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162-760: Fishing is the activity of trying to catch fish . Fish are often caught as wildlife from the natural environment ( fresh water or marine ), but may also be caught from stocked bodies of water such as ponds , canals , park wetlands and reservoirs . Fishing techniques include trawling , longlining , jigging , hand-gathering , spearing , netting , angling , shooting and trapping , as well as more destructive and often illegal techniques such as electrocution , blasting and poisoning . The term fishing broadly includes catching aquatic animals other than fish, such as crustaceans ( shrimp / lobsters / crabs ), shellfish , cephalopods ( octopus / squid ) and echinoderms ( starfish / sea urchins ). The term

324-1054: A dagger (†); groups of uncertain placement are labelled with a question mark (?) and dashed lines (- - - - -). Jawless fishes (118 species: hagfish , lampreys ) [REDACTED] † Thelodonti , † Conodonta , † Anaspida [REDACTED] [REDACTED] [REDACTED] † Galeaspida [REDACTED] † Osteostraci [REDACTED] † Placodermi [REDACTED] † Acanthodii [REDACTED]  (>1,100 species: sharks , rays , chimaeras ) [REDACTED]  (2 species: coelacanths ) [REDACTED] Dipnoi (6 species: lungfish ) [REDACTED] Tetrapoda (>38,000 species, not considered fish: amphibians, reptiles, birds, mammals) [REDACTED]  (14 species: bichirs , reedfish ) [REDACTED]  (27 species: sturgeons , paddlefish ) [REDACTED] Ginglymodi (7 species: gars , alligator gars ) [REDACTED] Halecomorphi (2 species: bowfin , eyetail bowfin ) [REDACTED]  (>32,000 species) [REDACTED] Fishes (without tetrapods) are

486-579: A paraphyletic group and for this reason, the class Pisces seen in older reference works is no longer used in formal classifications. Traditional classification divides fish into three extant classes (Agnatha, Chondrichthyes, and Osteichthyes), and with extinct forms sometimes classified within those groups, sometimes as their own classes. Fish account for more than half of vertebrate species. As of 2016, there are over 32,000 described species of bony fish, over 1,100 species of cartilaginous fish, and over 100 hagfish and lampreys. A third of these fall within

648-405: A subspecies of modern human as Homo sapiens neanderthalensis . This would necessitate the classification of modern humans as H. sapiens sapiens . A large part of the controversy stems from the vagueness of the term "species", as it is generally used to distinguish two genetically isolated populations, but admixture between modern humans and Neanderthals is known to have occurred. However,

810-644: A Neanderthal in a modern business suit and hat to emphasise that they would be, more or less, indistinguishable from modern humans had they survived into the present. William Golding 's 1955 novel The Inheritors depicts Neanderthals as much more emotional and civilised. However, Boule's image continued to influence works until the 1960s. In modern-day, Neanderthal reconstructions are often very humanlike. Hybridisation between Neanderthals and early modern humans had been suggested early on, such as by English anthropologist Thomas Huxley in 1890, Danish ethnographer Hans Peder Steensby in 1907, and Coon in 1962. In

972-650: A Neanderthal/ H. heidelbergensis lineage. Eight hundred thousand years ago has H. antecessor as the LCA, but different variations of this model would push the date back to 1 million years ago. However, a 2020 analysis of H. antecessor enamel proteomes suggests that H. antecessor is related but not a direct ancestor. DNA studies have yielded various results for the Neanderthal/human divergence time, such as 538–315, 553–321, 565–503, 654–475, 690–550, 765–550, 741–317, and 800–520,000 years ago; and

1134-636: A cold climate, but they may also have been adaptations for sprinting in the warmer, forested landscape that Neanderthals often inhabited. They had cold-specific adaptations, such as specialised body-fat storage and an enlarged nose to warm air (although the nose could have been caused by genetic drift ). Average Neanderthal men stood around 165 cm (5 ft 5 in) and women 153 cm (5 ft 0 in) tall, similar to pre-industrial modern Europeans. The braincases of Neanderthal men and women averaged about 1,600 cm (98 cu in) and 1,300 cm (79 cu in), respectively, which

1296-489: A combination of these factors. Neanderthals lived in a high-stress environment with high trauma rates, and about 80% died before the age of 40. The total population of Neanderthals remained low, and interbreeding with humans tended toward a loss of Neanderthal genes over time. They lacked effective long-distance networks. Despite this, there is evidence of regional cultures and regular communication between communities, possibly moving between caves seasonally. For much of

1458-543: A dental analysis concluded before 800,000 years ago. Neanderthals and Denisovans are more closely related to each other than they are to modern humans, meaning the Neanderthal/Denisovan split occurred after their split with modern humans. Assuming a mutation rate of 1 × 10 or 0.5 × 10 per base pair (bp) per year, the Neanderthal/Denisovan split occurred around either 236–190,000 or 473–381,000 years ago, respectively. Using 1.1 × 10 per generation with

1620-562: A different group of archaic humans, in Siberia. Around 1–4% of genomes of Eurasians , Indigenous Australians , Melanesians , Native Americans and North Africans is of Neanderthal ancestry, while most inhabitants of sub-Saharan Africa have around 0.3% of Neanderthal genes, save possible traces from early sapiens-to-Neanderthal gene flow and/or more recent back-migration of Eurasians to Africa. In all, about 20% of distinctly Neanderthal gene variants survive in modern humans. Although many of

1782-621: A drifter to steam power. In 1877, he built the first screw propelled steam trawler in the world. Steam trawlers were introduced at Grimsby and Hull in the 1880s. In 1890 it was estimated that there were 20,000 men on the North Sea. The steam drifter was not used in the herring fishery until 1897. The last sailing fishing trawler was built in 1925 in Grimsby. Trawler designs adapted as the way they were powered changed from sail to coal-fired steam by World War I to diesel and turbines by

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1944-581: A great leap forward after the English Civil War , where a newly found interest in the activity left its mark on the many books and treatises that were written on the subject at the time. Leonard Mascall in 1589 wrote A booke of Fishing with Hooke and Line along with many others he produced in his life on game and wildlife in England at the time. The Compleat Angler was written by Izaak Walton in 1653 (although Walton continued to add to it for

2106-442: A high-quality Neanderthal genome preserved in a toe bone from Denisova Cave, Siberia. Homo sapiens Denisovan from Denisova Cave Denisovan from Baishiya Karst Cave Neanderthal from Denisova Cave Neanderthal from Sidrón Cave Neanderthal from Vindija Cave Neanderthals are hominids in the genus Homo , humans, and generally classified as a distinct species , H. neanderthalensis , although sometimes as

2268-430: A laborious and time-consuming process – the new textile spinning machines allowed for a variety of tapered lines to be easily manufactured and marketed. British fly fishing continued to develop in the 19th century, with the emergence of fly fishing clubs, along with the appearance of several books on the subject of fly tying and fly fishing techniques. By the mid to late 19th century, expanding leisure opportunities for

2430-609: A lack of evidence of Southern Scandinavian occupation is (at least during the Eemian interglacial) due to the former explanation and a lack of research in the area. Middle Palaeolithic artefacts have been found up to 60°N on the Russian plains, but these are more likely attributed to modern humans. A 2017 study claimed the presence of Homo at the 130,000-year-old Californian Cerutti Mastodon site in North America, but this

2592-958: A larger cerebellum . Neanderthal brains also have larger occipital lobes (relating to the classic occurrence of an occipital bun in Neanderthal skull anatomy, as well as the greater width of their skulls), which implies internal differences in the proportionality of brain-internal regions, relative to Homo sapiens, consistent with external measurements obtained with fossil skulls. Their brains also have larger temporal lobe poles, wider orbitofrontal cortex, and larger olfactory bulbs, suggesting potential differences in language comprehension and associations with emotions ( temporal functions ), decision making (the orbitofrontal cortex ) and sense of smell ( olfactory bulbs ). Their brains also show different rates of brain growth and development. Such differences, while slight, would have been visible to natural selection and may underlie and explain differences in

