Is noted for its particular skill at caching. There are two types of caching behavior: larder hoarding, where a species creates a few large caches which it often defends, and scatter hoarding, where a species will create multiple caches, often with each individual food item stored in a unique place. Both types of caching have their advantage. Caching behavior is typically a way to save excess edible food for later consumption—either soon to be eaten food, such as when

1978-614: Is now Colombia. They established the Muisca Confederation of many clans, or cacicazgos , that had a free trade network among themselves. Many were goldsmiths and farmers. Other important Pre-Columbian cultures include: the Cañaris (in south central Ecuador), Chimú Empire (1300–1470, Peruvian northern coast), Chachapoyas , and the Aymaran kingdoms (1000–1450, Western Bolivia and southern Peru). Holding their capital at

2064-480: Is one of the most biodiverse continents on Earth. It is home to many unique species of animals including the llama , anaconda , piranha , jaguar , vicuña , and tapir , and to one of the largest known insects in the world, the Titan beetle . The Amazon rainforests possess high biodiversity , with Brazil estimated to contain 10% of Earth's species . 83% of South America's large mammals ( megafauna ) became extinct at

2150-470: Is still the question of why they would store the food outside of their bodies in the first place. The reason for this is that scatter hoarders must remain active during the caching period in order to hide the most food in the most places possible. Storing the food inside their body would reduce their mobility and be counterproductive to this objective. Cache spacing is the primary technique that scatter hoarders use to protect food from pilferers. By spreading

2236-469: Is straight and commonly without branches for well over half the tree's height, with a large, emergent crown of long branches above the surrounding canopy of other trees. The bark is grayish and smooth. The leaves are dry-season deciduous , alternate, simple, entire or crenate, oblong, 20–35 centimetres (8–14 in) long, and 10–15 cm (4–6 in) broad. The flowers are small, greenish-white, in panicles 5–10 cm (2–4 in) long; each flower has

2322-582: Is the largest country in South America, covering a little less than half of the continent's land area and encompassing around half of the continent's population. The remaining countries and territories are divided among four subregions : the Andean states , Caribbean South America , The Guianas , and the Southern Cone . Physiographically, South America also includes some of the nearby islands. The Dutch ABC islands ( Aruba , Bonaire , and Curaçao ),

2408-519: The Altiplano , Tiahuanaco or Tiwanaku (100 BC – 1200 AD, Bolivia) managed a large commercial network based on religion. Around the 7th century, both Tiahuanaco and Wari or Huari Empire (600–1200, Central and northern Peru) expanded its influence to all the Andean region, imposing the Huari urbanism and Tiahuanaco religious iconography. The Muisca were the main indigenous civilization in what

2494-586: The Archipelago of San Andrés, Providencia and Santa Catalina ( San Andrés Island , Providencia Island , and Santa Catalina Island etc., which are administered by Colombia ) are politically parts of South American countries but physiographically parts of North America. Other islands often associated with geopolitical South America are the Chiloé Archipelago and Robinson Crusoe Island (both administered by Chile), Easter Island (culturally

2580-820: The Atacama Desert ; the wettest place on earth, López de Micay in Colombia; the largest rainforest, the Amazon rainforest ; the highest capital city, La Paz , Bolivia ; the highest commercially navigable lake in the world, Lake Titicaca ; and, excluding research stations in Antarctica , the world's southernmost permanently inhabited community, Puerto Toro , Chile. South America's major mineral resources are gold, silver, copper, iron ore, tin , and petroleum. These resources have brought high income to its countries, especially in times of war or of rapid economic growth by industrialized countries elsewhere. However,

2666-717: The Bering Land Bridge (now the Bering Strait ) at least 15,000 years ago from the territory that is present-day Russia. They migrated south through North America, and eventually reached South America through the Isthmus of Panama . Amongst the oldest evidence for human presence in South America is the Monte Verde II site in Chile, suggested to date to around 14,500 years ago. From around 13,000 years ago,

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2752-634: The Caribbean colonies and Brazil, as European nations built up economically slave-dependent colonies in the New World . Nearly 40% of all African slaves trafficked to the Americas went to Brazil. An estimated 4.9 million slaves from Africa came to Brazil during the period from 1501 to 1866. In contrast to other European colonies in the Americas which mainly used the labor of African slaves, Spanish colonists mainly enslaved indigenous Americans. In 1750,

