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68-489: The Drosophilidae are a diverse, cosmopolitan family of flies , which includes species called fruit flies, although they are more accurately referred to as vinegar or pomace flies. Another distantly related family of flies, Tephritidae , are true fruit flies because they are frugivorous, and include apple maggot flies and many pests. The best known species of the Drosophilidae is Drosophila melanogaster , within

136-645: A broad range of environmental tolerances or from rapid dispersal compared to the time needed for speciation . Biogeography Knowledge of spatial variation in the numbers and types of organisms is as vital to us today as it was to our early human ancestors , as we adapt to heterogeneous but geographically predictable environments . Biogeography is an integrative field of inquiry that unites concepts and information from ecology , evolutionary biology , taxonomy , geology , physical geography , palaeontology , and climatology . Modern biogeographic research combines information and ideas from many fields, from

204-476: A combination of historical factors such as: speciation , extinction , continental drift , and glaciation . Through observing the geographic distribution of species, we can see associated variations in sea level , river routes, habitat, and river capture . Additionally, this science considers the geographic constraints of landmass areas and isolation, as well as the available ecosystem energy supplies. Over periods of ecological changes, biogeography includes

272-417: A common vector in propagating acetic acid bacteria in nature. This often ruins the alcohol fermentation process and can ruin beer or wine by turning it into vinegar . There are sinking traps available on the market for this nuisance, but one quick way to strongly limit the extent of it is to vacuum clean the flies both at rest and in their slow flight. The diagnostic characteristics for Drosophilidae include

340-509: A comprehensive phylogenetic classification of the genera based on both molecular and morphological characters has been published. Cosmopolitan distribution In biogeography , a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats ; most cosmopolitan species are known to be highly adaptable to a range of climatic and environmental conditions, though this

408-421: A global scale. GIS can show certain processes on the earth's surface like whale locations, sea surface temperatures , and bathymetry. Current scientists also use coral reefs to delve into the history of biogeography through the fossilized reefs. Two global information systems are either dedicated to, or have strong focus on, biogeography (in the form of the spatial location of observations of organisms), namely

476-419: A great impact on Charles Darwin , who was inspired to consider species adaptations and evolution after learning about botanical geography. De Candolle was the first to describe the differences between the small-scale and large-scale distribution patterns of organisms around the globe. Several additional scientists contributed new theories to further develop the concept of biogeography. Charles Lyell developed

544-413: A number of methods have been developed to produce arguably more complete "predictive" or "modelled" distributions for species based on their associated environmental or other preferences (such as availability of food or other habitat requirements); this approach is known as either Environmental niche modelling (ENM) or Species distribution modelling (SDM). Depending on the reliability of the source data and

612-659: A planet too large for local populations to interbreed routinely with each other include genetic effects such as ring species , such as in the Larus gulls, and the formation of clines such as in Drosophila . Cosmopolitan distributions can be observed both in extinct and extant species. For example, Lystrosaurus was cosmopolitan in the Early Triassic after the Permian-Triassic extinction event . In

680-987: A result of tectonic uplift (or subsidence ), natural damming created by a landslide , or headward or lateral erosion of the watershed between adjacent basins. Biogeography is a synthetic science, related to geography , biology , soil science , geology , climatology , ecology and evolution . Some fundamental concepts in biogeography include: The study of comparative biogeography can follow two main lines of investigation: There are many types of biogeographic units used in biogeographic regionalisation schemes, as there are many criteria ( species composition , physiognomy , ecological aspects) and hierarchization schemes: biogeographic realms (ecozones), bioregions ( sensu stricto ), ecoregions , zoogeographical regions , floristic regions , vegetation types, biomes , etc. The terms biogeographic unit, biogeographic area can be used for these categories, regardless of rank. In 2008, an International Code of Area Nomenclature

748-535: A serious agricultural pest. The leaf mining Scaptomyza flava , which is nested in the genus Drosophila phylogenetically , is an obligate leaf miner of mustard plants, including the model plant Arabidopsis thaliana and is a major pest of salad brassicas in New Zealand and an emerging pest of canola in the UK. Drosophila repleta larvae inhabit drains and spread bacteria. Fruit flies in general are considered as