2754-408: A larger diaphragm and possibly greater lung capacity . The lung capacity of Kebara 2 was estimated to have been 9.04 L (2.39 US gal), compared to the average human capacity of 6 L (1.6 US gal) for males and 4.7 L (1.2 US gal) for females. The Neanderthal chest was also more pronounced (expanded front-to-back, or antero-posteriorly). The sacrum (where

2916-548: A legitimate species. The most influential specimen was La Chapelle-aux-Saints 1 ("The Old Man") from La Chapelle-aux-Saints , France. French palaeontologist Marcellin Boule authored several publications, among the first to establish palaeontology as a science, detailing the specimen, but reconstructed him as slouching, ape-like, and only remotely related to modern humans. The 1912 'discovery' of Piltdown Man (a hoax), appearing much more similar to modern humans than Neanderthals,

3078-568: A lengthening of the head (phase 1), which then led to other changes in skull anatomy (phase 2). However, Neanderthal anatomy may not have been driven entirely by adapting to cold weather. Accretion holds that Neanderthals slowly evolved over time from the ancestral H. heidelbergensis , divided into four stages: early-pre-Neanderthals ( MIS 12 , Elster glaciation ), pre-Neanderthals (MIS 11 – 9 , Holstein interglacial ), early Neanderthals (MIS 7– 5 , Saale glaciation – Eemian ), and classic Neanderthals (MIS 4–3, Würm glaciation ). Numerous dates for

3240-772: A line pickup, a device which orbits around the stationary spool. Because the line did not have to pull against a rotating spool, much lighter lures could be cast than with conventional reels. The development of inexpensive fiberglass rods, synthetic fly lines, and monofilament leaders in the early 1950s revived the popularity of fly fishing. There are many fishing techniques and tactics for catching fish. The term can also be applied to methods for catching other aquatic animals such as molluscs ( shellfish , squid , octopus) and edible marine invertebrates . Fishing techniques include hand gathering , spearfishing , netting , angling , bowfishing and trapping , as well as less common techniques such as gaffing , snagging , clubbing and

3402-530: A major source of food. The British dogger was a very early type of sailing trawler from the 17th century, but the modern fishing trawler was developed in the 19th century, at the English fishing port of Brixham . By the early 19th century, the fishers at Brixham needed to expand their fishing area further than ever before due to the ongoing depletion of stocks that was occurring in the overfished waters of South Devon . The Brixham trawler that evolved there

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3564-436: A more spherical lens . Their retinas generally have both rods and cones (for scotopic and photopic vision ); many species have colour vision , often with three types of cone. Teleosts can see polarized light ; some such as cyprinids have a fourth type of cone that detects ultraviolet . Amongst jawless fish , the lamprey has well-developed eyes, while the hagfish has only primitive eyespots. Hearing too

3726-542: A more forward direction, by comparison. Neanderthal eyeballs are larger than those of modern humans. One study proposed that this was due to Neanderthals having enhanced visual abilities, at the expense of neocortical and social development. However, this study was rejected by other researchers who concluded that eyeball size does not offer any evidence for the cognitive abilities of Neanderthal or modern humans. The projected Neanderthal nose and paranasal sinuses have generally been explained as having warmed air as it entered

3888-400: A more recent mitochondrial LCA (observable by studying mtDNA) and Y chromosome LCA. This likely resulted from an interbreeding event subsequent to the Neanderthal/Denisovan split. This involved either introgression coming from an unknown archaic human into Denisovans, or introgression from an earlier unidentified modern human wave from Africa into Neanderthals. The fact that the mtDNA of

4050-544: A new generation every 29 years, the time is 744,000 years ago. Using 5 × 10 nucleotide sites per year, it is 616,000 years ago. Using the latter dates, the split had likely already occurred by the time hominins spread out across Europe, and unique Neanderthal features had begun evolving by 600–500,000 years ago. Before splitting, Neanderthal/Denisovans (or "Neandersovans") migrating out of Africa into Europe apparently interbred with an unidentified "superarchaic" human species who were already present there; these superarchaics were

4212-411: A novel reel and fly design in 1874, described by reel historian Jim Brown as the "benchmark of American reel design," and the first fully modern fly reel. Albert Illingworth, 1st Baron Illingworth a textiles magnate, patented the modern form of fixed-spool spinning reel in 1905. When casting Illingworth's reel design, the line was drawn off the leading edge of the spool but was restrained and rewound by

4374-419: A protective bony cover or operculum . They are able to oxygenate their gills using muscles in the head. Some 400 species of fish in 50 families can breathe air, enabling them to live in oxygen-poor water or to emerge on to land. The ability of fish to do this is potentially limited by their single-loop circulation, as oxygenated blood from their air-breathing organ will mix with deoxygenated blood returning to

4536-504: A protrusion on the back of the skull, although it is within the range of variation for modern humans who have it. It is caused by the cranial base and temporal bones being placed higher and more towards the front of the skull, and a flatter skullcap . The Neanderthal face is characterised by subnasal as well as mid-facial prognathism , where the zygomatic arches are positioned in a rearward location relative to modern humans, while their maxillary bones and nasal bones are positioned in

4698-559: A quantum radical pair mechanism . Neanderthal Neanderthals ( / n i ˈ æ n d ər ˌ t ɑː l , n eɪ -, - ˌ θ ɑː l / nee- AN -də(r)- TAHL , nay-, -⁠ THAHL ; Homo neanderthalensis or H. sapiens neanderthalensis ) are an extinct group of archaic humans (generally regarded as a distinct species, though some regard it as a subspecies of Homo sapiens ) who lived in Eurasia until about 40,000 years ago. The type specimen , Neanderthal 1 ,

4860-567: A quarter of a century) and described the fishing in the Derbyshire Wye . It was a celebration of the art and spirit of fishing in prose and verse. A second part to the book was added by Walton's friend Charles Cotton . Charles Kirby designed an improved fishing hook in 1655 that remains relatively unchanged to this day. He went on to invent the Kirby bend, a distinctive hook with an offset point, still commonly used today. The 18th century

5022-603: A rather large jaw which was once cited as a response to a large bite force evidenced by heavy wearing of Neanderthal front teeth (the "anterior dental loading" hypothesis), but similar wearing trends are seen in contemporary humans. It could also have evolved to fit larger teeth in the jaw, which would better resist wear and abrasion, and the increased wear on the front teeth compared to the back teeth probably stems from repetitive use. Neanderthal dental wear patterns are most similar to those of modern Inuit. The incisors are large and shovel-shaped, and, compared to modern humans, there

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5184-495: A recreational fishing boat, so long as a fisher periodically climbs aboard with the intent to catch a fish. Fish are caught for recreational purposes from boats which range from dugout canoes , float tubes , kayaks , rafts , stand up paddleboards, pontoon boats and small dinghies to runabouts , cabin cruisers and cruising yachts to large, hi-tech and luxurious big game rigs. Larger boats, purpose-built with recreational fishing in mind, usually have large, open cockpits at

5346-464: A second variation of BNC2 was also present, which in modern populations is associated with darker skin colour in the UK Biobank . DNA analysis of three Neanderthal females from southeastern Europe indicates that they had brown eyes, dark skin colour and brown hair, with one having red hair. In modern humans, skin and hair colour is regulated by the melanocyte-stimulating hormone —which increases

5508-441: A sense of touch and of hearing . Blind cave fish navigate almost entirely through the sensations from their lateral line system. Some fish, such as catfish and sharks, have the ampullae of Lorenzini , electroreceptors that detect weak electric currents on the order of millivolt. Vision is an important sensory system in fish. Fish eyes are similar to those of terrestrial vertebrates like birds and mammals, but have

5670-524: A skull, shares several physical attributes with Neanderthals, although these may be the result of convergent evolution rather than Neanderthals extending their range to the Pacific Ocean. The northernmost bound is generally accepted to have been 55°N , with unambiguous sites known between 50 – 53°N , although this is difficult to assess because glacial advances destroy most human remains, and palaeoanthropologist Trine Kellberg Nielsen has argued that