2838-567: The European Union imposed strict regulations on the import of Brazilian-harvested Brazil nuts in their shells, as the shells are considered to contain unsafe levels of aflatoxins , a potential cause of liver cancer . Brazil nuts are susceptible to contamination by aflatoxins , produced by fungi, once they fall to the ground. Aflatoxins can cause liver damage , including possible cancer, if consumed. Aflatoxin levels have been found in Brazil nuts during inspections that were far higher than

2924-470: The Fishtail projectile point style became widespread across South America, with its disappearance around 11,000 years ago coincident with the disappearance of South America's megafauna. Maize was present in northern South America by around 6,000 years ago. By 2000 BC, many agrarian communities had been settled throughout the Andes and the surrounding regions. Fishing became a widespread practice along

3010-472: The Fortaleza area, contemporaneous with the pyramids of Ancient Egypt , one of the oldest-known civilizations in the Americas and one of the six sites where civilization separately originated in the ancient world. Caral–Supe governing class established a trade network and developed agriculture then followed by Chavín by 900 BC, according to some estimates and archaeological finds. Artifacts were found at

3096-715: The Maracaibo Lake and the mouth of the Orinoco, predominates an equatorial climate of the type Congolese, that also includes parts of the Brazilian territory. The east-central Brazilian plateau has a humid and warm tropical climate. The northern and eastern parts of the Argentine pampas have a humid subtropical climate with dry winters and humid summers of the Chinese type, while the western and eastern ranges have

3182-599: The Orinoco Llanos and in the Guianas Plateau , the precipitation levels go from moderate to high. The Pacific coast of Colombia and northern Ecuador are rainy regions, with Chocó in Colombia being the rainiest place in the world along with the northern slopes of Indian Himalayas. The Atacama Desert, along this stretch of coast, is one of the driest regions in the world. The central and southern parts of Chile are subject to extratropical cyclones , and most of

3268-547: The Portuguese Crown abolished the enslavement of indigenous peoples in colonial Brazil , under the belief that they were unfit for labor and less effective than enslaved Africans. Enslaved Africans were brought to the Americas on slave ships , under inhuman conditions and ill-treatment, and those who survived were sold in slave markets . After independence, all South American countries maintained slavery for some time. The first South American country to abolish slavery

3354-738: The Portuguese Kingdom of Brazil , Dom Pedro I (also Pedro IV of Portugal), son of the Portuguese King Dom João VI , proclaimed the independent Kingdom of Brazil in 1822, which later became the Empire of Brazil . Despite the Portuguese loyalties of garrisons in Bahia , Cisplatina and Pará , independence was diplomatically accepted by Portugal in 1825, on condition of a high compensation paid by Brazil mediated by

3440-539: The Riograndense Republic ). Cache (biology) Hoarding is done either on a long-term basis—cached on a seasonal cycle, with food to be consumed months down the line—or on a short-term basis, in which case the food will be consumed over a period of one or several days. Some common animals that cache their food are rodents such as hamsters and squirrels , and many different bird species, such as rooks and woodpeckers . The western scrub jay

3526-605: The South American Plate . South America is home to several superlatives, including the world's highest uninterrupted waterfall , Angel Falls in Venezuela; the highest single-drop waterfall Kaieteur Falls in Guyana; the largest river by volume, the Amazon River ; the longest mountain range, the Andes (whose highest mountain is Aconcagua at 6,962 m or 22,841 ft); the driest non-polar place on earth,

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3612-613: The Spanish American wars of independence between the patriots, who promoted such autonomy, and the royalists , who supported Spanish authority over the Americas. The Juntas, in both Spain and the Americas, promoted the ideas of the Enlightenment . Five years after the beginning of the war, Ferdinand VII returned to the throne and began the Absolutist Restoration , as the royalists got the upper hand in

3698-444: The eastern gray squirrel , fox squirrel , and wood mouse . Specifically, those who do not migrate to warmer climates or hibernate for winter are most likely to scatter hoard. This behavior plays an important part in seed dispersal , as those seeds that are left uneaten will have a chance to germinate , thus enabling plants to spread their populations effectively. While it is clear why some animals scatter their food caches, there

