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130-547: Fins first evolved on fish as a means of locomotion. Fish fins are used to generate thrust and control the subsequent motion. Fish and other aquatic animals, such as cetaceans , actively propel and steer themselves with pectoral and tail fins . As they swim, they use other fins, such as dorsal and anal fins , to achieve stability and refine their maneuvering. The fins on the tails of cetaceans, ichthyosaurs , metriorhynchids , mosasaurs and plesiosaurs are called flukes . Foil shaped fins generate thrust when moved,

260-593: A crown group of ray-finned fish that can protrude their jaws . The tetrapods , a mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since the Late Paleozoic , evolved from lobe-finned fish during the Carboniferous , developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish"

390-414: A differential gear to twin contra-rotating propellers. If one drum was rotated faster than the other, then the rudder was activated. The other ends of the wires were connected to steam-powered winding engines, which were arranged so that speeds could be varied within fine limits, giving sensitive steering control for the torpedo. The torpedo attained a speed of 20 knots (37 km/h; 23 mph) using

520-579: A paraphyletic group and for this reason, the class Pisces seen in older reference works is no longer used in formal classifications. Traditional classification divides fish into three extant classes (Agnatha, Chondrichthyes, and Osteichthyes), and with extinct forms sometimes classified within those groups, sometimes as their own classes. Fish account for more than half of vertebrate species. As of 2016, there are over 32,000 described species of bony fish, over 1,100 species of cartilaginous fish, and over 100 hagfish and lampreys. A third of these fall within

650-426: A paraphyletic group. Fish have been an important natural resource for humans since prehistoric times, especially as food . Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in the ocean. Fish are caught for recreation , or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds . Fish have had

780-545: A propulsive efficiency greater than 90%. Fish can accelerate and maneuver much more effectively than boats or submarine , and produce less water disturbance and noise. This has led to biomimetic studies of underwater robots which attempt to emulate the locomotion of aquatic animals. An example is the Robot Tuna built by the Institute of Field Robotics , to analyze and mathematically model thunniform motion . In 2005,

910-428: A swimmer or underwater diver Surfboard fins provide surfers with means to maneuver and control their boards. Contemporary surfboards often have a centre fin and two cambered side fins. The bodies of reef fishes are often shaped differently from open water fishes . Open water fishes are usually built for speed, streamlined like torpedoes to minimise friction as they move through the water. Reef fish operate in

1040-489: A "torpedo" is an underwater self-propelled explosive, but historically, the term also applied to primitive naval mines and spar torpedoes . These were used on an ad hoc basis during the early modern period up to the late 19th century. In the early 17th century, the Dutchman Cornelius Drebbel , in the employ of King James I of England , invented the spar torpedo; he attached explosives to the end of

1170-431: A batch of 50 as torpedo production at home and Rijeka could not meet demand. By World War I, Whitehead's torpedo remained a worldwide success, and his company was able to maintain a monopoly on torpedo production. By that point, his torpedo had grown to a diameter of 18 inches with a maximum speed of 30.5 knots (56.5 km/h; 35.1 mph) with a warhead weighing 170 pounds (77 kg). Whitehead faced competition from

1300-763: A beam affixed to one of his submarines. These were used (to little effect) during the English expeditions to La Rochelle in 1626. The first use of a torpedo by a submarine was in 1775, by the American Turtle , which attempted to lay a bomb with a timed fuse on the hull of HMS  Eagle during the American Revolutionary War , but failed in the attempt. In the early 1800s, the American inventor Robert Fulton , while in France, "conceived

1430-645: A demonstration in late 1869, and in 1870 a batch of torpedoes was ordered. In 1871, the British Admiralty paid Whitehead £ 15,000 for certain of his developments and production started at the Royal Laboratories in Woolwich the following year. In the 1880s, a British committee, informed by hydrodynamicist Dr. R. E. Froude , conducted comparative tests and determined that a blunt nose, contrary to prior assumptions, did not hinder speed: in fact,

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1560-608: A depth just beneath the ship, relying on a magnetic exploder to activate at the appropriate time. Germany, Britain, and the U.S. independently devised ways to do this; German and American torpedoes, however, suffered problems with their depth-keeping mechanisms, coupled with faults in magnetic pistols shared by all designs. Inadequate testing had failed to reveal the effect of the Earth's magnetic field on ships and exploder mechanisms, which resulted in premature detonation. The Kriegsmarine and Royal Navy promptly identified and eliminated

1690-429: A different reason. Unlike dolphins, these fish do not feel the bubbles, because they have bony fins without nerve endings. Nevertheless, they cannot swim faster because the cavitation bubbles create a vapor film around their fins that limits their speed. Lesions have been found on tuna that are consistent with cavitation damage. Scombrid fishes (tuna, mackerel and bonito) are particularly high-performance swimmers. Along

1820-529: A factory at St Tropez in 1890 that exported torpedoes to Brazil, The Netherlands, Turkey, and Greece. Whitehead purchased rights to the gyroscope of Ludwig Obry in 1888 but it was not sufficiently accurate, so in 1890 he purchased a better design to improve control of his designs, which came to be called the "Devil's Device". The firm of L. Schwartzkopff in Germany also produced torpedoes and exported them to Russia, Japan, and Spain. In 1885, Britain ordered

1950-552: A fish-like body shape through convergent evolution . Fishes of the World comments that "it is increasingly widely accepted that tetrapods, including ourselves, are simply modified bony fishes, and so we are comfortable with using the taxon Osteichthyes as a clade, which now includes all tetrapods". The biodiversity of extant fish is unevenly distributed among the various groups; teleosts , bony fishes able to protrude their jaws , make up 96% of fish species. The cladogram shows

2080-427: A hard skull , but lacking limbs with digits . Fish can be grouped into the more basal jawless fish and the more common jawed fish , the latter including all living cartilaginous and bony fish , as well as the extinct placoderms and acanthodians . Most fish are cold-blooded , their body temperature varying with the surrounding water, though some large active swimmers like white shark and tuna can hold

2210-613: A higher core temperature . Many fish can communicate acoustically with each other, such as during courtship displays . The earliest fish appeared during the Cambrian as small filter feeders ; they continued to evolve through the Paleozoic , diversifying into many forms. The earliest fish with dedicated respiratory gills and paired fins , the ostracoderms , had heavy bony plates that served as protective exoskeletons against invertebrate predators . The first fish with jaws ,

