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119-763: Kielder Forest is a large forestry plantation in Northumberland , England , surrounding Kielder village and the Kielder Water reservoir. It is the largest man-made woodland in England with three-quarters of its 250 square miles (650 km) covered by forest. The majority of the forest lies within Kielder Water and Forest Park, with the southern tip known as Wark Forest lying within Northumberland National Park . The forest

238-604: A fossil record extending back about 300 million years to the Paleozoic in the late Carboniferous period; even many of the modern genera are recognizable from fossils 60–120 million years old. Other classes and orders, now long extinct, also occur as fossils, particularly from the late Paleozoic and Mesozoic eras. Fossil conifers included many diverse forms, the most dramatically distinct from modern conifers being some herbaceous conifers with no woody stems. Major fossil orders of conifers or conifer-like plants include

357-697: A sister group to Pinales (the 'gnepine' hypothesis) or as being more derived than Pinales but sister to the rest of the group. Most recent studies favor the 'gnepine' hypothesis. The earliest conifers appear in the fossil record during the Late Carboniferous ( Pennsylvanian ), over 300 million years ago. Conifers are thought to be most closely related to the Cordaitales , a group of extinct Carboniferous-Permian trees and clambering plants whose reproductive structures had some similarities to those of conifers. The most primitive conifers belong to

476-532: A border. In the Roman era most of the county lay north of Hadrian's Wall , and the region was contested between England and Scotland into the Early Modern era, leading to the construction of many castles, peel towers and bastle houses , and the early modern fortifications at Berwick-upon-Tweed . Northumberland is also associated with Celtic Christianity , particularly the tidal island of Lindisfarne . During

595-462: A diverse physical geography. It is low and flat near the North Sea coast and increasingly mountainous towards the northwest. Being in the far north of England, above 55° latitude , and having many areas of high land, Northumberland is one of the coldest areas of the country. But as the county lies on the east coast, it has relatively low rainfall, with the highest amounts falling on the high land in

714-422: A female multicellular gametophyte. The female gametophytes grow to produce two or more archegonia , each of which contains an egg. Upon fertilization, the diploid egg will give rise to the embryo, and a seed is produced. The female cone then opens, releasing the seeds which grow to a young seedling . Conifer reproduction is synchronous with seasonal changes in temperate zones. Reproductive development slows to

833-399: A female cone and are drawn into a tiny opening on the ovule called the micropyle . It is within the ovule that pollen-germination occurs. From here, a pollen tube seeks out the female gametophyte, which contains archegonia each with an egg, and if successful, fertilization occurs. The resulting zygote develops into an embryo , which along with the female gametophyte (nutritional material for

952-665: A halt during each winter season and then resumes each spring. The male strobilus development is completed in a single year. Conifers are classified by three reproductive cycles that refer to the completion of female strobilus development from initiation to seed maturation. All three types of reproductive cycle have a long gap between pollination and fertilization . One year reproductive cycle : The genera include Abies , Picea , Cedrus , Pseudotsuga , Tsuga , Keteleeria ( Pinaceae ) and Cupressus , Thuja , Cryptomeria , Cunninghamia and Sequoia ( Cupressaceae ) . Female strobili are initiated in late summer or fall of

1071-536: A key role in the Industrial Revolution from the 18th century on. Many coal mines operated in Northumberland until the widespread closures in the 1970s and 1980s. Collieries operated at Ashington , Bedlington, Blyth , Choppington, Netherton, Ellington , and Pegswood . The region's coalfields fuelled industrial expansion in other areas of Britain, and the need to transport the coal from

1190-413: A maximum of energy from weak sunshine at high latitudes or under forest canopy shade. Conifers from hotter areas with high sunlight levels (e.g. Turkish pine Pinus brutia ) often have yellower-green leaves, while others (e.g. blue spruce , Picea pungens ) may develop blue or silvery leaves to reflect ultraviolet light. In the great majority of genera the leaves are evergreen , usually remaining on