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816-443: A species; another example, the rock dove (commonly referred to as a ' pigeon '), in addition to having been bred domestically for centuries, now occurs in most urban areas around the world. The extreme opposite of a cosmopolitan species is an endemic (native) species, or one that is found only in a single geographical location. Endemism usually results in organisms with specific adaptations to one particular climate or region, and

884-440: A whole suite of predictor variables for biogeographic analysis, including satellite imaging and processing of the Earth. Two main types of satellite imaging that are important within modern biogeography are Global Production Efficiency Model (GLO-PEM) and Geographic Information Systems (GIS). GLO-PEM uses satellite-imaging gives "repetitive, spatially contiguous, and time specific observations of vegetation". These observations are on

952-424: Is an alternate view than that of Linnaeus. Buffon's law eventually became a principle of biogeography by explaining how similar environments were habitats for comparable types of organisms. Buffon also studied fossils which led him to believe that the Earth was over tens of thousands of years old, and that humans had not lived there long in comparison to the age of the Earth. Following the period of exploration came

1020-467: Is evidence to support that pathogens living within certain flies are beneficial to the behavior and survival of the host. One such example of this is in the fly Scaptomyza flava, which carries the pathogen Pseudomonas syringae in exchange for the pathogen damaging the anti-herbivore defenses of plants in the family Brassicaceae , which are the main food source for the fly. The family contains more than 4,000 species classified under 75 genera. Recently,

1088-539: Is home to some indigenous species within the Myrmeleontidae, but nonetheless no one species, nor even genus, of the Myrmeleontidae is cosmopolitan. Conversely, partly as a result of human introduction of unnatural apiculture to the New World, Apis mellifera probably is the only cosmopolitan member of its family; the rest of the family Apidae have modest distributions. Even where a cosmopolitan population

1156-404: Is in the geological similarities between varying locations around the globe, the geographic distribution of some fossils (including the mesosaurs ) on various continents, and the jigsaw puzzle shape of the landmasses on Earth. Though Wegener did not know the mechanism of this concept of Continental Drift, this contribution to the study of biogeography was significant in the way that it shed light on

1224-501: Is not always so. Killer whales ( orcas ) are among the most well-known cosmopolitan species on the planet, as they maintain several different resident and transient (migratory) populations in every major oceanic body on Earth, from the Arctic Circle to Antarctica and every coastal and open-water region in-between. Such a taxon (usually a species ) is said to have a cosmopolitan distribution, or exhibit cosmopolitanism , as

1292-473: Is on how the environment and humans affect the distribution of species as well as other manifestations of Life such as species or genetic diversity. Biogeography is being applied to biodiversity conservation and planning, projecting global environmental changes on species and biomes, projecting the spread of infectious diseases, invasive species, and for supporting planning for the establishment of crops. Technological evolving and advances have allowed for generating

1360-541: Is recognised as a single species, such as indeed Apis mellifera , there generally will be variation between regional sub-populations. Such variation commonly is at the level of subspecies , varieties or morphs , whereas some variation is too slight or inconsistent for formal recognition. For an example of subspecific variation, consider the East African lowland honey bee ( Apis mellifera scutellata )—best known for being hybridized with various European subspecies of

1428-614: The Age of Enlightenment in Europe, which attempted to explain the patterns of biodiversity observed by Buffon and Linnaeus. At the birth of the 19th century, Alexander von Humboldt, known as the "founder of plant geography", developed the concept of physique generale to demonstrate the unity of science and how species fit together. As one of the first to contribute empirical data to the science of biogeography through his travel as an explorer, he observed differences in climate and vegetation. The Earth

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1496-569: The Arctic tern occur from the Arctic to the Southern Ocean , but at any one season of the year they are likely to be largely in passage or concentrated at only one end of the range. Also, some such species breed only at one end of the range. Seen purely as an aspect of cosmopolitanism, such distributions could be seen as temporal, seasonal variations. Other complications of cosmopolitanism on

1564-713: The Global Biodiversity Information Facility (GBIF: 2.57 billion species occurrence records reported as at August 2023) and, for marine species only, the Ocean Biodiversity Information System (OBIS, originally the Ocean Biogeographic Information System : 116 million species occurrence records reported as at August 2023), while at a national scale, similar compilations of species occurrence records also exist such as