2340-518: A liquid, which then promptly and violently collapse. It can cause significant damage and wear. Cavitation damage can also occur to the tail fins of powerful swimming marine animals, such as dolphins and tuna. Cavitation is more likely to occur near the surface of the ocean, where the ambient water pressure is relatively low. Even if they have the power to swim faster, dolphins may have to restrict their speed because collapsing cavitation bubbles on their tail are too painful. Cavitation also slows tuna, but for

2470-436: A more spherical lens . Their retinas generally have both rods and cones (for scotopic and photopic vision ); many species have colour vision , often with three types of cone. Teleosts can see polarized light ; some such as cyprinids have a fourth type of cone that detects ultraviolet . Amongst jawless fish , the lamprey has well-developed eyes, while the hagfish has only primitive eyespots. Hearing too

2600-462: A night-time approach so that the target ship would be less able to defend itself. Fiske determined that the notional torpedo bomber should descend rapidly in a sharp spiral to evade enemy guns, then when about 10 to 20 feet (3 to 6 m) above the water the aircraft would straighten its flight long enough to line up with the torpedo's intended path. The aircraft would release the torpedo at a distance of 1,500 to 2,000 yards (1,400 to 1,800 m) from

2730-512: A number of rotating fins, also called foils, wings, arms or blades. Propellers use the fins to translate torquing force to lateral thrust, thus propelling an aircraft or ship. Turbines work in reverse, using the lift of the blades to generate torque and power from moving gases or water. Cavitation can be a problem with high power applications, resulting in damage to propellers or turbines, as well as noise and loss of power. Cavitation occurs when negative pressure causes bubbles (cavities) to form in

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2860-478: A pair of 24.5-inch torpedoes from her port-side tube and claimed one hit. According to Ludovic Kennedy , "if true, [this is] the only instance in history of one battleship torpedoing another". The Royal Navy continued the development of oxygen-enriched air torpedoes with the 21 in. Mk. VII of the 1920s designed for the County-class cruisers although once again these were converted to run on normal air at

2990-407: A pancake, and will fit into fissures in rocks. Their pelvic and pectoral fins are designed differently, so they act together with the flattened body to optimise maneuverability. Some fishes, such as puffer fish , filefish and trunkfish , rely on pectoral fins for swimming and hardly use tail fins at all. Aristotle recognised the distinction between analogous and homologous structures , and made

3120-594: A production facility at the Royal Naval Torpedo Factory, Greenock , in 1910. These are now closed. Whitehead opened a new factory adjacent to Portland Harbour , England, in 1890, which continued making torpedoes until the end of World War II . Because orders from the RN were not as large as expected, torpedoes were mostly exported. A series of devices was produced at Rijeka, with diameters from 14 in (36 cm) upward. The largest Whitehead torpedo

3250-419: A protective bony cover or operculum . They are able to oxygenate their gills using muscles in the head. Some 400 species of fish in 50 families can breathe air, enabling them to live in oxygen-poor water or to emerge on to land. The ability of fish to do this is potentially limited by their single-loop circulation, as oxygenated blood from their air-breathing organ will mix with deoxygenated blood returning to

3380-450: A quantum radical pair mechanism . Torpedo A modern torpedo is an underwater ranged weapon launched above or below the water surface, self-propelled towards a target, and with an explosive warhead designed to detonate either on contact with or in proximity to the target. Historically, such a device was called an automotive, automobile, locomotive, or fish torpedo; colloquially a fish . The term torpedo originally applied to

3510-636: A radio guidance system for Allied torpedoes, it intended to use frequency-hopping technology to defeat the threat of jamming by the Axis powers . As radio guidance had been abandoned some years earlier, it was not pursued. Although the US Navy never adopted the technology, it did, in the 1960s, investigate various spread-spectrum techniques. Spread-spectrum techniques are incorporated into Bluetooth technology and are similar to methods used in legacy versions of Wi-Fi . This work led to their induction into

3640-582: A role in human culture through the ages, serving as deities , religious symbols, and as the subjects of art, books and movies. The word fish is inherited from Proto-Germanic , and is related to German Fisch , the Latin piscis and Old Irish īasc , though the exact root is unknown; some authorities reconstruct a Proto-Indo-European root * peysk- , attested only in Italic , Celtic , and Germanic . About 530 million years ago during

3770-441: A sense of touch and of hearing . Blind cave fish navigate almost entirely through the sensations from their lateral line system. Some fish, such as catfish and sharks, have the ampullae of Lorenzini , electroreceptors that detect weak electric currents on the order of millivolt. Vision is an important sensory system in fish. Fish eyes are similar to those of terrestrial vertebrates like birds and mammals, but have

3900-542: A success against an Austrian-Hungarian squadron , sinking the battleship SMS  Szent István with two torpedoes. The Royal Navy had been experimenting with ways to further increase the range of torpedoes during World War 1 using pure oxygen instead of compressed air, this work ultimately leading to the development of the oxygen-enriched air 24.5 in. Mk. I intended originally for the G3-class battlecruisers and N3 class battleships of 1921, both being cancelled due to

4030-485: A tail fin, jaws, skin covered with scales , and lays eggs. Each criterion has exceptions, creating a wide diversity in body shape and way of life. For example, some fast-swimming fish are warm-blooded, while some slow-swimming fish have abandoned streamlining in favour of other body shapes. Fish species are roughly divided equally between freshwater and marine (oceanic) ecosystems; there are some 15,200 freshwater species and around 14,800 marine species. Coral reefs in

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4160-616: A torpedo against the Ottoman cruiser "Medjidieh". The end of the Russo-Japanese War fuelled new theories, and the idea of dropping lightweight torpedoes from aircraft was conceived in the early 1910s by Bradley A. Fiske , an officer in the United States Navy . Awarded a patent in 1912, Fiske worked out the mechanics of carrying and releasing the aerial torpedo from a bomber , and defined tactics that included

4290-589: A true "land fish" as this worm-like catfish strictly lives among waterlogged leaf litter . Cavefish of multiple families live in underground lakes , underground rivers or aquifers . Like other animals, fish suffer from parasitism . Some species use cleaner fish to remove external parasites. The best known of these are the bluestreak cleaner wrasses of coral reefs in the Indian and Pacific oceans. These small fish maintain cleaning stations where other fish congregate and perform specific movements to attract