5832-485: A tail fin, jaws, skin covered with scales , and lays eggs. Each criterion has exceptions, creating a wide diversity in body shape and way of life. For example, some fast-swimming fish are warm-blooded, while some slow-swimming fish have abandoned streamlining in favour of other body shapes. Fish species are roughly divided equally between freshwater and marine (oceanic) ecosystems; there are some 15,200 freshwater species and around 14,800 marine species. Coral reefs in

5994-589: A true "land fish" as this worm-like catfish strictly lives among waterlogged leaf litter . Cavefish of multiple families live in underground lakes , underground rivers or aquifers . Like other animals, fish suffer from parasitism . Some species use cleaner fish to remove external parasites. The best known of these are the bluestreak cleaner wrasses of coral reefs in the Indian and Pacific oceans. These small fish maintain cleaning stations where other fish congregate and perform specific movements to attract

6156-422: A typical fish is adapted for efficient swimming by alternately contracting paired sets of muscles on either side of the backbone. These contractions form S-shaped curves that move down the body. As each curve reaches the tail fin, force is applied to the water, moving the fish forward. The other fins act as control surfaces like an aircraft's flaps, enabling the fish to steer in any direction. Since body tissue

6318-573: A wide array of food, mainly hoofed mammals , but also megafauna , plants, small mammals, birds, and aquatic and marine resources. Although they were probably apex predators , they still competed with cave lions , cave hyenas and other large predators. A number of examples of symbolic thought and Palaeolithic art have been inconclusively attributed to Neanderthals, namely possible ornaments made from bird claws and feathers, shells, collections of unusual objects including crystals and fossils, engravings, music production (possibly indicated by

6480-446: A worldwide catch of well over a million tonnes in 1999, with herring and sardines together providing a catch of over 22 million metric tons in 1999. Many other species as well are fished in smaller numbers. Fish farming is the principal form of aquaculture , while other methods may fall under mariculture . It involves raising fish commercially in tanks or enclosures, usually for food. A facility that releases juvenile fish into

6642-631: A ~430,000 years old early Neanderthal-line archaic human from Sima de los Huesos in Spain is more closely related to those of Denisovans than to other Neanderthals or modern humans has been cited as evidence in favour of the latter hypothesis. It is largely thought that H. heidelbergensis was the last common ancestor of Neanderthals, Denisovans and modern humans before populations became isolated in Europe, Asia and Africa, respectively. The taxonomic distinction between H. heidelbergensis and Neanderthals

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6804-463: Is a boat or ship used to catch fish in the sea, or on a lake or river. Many different kinds of vessels are used in commercial , artisanal , and recreational fishing . According to the FAO , in 2004 there were four million commercial fishing vessels. About 1.3 million of these are decked vessels with enclosed areas. Nearly all of these decked vessels are mechanised, and 40,000 of them are over 100 tons. At

6966-567: Is always Neandertaler ("inhabitant of the Neander Valley"), whereas Neandertal always refers to the valley. The valley itself was named after the late 17th century German theologian and hymn writer Joachim Neander , who often visited the area. His name in turn means 'new man', being a learned Graecisation of the German surname Neumann . Neanderthal can be pronounced using the /t/ (as in / n i ˈ æ n d ər t ɑː l / ) or

7128-423: Is an aquatic , anamniotic , gill -bearing vertebrate animal with swimming fins and a hard skull , but lacking limbs with digits . Fish can be grouped into the more basal jawless fish and the more common jawed fish , the latter including all living cartilaginous and bony fish , as well as the extinct placoderms and acanthodians . Most fish are cold-blooded , their body temperature varying with

7290-597: Is an ancient practice that dates back to at least the beginning of the Upper Paleolithic period about 40,000 years ago. Isotopic analysis of the remains of Tianyuan man , a 40,000-year-old modern human from eastern Asia, has shown that he regularly consumed freshwater fish. Archaeology features such as shell middens , discarded fish bones, and cave paintings show that seafood was important for survival and consumed in significant quantities. Fishing in Africa

7452-465: Is an important sensory system in fish. Fish sense sound using their lateral lines and otoliths in their ears, inside their heads. Some can detect sound through the swim bladder. Some fish, including salmon, are capable of magnetoreception ; when the axis of a magnetic field is changed around a circular tank of young fish, they reorient themselves in line with the field. The mechanism of fish magnetoreception remains unknown; experiments in birds imply

7614-623: Is considerably larger than the modern human average (1,260 cm (77 cu in) and 1,130 cm (69 cu in), respectively). The Neanderthal skull was more elongated and the brain had smaller parietal lobes and cerebellum, but larger temporal, occipital and orbitofrontal regions. The 2010 Neanderthal genome project 's draft report presented evidence for interbreeding between Neanderthals and modern humans . It possibly occurred 316,000 to 219,000 years ago, but more likely 100,000 years ago and again 65,000 years ago. Neanderthals also appear to have interbred with Denisovans ,

7776-476: Is defined by the FAO as including recreational , subsistence and commercial fishing , and the harvesting, processing , and marketing sectors. The commercial activity is aimed at the delivery of fish and other seafood products for human consumption or use as raw material in other industrial processes. In 2022 24% of fishers and fish farmers and 62% of workers in post-harvest sector were women. There are three principal industry sectors: Commercial fishing

7938-431: Is denser than water, fish must compensate for the difference or they will sink. Many bony fish have an internal organ called a swim bladder that allows them to adjust their buoyancy by increasing or decreasing the amount of gas it contains. The scales of fish provide protection from predators at the cost of adding stiffness and weight. Fish scales are often highly reflective; this silvering provides camouflage in

8100-547: Is evident very early on in human history. Neanderthals were fishing by about 200,000 BC. People could have developed basketry for fish traps, using spinning and early forms of knitting to make fishing nets able to catch more fish. During this period, most people lived a hunter-gatherer lifestyle and were, of necessity, constantly on the move. However, where there are early examples of permanent settlements (though not necessarily permanently occupied) such as those at Lepenski Vir , they are almost always associated with fishing as

8262-584: Is fishing from boats to catch large open-water species such as swordfish , tuna , sharks , and marlin . Sportfishing (sometimes game fishing) is recreational fishing where the primary reward is the challenge of finding and catching the fish rather than the culinary or financial value of the fish's flesh. Fish sought after include tarpon , sailfish , mackerel , grouper and many others. The fishing industry includes any industry or activity concerned with taking, culturing, processing, preserving, storing, transporting, marketing or selling fish or fish products. It

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8424-467: Is largely considered implausible. It is unknown how the rapidly fluctuating climate of the last glacial period ( Dansgaard–Oeschger events ) impacted Neanderthals, as warming periods would produce more favourable temperatures but encourage forest growth and deter megafauna, whereas frigid periods would produce the opposite. However, Neanderthals may have preferred a forested landscape. Stable environments with mild mean annual temperatures may have been

8586-484: Is limited by the amount of food it can obtain, which in turn is limited by its technology. Innovation increases with population, but if the population is too low, innovation will not occur very rapidly and the population will remain low. This is consistent with the apparent 150,000 year stagnation in Neanderthal lithic technology. In a sample of 206 Neanderthals, based on the abundance of young and mature adults in comparison to other age demographics, about 80% of them above

8748-442: Is most similar to that of Inuit and Siberian Yupiks among modern humans —and shorter limbs result in higher retention of body heat. Nonetheless, Neanderthals from more temperate climates—such as Iberia—still retain the "hyperarctic" physique. In 2019, English anthropologist John Stewart and colleagues suggested Neanderthals instead were adapted for sprinting, because of evidence of Neanderthals preferring warmer wooded areas over

8910-513: Is mostly based on a fossil gap in Europe between 300 and 243,000 years ago during marine isotope stage 8. "Neanderthals", by convention, are fossils which date to after this gap. DNA from archaic humans from the 430,000-year-old Sima de los Huesos site in Spain indicate that they are more closely related to Neanderthals than to Denisovans, indicating that the split between Neanderthals and Denisovans must predate this time. The 400,000-year-old Aroeira 3 skull may also represent an early member of

9072-424: Is not normally applied to harvesting fish raised in controlled cultivations ( fish farming ). Nor is it normally applied to hunting aquatic mammals , where terms like whaling and sealing are used instead. Fishing has been an important part of human culture since hunter-gatherer times. It is one of the few food production activities that has persisted from prehistory into the modern age , surviving both