1309-880: A needle-like appearance, but others, including most of the Cupressaceae and some of the Podocarpaceae , have flat, triangular scale-like leaves. Some, notably Agathis in Araucariaceae and Nageia in Podocarpaceae, have broad, flat strap-shaped leaves. Others such as Araucaria columnaris have leaves that are awl-shaped. In the majority of conifers, the leaves are arranged spirally, the exceptions being most of Cupressaceae and one genus in Podocarpaceae, where they are arranged in decussate opposite pairs or whorls of 3 (−4). In many species with spirally arranged leaves, such as Abies grandis (pictured),

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1428-472: A number of sites of special scientific interest , primarily associated with the upland moorland environment. A programme of restoration of Border mires is ongoing. The forest is also home to around 50% of England's red squirrel population, and is their biggest remaining stronghold in the country. The best place to catch a glimpse of one is at the squirrel hide at Kielder waterside. It also provides excellent habitat for many species of birds of prey . In 2009

1547-498: A number of art and architectural installations including a Skyspace designed by James Turrell and Wave Chamber , a camera obscura in a stone cairn by Chris Drury. The forest also contains Kielder Observatory which is an astronomical observatory . In 2010, former British distance runner Steve Cram inaugurated the Kielder Marathon which is a circuit around the lake taking in the surrounding gentle contours. In

1666-409: A pair of osprey ( Pandion haliaetus ) nested successfully in the forest. This pair have continued to nest there each year since, and a second pair nested in 2011. In 2014 eight young were fledged and by 2016 there were four nesting osprey pairs fledging eleven young. A large population of roe deer is actively managed. Due to the low human population and scarcity of roads and railways, Kielder Forest

1785-402: A protective cone called a strobilus . The cones take from four months to three years to reach maturity, and vary in size from 2 to 600 millimetres ( 1 ⁄ 8 to 23 + 5 ⁄ 8  in) long. In Pinaceae , Araucariaceae , Sciadopityaceae and most Cupressaceae , the cones are woody , and when mature the scales usually spread open allowing the seeds to fall out and be dispersed by

1904-487: A serious environmental issue causing problems for pastoral farming and for conservation . Radiata pine was introduced to Australia in the 1870s. It is "the dominant tree species in the Australian plantation estate" – so much so that many Australians are concerned by the resulting loss of native wildlife habitat. The species is widely regarded as an environmental weed across southeastern and southwestern Australia and

2023-735: A shire into the unified Kingdom of England by the House of Wessex. However, the West Saxon governmental structures were not extended beyond Tees, leaving the Earldom of Bamburgh and the Community of St. Cuthbert as contested buffer states with the emerging Kingdom of Scotland. After the battle of Nechtansmere , Northumbrian influence north of the Tweed began to decline as the Picts gradually reclaimed

2142-438: A single extant class , Pinopsida . All extant conifers are perennial woody plants with secondary growth . The great majority are trees , though a few are shrubs . Examples include cedars , Douglas-firs , cypresses , firs , junipers , kauri , larches , pines , hemlocks , redwoods , spruces , and yews . As of 2002, Pinophyta contained seven families, 60 to 65 genera, and more than 600 living species. Although

2261-406: A single white spruce tree from 1926 to 1961. Apical growth of the stem was slow from 1926 through 1936 when the tree was competing with herbs and shrubs and probably shaded by larger trees. Lateral branches began to show reduced growth and some were no longer in evidence on the 36-year-old tree. Apical growth totaling about 340 m, 370 m, 420 m, 450 m, 500 m, 600 m, and 600 m was made by the tree in

2380-501: A single-objective plantation to a multi-purpose forest mirrors the development of plantation forestry across the United Kingdom. Many archaeological remains can be found within the forest and are an important cultural link to the often turbulent history of the area. The name "Kielder Forest" is often also applied to the area of hills and remote moorland that surround the forestry plantations. The group of hills merges into

2499-771: A site formerly owned by Shultons, who originated Old Spice and were bought by P&G in 1990. McQuay UK makes air conditioning systems on the Bassington Industrial Estate at the A1068/A1172 junction in Cramlington , and Avery Dennison UK make labels on the Nelson Industrial Estate off of the A192. Schweppes' Abbey Well mineral water is made by Coca-Cola in the east of Morpeth . The National Renewable Energy Centre (Narec)