1632-500: The distribution , say, complete absence from Australia. Terminology varies, and there is some debate whether the true opposite of endemism is pandemism or cosmopolitanism. A related concept in biogeography is that of oceanic cosmopolitanism and endemism. Rather than allow ubiquitous travel, the World Ocean is complicated by physical obstacles such as temperature gradients. These prevent migration of tropical species between

1700-428: The genus Drosophila , also called the "fruit fly." Drosophila melanogaster is used extensively for studies concerning genetics , development, physiology , ecology and behaviour . Many fundamental biological mechanisms were discovered first in D. melanogaster. The fruit fly is mostly composed of post-mitotic cells, has a very short lifespan, and shows gradual aging. As in other species, temperature influences

1768-462: The housefly is highly cosmopolitan, yet is neither oceanic nor polar in its distribution. The term pandemism also is in use, but not all authors are consistent in the sense in which they use the term; some speak of pandemism mainly in referring to diseases and pandemics , and some as a term intermediate between endemism and cosmopolitanism, in effect regarding pandemism as subcosmopolitanism . This means near cosmopolitanism, but with major gaps in

1836-483: The western honey bee to create the so-called " African killer bee "—and the Cape bee , which is the subspecies Apis mellifera capensis ; both of them are in the same cosmopolitan species Apis mellifera , but their ranges barely overlap. Other cosmopolitan species, such as the house sparrow and osprey , present similar examples, but in yet other species there are less familiar complications: some migratory birds such as

1904-479: The 36 volume Histoire Naturelle, générale et particulière , in which he argued that varying geographical regions would have different forms of life. This was inspired by his observations comparing the Old and New World, as he determined distinct variations of species from the two regions. Buffon believed there was a single species creation event, and that different regions of the world were homes for varying species, which

1972-509: The Amazon basin, Orinoco basin, and Guianas ) with an exceptionally low (flat) topographic relief, the many waterways have had a highly reticulated history over geological time . In such a context, stream capture is an important factor affecting the evolution and distribution of freshwater organisms. Stream capture occurs when an upstream portion of one river drainage is diverted to the downstream portion of an adjacent basin. This can happen as

2040-664: The Atlantic and Indian/Pacific oceans. Conversely, the Northern marine regions and Southern Ocean are separated by the tropics , too warm for many species to traverse. Another aspect of cosmopolitanism is that of ecological limitations. A species that is apparently cosmopolitan because it occurs in all oceans might in fact occupy only littoral zones , or only particular ranges of depths, or only estuaries , for example. Analogously, terrestrial species might be present only in forests, or mountainous regions, or sandy arid regions or

2108-597: The Gómez Farias Region, Tamaulipas, Mexico , which has been described as "ground-breaking" and "a classic treatise in historical biogeography". Martin applied several disciplines including ecology , botany , climatology , geology , and Pleistocene dispersal routes to examine the herpetofauna of a relatively small and largely undisturbed area, but ecologically complex, situated on the threshold of temperate – tropical (nearctic and neotropical) regions, including semiarid lowlands at 70 meters elevation and

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2176-619: The Indian Ocean was much narrower than it is today, and that South America was closer to the Antarctic, one would be hard pressed to explain the presence of many "ancient" lineages of perching birds in Africa, as well as the mainly South American distribution of the suboscines . Paleobiogeography also helps constrain hypotheses on the timing of biogeographic events such as vicariance and geodispersal , and provides unique information on

2244-452: The Origin of Species were devoted to geographical distribution. The first discoveries that contributed to the development of biogeography as a science began in the mid-18th century, as Europeans explored the world and described the biodiversity of life. During the 18th century most views on the world were shaped around religion and for many natural theologists, the bible. Carl Linnaeus , in

2312-482: The Theory of Continental Drift in 1912, though it was not widely accepted until the 1960s. This theory was revolutionary because it changed the way that everyone thought about species and their distribution around the globe. The theory explained how continents were formerly joined in one large landmass, Pangea , and slowly drifted apart due to the movement of the plates below Earth's surface. The evidence for this theory