9234-465: Is the biggest part of the brain; it is small in hagfish and lampreys , but very large in mormyrids , processing their electrical sense . The brain stem or myelencephalon controls some muscles and body organs, and governs respiration and osmoregulation . The lateral line system is a network of sensors in the skin which detects gentle currents and vibrations, and senses the motion of nearby fish, whether predators or prey. This can be considered both

9396-512: Is the capture of fish for commercial purposes. Those who practice it must often pursue fish far from the land under adverse conditions. Commercial fishermen harvest a wide range of aquatic species, from tuna , cod and salmon to shrimp , krill , lobster , clams , squid and crab , in various fisheries for these species. Commercial fishing methods have become very efficient using large nets and sea-going processing factories. Individual fishing quotas and international treaties seek to control

9558-483: The Altai Mountains in southern Siberia. Pre- and early Neanderthals, on the other hand, seem to have continuously occupied only France, Spain and Italy, although some appear to have moved out of this "core-area" to form temporary settlements eastward (although without leaving Europe). Nonetheless, southwestern France has the highest density of sites for pre-, early and classic Neanderthals. The Neanderthals were

9720-512: The Divje Babe flute ), and Spanish cave paintings contentiously dated to before 65,000 years ago. Some claims of religious beliefs have been made. Neanderthals were likely capable of speech, possibly articulate, although the complexity of their language is not known. Compared with modern humans, Neanderthals had a more robust build and proportionally shorter limbs. Researchers often explain these features as adaptations to conserve heat in

9882-632: The Food and Agriculture Organization of the United Nations, the total world capture fisheries production in 2000 was 86 million tons (FAO 2002). The top producing countries were, in order, the People's Republic of China (excluding Hong Kong and Taiwan), Peru, Japan, the United States, Chile, Indonesia, Russia, India, Thailand, Norway, and Iceland. Those countries accounted for more than half of

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10044-617: The Gnathostomata or (for bony fish) Osteichthyes , also contains the clade of tetrapods (four-limbed vertebrates, mostly terrestrial), which are usually not considered fish. Some tetrapods, such as cetaceans and ichthyosaurs , have secondarily acquired a fish-like body shape through convergent evolution . Fishes of the World comments that "it is increasingly widely accepted that tetrapods, including ourselves, are simply modified bony fishes, and so we are comfortable with using

10206-723: The Indo-Pacific constitute the center of diversity for marine fishes, whereas continental freshwater fishes are most diverse in large river basins of tropical rainforests , especially the Amazon , Congo , and Mekong basins. More than 5,600 fish species inhabit Neotropical freshwaters alone, such that Neotropical fishes represent about 10% of all vertebrate species on the Earth. Fish are abundant in most bodies of water. They can be found in nearly all aquatic environments, from high mountain streams (e.g., char and gudgeon ) to

10368-546: The Neolithic Revolution and successive Industrial Revolutions . In addition to fishing for food , people commonly fish as a recreational pastime . Fishing tournaments are held, and caught fish are sometimes kept long-term as preserved or living trophies . When bioblitzes occur, fish are typically caught, identified, and then released . According to the United Nations FAO statistics,

10530-681: The Silurian , with giant armoured placoderms such as Dunkleosteus . Jawed fish, too, appeared during the Silurian: the cartilaginous Chondrichthyes and the bony Osteichthyes . During the Devonian , fish diversity greatly increased, including among the placoderms, lobe-finned fishes, and early sharks, earning the Devonian the epithet "the age of fishes". Fishes are a paraphyletic group, since any clade containing all fish, such as

10692-465: The West Indies . Bamboo rods became the generally favoured option from the mid-19th century, and several strips of the material were cut from the cane, milled into shape, and then glued together to form the light, strong, hexagonal rods with a solid core that were superior to anything that preceded them. George Cotton and his predecessors fished their flies with long rods, and light lines allowing

10854-477: The abyssal and even hadal depths of the deepest oceans (e.g., cusk-eels and snailfish ), although none have been found in the deepest 25% of the ocean. The deepest living fish in the ocean so far found is a cusk-eel, Abyssobrotula galatheae , recorded at the bottom of the Puerto Rico Trench at 8,370 m (27,460 ft). In terms of temperature, Jonah's icefish live in cold waters of

11016-402: The body mass index for Neanderthal males was calculated to be 26.9–28.2, which in modern humans correlates to being overweight . This indicates a very robust build. The Neanderthal LEPR gene concerned with storing fat and body heat production is similar to that of the woolly mammoth , and so was likely an adaptation for cold climate. The neck vertebrae of Neanderthals are thicker from

11178-425: The end-Devonian extinction wiped out the apex placoderms. Bony fish are further divided into the lobe-finned and ray-finned fish . About 96% of all living fish species today are teleosts , a crown group of ray-finned fish that can protrude their jaws . The tetrapods , a mostly terrestrial clade of vertebrates that have dominated the top trophic levels in both aquatic and terrestrial ecosystems since

11340-400: The intertidal zone , are facultative air breathers, able to breathe air when out of water, as may occur daily at low tide , and to use their gills when in water. Some coastal fish like rockskippers and mudskippers choose to leave the water to feed in habitats temporarily exposed to the air. Some catfish absorb air through their digestive tracts. The digestive system consists of a tube,

11502-405: The ostracoderms , had heavy bony plates that served as protective exoskeletons against invertebrate predators . The first fish with jaws , the placoderms, appeared in the Silurian and greatly diversified during the Devonian , the "Age of Fishes". Bony fish, distinguished by the presence of swim bladders and later ossified endoskeletons , emerged as the dominant group of fish after

11664-462: The pelvis connects to the spine ) was more vertically inclined, and was placed lower in relation to the pelvis, causing the spine to be less curved (exhibit less lordosis ) and to fold in on itself somewhat (to be invaginated). In modern populations, this condition affects just a proportion of the population, and is known as a lumbarised sacrum. Such modifications to the spine would have enhanced side-to-side (mediolateral) flexion , better supporting

11826-489: The stern , designed for convenient fishing. Traditional fishing is any kind of small scale, commercial or subsistence fishing practices using traditional techniques such as rod and tackle , arrows and harpoons , throw nets and drag nets, etc. Recreational and sport fishing refer to fishing primarily for pleasure or competition. Recreational fishing has conventions, rules, licensing restrictions and laws that limit how fish may be caught; typically, these prohibit

11988-425: The stout infantfish . Swimming performance varies from fish such as tuna, salmon , and jacks that can cover 10–20 body-lengths per second to species such as eels and rays that swim no more than 0.5 body-lengths per second. A typical fish is cold-blooded , has a streamlined body for rapid swimming, extracts oxygen from water using gills, has two sets of paired fins, one or two dorsal fins, an anal fin and

12150-533: The 140- to 120,000-year-old Israeli Nesher Ramla remains, which feature a mix of Neanderthal and more ancient H. erectus traits, represent one such source population which recolonised Europe following a glacial period. Like modern humans, Neanderthals probably descended from a very small population with an effective population —the number of individuals who can bear or father children—of 3,000 to 12,000 approximately. However, Neanderthals maintained this very low population, proliferating weakly harmful genes due to

12312-478: The 1911 The Quest for Fire by J.-H. Rosny aîné and the 1927 The Grisly Folk by H. G. Wells in which they are depicted as monsters. In 1911, Scottish anthropologist Arthur Keith reconstructed La Chapelle-aux-Saints 1 as an immediate precursor to modern humans, sitting next to a fire, producing tools, wearing a necklace, and having a more humanlike posture, but this failed to garner much scientific rapport, and Keith later abandoned his thesis in 1915. By

12474-595: The German fishing fleet. The earliest steam-powered fishing boats first appeared in the 1870s and used the trawl system of fishing as well as lines and drift nets. These were large boats, usually 80–90 feet (24–27 m) in length with a beam of around 20 feet (6 m). They weighed 40–50 tons and travelled at 9–11 knots (17–20 km/h; 10–13 mph). David Allen designed and made the earliest purpose-built fishing vessels in Leith , Scotland in March 1875, when he converted