4420-600: A tubular device, designed to run underwater on its own, and powered by compressed air. The result was a submarine weapon, the Minenschiff (mine ship), the first modern self-propelled torpedo, officially presented to the Austrian Imperial Naval commission on 21 December 1866. The first trials were not successful as the weapon was unable to maintain a course at a steady depth. After much work, Whitehead introduced his "secret" in 1868 which overcame this. It

4550-422: A typical fish is adapted for efficient swimming by alternately contracting paired sets of muscles on either side of the backbone. These contractions form S-shaped curves that move down the body. As each curve reaches the tail fin, force is applied to the water, moving the fish forward. The other fins act as control surfaces like an aircraft's flaps, enabling the fish to steer in any direction. Since body tissue

4680-581: A variety of devices, most of which would today be called mines . From about 1900, torpedo has been used strictly to designate a self-propelled underwater explosive device. While the 19th-century battleship had evolved primarily with a view to engagements between armored warships with large-caliber guns , the invention and refinement of torpedoes from the 1860s onwards allowed small torpedo boats and other lighter surface vessels , submarines / submersibles , even improvised fishing boats or frogmen , and later light aircraft , to destroy large ships without

4810-506: A wire 1.0 millimetre (0.04 in) in diameter but later this was changed to 1.8 mm (0.07 in) to increase the speed to 27 knots (50 km/h; 31 mph). The torpedo was fitted with elevators controlled by a depth-keeping mechanism, and the fore and aft rudders operated by the differential between the drums. Brennan traveled to Britain, where the Admiralty examined the torpedo and found it unsuitable for shipboard use. However,

4940-465: Is an important sensory system in fish. Fish sense sound using their lateral lines and otoliths in their ears, inside their heads. Some can detect sound through the swim bladder. Some fish, including salmon, are capable of magnetoreception ; when the axis of a magnetic field is changed around a circular tank of young fish, they reorient themselves in line with the field. The mechanism of fish magnetoreception remains unknown; experiments in birds imply

5070-431: Is denser than water, fish must compensate for the difference or they will sink. Many bony fish have an internal organ called a swim bladder that allows them to adjust their buoyancy by increasing or decreasing the amount of gas it contains. The scales of fish provide protection from predators at the cost of adding stiffness and weight. Fish scales are often highly reflective; this silvering provides camouflage in

5200-522: Is still in limited service in the 21st century. The improved Mark VIII** was used in two particularly notable incidents; on 6 February 1945 the only intentional wartime sinking of one submarine by another while both were submerged took place when HMS Venturer sank the German submarine U-864 with four Mark VIII** torpedoes and on 2 May 1982 when the Royal Navy submarine HMS  Conqueror sank

5330-465: Is the biggest part of the brain; it is small in hagfish and lampreys , but very large in mormyrids , processing their electrical sense . The brain stem or myelencephalon controls some muscles and body organs, and governs respiration and osmoregulation . The lateral line system is a network of sensors in the skin which detects gentle currents and vibrations, and senses the motion of nearby fish, whether predators or prey. This can be considered both

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5460-474: The American Civil War , the term torpedo was used for what is today called a contact mine , floating on or below the water surface using an air-filled demijohn or similar flotation device. These devices were very primitive and apt to prematurely explode. They would be detonated on contact with the ship or after a set time, although electrical detonators were also occasionally used. USS  Cairo

5590-587: The Battle of Tsushima , Admiral Rozhestvensky 's flagship , the battleship Knyaz Suvorov , had been gunned to a wreck by Admiral Tōgō 's 12-inch gunned battleline . With the Russians sunk and scattering, Tōgō prepared for pursuit, and while doing so ordered his torpedo boat destroyers (TBDs) (mostly referred to as just destroyers in most written accounts) to finish off the Russian battleship. Knyaz Suvorov

5720-631: The Battle of the Atlantic and the Pacific War. Torpedo boats, such as MTBs , PT boats , or S-boats , enabled the relatively small but fast craft to carry enough firepower, in theory, to destroy a larger ship, though this rarely occurred in practice. The largest warship sunk by torpedoes from small craft in World War II was the British cruiser Manchester , sunk by Italian MAS boats on

5850-621: The Battle of the North Cape in December 1943, torpedo hits from British destroyers Savage and Saumarez slowed the German battleship Scharnhorst enough for the British battleship Duke of York to catch and sink her, and in May 1945 the British 26th Destroyer Flotilla (coincidentally led by Saumarez again) ambushed and sank Japanese heavy cruiser Haguro . During World War II , Hedy Lamarr and composer George Antheil developed

5980-581: The British Empire for more than fifteen years. The Royal Navy frigate HMS  Shah was the first naval vessel to fire a self-propelled torpedo in anger during the Battle of Pacocha against rebel Peruvian ironclad Huáscar on 29 May 1877. The Peruvian ship successfully outran the device. On 16 January 1878, the Turkish steamer Intibah became the first vessel to be sunk by self-propelled torpedoes, launched from torpedo boats operating from

6110-541: The Cambrian explosion , fishlike animals with a notochord and eyes at the front of the body, such as Haikouichthys , appear in the fossil record . During the late Cambrian , other jawless forms such as conodonts appear. Jawed vertebrates appear in the Silurian , with giant armoured placoderms such as Dunkleosteus . Jawed fish, too, appeared during the Silurian: the cartilaginous Chondrichthyes and

6240-544: The First Sino-Japanese War in 1894. At this time torpedo attacks were still very close range and very dangerous to the attackers. Several western sources reported that the Qing dynasty Imperial Chinese military, under the direction of Li Hongzhang , acquired electric torpedoes, which they deployed in numerous waterways, along with fortresses and numerous other modern military weapons acquired by China. At

6370-671: The Indo-Pacific constitute the center of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large river basins of tropical rainforests , especially the Amazon , Congo , and Mekong basins. More than 5,600 fish species inhabit Neotropical freshwaters alone, such that Neotropical fishes represent about 10% of all vertebrate species on the Earth. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams (e.g., char and gudgeon ) to

6500-557: The Sea Life London Aquarium displayed three robotic fish created by the computer science department at the University of Essex . The fish were designed to be autonomous, swimming around and avoiding obstacles like real fish. Their creator claimed that he was trying to combine "the speed of tuna, acceleration of a pike, and the navigating skills of an eel". The AquaPenguin , developed by Festo of Germany, copies