3784-712: The native populations had no immune resistance – caused large-scale depopulation of the native population under Spanish control. Systems of forced labor, such as the haciendas and mining industry's mit'a also contributed to the depopulation. After this, enslaved Africans , who had developed immunities to these diseases, were quickly brought in to replace them. The Spaniards were committed to converting their native subjects to Christianity and were quick to purge any native cultural practices that hindered this end; however, many initial attempts at this were only partially successful, as native groups simply blended Catholicism with their established beliefs and practices. Furthermore,

3870-512: The phytosterol , beta-sitosterol , and fat-soluble vitamin E . The following table presents the composition of fatty acids in Brazil nut oil: The lumber from Brazil nut trees (not to be confused with Brazilwood ) is of excellent quality, having diverse uses from flooring to heavy construction. Logging the trees is prohibited by law in all three producing countries (Brazil, Bolivia, and Peru). Illegal extraction of timber and land clearances present continuing threats. In Brazil, cutting down

3956-417: The slang term " nigger toes", a vulgarity that fell out of use after the racial slur became more socially unacceptable . The Brazil nut is a large tree, reaching 50 metres (160 feet) tall, and with a trunk 1 to 2 m (3 to 7 ft) in diameter , making it among the largest of trees in the Amazon rainforest . It may live for 500 years or more, and can often reach a thousand years of age. The stem

4042-755: The Americas. The continent is generally delimited on the northwest by the Darién watershed along the Colombia–Panama border , although some may consider the border instead to be the Panama Canal . Geopolitically and geographically, all of Panama – including the segment east of the Panama Canal in the isthmus – is typically included in North America alone and among the countries of Central America . Almost all of mainland South America sits on

4128-692: The Argentine Patagonia is desert . In the Pampas of Argentina, Uruguay and South of Brazil the rainfall is moderate, with rains well distributed during the year. The moderately dry conditions of the Chaco oppose the intense rainfall of the eastern region of Paraguay. In the semiarid coast of the Brazilian Northeast the rains are linked to a monsoon regime. Important factors in the determination of climates are sea currents, such as

4214-748: The Breton type and, already in Tierra del Fuego , cold climate of the Siberian type. The distribution of rainfall is related to the regime of winds and air masses. In most of the tropical region east of the Andes, winds blowing from the northeast, east and southeast carry moisture from the Atlantic, causing abundant rainfall. However, due to a consistently strong wind shear and a weak Intertropical Convergence Zone , South Atlantic tropical cyclones are rare. In

4300-574: The South Sandwich Islands ( biogeographically and hydrologically associated with Antarctica) have been administered as two British Overseas Territories under the Crown , whose sovereignty over the islands is disputed by Argentina . An isolated volcanic island on the South American Plate , Ascension Island is geologically a part of South America. Administered as a dependency of Saint Helena, Ascension and Tristan da Cunha ,

4386-909: The South Sandwich Islands ; and one internal territory : French Guiana . The ABC islands of the Dutch Caribbean (north of Venezuela in the Leeward Antilles ), Ascension Island (a part of Saint Helena, Ascension and Tristan da Cunha ), Bouvet Island (a dependency of Norway ), Panama , and the Caribbean island country of Trinidad and Tobago may also be considered parts of South America. South America has an area of 17,840,000 square kilometers (6,890,000 sq mi). Its population as of 2021 has been estimated at more than 434 million. South America ranks fourth in area (after Asia, Africa, and North America) and fifth in population (after Asia, Africa, Europe, and North America). Brazil

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4472-1014: The Spaniards brought their language to the degree they did with their religion, although the Roman Catholic Church 's evangelization in Quechua , Aymara , and Guaraní actually contributed to the continuous use of these native languages albeit only in the oral form. Eventually, the natives and the Spaniards interbred, forming a mestizo class. At the beginning, many mestizos of the Andean region were offspring of Amerindian mothers and Spanish fathers. After independence, most mestizos had native fathers and European or mestizo mothers. Many native artworks were considered pagan idols and destroyed by Spanish explorers; this included many gold and silver sculptures and other artifacts found in South America, which were melted down before their transport to Spain or Portugal. Spaniards and Portuguese brought

4558-534: The Spanish and Portuguese colonies. First, Napoleon invaded Portugal, but the House of Braganza avoided capture by escaping to Brazil . Napoleon captured King Ferdinand VII of Spain, and appointed his own brother instead. This appointment provoked popular resistance, which created Juntas to rule in the name of the captured king. Many cities in the Spanish colonies, however, considered themselves equally authorized to appoint local Juntas like those of Spain. This began