12636-689: The Italian Visogliano and Fontana Ranuccio sites indicate that Neanderthal dental features had evolved by around 450–430,000 years ago during the Middle Pleistocene . There are two main hypotheses regarding the evolution of Neanderthals following the Neanderthal/human split: two-phase and accretion. Two-phase argues that a single major environmental event—such as the Saale glaciation —caused European H. heidelbergensis to increase rapidly in body size and robustness, as well as undergoing

12798-624: The Late Paleozoic , evolved from lobe-finned fish during the Carboniferous , developing air-breathing lungs homologous to swim bladders. Despite the cladistic lineage, tetrapods are usually not considered to be fish, making "fish" a paraphyletic group. Fish have been an important natural resource for humans since prehistoric times, especially as food . Commercial and subsistence fishers harvest fish in wild fisheries or farm them in ponds or in breeding cages in

12960-456: The Neanderthal braincase to that of a chimpanzee and argued that they were "incapable of moral and [ theistic ] conceptions". The first Neanderthal remains— Engis 2 (a skull)—were discovered in 1829 by Dutch/Belgian prehistorian Philippe-Charles Schmerling in the Grottes d'Engis , Belgium. He concluded that these "poorly developed" human remains must have been buried at the same time and by

13122-525: The Neanderthal line. It is possible that gene flow between Western Europe and Africa during the Middle Pleistocene, may have obscured Neanderthal characteristics in some Middle Pleistocene European hominin specimens, such those from Ceprano , Italy, and Sićevo Gorge , Serbia. The fossil record is much more complete from 130,000 years ago onwards, and specimens from this period make up the bulk of known Neanderthal skeletons. Dental remains from

13284-484: The Neanderthal/human split have been suggested. The date of around 250,000 years ago cites " H. helmei " as being the last common ancestor (LCA), and the split is associated with the Levallois technique of making stone tools. The date of about 400,000 years ago uses H. heidelbergensis as the LCA. Estimates of 600,000 years ago assume that " H. rhodesiensis " was the LCA, which split off into modern human lineage and

13446-601: The Southern Ocean, including under the Filchner–Ronne Ice Shelf at a latitude of 79°S, while desert pupfish live in desert springs, streams, and marshes, sometimes highly saline, with water temperatures as high as 36 C. A few fish live mostly on land or lay their eggs on land near water. Mudskippers feed and interact with one another on mudflats and go underwater to hide in their burrows. A single undescribed species of Phreatobius has been called

13608-547: The abilities to create fire , build cave hearths (to cook food, keep warm, defend themselves from animals, placing it at the centre of their homes), make adhesive birch bark tar , craft at least simple clothes similar to blankets and ponchos, weave, go seafaring through the Mediterranean, make use of medicinal plants , treat severe injuries, store food, and use various cooking techniques such as roasting , boiling , and smoking . Neanderthals consumed

13770-429: The absence of Neanderthal-derived patrilineal Y-chromosome and matrilineal mitochondrial DNA (mtDNA) in modern humans, along with the underrepresentation of Neanderthal X chromosome DNA, could imply reduced fertility or frequent sterility of some hybrid crosses, representing a partial biological reproductive barrier between the groups, and therefore species distinction. In 2014 geneticist Svante Pääbo summarised

13932-401: The actual function would be unclear, so they may not be a good indicator of evolutionary pressures to evolve such a nose. Further, a computer reconstruction of the Neanderthal nose and predicted soft tissue patterns shows some similarities to those of modern Arctic peoples, potentially meaning the noses of both populations convergently evolved for breathing cold, dry air. Neanderthals featured

14094-597: The age of 20 died before reaching 40. This high mortality rate was probably due to their high-stress environment. However, it has also been estimated that the age pyramids for Neanderthals and contemporary modern humans were the same. Infant mortality was estimated to have been very high for Neanderthals, about 43% in northern Eurasia. Neanderthals had more robust and stockier builds than typical modern humans, wider and barrel-shaped rib cages; wider pelvises; and proportionally shorter forearms and forelegs. Based on 45 Neanderthal long bones from 14 men and 7 women,

14256-592: The assistance of a drone . Why a fish bites a baited hook or lure involves several factors related to the sensory physiology, behaviour, feeding ecology, and biology of the fish as well as the environment and characteristics of the bait/hook/lure. There is an intricate link between various fishing techniques and knowledge about the fish and their behaviour including migration , foraging and habitat . The effective use of fishing techniques often depends on this additional knowledge. Some fishers follow fishing folklores which claim that fish feeding patterns are influenced by

14418-556: The attention of the cleaners. Cleaning behaviors have been observed in a number of fish groups, including an interesting case between two cichlids of the same genus, Etroplus maculatus , the cleaner, and the much larger E. suratensis . Fish occupy many trophic levels in freshwater and marine food webs . Fish at the higher levels are predatory , and a substantial part of their prey consists of other fish. In addition, mammals such as dolphins and seals feed on fish, alongside birds such as gannets and cormorants . The body of

14580-422: The average height was 164 to 168 cm (5 ft 5 in to 5 ft 6 in) for males and 152 to 156 cm (5 ft 0 in to 5 ft 1 in) for females. For comparison, the average height of 20 males and 10 females Upper Palaeolithic humans is, respectively, 176.2 cm (5 ft 9.4 in) and 162.9 cm (5 ft 4.1 in), although this decreases by 10 cm (4 in) nearer

14742-489: The average was about 1,478 cm (90.2 cu in) disregarding sex, and modern human brain size is suggested to have decreased since the Upper Palaeolithic. The largest Neanderthal brain, Amud 1 , was calculated to be 1,736 cm (105.9 cu in), one of the largest ever recorded in hominids. Both Neanderthal and human infants measure about 400 cm (24 cu in). When viewed from

14904-522: The bait to drift a long way out with the current. Geared multiplying reels never successfully caught on in Britain, but had more success in the United States, where George Snyder of Kentucky modified similar models into his bait-casting reel, the first American-made design in 1810. The material used for the rod itself changed from the heavy woods native to England to lighter and more elastic varieties imported from abroad, especially from South America and

15066-550: The body, and produce a concentrated urine. The reverse happens in freshwater fish : they tend to gain water osmotically, and produce a dilute urine. Some fish have kidneys able to operate in both freshwater and saltwater. Fish have small brains relative to body size compared with other vertebrates, typically one-fifteenth the brain mass of a similarly sized bird or mammal. However, some fish have relatively large brains, notably mormyrids and sharks , which have brains about as large for their body weight as birds and marsupials . At

15228-403: The classification remains uncertain. Neanderthals are known from numerous fossils, especially from after 130,000 years ago. The reasons for Neanderthal extinction are disputed. Theories for their extinction include demographic factors such as small population size and inbreeding, competitive replacement, interbreeding and assimilation with modern humans, change of climate, disease, or

15390-637: The colder mammoth steppe , and DNA analysis indicating a higher proportion of fast-twitch muscle fibres in Neanderthals than in modern humans. He explained their body proportions and greater muscle mass as adaptations to sprinting as opposed to the endurance-oriented modern human physique , as persistence hunting may only be effective in hot climates where the hunter can run prey to the point of heat exhaustion ( hyperthermia ). They had longer heel bones , reducing their ability for endurance running, and their shorter limbs would have reduced moment arm at

15552-429: The controversy, describing such " taxonomic wars" as unresolvable, "since there is no definition of species perfectly describing the case". Neanderthals are thought to have been more closely related to Denisovans than to modern humans. Likewise, Neanderthals and Denisovans share a more recent last common ancestor (LCA) than to modern humans, based on nuclear DNA (nDNA). However, Neanderthals and modern humans share

15714-581: The cranium, thigh bones, right arm, left humerus and ulna , left ilium (hip bone), part of the right shoulder blade , and pieces of the ribs . Following Charles Darwin 's On the Origin of Species , Fuhlrott and Schaaffhausen argued the bones represented an ancient modern human form; Schaaffhausen, a social Darwinist , believed that humans linearly progressed from savage to civilised, and so concluded that Neanderthals were barbarous cave-dwellers. Fuhlrott and Schaaffhausen met opposition namely from