2380-603: The Theory of Uniformitarianism after studying fossils. This theory explained how the world was not created by one sole catastrophic event, but instead from numerous creation events and locations. Uniformitarianism also introduced the idea that the Earth was actually significantly older than was previously accepted. Using this knowledge, Lyell concluded that it was possible for species to go extinct. Since he noted that Earth's climate changes, he realized that species distribution must also change accordingly. Lyell argued that climate changes complemented vegetation changes, thus connecting

2448-639: The U.K. National Biodiversity Network , the Atlas of Living Australia , and many others. In the case of the oceans, in 2017 Costello et al. analyzed the distribution of 65,000 species of marine animals and plants as then documented in OBIS, and used the results to distinguish 30 distinct marine realms, split between continental-shelf and offshore deep-sea areas. Since it is self evident that compilations of species occurrence records cannot cover with any completeness, areas that have received either limited or no sampling,

2516-414: The anticipated effects of climate change can also be used to show potential changes in species distributions that may occur in the future based on such scenarios. Paleobiogeography goes one step further to include paleogeographic data and considerations of plate tectonics . Using molecular analyses and corroborated by fossils , it has been possible to demonstrate that perching birds evolved first in

2584-421: The biotic and abiotic features of the Earth in his book, Cosmos . Augustin de Candolle contributed to the field of biogeography as he observed species competition and the several differences that influenced the discovery of the diversity of life. He was a Swiss botanist and created the first Laws of Botanical Nomenclature in his work, Prodromus. He discussed plant distribution and his theories eventually had

2652-475: The development of biogeography as a science. The scientific theory of biogeography grows out of the work of Alexander von Humboldt (1769–1859), Francisco Jose de Caldas (1768–1816), Hewett Cottrell Watson (1804–1881), Alphonse de Candolle (1806–1893), Alfred Russel Wallace (1823–1913), Philip Lutley Sclater (1829–1913) and other biologists and explorers. The patterns of species distribution across geographical areas can usually be explained through

2720-420: The development of the fields of conservation biology and landscape ecology . Classic biogeography has been expanded by the development of molecular systematics , creating a new discipline known as phylogeography . This development allowed scientists to test theories about the origin and dispersal of populations, such as island endemics . For example, while classic biogeographers were able to speculate about

2788-504: The environmental surroundings to varying species. This largely influenced Charles Darwin in his development of the theory of evolution. Charles Darwin was a natural theologist who studied around the world, and most importantly in the Galapagos Islands . Darwin introduced the idea of natural selection, as he theorized against previously accepted ideas that species were static or unchanging. His contributions to biogeography and

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2856-511: The evolution of herbivory and specialist on toxic mustard leaves. Generally, drosophilids are considered to be nuisance flies rather than pests, since most species breed in rotting material. Zaprionus indianus is unusual among Drosophilidae species in being a serious, primary pest of at least one commercial fruit, figs in Brazil. Another species, Drosophila suzukii , infests thin-skinned fruit such as raspberries and cherries and can be

2924-597: The factors affecting organism distribution, and to predict future trends in organism distribution. Often mathematical models and GIS are employed to solve ecological problems that have a spatial aspect to them. Biogeography is most keenly observed on the world's islands . These habitats are often much more manageable areas of study because they are more condensed than larger ecosystems on the mainland. Islands are also ideal locations because they allow scientists to look at habitats that new invasive species have only recently colonized and can observe how they disperse throughout

2992-458: The formation of regional biotas. For example, data from species-level phylogenetic and biogeographic studies tell us that the Amazonian teleost fauna accumulated in increments over a period of tens of millions of years, principally by means of allopatric speciation, and in an arena extending over most of the area of tropical South America (Albert & Reis 2011). In other words, unlike some of

3060-483: The former Lifemapper project at the University of Kansas (now continued as a part of BiotaPhy ) and AquaMaps , which as at 2023 contain modelled distributions for around 200,000 terrestrial, and 33,000 species of teleosts , marine mammals and invertebrates, respectively. One advantage of ENM/SDM is that in addition to showing current (or even past) modelled distributions, insertion of changed parameters such as