6630-645: The Tientsin Arsenal in 1876, the Chinese developed the capacity to manufacture these "electric torpedoes" on their own. Although a form of Chinese art, the Nianhua , depict such torpedoes being used against Russian ships during the Boxer Rebellion , whether they were actually used in battle against them is undocumented and unknown. The Russo-Japanese War (1904–1905) was the first great war of

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6760-750: The War Office proved more amenable, and in early August 1881, a special Royal Engineer committee was instructed to inspect the torpedo at Chatham and report back directly to the Secretary of State for War, Hugh Childers . The report strongly recommended that an improved model be built at government expense. In 1883 an agreement was reached between the Brennan Torpedo Company and the government. The newly appointed Inspector-General of Fortifications in England, Sir Andrew Clarke , appreciated

6890-702: The War of 1812 broke out, the Royal Navy established a blockade of the East Coast of the United States . During the war, American forces unsuccessfully attempted to destroy the British ship of the line HMS Ramillies while it was lying at anchor in New London, Connecticut 's harbor with torpedoes launched from small boats. This prompted the captain of Ramillies , Sir Thomas Hardy, 1st Baronet , to warn

7020-549: The Washington Naval Treaty . Initially, the Imperial Japanese Navy purchased Whitehead or Schwartzkopf torpedoes but by 1917, like the Royal Navy, they were conducting experiments with pure oxygen instead of compressed air. Because of explosions they abandoned the experiments but resumed them in 1926 and by 1933 had a working torpedo. They also used conventional wet heater torpedoes. In

7150-477: The abyssal and even hadal depths of the deepest oceans (e.g., cusk-eels and snailfish ), although none have been found in the deepest 25% of the ocean. The deepest living fish in the ocean so far found is a cusk-eel, Abyssobrotula galatheae , recorded at the bottom of the Puerto Rico Trench at 8,370 m (27,460 ft). In terms of temperature, Jonah's icefish live in cold waters of

7280-1206: The evolutionary relationships of all groups of living fishes (with their respective diversity ) and the tetrapods. Extinct groups are marked with a dagger (†); groups of uncertain placement are labelled with a question mark (?) and dashed lines (- - - - -). Jawless fishes (118 species: hagfish , lampreys ) [REDACTED] † Thelodonti , † Conodonta , † Anaspida [REDACTED] [REDACTED] [REDACTED] † Galeaspida [REDACTED] † Osteostraci [REDACTED] † Placodermi [REDACTED] † Acanthodii [REDACTED]  (>1,100 species: sharks , rays , chimaeras ) [REDACTED]  (2 species: coelacanths ) [REDACTED] Dipnoi (6 species: lungfish ) [REDACTED] Tetrapoda (>38,000 species, not considered fish: amphibians, reptiles, birds, mammals) [REDACTED]  (14 species: bichirs , reedfish ) [REDACTED]  (27 species: sturgeons , paddlefish ) [REDACTED] Ginglymodi (7 species: gars , alligator gars ) [REDACTED] Halecomorphi (2 species: bowfin , eyetail bowfin ) [REDACTED]  (>32,000 species) [REDACTED] Fishes (without tetrapods) are

7410-503: The inter-war years , financial stringency caused nearly all navies to skimp on testing their torpedoes. Only the British and Japanese had fully tested new technologies for torpedoes (in particular the Type 93 , nicknamed Long Lance postwar by the US official historian Samuel E. Morison ) at the start of World War II. Unreliable torpedoes caused many problems for the American submarine force in

7540-400: The intertidal zone , are facultative air breathers, able to breathe air when out of water, as may occur daily at low tide , and to use their gills when in water. Some coastal fish like rockskippers and mudskippers choose to leave the water to feed in habitats temporarily exposed to the air. Some catfish absorb air through their digestive tracts. The digestive system consists of a tube,

7670-425: The stout infantfish . Swimming performance varies from fish such as tuna, salmon , and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. A typical fish is cold-blooded , has a streamlined body for rapid swimming, extracts oxygen from water using gills, has two sets of paired fins, one or two dorsal fins, an anal fin and

7800-544: The 20th century. During the war the Imperial Russian and Imperial Japanese navies launched nearly 300 torpedoes at each other, all of them of the "self-propelled automotive" type. The deployment of these new underwater weapons resulted in one battleship, two armored cruisers, and two destroyers being sunk in action, with the remainder of the roughly 80 warships being sunk by the more conventional methods of gunfire, mines, and scuttling . On 27 May 1905, during

7930-604: The American Lieutenant Commander John A. Howell , whose design , driven by a flywheel , was simpler and cheaper. It was produced from 1885 to 1895, and it ran straight, leaving no wake. A Torpedo Test Station was set up in Rhode Island in 1870. The Howell torpedo was the only United States Navy model until an American company, Bliss and Williams secured manufacturing rights to produce Whitehead torpedoes. These were put into service for

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8060-683: The Americans to cease using this "cruel and unheard-of warfare" or he would "order every house near the shore to be destroyed". The fact that Hardy had been previously so lenient and considerate to the Americans led them to abandon such attempts with immediate effect. Torpedoes were used by the Russian Empire during the Crimean War in 1855 against British warships in the Gulf of Finland . They used an early form of chemical detonator. During

8190-705: The Argentine cruiser ARA  General Belgrano with two Mark VIII** torpedoes during the Falklands War . This is the only sinking of a surface ship by a nuclear-powered submarine in wartime and the second (of three) sinkings of a surface ship by any submarine since the end of World War II). The other two sinkings were of the Indian frigate INS  Khukri and the South Korean corvette ROKS Cheonan . Many classes of surface ships, submarines, and aircraft were armed with torpedoes. Naval strategy at

8320-622: The British government to employ his 'catamaran' against the French. An April 1804 torpedo attack on French ships anchored at Boulogne, and a follow-up attack in October, produced several explosions but no significant damage and the weapon was abandoned. Fulton carried out a demonstration for the US government on 20 July 1807, destroying a vessel in New York's harbor. Further development languished as Fulton focused on his "steam-boat matters". After

8450-601: The Southern Ocean, including under the Filchner–Ronne Ice Shelf at a latitude of 79°S, while desert pupfish live in desert springs, streams, and marshes, sometimes highly saline, with water temperatures as high as 36 C. A few fish live mostly on land or lay their eggs on land near water. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. A single undescribed species of Phreatobius has been called