15876-470: The descendants of a very early migration out of Africa around 1.9 mya. Pre- and early Neanderthals, living before the Eemian interglacial (130,000 years ago), are poorly known and come mostly from Western European sites. From 130,000 years ago onwards, the quality of the fossil record increases dramatically with classic Neanderthals, who are recorded from Western, Central, Eastern and Mediterranean Europe, as well as Southwest , Central and Northern Asia up to

16038-558: The early 2000s, supposed hybrid specimens were discovered: Lagar Velho 1 and Muierii 1 . However, similar anatomy could also have been caused by adapting to a similar environment rather than interbreeding. Neanderthal admixture was found to be present in modern populations in 2010 with the mapping of the first Neanderthal genome sequence. This was based on three specimens in Vindija Cave , Croatia, which contained almost 4% archaic DNA (allowing for near complete sequencing of

16200-475: The early 20th century, European researchers depicted Neanderthals as primitive, unintelligent and brutish. Although knowledge and perception of them has markedly changed since then in the scientific community, the image of the unevolved caveman archetype remains prevalent in popular culture. In truth, Neanderthal technology was quite sophisticated. It includes the Mousterian stone-tool industry as well as

16362-449: The end of World War II . In 1931, the first powered drum was created by Laurie Jarelainen. The drum was a circular device that was set to the side of the boat and would draw in the nets. Since World War II , radio navigation aids and fish finders have been widely used. The first trawlers fished over the side, rather than over the stern . The first purpose-built stern trawler was Fairtry built in 1953 at Aberdeen , Scotland. The ship

16524-659: The end of the period based on 21 males and 15 females; and the average in the year 1900 was 163 cm (5 ft 4 in) and 152.7 cm (5 ft 0 in), respectively. The fossil record shows that adult Neanderthals varied from about 147.5 to 177 cm (4 ft 10 in to 5 ft 10 in) in height, although some may have grown much taller (73.8 to 184.8 cm based on footprint length and from 65.8 to 189.3 cm based on footprint width). For Neanderthal weight, samples of 26 specimens found an average of 77.6 kg (171 lb) for males and 66.4 kg (146 lb) for females. Using 76 kg (168 lb),

16686-570: The exact root is unknown; some authorities reconstruct a Proto-Indo-European root * peysk- , attested only in Italic , Celtic , and Germanic . About 530 million years ago during the Cambrian explosion , fishlike animals with a notochord and eyes at the front of the body, such as Haikouichthys , appear in the fossil record . During the late Cambrian , other jawless forms such as conodonts appear. Jawed vertebrates appear in

16848-494: The first human species to permanently occupy Europe as the continent was only sporadically occupied by earlier humans. The southernmost find was recorded at Shuqba Cave , Levant; reports of Neanderthals from the North African Jebel Irhoud and Haua Fteah have been reidentified as H. sapiens . Their easternmost presence is recorded at Denisova Cave , Siberia 85°E ; the southeast Chinese Maba Man ,

17010-462: The first modern fishing port. The elegant Brixham trawler spread across the world, influencing fishing fleets everywhere. By the end of the 19th century, there were over 3,000 fishing trawlers in commission in Britain, with almost 1,000 at Grimsby. These trawlers were sold to fishers around Europe, including from the Netherlands and Scandinavia . Twelve trawlers went on to form the nucleus of

17172-405: The front of the brain are the olfactory lobes , a pair of structures that receive and process signals from the nostrils via the two olfactory nerves . Fish that hunt primarily by smell, such as hagfish and sharks, have very large olfactory lobes. Behind these is the telencephalon , which in fish deals mostly with olfaction. Together these structures form the forebrain. Connecting the forebrain to

17334-399: The front to the rear and transversely than those of (most) modern humans, leading to stability, possibly to accommodate a different head shape and size. Although the Neanderthal thorax (where the ribcage is) was similar in size to modern humans, the longer and straighter ribs would have equated to a widened mid-lower thorax and stronger breathing in the lower thorax, which are indicative of

17496-466: The gene variants inherited from Neanderthals may have been detrimental and selected out, Neanderthal introgression appears to have affected the modern human immune system , and is also implicated in several other biological functions and structures, but a large portion appears to be non-coding DNA . Neanderthals are named after the Neander Valley in which the first identified specimen

17658-400: The genome). However, there was approximately 1 error for every 200 letters ( base pairs ) based on the implausibly high mutation rate, probably due to the preservation of the sample. In 2012, British-American geneticist Graham Coop hypothesised that they instead found evidence of a different archaic human species interbreeding with modern humans, which was disproven in 2013 by the sequencing of

17820-412: The gills flows in the opposite direction to the water, resulting in efficient countercurrent exchange . The gills push the oxygen-poor water out through openings in the sides of the pharynx. Cartilaginous fish have multiple gill openings: sharks usually have five, sometimes six or seven pairs; they often have to swim to oxygenate their gills. Bony fish have a single gill opening on each side, hidden beneath

17982-532: The gills. Oxygen-rich blood then flows without further pumping, unlike in mammals, to the body tissues. Finally, oxygen-depleted blood returns to the heart. Fish exchange gases using gills on either side of the pharynx . Gills consist of comblike structures called filaments. Each filament contains a capillary network that provides a large surface area for exchanging oxygen and carbon dioxide . Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gills. Capillary blood in

18144-447: The gut, leading from the mouth to the anus. The mouth of most fishes contains teeth to grip prey, bite off or scrape plant material, or crush the food. An esophagus carries food to the stomach where it may be stored and partially digested. A sphincter, the pylorus, releases food to the intestine at intervals. Many fish have finger-shaped pouches, pyloric caeca , around the pylorus, of doubtful function. The pancreas secretes enzymes into

18306-457: The heart from the rest of the body. Lungfish, bichirs, ropefish, bowfins, snakefish, and the African knifefish have evolved to reduce such mixing, and to reduce oxygen loss from the gills to oxygen-poor water. Bichirs and lungfish have tetrapod-like paired lungs, requiring them to surface to gulp air, and making them obligate air breathers. Many other fish, including inhabitants of rock pools and

18468-526: The individual reconstructed on the basis of the skull was occasionally called "the Neanderthal man". The binomial name Homo neanderthalensis —extending the name "Neanderthal man" from the individual specimen to the entire species, and formally recognising it as distinct from humans—was first proposed by Irish geologist William King in a paper read to the 33rd British Science Association in 1863. However, in 1864, he recommended that Neanderthals and modern humans be classified in different genera as he compared

18630-455: The intestine to digest the food; other enzymes are secreted directly by the intestine itself. The liver produces bile which helps to break up fat into an emulsion which can be absorbed in the intestine. Most fish release their nitrogenous wastes as ammonia . This may be excreted through the gills or filtered by the kidneys . Salt is excreted by the rectal gland. Saltwater fish tend to lose water by osmosis ; their kidneys return water to

18792-522: The limbs, allowing for greater net rotational force at the wrists and ankles, causing faster acceleration. In 1981, American palaeoanthropologist Erik Trinkaus made note of this alternate explanation, but considered it less likely. Neanderthals had less developed chins, sloping foreheads, and longer, broader, more projecting noses. The Neanderthal skull is typically more elongated, but also wider, and less globular than that of most modern humans, and features much more of an occipital bun , or "chignon",

18954-419: The lungs and retained moisture ("nasal radiator" hypothesis); if their noses were wider, it would differ to the generally narrowed shape in cold-adapted creatures, and that it would have been caused instead by genetic drift . Also, the sinuses reconstructed wide are not grossly large, being comparable in size to those of modern humans. However, if sinus size is not an important factor for breathing cold air, then

19116-494: The material record in things like social behaviours, technological innovation and artistic output. The lack of sunlight most likely led to the proliferation of lighter skin in Neanderthals; however, it has been recently claimed that light skin in modern Europeans was not particularly prolific until perhaps the Bronze Age . Genetically, BNC2 was present in Neanderthals, which is associated with light skin colour; however,