4644-607: The United Kingdom. The newly independent nations began a process of fragmentation, with several civil and international wars. However, it was not as strong as in Central America. Some countries created from provinces of larger countries stayed as such up to modern times (such as Paraguay or Uruguay), while others were reconquered and reincorporated into their former countries (such as the Republic of Entre Ríos and

4730-571: The accumulation, use and rotting of the stored food. This phenomenon is referenced in the fable The Ant and the Grasshopper . However, in ripening caching behavior, animals collect and cache food which is immediately inedible but will become "ripe" and edible after a short while. For instance, tayras (a Central American weasel) have been observed to harvest whole green plantains , hide them, and then come back to eat them after they have ripened. Leafcutter ants harvest pieces of inedible leaves and then cache them in underground chambers to ripen with

4816-679: The amount present in Brazil nuts is orders of magnitude too small to have noticeable health effects. Brazil nuts are 3% water, 14% protein , 12% carbohydrates , and 66% fats . The fat components are 16% saturated , 24% monounsaturated , and 24% polyunsaturated . In a 100 grams (3.5 ounces) reference amount, Brazil nuts supply 659 calories , and are a rich source (20% or more of the Daily Value , DV) of dietary fiber (30% DV), thiamin (54% DV), vitamin E (38% DV), magnesium (106% DV), phosphorus (104% DV), manganese (57% DV), and zinc (43% DV). Calcium , iron , and potassium are present in moderate amounts (10–19% DV). Brazil nuts are

4902-478: The cache locations are secret . Not all caches are concealed however, for example shrikes store prey items on thorns on branches in the open. Although a small handful of species share food stores, food hoarding is a solo endeavor for most species, including almost all rodents and birds. For example, a number of jays live in large family groups, but they don't demonstrate sharing of cached food. Rather, they hoard their food supply selfishly, caching and retrieving

4988-437: The coast, helping establish fish as a primary source of food. Irrigation systems were also developed at this time, which aided in the rise of an agrarian society . South American cultures began domesticating llamas , vicuñas , guanacos , and alpacas in the highlands of the Andes circa 3500 BC. Besides their use as sources of meat and wool, these animals were used for transportation of goods. The rise of plant growing and

5074-461: The concentration in producing one, or few, major export commodities has often hindered the development and diversification of its economies. The fluctuation in the price of commodities in international markets has led historically to major highs and lows, booms and busts, in the economies of South American states, often causing political instability. This has led for calls to diversify production and increase trade within South America itself. Brazil

5160-424: The conflict. The independence of South America was secured by Simón Bolívar (Venezuela) and José de San Martín (Argentina), the two most important Libertadores . Bolívar led a great uprising in the north, then led his army south towards Lima , the capital of the Viceroyalty of Peru . Meanwhile, San Martín led an army across the Andes Mountains, along with Chilean expatriates, and liberated Chile. He organized

5246-405: The current Humboldt and Falklands . The equatorial current of the South Atlantic strikes the coast of the Northeast and there is divided into two others: the current of Brazil and a coastal current that flows to the northwest towards the Antilles , where there it moves towards northeast course thus forming the most Important and famous ocean current in the world, the Gulf Stream . South America

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5332-427: The end of the Pleistocene around 11,000 years ago as part of the Quaternary extinction event , among the highest of any continent, with the casualties including saber-toothed cats , ground sloths , glyptodonts , gomphotheres , the equines Hippidion and Equus neogeus , and all remaining South American native ungulates . South America is thought to have been first inhabited by humans when people were crossing

5418-569: The food supply around geographically, hoarders discourage competitors who happen upon a cache from conducting area-restricted searching for more of the supply. Despite cache spacing, hoarders are still unable to eliminate the threat of pilferage . However, having multiple cache sites is costly because it requires a good spatial memory. Scatter-hoarders generally have larger hippocampi than animals that do not participate in scatter-hoarding behavior. Additionally, studies have shown that hippocampal volume in scatter-hoarders varies seasonally and based on