8580-617: The U.S. Navy in 1892. Five varieties were produced, all 18-inch diameter. The Royal Navy introduced the Brotherhood wet heater engine in 1907 with the 18 in. Mk. VII & VII* which greatly increased the speed and/or range over compressed air engines and wet heater type engines became the standard in many major navies up to and during the Second World War. Ships of the line were superseded by ironclads , large steam-powered ships with heavy gun armament and heavy armor, in

8710-552: The USN twenty-one months into the Pacific War. British submarines used torpedoes to interdict the Axis supply shipping to North Africa , while Fleet Air Arm Swordfish sank three Italian battleships at Taranto by a torpedo and (after a mistaken, but abortive, attack on Sheffield ) scored one crucial hit in the hunt for the German battleship Bismarck . Large tonnages of merchant shipping were sunk by submarines with torpedoes in both

8840-478: The University of Chicago found bottom-walking lungfishes had already evolved characteristics of the walking gaits of terrestrial tetrapods. In a classic example of convergent evolution , the pectoral limbs of pterosaurs , birds and bats further evolved along independent paths into flying wings. Even with flying wings there are many similarities with walking legs, and core aspects of the genetic blueprint of

8970-556: The attention of the cleaners. Cleaning behaviors have been observed in a number of fish groups, including an interesting case between two cichlids of the same genus, Etroplus maculatus , the cleaner, and the much larger E. suratensis . Fish occupy many trophic levels in freshwater and marine food webs . Fish at the higher levels are predatory , and a substantial part of their prey consists of other fish. In addition, mammals such as dolphins and seals feed on fish, alongside birds such as gannets and cormorants . The body of

9100-475: The blunt nose provided a speed advantage of approximately one knot compared to the traditional pointed nose design. This discovery allowed for larger explosive payloads and increased air storage for propulsion without compromising speed. In 1893, RN torpedo production was transferred to the Royal Gun Factory . The British later established a Torpedo Experimental Establishment at HMS  Vernon and

9230-550: The body, and produce a concentrated urine. The reverse happens in freshwater fish : they tend to gain water osmotically, and produce a dilute urine. Some fish have kidneys able to operate in both freshwater and saltwater. Fish have small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, notably mormyrids and sharks , which have brains about as large for their body weight as birds and marsupials . At

9360-733: The bony Osteichthyes . During the Devonian , fish diversity greatly increased, including among the placoderms, lobe-finned fishes, and early sharks, earning the Devonian the epithet "the age of fishes". Fishes are a paraphyletic group, since any clade containing all fish, such as the Gnathostomata or (for bony fish) Osteichthyes , also contains the clade of tetrapods (four-limbed vertebrates, mostly terrestrial), which are usually not considered fish. Some tetrapods, such as cetaceans and ichthyosaurs , have secondarily acquired

9490-482: The case of humans) and the pelvic fins developed into hind legs. Much of the genetic machinery that builds a walking limb in a tetrapod is already present in the swimming fin of a fish. In 2011, researchers at Monash University in Australia used primitive but still living lungfish "to trace the evolution of pelvic fin muscles to find out how the load-bearing hind limbs of the tetrapods evolved." Further research at

9620-727: The construction of the first purpose-built operational torpedo aircraft, the Short Type 184 , built-in 1915. An order for ten aircraft was placed, and 936 aircraft were built by ten different British aircraft companies during the First World War . The two prototype aircraft were embarked upon HMS  Ben-my-Chree , which sailed for the Aegean on 21 March 1915 to take part in the Gallipoli campaign . On 12 August 1915 one of these, piloted by Flight Commander Charles Edmonds ,

9750-581: The early years of the war, primarily in the Pacific Theater . One possible exception to the pre-war neglect of torpedo development was the 45-cm caliber, 1931-premiered Japanese Type 91 torpedo , the sole aerial torpedo ( Koku Gyorai ) developed and brought into service by the Japanese Empire before the war. The Type 91 had an advanced PID controller and jettisonable, wooden Kyoban aerial stabilizing surfaces which released upon entering

9880-403: The factory was exporting torpedoes to ten other countries. The torpedo was powered by compressed air and had an explosive charge of gun-cotton . Whitehead went on to develop more efficient devices, demonstrating torpedoes capable of 18 knots (33 km/h) in 1876, 24 knots (44 km/h) in 1886, and, finally, 30 knots (56 km/h) in 1890. Royal Navy (RN) representatives visited Rijeka for

10010-406: The female cichlid , Pelvicachromis taeniatus , displays a large and visually arresting purple pelvic fin . "The researchers found that males clearly preferred females with a larger pelvic fin and that pelvic fins grew in a more disproportionate way than other fins on female fish." Reshaping human feet with swim fins , rather like the tail fin of a fish, add thrust and efficiency to the kicks of

10140-477: The first practical guided missile was patented by Louis Brennan , an emigre to Australia, in 1877. It was designed to run at a consistent depth of 12 feet (3.7 m), and was fitted with an indicator mast that just broke the surface of the water. At night the mast had a small light, only visible from the rear. Two steel drums were mounted one behind the other inside the torpedo, each carrying several thousand yards of high-tensile steel wire. The drums connected via

10270-491: The first self-propelled torpedo. Torpedo-like weapons were first proposed many centuries before they were successfully developed. For example, in 1275, Arab engineer Hasan al-Rammah – who worked as a military scientist for the Mamluk Sultanate of Egypt – wrote that it might be possible to create a projectile resembling "an egg", which propelled itself through water, whilst carrying "fire". In modern language,

10400-525: The following prophetic comparison: "Birds in a way resemble fishes. For birds have their wings in the upper part of their bodies and fishes have two fins in the front part of their bodies. Birds have feet on their underpart and most fishes have a second pair of fins in their under-part and near their front fins." – Aristotle, De incessu animalium There is an old theory, proposed by anatomist Carl Gegenbaur , which has been often disregarded in science textbooks, "that fins and (later) limbs evolved from

10530-464: The formation of a linked chain of vortex rings" and that "the dorsal and anal fin wakes are rapidly entrained by the caudal fin wake, approximately within the timeframe of a subsequent tail beat". Once motion has been established, the motion itself can be controlled with the use of other fins. Boats control direction (yaw) with fin-like rudders, and roll with stabilizer and keel fins. Airplanes achieve similar results with small specialised fins that change