19278-419: The midbrain is the diencephalon ; it works with hormones and homeostasis . The pineal body is just above the diencephalon; it detects light, maintains circadian rhythms, and controls color changes. The midbrain contains the two optic lobes . These are very large in species that hunt by sight, such as rainbow trout and cichlids . The hindbrain controls swimming and balance.The single-lobed cerebellum

19440-426: The middle and lower classes began to have an effect on fly fishing, which steadily grew in mass appeal. The expansion of the railway network in Britain allowed the less affluent for the first time to take weekend trips to the seaside or rivers for fishing. Richer hobbyists ventured further abroad. The large rivers of Norway replete with large stocks of salmon began to attract fishers from England in large numbers in

19602-416: The middle of the century – Jones's guide to Norway, and salmon-fisher's pocket companion , published in 1848, was written by Frederic Tolfrey and was a popular guide to the country. Modern reel design had begun in England during the latter part of the 18th century, and the predominant model in use was known as the ' Nottingham reel'. The reel was a wide drum that spooled out freely and was ideal for allowing

19764-475: The middle of the century, based on the exposure of Piltdown Man as a hoax as well as a reexamination of La Chapelle-aux-Saints 1 (who had osteoarthritis which caused slouching in life) and new discoveries, the scientific community began to rework its understanding of Neanderthals. Ideas such as Neanderthal behaviour, intelligence and culture were being discussed, and a more humanlike image of them emerged. In 1939, American anthropologist Carleton Coon reconstructed

19926-528: The most suitable Neanderthal habitats. Populations may have peaked in cold but not extreme intervals, such as marine isotope stages 8 and 6 (respectively, 300,000 and 191,000 years ago during the Saale glaciation). It is possible their range expanded and contracted as the ice retreated and grew, respectively, to avoid permafrost areas, residing in certain refuge zones during glacial maxima. In 2021, Israeli anthropologist Israel Hershkovitz and colleagues suggested

20088-501: The next century. He received a royal warrant from three successive monarchs starting with King George IV . He also invented the multiplying winch . The commercialization of the industry came at a time of expanded interest in fishing as a recreational hobby for members of the aristocracy . The impact of the Industrial Revolution was first felt in the manufacture of fly lines. Instead of anglers twisting their lines –

20250-443: The nine largest families; from largest to smallest, these are Cyprinidae , Gobiidae , Cichlidae , Characidae , Loricariidae , Balitoridae , Serranidae , Labridae , and Scorpaenidae . About 64 families are monotypic , containing only one species. Fish range in size from the huge 16-metre (52 ft) whale shark to some tiny teleosts only 8-millimetre (0.3 in) long, such as the cyprinid Paedocypris progenetica and

20412-494: The ocean possible for the first time, resulting in a massive migration of fishers from the ports in the south of England, to villages further north, such as Scarborough , Hull , Grimsby , Harwich and Yarmouth , that were points of access to the large fishing grounds in the Atlantic Ocean . The small village of Grimsby grew to become the largest fishing port in the world by the mid 19th century. An Act of Parliament

20574-500: The ocean. Fish are caught for recreation , or raised by fishkeepers as ornaments for private and public exhibition in aquaria and garden ponds . Fish have had a role in human culture through the ages, serving as deities , religious symbols, and as the subjects of art, books and movies. The word fish is inherited from Proto-Germanic , and is related to German Fisch , the Latin piscis and Old Irish īasc , though

20736-402: The open ocean. Because the water all around is the same colour, reflecting an image of the water offers near-invisibility. Fish have a closed-loop circulatory system . The heart pumps the blood in a single loop throughout the body; for comparison, the mammal heart has two loops, one for the lungs to pick up oxygen, one for the body to deliver the oxygen. In fish, the heart pumps blood through

20898-549: The other extreme, two-thirds (1.8 million) of the undecked boats are traditional craft of various types, powered only by sail and oars. These boats are used by artisan fishers . It is difficult to estimate how many recreational fishing boats there are, although the number is high. The term is fluid since some recreational boats may also be used for fishing from time to time. Unlike most commercial fishing vessels, recreational fishing boats are often not dedicated just to fishing. Just about anything that will stay afloat can be called

21060-487: The period of the Neanderthal/modern human transition, and Neanderthals may have been at a demographic disadvantage due to a lower fertility rate, a higher infant mortality rate, or a combination of the two. Estimates giving a total population in the higher tens of thousands are contested. A consistently low population may be explained in the context of the " Boserupian Trap ": a population's carrying capacity

21222-444: The position of the sun and the moon. Fishing tackles are the equipment used by fishers when fishing. Almost any equipment or gear used for fishing can be called a fishing tackle, although the term is most commonly associated with gear used in angling . Some examples are hooks , lines , sinkers , floats , rods , reels , baits , lures , spears , nets , gaffs , traps , waders , and tackle boxes. Fishing techniques refer to

21384-440: The prolific pathologist Rudolf Virchow who argued against defining new species based on only a single find. In 1872, Virchow erroneously interpreted Neanderthal characteristics as evidence of senility , disease and malformation instead of archaicness, which stalled Neanderthal research until the end of the century. By the early 20th century, numerous other Neanderthal discoveries were made, establishing H. neanderthalensis as

21546-510: The proportion of eumelanin (black pigment) to phaeomelanin (red pigment)—which is encoded by the MC1R gene. There are five known variants in modern humans of the gene which cause loss-of-function and are associated with light skin and hair colour, and another unknown variant in Neanderthals (the R307G variant) which could be associated with pale skin and red hair. The R307G variant was identified in

21708-455: The rear, the Neanderthal braincase has lower, wider, rounder appearance than in anatomically modern humans. This characteristic shape is referred to as "en bombe" (bomb-like), and is unique to Neanderthals, with all other hominid species (including most modern humans) generally having narrow and relatively upright cranial vaults, when viewed from behind. The Neanderthal brain would have been characterised by relatively smaller parietal lobes and

21870-480: The reduced effectivity of natural selection . Various studies, using mtDNA analysis, yield varying effective populations, such as about 1,000 to 5,000; 5,000 to 9,000 remaining constant; or 3,000 to 25,000 steadily increasing until 52,000 years ago before declining until extinction. Archaeological evidence suggests that there was a tenfold increase in the modern human population in Western Europe during

22032-462: The reel and dried every four hours or so to prevent them from becoming waterlogged. Another negative consequence was that it became easy for the much longer line to get into a tangle – this was called a 'tangle' in Britain, and a 'backlash' in the US. This problem spurred the invention of the regulator to evenly spool the line out and prevent tangling. The American, Charles F. Orvis, designed and distributed

22194-464: The rod, giving it much greater strength and flexibility. The industry also became commercialised – rods and tackle were sold at the haberdashers store. After the Great Fire of London in 1666, artisans moved to Redditch which became a centre of production of fishing-related products from the 1730s. Onesimus Ustonson established his shop in 1761, and his establishment remained a market leader for

22356-702: The same causes as the co-existing remains of extinct animal species. In 1848, Gibraltar 1 from Forbes' Quarry was presented to the Gibraltar Scientific Society by their Secretary Lieutenant Edmund Henry Réné Flint, but was thought to be a modern human skull. In 1856, local schoolteacher Johann Carl Fuhlrott recognised bones from Kleine Feldhofer Grotte in Neander Valley—Neanderthal 1 (the holotype specimen )—as distinct from modern humans, and gave them to German anthropologist Hermann Schaaffhausen to study in 1857. It comprised

22518-414: The same techniques. Recreational fishers fish for pleasure, sport, or to provide food for themselves, while commercial fishers fish for profit. Artisanal fishers use traditional, low-tech methods, for survival in third-world countries, and as a cultural heritage in other countries. Usually, recreational fishers use angling methods and commercial fishers use netting methods. A modern development is to fish with

22680-540: The same, about 285  N (64 lbf) and 255 N (57 lbf) in modern human males and females, respectively. The Neanderthal braincase averages 1,640 cm (100 cu in) for males and 1,460 cm (89 cu in) for females, which is significantly larger than the averages for all groups of extant humans; for example, modern European males average 1,362 cm (83.1 cu in) and females 1,201 cm (73.3 cu in). For 28 modern human specimens from 190,000 to 25,000 years ago,