3128-655: The geographical distribution of organisms around the globe. Alfred Russel Wallace studied the distribution of flora and fauna in the Amazon Basin and the Malay Archipelago in the mid-19th century. His research was essential to the further development of biogeography, and he was later nicknamed the "father of Biogeography". Wallace conducted fieldwork researching the habits, breeding and migration tendencies, and feeding behavior of thousands of species. He studied butterfly and bird distributions in comparison to

3196-411: The importance of environmental and geographic similarities or differences as a result of climate and other pressures on the planet. Importantly, late in his career Wegener recognised that testing his theory required measurement of continental movement rather than inference from fossils species distributions. In 1958 paleontologist Paul S. Martin published A Biogeography of Reptiles and Amphibians in

3264-559: The island and change it. They can then apply their understanding to similar but more complex mainland habitats. Islands are very diverse in their biomes , ranging from the tropical to arctic climates. This diversity in habitat allows for a wide range of species study in different parts of the world. One scientist who recognized the importance of these geographic locations was Charles Darwin , who remarked in his journal "The Zoology of Archipelagoes will be well worth examination". Two chapters in On

3332-402: The life history of the animal. Several genes have been identified that can be manipulated to extend the lifespan of these insects. Additionally, Drosophila subobscura , also within the genus Drosophila , has been reputed as a model organism for evolutionary-biological studies, along with D. sechellia for the evolution of host specialization on the toxic noni fruit and Scaptomyza flava for

3400-430: The like. Such distributions might be patchy, or extended, but narrow. Factors of such a nature are taken widely for granted, so they seldom are mentioned explicitly in mentioning cosmopolitan distributions. Cosmopolitanism of a particular species or variety should not be confused with cosmopolitanism of higher taxa. For example, the family Myrmeleontidae is cosmopolitan in the sense that every continent except Antarctica

3468-488: The mid-18th century, improved our classifications of organisms through the exploration of undiscovered territories by his students and disciples. When he noticed that species were not as perpetual as he believed, he developed the Mountain Explanation to explain the distribution of biodiversity; when Noah's ark landed on Mount Ararat and the waters receded, the animals dispersed throughout different elevations on

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3536-474: The modern world, the orca , the blue whale , and the great white shark all have cosmopolitan distribution, extending over most of the Earth's oceans . The wasp Copidosoma floridanum is another example, as it is found around the world. Other examples include humans , cats , dogs , the western honey bee , brown rats , the foliose lichen Parmelia sulcata , and the mollusc genus Mytilus . The term can also apply to some diseases. It may result from

3604-515: The most important and consequential developments in biogeography has been to show how multiple organisms, including mammals like monkeys and reptiles like squamates , overcame barriers such as large oceans that many biogeographers formerly believed were impossible to cross. See also Oceanic dispersal . Biogeography now incorporates many different fields including but not limited to physical geography, geology, botany and plant biology, zoology, general biology, and modelling. A biogeographer's main focus

3672-505: The mountain. This showed different species in different climates proving species were not constant. Linnaeus' findings set a basis for ecological biogeography. Through his strong beliefs in Christianity, he was inspired to classify the living world, which then gave way to additional accounts of secular views on geographical distribution. He argued that the structure of an animal was very closely related to its physical surroundings. This

3740-683: The nature of the models employed (including the scales for which data are available), maps generated from such models may then provide better representations of the "real" biogeographic distributions of either individual species, groups of species, or biodiversity as a whole, however it should also be borne in mind that historic or recent human activities (such as hunting of great whales , or other human-induced exterminations) may have altered present-day species distributions from their potential "full" ecological footprint. Examples of predictive maps produced by niche modelling methods based on either GBIF (terrestrial) or OBIS (marine, plus some freshwater) data are

3808-476: The northernmost cloud forest in the western hemisphere at over 2200 meters. The publication of The Theory of Island Biogeography by Robert MacArthur and E.O. Wilson in 1967 showed that the species richness of an area could be predicted in terms of such factors as habitat area, immigration rate and extinction rate. This added to the long-standing interest in island biogeography . The application of island biogeography theory to habitat fragments spurred

3876-583: The origins of species in the Hawaiian Islands , phylogeography allows them to test theories of relatedness between these populations and putative source populations on various continents, notably in Asia and North America . Biogeography continues as a point of study for many life sciences and geography students worldwide, however it may be under different broader titles within institutions such as ecology or evolutionary biology. In recent years, one of