10660-405: The front of the brain are the olfactory lobes , a pair of structures that receive and process signals from the nostrils via the two olfactory nerves . Fish that hunt primarily by smell, such as hagfish and sharks, have very large olfactory lobes. Behind these is the telencephalon , which in fish deals mostly with olfaction. Together these structures form the forebrain. Connecting the forebrain to

10790-412: The gills flows in the opposite direction to the water, resulting in efficient countercurrent exchange . The gills push the oxygen-poor water out through openings in the sides of the pharynx. Cartilaginous fish have multiple gill openings: sharks usually have five, sometimes six or seven pairs; they often have to swim to oxygenate their gills. Bony fish have a single gill opening on each side, hidden beneath

10920-447: The gills of an extinct vertebrate". Gaps in the fossil record had not allowed a definitive conclusion. In 2009, researchers from the University of Chicago found evidence that the "genetic architecture of gills, fins and limbs is the same", and that "the skeleton of any appendage off the body of an animal is probably patterned by the developmental genetic program that we have traced back to formation of gills in sharks". Recent studies support

11050-532: The gills. Oxygen-rich blood then flows without further pumping, unlike in mammals, to the body tissues. Finally, oxygen-depleted blood returns to the heart. Fish exchange gases using gills on either side of the pharynx . Gills consist of comblike structures called filaments. Each filament contains a capillary network that provides a large surface area for exchanging oxygen and carbon dioxide . Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gills. Capillary blood in

11180-447: The gut, leading from the mouth to the anus. The mouth of most fishes contains teeth to grip prey, bite off or scrape plant material, or crush the food. An esophagus carries food to the stomach where it may be stored and partially digested. A sphincter, the pylorus, releases food to the intestine at intervals. Many fish have finger-shaped pouches, pyloric caeca , around the pylorus, of doubtful function. The pancreas secretes enzymes into

11310-457: The heart from the rest of the body. Lungfish, bichirs, ropefish, bowfins, snakefish, and the African knifefish have evolved to reduce such mixing, and to reduce oxygen loss from the gills to oxygen-poor water. Bichirs and lungfish have tetrapod-like paired lungs, requiring them to surface to gulp air, and making them obligate air breathers. Many other fish, including inhabitants of rock pools and

11440-485: The idea of destroying ships by introducing floating mines under their bottoms in submarine boats". He employed the term "torpedo" for the explosive charges with which he outfitted his submarine Nautilus . However, both the French and the Dutch governments were uninterested in the submarine. Fulton then concentrated on developing the torpedo-like weapon independent of a submarine deployment, and in 1804 succeeded in convincing

11570-442: The idea that gill arches and paired fins are serially homologous and thus that fins may have evolved from gill tissues. Fish are the ancestors of all mammals, reptiles, birds and amphibians. In particular, terrestrial tetrapods (four-legged animals) evolved from fish and made their first forays onto land 400 million years ago. They used paired pectoral and pelvic fins for locomotion. The pectoral fins developed into forelegs (arms in

11700-455: The intestine to digest the food; other enzymes are secreted directly by the intestine itself. The liver produces bile which helps to break up fat into an emulsion which can be absorbed in the intestine. Most fish release their nitrogenous wastes as ammonia . This may be excreted through the gills or filtered by the kidneys . Salt is excreted by the rectal gland. Saltwater fish tend to lose water by osmosis ; their kidneys return water to

11830-401: The lift of the fin sets water or air in motion and pushes the fin in the opposite direction. Aquatic animals get significant thrust by moving fins back and forth in water. Often the tail fin is used, but some aquatic animals generate thrust from pectoral fins . Fins can also generate thrust if they are rotated in air or water. Turbines and propellers (and sometimes fans and pumps ) use

11960-426: The location of the locomotor surface can be known accurately. And, individual components of a natural motion (such as outstroke vs. instroke of a flapping appendage) can be programmed separately, which is certainly difficult to achieve when working with a live animal." Fish A fish ( pl. : fish or fishes ) is an aquatic , anamniotic , gill -bearing vertebrate animal with swimming fins and

12090-458: The margin at the rear of their bodies is a line of small rayless, non-retractable fins, known as finlets . There has been much speculation about the function of these finlets. Research done in 2000 and 2001 by Nauen and Lauder indicated that "the finlets have a hydrodynamic effect on local flow during steady swimming" and that "the most posterior finlet is oriented to redirect flow into the developing tail vortex, which may increase thrust produced by

12220-451: The mid 19th century. Ultimately this line of development led to the dreadnought category of all-big-gun battleships, starting with HMS  Dreadnought . Although these ships were incredibly powerful, the new weight of armor slowed them down, and the huge guns needed to penetrate that armor fired at very slow rates. The development of torpedoes allowed for the possibility that small and fast vessels could credibly threaten if not sink even

12350-419: The midbrain is the diencephalon ; it works with hormones and homeostasis . The pineal body is just above the diencephalon; it detects light, maintains circadian rhythms, and controls color changes. The midbrain contains the two optic lobes . These are very large in species that hunt by sight, such as rainbow trout and cichlids . The hindbrain controls swimming and balance.The single-lobed cerebellum

12480-405: The more remarkable because they evolved from nothing — the ancestral terrestrial reptile had no hump on its back or blade on its tail to serve as a precursor." The biologist Stephen Jay Gould said the ichthyosaur was his favorite example of convergent evolution . The use of fins for the propulsion of aquatic animals can be remarkably effective. It has been calculated that some fish can achieve

12610-422: The most powerful battleships. While such attacks would carry enormous risks to the attacking boats and their crews (which would likely need to expose themselves to artillery fire which their small vessels were not designed to withstand) this was offset by the ability to construct large numbers of small vessels far more quickly and for a much lower unit cost compared to a capital ship. The first boat designed to fire

12740-414: The muscle fibers with electricity. Robotic fish offer some research advantages, such as the ability to examine part of a fish design in isolation from the rest, and variance of a single parameter, such as flexibility or direction. Researchers can directly measure forces more easily than in live fish. "Robotic devices also facilitate three-dimensional kinematic studies and correlated hydrodynamic analyses, as