22842-483: The species and quantities caught. A commercial fishing enterprise may vary from one person with a small boat with hand-casting nets or a few pot traps, to a huge fleet of trawlers processing tons of fish every day. Commercial fishing gear includes weights, nets (e.g. purse seine ), seine nets (e.g. beach seine), trawls (e.g. bottom trawl ), dredges, hooks and line (e.g. long line and handline ), lift nets, gillnets , entangling nets and traps . According to

23004-413: The standard English pronunciation of th with the fricative / θ / (as / n i ˈ æ n d ər θ ɔː l / ). The latter pronunciation, nevertheless, has no basis in the original German word which is pronounced always with a t regardless of the historical spelling. Neanderthal 1 , the type specimen , was known as the "Neanderthal cranium" or "Neanderthal skull" in anthropological literature, and

23166-507: The surrounding water, though some large active swimmers like white shark and tuna can hold a higher core temperature . Many fish can communicate acoustically with each other, such as during courtship displays . The earliest fish appeared during the Cambrian as small filter feeders ; they continued to evolve through the Paleozoic , diversifying into many forms. The earliest fish with dedicated respiratory gills and paired fins ,

23328-415: The taxon Osteichthyes as a clade, which now includes all tetrapods". The biodiversity of extant fish is unevenly distributed among the various groups; teleosts , bony fishes able to protrude their jaws , make up 96% of fish species. The cladogram shows the evolutionary relationships of all groups of living fishes (with their respective diversity ) and the tetrapods. Extinct groups are marked with

23490-431: The total number of commercial fishers and fish farmers is estimated to be 39.0 million. Fishing industries and aquaculture provide direct and indirect employment to over 500 million people in developing countries . In 2005, the worldwide per capita consumption of fish captured from wild fisheries was 14.4 kilograms (32 lb), with an additional 7.4 kilograms (16 lb) harvested from fish farms . Fishing

23652-410: The use of nets and the catching of fish with hooks not in the mouth. The most common form of recreational fishing is done with a rod , reel , line , hooks and any one of a wide range of baits or lures such as artificial flies . The practice of catching or attempting to catch fish with a hook is generally known as angling . In angling, it is sometimes expected or required that fish be returned to

23814-552: The use of specially trained animals such as cormorants and otters . There are also destructive fishing techniques (such as electrocution , blasting and poisoning ) that can do irreversible damage to the local ecosystems by killing/ sterilizing entire fish stocks , habitat destruction and/or upsetting the equilibrium of interspecific competitions , and such practices are often deemed illegal and liable to criminal punishments . Recreational , commercial and artisanal fishers use different techniques, and also, sometimes,

23976-514: The water ( catch and release ). Recreational or sport fishermen may log their catches or participate in fishing competitions. The estimated global number of recreational fishers varies from 220 million to a maximum number of 700 million fishers globally, which is thought to be double the number of individuals working as commercial fishers. In the United States alone it was estimated that 50.1 million people engaged in fishing activities in both saltwater and freshwater environments. Big-game fishing

24138-430: The ways the tackles are used when fishing. Tackles that are attached to the end of a fishing line are collectively called terminal tackles . These include hooks, sinkers, floats, leader lines, swivels , split rings, and any wires, snaps, beads, spoons, blades, spinners and clevises used to attach spinner blades to fishing lures. People also tend to use dead or live bait fish as another form of bait . A fishing vessel

24300-442: The wider lower thorax. It is claimed by some that this feature would be normal for all Homo , even tropically-adapted Homo ergaster or erectus , with the condition of a narrower thorax in most modern humans being a unique characteristic. Body proportions are usually cited as being "hyperarctic" as adaptations to the cold, because they are similar to those of human populations which developed in cold climates —the Neanderthal build

24462-484: The wild for recreational fishing or to supplement a species' natural population is generally referred to as a fish hatchery . Fish species raised by fish farms include salmon , carp , tilapia , catfish , white seabass and trout . Increased demands on wild fisheries by commercial fishing has caused widespread overfishing . Fish farming offers an alternative solution to the increasing market demand for fish. Fish A fish ( pl. : fish or fishes )

24624-457: The wind to do most of the work of getting the fly to the fish. Tackle design began to improve in the 1880s. The introduction of new woods to the manufacture of fly rods made it possible to cast flies into the wind on silk lines, instead of horse hair . These lines allowed for a much greater casting distance. However, these early fly lines proved troublesome as they had to be coated with various dressings to make them float and needed to be taken off

24786-433: The world's production; China alone accounted for a third of the world's production. Of that production, over 90% was marine and less than 10% was inland. A small number of species support the majority of the world's fisheries. Some of these species are herring , cod , sardine , anchovy , tuna , flounder , mullet , squid , shrimp , salmon , crab, lobster , oyster and scallops . All except these last four provided

24948-425: Was an unusually high frequency of taurodontism , a condition where the molars are bulkier due to an enlarged pulp (tooth core). Taurodontism was once thought to have been a distinguishing characteristic of Neanderthals which lent some mechanical advantage or stemmed from repetitive use, but was more likely simply a product of genetic drift. The bite force of Neanderthals and modern humans is now thought to be about

25110-660: Was first obtained in 1796, which authorised the construction of new quays and dredging of the Haven to make it deeper. It was only in 1846, with the tremendous expansion in the fishing industry , that the Grimsby Dock Company was formed. The foundation stone for the Royal Dock was laid by Albert the Prince consort in 1849. The dock covered 25 acres (10 ha) and was formally opened by Queen Victoria in 1854 as

25272-409: Was found in 1856 in the Neander Valley in present-day Germany. It is not clear when the line of Neanderthals split from that of modern humans ; studies have produced various times ranging from 315,000 to more than 800,000 years ago. The date of divergence of Neanderthals from their ancestor H. heidelbergensis is also unclear. The oldest potential Neanderthal bones date to 430,000 years ago, but

25434-410: Was found. The valley was spelled Neanderthal and the species was spelled Neanderthaler in German until the spelling reform of 1901 . The spelling Neandertal for the species is occasionally seen in English, even in scientific publications, but the scientific name, H. neanderthalensis , is always spelled with th according to the principle of priority . The vernacular name of the species in German

25596-531: Was likely to have been a means of survival, rather than recreation. The earliest English essay on recreational fishing was published in 1496, by Dame Juliana Berners , the prioress of the Benedictine Sopwell Nunnery . The essay was titled Treatyse of Fysshynge wyth an Angle , and included detailed information on fishing waters, the construction of rods and lines, and the use of natural baits and artificial flies. Recreational fishing took

25758-409: Was mainly an era of consolidation of the techniques developed in the previous century. Running rings began to appear along the fishing rods, which gave anglers greater control over the cast line. The rods themselves were also becoming increasingly sophisticated and specialised for different roles. Jointed rods became common from the middle of the century and bamboo came to be used for the top section of

25920-516: Was much larger than any other trawlers then in operation and inaugurated the era of the 'super trawler'. As the ship pulled its nets over the stern, it could lift out a much greater haul of up to 60 tons. The ship served as a basis for the expansion of 'super trawlers' around the world in the following decades. The early evolution of fishing as recreation is not clear. For example, there is anecdotal evidence for fly fishing in Japan. However, fly fishing

26082-407: Was of a sleek build and had a tall gaff rig , which gave the vessel sufficient speed to make long-distance trips out to the fishing grounds in the ocean. They were also sufficiently robust to be able to tow large trawls in deep water. The great trawling fleet that built up at Brixham earned the village the title of 'Mother of Deep-Sea Fisheries'. This revolutionary design made large-scale trawling in

26244-412: Was used as evidence that multiple different and unrelated branches of primitive humans existed, and supported Boule's reconstruction of H. neanderthalensis as a far distant relative and an evolutionary dead-end . He fuelled the popular image of Neanderthals as barbarous, slouching, club-wielding primitives; this image was reproduced for several decades and popularised in science fiction works, such as

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