5504-502: The great city of Cusco , the Inca civilization dominated the Andes region from 1438 to 1533. Known as Tawantin suyu , and "the land of the four regions", in Quechua , the Inca Empire was highly distinct and developed. Inca rule extended to nearly a hundred linguistic or ethnic communities, some nine to fourteen million people connected by a 25,000 kilometer road system . Cities were built with precise, unmatched stonework, constructed over many levels of mountain terrain. Terrace farming

5590-486: The harshness of the climate that the animal lives in. In larder hoarding , the hoard is large and is found in a single place termed a larder , which usually also serves as the nest where the animal lives. Hamsters are famous larder hoarders. Indeed, the German verb "hamstern" (to hoard) is derived from the noun "Hamster" which refers to the rodent; similar verbs are found in various related languages ( Dutch hamsteren , and Swedish hamstra ). Other languages also draw

5676-417: The island is geopolitically a part of Africa. All of the world's major climate zones are present in South America. The distribution of the average temperatures in the region presents a constant regularity from the 30° of latitude south , when the isotherms tend, more and more, to be confused with the degrees of latitude. In temperate latitudes, winters and summers are milder than in North America. This

5762-487: The islands of Trinidad and Tobago ( Trinidad Island and Tobago Island etc.), the State of Nueva Esparta , and the Federal Dependencies of Venezuela sit on the northern portion of the South American continental shelf and are sometimes considered parts of the continent. Geopolitically, all the island countries and territories in the Caribbean have generally been grouped as a subregion of North America instead. By contrast, Aves Island (administered by Venezuela ) and

5848-466: The land outside Europe should be an exclusive duopoly between the two countries. The treaty established an imaginary line along a north–south meridian 370 leagues west of the Cape Verde Islands , roughly 46° 37' W. In terms of the treaty, all land to the west of the line (known to comprise most of the South American soil) would belong to Spain, and all land to the east, to Portugal. As accurate measurements of longitude were impossible at that time,

5934-512: The limits set by the EU. However, mechanical sorting and drying was found to eliminate 98% of aflatoxins; a 2003 EU ban on importation was rescinded after new tolerance levels were set. The nuts may contain traces of radium , a radioactive element, with a kilogram of nuts containing an activity between 40 and 260 becquerels (1 and 7 nanocuries ). This level of radium is small, although higher than in other common foods. According to Oak Ridge Associated Universities , elevated levels of radium in

6020-476: The line was not strictly enforced, resulting in a Portuguese expansion of Brazil across the meridian. Beginning in the 1530s, the people and natural resources of South America were repeatedly exploited by foreign conquistadors , first from Spain and later from Portugal. These competing colonial nations claimed the land and resources as their own and divided it into colonies. European infectious diseases ( smallpox , influenza , measles , and typhus ) – to which

6106-402: The mid-17th century. It became a Dutch colony in 1667. The indigenous peoples of the Americas in various European colonies were forced to work in European plantations and mines; along with enslaved Africans who were also introduced in the proceeding centuries via the slave trade . European colonists were heavily dependent on indigenous labor during the initial phases of settlement to maintain

6192-436: The primary pollinators in different areas, and different times of year. Brazil nuts have been harvested from plantations, but production is low and is currently not economically viable. The fruit takes 14 months to mature after pollination of the flowers. The fruit itself is a large capsule 10–15 cm (4–6 in) in diameter, resembling a coconut endocarp in size and weighing up to 2 kg (4 lb 7 oz). It has

6278-665: The same way as jeweler's rouge , while charcoal from the shells can be used to purify water. South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere , with a considerably smaller portion in the Northern Hemisphere . It can also be described as the southern subregion of the Americas . South America is bordered on

6364-494: The seeds were found in three sites, 9.4% were found in four sites and 1.3% were found in five sites over a 3-month period. These results, and those from other studies, demonstrate the dynamic nature of the food supplies of scatter hoarding animals. Group-foraging common ravens , ( Corvus corax ), scatter hoard their food and also raid the caches made by others. Cachers withdraw from conspecifics when hiding their food and most often place their caches behind structures, obstructing

6450-451: The soil does not directly cause the concentration of radium, but "the very extensive root system of the tree" can concentrate naturally occurring radioactive material, when present in the soil. Radium can be concentrated in nuts only if it is present in the soil. Brazil nuts also contain barium , a metal with a chemical behavior quite similar to radium. While barium, if ingested, can have toxic effects, such as weakness, vomiting, or diarrhea,