12870-449: The need of large guns, though sometimes at the risk of being hit by longer-range artillery fire. Modern torpedoes are classified variously as lightweight or heavyweight; straight-running, autonomous homers, and wire-guided types. They can be launched from a variety of platforms. In modern warfare, a submarine-launched torpedo is almost certain to hit its target; the best defense is a counterattack using another torpedo. The word torpedo

13000-423: The night of 12/13 August 1942 during Operation Pedestal . Destroyers of all navies were also armed with torpedoes to attack larger ships. In the Battle off Samar , destroyer torpedoes from the escorts of the American task force "Taffy 3" showed effectiveness at defeating armor. Damage and confusion caused by torpedo attacks were instrumental in beating back a superior Japanese force of battleships and cruisers. In

13130-443: The nine largest families; from largest to smallest, these are Cyprinidae , Gobiidae , Cichlidae , Characidae , Loricariidae , Balitoridae , Serranidae , Labridae , and Scorpaenidae . About 64 families are monotypic , containing only one species. Fish range in size from the huge 16-metre (52 ft) whale shark to some tiny teleosts only 8-millimetre (0.3 in) long, such as the cyprinid Paedocypris progenetica and

13260-402: The open ocean. Because the water all around is the same colour, reflecting an image of the water offers near-invisibility. Fish have a closed-loop circulatory system . The heart pumps the blood in a single loop throughout the body; for comparison, the mammal heart has two loops, one for the lungs to pick up oxygen, one for the body to deliver the oxygen. In fish, the heart pumps blood through

13390-504: The pectoral fin have been retained. About 200 million years ago the first mammals appeared. A group of these mammals started returning to the sea about 52 million years ago, thus completing a circle. These are the cetaceans (whales, dolphins and porpoises). Recent DNA analysis suggests that cetaceans evolved from within the even-toed ungulates , and that they share a common ancestor with the hippopotamus . About 23 million years ago another group of bearlike land mammals started returning to

13520-454: The placoderms, appeared in the Silurian and greatly diversified during the Devonian , the "Age of Fishes". Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons , emerged as the dominant group of fish after the end-Devonian extinction wiped out the apex placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish . About 96% of all living fish species today are teleosts ,

13650-551: The plans of the Salvacoste ("Coastsaver"), a floating weapon driven by ropes from the land that had been dismissed by the naval authorities due to the impractical steering and propulsion mechanisms. In 1866, Whitehead invented the first effective self-propelled torpedo, the eponymous Whitehead torpedo , the first modern torpedo. French and German inventions followed closely, and the term torpedo came to describe self-propelled projectiles that traveled under or on water. By 1900,

13780-653: The problems. In the United States Navy (USN), there was an extended wrangle over the problems plaguing the Mark 14 torpedo (and its Mark 6 exploder ). Cursory trials had allowed bad designs to enter service. Both the Navy Bureau of Ordnance and the United States Congress were too busy protecting their interests to correct the errors, and fully functioning torpedoes only became available to

13910-664: The relatively confined spaces and complex underwater landscapes of coral reefs . For this manoeuvrability is more important than straight line speed, so coral reef fish have developed bodies which optimize their ability to dart and change direction. They outwit predators by dodging into fissures in the reef or playing hide and seek around coral heads. The pectoral and pelvic fins of many reef fish, such as butterflyfish , damselfish and angelfish , have evolved so they can act as brakes and allow complex maneuvers. Many reef fish, such as butterflyfish , damselfish and angelfish , have evolved bodies which are deep and laterally compressed like

14040-417: The same way as in other mammals. Ichthyosaurs are ancient reptiles that resembled dolphins. They first appeared about 245 million years ago and disappeared about 90 million years ago. "This sea-going reptile with terrestrial ancestors converged so strongly on fishes that it actually evolved a dorsal fin and tail in just the right place and with just the right hydrological design. These structures are all

14170-516: The sea. These were the pinnipeds (seals). What had become walking limbs in cetaceans and seals evolved further, independently in a reverse form of convergent evolution, back to new forms of swimming fins. The forelimbs became flippers and, in pinnipeds, the hind limbs became a tail terminating in two fins (the cetacean fluke , conversely, is an entirely new organ). Fish tails are usually vertical and move from side to side. Cetacean flukes are horizontal and move up and down, because cetacean spines bend

14300-493: The self-propelled Whitehead torpedo was HMS  Lightning , completed in 1877. The French Navy followed suit in 1878 with Torpilleur No 1 , launched in 1878 though she had been ordered in 1875. The first torpedo boats were built at the shipyards of Sir John Thornycroft and gained recognition for their effectiveness. At the same time, inventors were working on building a guided torpedo. Prototypes were built by John Ericsson , John Louis Lay , and Victor von Scheliha, but

14430-567: The shape of their wings and tail fins. Stabilising fins are used as fletching on arrows and some darts , and at the rear of some bombs , missiles , rockets and self-propelled torpedoes . These are typically planar and shaped like small wings, although grid fins are sometimes used. Static fins have also been used for one satellite, GOCE . Engineering fins are also used as heat transfer fins to regulate temperature in heat sinks or fin radiators . In biology, fins can have an adaptive significance as sexual ornaments. During courtship,

14560-428: The start of World War II. Around this time too the Royal Navy were perfecting the Brotherhood burner cycle engine which offered a performance as good as the oxygen-enriched air engine but without the issues arising from the oxygen equipment and which was first used in the extremely successful and long-lived 21 in. Mk. VIII torpedo of 1925. This torpedo served throughout WW II (with 3,732 being fired by September 1944) and

14690-411: The streamlined shape and propulsion by front flippers of penguins . Festo also developed AquaRay , AquaJelly and AiraCuda , respectively emulating the locomotion of manta rays, jellyfish and barracuda. In 2004, Hugh Herr at MIT prototyped a biomechatronic robotic fish with a living actuator by surgically transplanting muscles from frog legs to the robot and then making the robot swim by pulsing

14820-549: The tail of swimming mackerel". Fish use multiple fins, so it is possible that a given fin can have a hydrodynamic interaction with another fin. In particular, the fins immediately upstream of the caudal (tail) fin may be proximate fins that can directly affect the flow dynamics at the caudal fin. In 2011, researchers using volumetric imaging techniques were able to generate "the first instantaneous three-dimensional views of wake structures as they are produced by freely swimming fishes". They found that "continuous tail beats resulted in