3944-495: The physiological and ecological constraints on organismal dispersal to geological and climatological phenomena operating at global spatial scales and evolutionary time frames. The short-term interactions within a habitat and species of organisms describe the ecological application of biogeography. Historical biogeography describes the long-term, evolutionary periods of time for broader classifications of organisms. Early scientists, beginning with Carl Linnaeus , contributed to

4012-611: The presence of an incomplete subcostal vein , two breaks in the costal vein, a small anal cell in the wing , convergent postocellar bristles; and usually three frontal bristles on each side of the head, one directed forward and the other two directed rearward. More extensive identification characteristics can be found in " Drosophila : A Guide to Species Identification and Use" by Therese A. Markow and Patrick O'Grady, (Academic Press, 2005) ISBN   0-12-473052-3 or " Drosophila : A Laboratory Handbook" by M. Ashburner , K. Golic, S. Hawley, (Cold Spring Harbor Laboratory Press, 2005). There

4080-448: The presence or absence of geographical barriers. His observations led him to conclude that the number of organisms present in a community was dependent on the amount of food resources in the particular habitat. Wallace believed species were dynamic by responding to biotic and abiotic factors. He and Philip Sclater saw biogeography as a source of support for the theory of evolution as they used Darwin's conclusion to explain how biogeography

4148-420: The region of Australia or the adjacent Antarctic (which at that time lay somewhat further north and had a temperate climate). From there, they spread to the other Gondwanan continents and Southeast Asia – the part of Laurasia then closest to their origin of dispersal – in the late Paleogene , before achieving a global distribution in the early Neogene . Not knowing that at the time of dispersal,

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4216-596: The species would likely face challenges if placed in a different environment. There are far more examples of endemic species than cosmopolitan species; one example being the snow leopard , a species found only in Central Asian mountain ranges, an environment to which the cats have adapted over millions of years. The caveat "in appropriate habitat" is used to qualify the term "cosmopolitan distribution", excluding in most instances polar regions, extreme altitudes, oceans, deserts, or small, isolated islands. For example,

4284-428: The study of plant and animal species in: their past and/or present living refugium habitat ; their interim living sites; and/or their survival locales. As writer David Quammen put it, "...biogeography does more than ask Which species? and Where . It also asks Why? and, what is sometimes more crucial, Why not? ." Modern biogeography often employs the use of Geographic Information Systems (GIS), to understand

4352-416: The theory of evolution were different from those of other explorers of his time, because he developed a mechanism to describe the ways that species changed. His influential ideas include the development of theories regarding the struggle for existence and natural selection. Darwin's theories started a biological segment to biogeography and empirical studies, which enabled future scientists to develop ideas about

4420-447: The well-known insular faunas ( Galapagos finches , Hawaiian drosophilid flies, African rift lake cichlids ), the species-rich Amazonian ichthyofauna is not the result of recent adaptive radiations . For freshwater organisms, landscapes are divided naturally into discrete drainage basins by watersheds , episodically isolated and reunited by erosional processes. In regions like the Amazon Basin (or more generally Greater Amazonia,

4488-400: Was divided into regions which he defined as tropical, temperate, and arctic and within these regions there were similar forms of vegetation. This ultimately enabled him to create the isotherm, which allowed scientists to see patterns of life within different climates. He contributed his observations to findings of botanical geography by previous scientists, and sketched this description of both

4556-529: Was important to a George Louis Buffon's rival theory of distribution. Closely after Linnaeus, Georges-Louis Leclerc, Comte de Buffon observed shifts in climate and how species spread across the globe as a result. He was the first to see different groups of organisms in different regions of the world. Buffon saw similarities between some regions which led him to believe that at one point continents were connected and then water separated them and caused differences in species. His hypotheses were described in his work,

4624-502: Was similar to a record of species inheritance. Key findings, such as the sharp difference in fauna either side of the Wallace Line , and the sharp difference that existed between North and South America prior to their relatively recent faunal interchange , can only be understood in this light. Otherwise, the field of biogeography would be seen as a purely descriptive one. Moving on to the 20th century, Alfred Wegener introduced

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