6536-421: The subsequent appearance of permanent human settlements allowed for the multiple and overlapping beginnings of civilizations in South America. One of the earliest known South American civilizations was Caral–Supe , on the central Peruvian coast. Though a pre-ceramic culture, the monumental architecture of Caral–Supe created one of the first cities of the world, generally dated to around 3500 BC, at Huaricanga in

6622-411: The subsistence economy, and natives were often captured by expeditions. The importation of African slaves began midway through the 16th century, but the enslavement of indigenous peoples continued well into the 17th and 18th centuries. The Atlantic slave trade brought enslaved Africans primarily to South American colonies, beginning with the Portuguese since 1502. The main destinations of this phase were

6708-647: The supply in secret. There are only two species in which kin selection has resulted in a shared food store, i.e. beavers ( Castor canadensis ) and acorn woodpeckers ( Melanerpes formicivorous ); the former live in family groups and construct winter larders of submerged branches, while the latter are unusual in that they construct a conspicuous communal larder. Wolves , foxes , and coyotes identify their food caches by scent-marking them, usually after they have been emptied. Pilferage occurs when one animal takes food from another animal's larder. Some species experience high levels of cache pilferage, up to 30% of

6794-560: The supply per day. Models of scatter hoarding suggested the value of cached food is equal to the hoarder's ability to retrieve it. It has been observed that members of certain species, such as rodents and chickadees , act as both hoarder and pilferer. In other words, pilfering can be reciprocal and, thus, tolerable. Animals recache the food that they've pilfered from other animal's caches. For example, 75% percent of mildly radioactive (thus traceable ) Jeffrey pine seeds cached by yellow pine chipmunks were found in two cache sites, 29% of

6880-431: The upper and lower whorls (see illustration above). The capsule contains a small hole at one end, which enables large rodents like the agouti to gnaw it open. They then eat some of the seeds inside while burying others for later use; some of these are able to germinate into new Brazil nut trees. Most of the seeds are "planted" by the agoutis in caches during wet season , and the young saplings may have to wait years, in

6966-607: The view of potential observers. Raiders watch inconspicuously and keep at a distance to cachers close to their cache sites. In response to the presence of potential raiders or because of their initial movements towards caches, the cachers frequently interrupt caching, change cache sites, or recover their food items. These behaviors suggest that ravens are capable of withholding information about their intentions, which may qualify as tactical deception . Similarly, Eurasian jays ( Garrulus glandarius ) when being watched by another jay, prefer to cache food behind an opaque barrier rather than

7052-594: The west by the Pacific Ocean and on the north and east by the Atlantic Ocean ; North America and the Caribbean Sea lie to the northwest. The continent includes twelve sovereign states : Argentina , Bolivia , Brazil , Chile , Colombia , Ecuador , Guyana , Paraguay , Peru , Suriname , Uruguay , and Venezuela ; two dependent territories : the Falkland Islands and South Georgia and

7138-573: The western European architectural style to the continent, and helped to improve infrastructures like bridges, roads, and the sewer system of the cities they discovered or conquered. They also significantly increased economic and trade relations, not just between the old and new world but between the different South American regions and peoples. Finally, with the expansion of the Portuguese and Spanish languages, many cultures that were previously separated became united through that of Latin American. Guyana

7224-472: Was Chile in 1823, Uruguay in 1830, Bolivia in 1831, Guyana in 1833, Colombia and Ecuador in 1851, Argentina in 1853, Peru and Venezuela in 1854, Suriname in 1863, Paraguay in 1869, and in 1888 Brazil was the last South American nation and the last country in western world to abolish slavery. The European Peninsular War (1807–14), a theater of the Napoleonic Wars , changed the political situation of

7310-689: Was a useful form of agriculture. The Mapuche in Central and Southern Chile resisted the European and Chilean settlers, waging the Arauco War for more than 300 years. In 1494, Portugal and Spain, the two great maritime European powers of that time, on the expectation of new lands being discovered in the west, signed the Treaty of Tordesillas , by which they agreed, with the support of the Pope, that all

7396-562: Was initially colonized by the Dutch before coming under British control , though there was a brief period during the Napoleonic Wars when it was occupied by the French . The region was initially partitioned between the Dutch, French and British before fully coming under the control of Britain. Suriname was first explored by the Spanish in the 16th century and then settled by the English in

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