14950-512: The target. Fiske reported in 1915 that, using this method, enemy fleets could be attacked within their harbors if there was enough room for the torpedo track. Meanwhile, the Royal Naval Air Service began actively experimenting with this possibility. The first successful aerial torpedo drop was performed by Gordon Bell in 1914 – dropping a Whitehead torpedo from a Short S.64 seaplane . The success of these experiments led to

15080-722: The tender Velikiy Knyaz Konstantin under the command of Stepan Osipovich Makarov during the Russo-Turkish War of 1877–78 . In another early use of the torpedo, during the War of the Pacific , the Peruvian ironclad Huáscar commanded by captain Miguel Grau attacked the Chilean corvette Abtao on 28 August 1879 at Antofagasta with a self-propelled Lay torpedo only to have it reverse course. The ship Huascar

15210-435: The term no longer included mines and booby-traps as the navies of the world added submarines, torpedo boats and torpedo boat destroyers to their fleets. Whitehead was unable to improve the machine substantially, since the clockwork motor, attached ropes, and surface attack mode all contributed to a slow and cumbersome weapon. However, he kept considering the problem after the contract had finished, and eventually developed

15340-542: The time was to use torpedoes, launched from submarines or warships, against enemy warships in a fleet action on the high seas. There were concerns torpedoes would be ineffective against warships' heavy armor; an answer to this was to detonate torpedoes underneath a ship, badly damaging its keel and the other structural members in the hull, commonly called "breaking its back". This was demonstrated by magnetic influence mines in World War I. The torpedo would be set to run at

15470-561: The tug. Without the weight of the torpedo Dacre was able to take off and return to Ben-My-Chree . Torpedoes were widely used in World War I , both against shipping and against submarines. Germany disrupted the supply lines to Britain largely by use of submarine torpedoes, though submarines also extensively used guns. Britain and its allies also used torpedoes throughout the war. U-boats themselves were often targeted, twenty being sunk by torpedo. Two Royal Italian Navy torpedo boats scored

15600-593: The value of the torpedo and in spring 1883 an experimental station was established at Garrison Point Fort , Sheerness , on the River Medway , and a workshop for Brennan was set up at the Chatham Barracks , the home of the Royal Engineers. Between 1883 and 1885 the Royal Engineers held trials and in 1886 the torpedo was recommended for adoption as a harbor defense torpedo. It was used throughout

15730-483: The water, making it a formidable anti-ship weapon; Nazi Germany considered manufacturing it as the Luftorpedo LT 850 after August 1942 . The Royal Navy's 24.5-inch oxygen-enriched air torpedo saw service in the two Nelson class battleships although by World War II the use of enriched oxygen had been discontinued due to safety concerns. In the final phase of the action against Bismarck , Rodney fired

15860-547: The weapon was apt to cause as much harm to its user as to its target. Rear Admiral David Farragut 's famous/apocryphal command during the Battle of Mobile Bay in 1864, " Damn the torpedoes, full speed ahead! " refers to a minefield laid at Mobile, Alabama . On 26 May 1877, during the Romanian War of Independence , the Romanian spar torpedo boat Rândunica attacked and sank the Ottoman river monitor Seyfi . This

15990-493: Was 18 in (46 cm) in diameter and 19 ft (5.8 m) long, made of polished steel or phosphor bronze , with a 200-pound (91 kg) gun-cotton warhead. It was propelled by a three-cylinder Brotherhood radial engine, using compressed air at around 1,300  psi (9.0  MPa ) and driving two contra-rotating propellers, and was designed to self-regulate its course and depth as far as possible. By 1881, nearly 1,500 torpedoes had been produced. Whitehead also opened

16120-523: Was a mechanism consisting of a hydrostatic valve and pendulum that caused the torpedo's hydroplanes to be adjusted to maintain a preset depth. After the Austrian government decided to invest in the invention, Whitehead started the first torpedo factory in Rijeka. In 1870, he improved the devices to travel up to approximately 1,000 yards (910 m) at a speed of up to 6 knots (11 km/h), and by 1881

16250-531: Was first used as a name for electric rays (in the order Torpediniformes ), which in turn comes from the Latin word torpēdō ("lethargy" or "sluggishness"). In naval usage, the American inventor David Bushnell was reported to have first used the term as the name of a submarine of his own design, the "American Turtle or Torpedo." This usage likely inspired Robert Fulton 's use of the term to describe his stationary mines , and later Robert Whitehead 's naming of

16380-556: Was saved when an officer jumped overboard to divert it. The Chilean ironclad Blanco Encalada was sunk on 23 April 1891 by a self-propelled torpedo from the Almirante Lynch , during the Chilean Civil War of 1891 , becoming the first ironclad warship sunk by this weapon. The Chinese turret ship Dingyuan was purportedly hit and disabled by a torpedo after numerous attacks by Japanese torpedo boats during

16510-488: Was set upon by 17 torpedo-firing warships, ten of which were destroyers and four torpedo boats. Twenty-one torpedoes were launched at the pre-dreadnought , and three struck home, one fired from the destroyer Murasame and two from torpedo boats No. 72 and No. 75 . The flagship slipped under the waves shortly thereafter, taking over 900 men with her to the bottom. On December 9, 1912, the Greek submarine "Dolphin" launched

16640-477: Was the first aircraft in the world to attack an enemy ship with an air-launched torpedo. On 17 August 1915 Flight Commander Edmonds torpedoed and sank an Ottoman transport ship a few miles north of the Dardanelles. His formation colleague, Flight Lieutenant G B Dacre, was forced to land on the water owing to engine trouble but, seeing an enemy tug close by, taxied up to it and released his torpedo, sinking

16770-529: Was the first instance in history when a torpedo boat sank its targets without also sinking. A prototype of the self-propelled torpedo was created on a commission placed by Giovanni Luppis , an Austro-Hungarian naval officer from Rijeka (modern-day Croatia ), at the time a port city of the Austro-Hungarian Monarchy and Robert Whitehead , an English engineer who was the manager of a town factory. In 1864, Luppis presented Whitehead with

16900-478: Was the first warship to be sunk in 1862 by an electrically-detonated mine. Spar torpedoes were also used; an explosive device was mounted at the end of a spar up to 30 feet (9.1 m) long projecting forward underwater from the bow of the attacking vessel, which would then ram the opponent with the explosives. These were used by the Confederate submarine H. L. Hunley to sink USS  Housatonic although

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