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2618-402: A two-tier council system for the county and its six districts , both responsible for different aspects of local government . These districts were Blyth Valley , Wansbeck , Castle Morpeth , Tynedale , Alnwick and Berwick-upon-Tweed . Elections for the unitary authority council first took place on 1 May 2008. The most recent elections, in 2021, had the following results: Northumberland

2737-469: A useful guide by which to assess the adequacy of particular nutrients, and the ratios among the major nutrients are helpful guides to nutritional imbalances. The softwood derived from conifers is of great economic value, providing about 45% of the world's annual lumber production. Other uses of the timber include the production of paper and plastic from chemically treated wood pulp. Some conifers also provide foods such as pine nuts and juniper berries ,

2856-488: A visitor and information centre. It is located on the edge of Kielder Village at the head of the River North Tyne valley. The Castle serves as a hub for the growing number of recreational facilities on offer, walking and cycling trails, picnic areas and a forest drive. A great football match is reported to have taken place between the men of Tynedale and Redesdale at Kielder Castle in 1790. The forest contains

2975-496: A whole. In the 2001 UK Census , 81% of the population reported their religion as Christianity, 0.8% as "other religion", and 12% as having no religion. Being primarily rural with significant areas of upland, the population density of Northumberland is only 62 persons per square kilometre, giving it the lowest population density in England . Northumberland's industry is dominated by some multinational corporations: Coca-Cola , MSD, GE and Drager all have significant facilities in

3094-591: A year, then they overwinter. Female strobili emerge followed by pollination in the following spring. Fertilization takes place in summer of the following year, only 3–4 months after pollination. Cones mature and seeds are then shed by the end of that same year. Pollination and fertilization occur in a single growing season. Two-year reproductive cycle : The genera includes Widdringtonia , Sequoiadendron ( Cupressaceae ) and most species of Pinus . Female strobilus initials are formed in late summer or fall then overwinter. Female strobili emerge and receive pollen in

3213-879: Is Hexham -based Egger (UK) Limited . Pharmaceutical , health care and emerging biotechnology companies form a very significant part of the county's economy. Many of these companies are part of the approximately 11,000-worker Northeast of England Process Industry Cluster (NEPIC) and include Aesica Pharmaceuticals, Arcinova, MSD , Piramal Healthcare , Procter & Gamble , Shire Plc (formerly SCM Pharma), Shasun Pharma Solutions, Specials Laboratory, and Thermo Fisher Scientific . The cluster also includes Cambridge Bioresearch, GlaxoSmithKline, Fujifilm Diosynth Biotech, Leica Bio, Data Trial, High Force Research, Non-Linear Dynamics, and Immuno Diagnostic Systems (IDS). The towns of Alnwick, Cramlington, Morpeth, Prudhoe all have significant pharmaceutical factories and laboratories. Newcastle University and Northumbria University are

3332-446: Is a coast redwood ( Sequoia sempervirens ), with a height of 115.55 metres (although one mountain ash, Eucalyptus regnans , allegedly grew to a height of 140 metres, the tallest living angiosperms are significantly smaller at around 100 metres. ) The thickest (that is, the tree with the greatest trunk diameter ) is a Montezuma cypress ( Taxodium mucronatum ), 11.42 metres in diameter. The largest tree by three-dimensional volume

3451-418: Is a giant sequoia ( Sequoiadendron giganteum ), with a volume 1486.9 cubic metres. The smallest is the pygmy pine ( Lepidothamnus laxifolius ) of New Zealand, which is seldom taller than 30 cm when mature. The oldest non-clonal living tree is a Great Basin bristlecone pine ( Pinus longaeva ), 4,700 years old. Since most conifers are evergreens, the leaves of many conifers are long, thin and have

3570-541: Is at Blyth. NOTE: New parishes have been added since 2001. These are missing from the list, see List of civil parishes in Northumberland . Although not on this list, the population of Cramlington is estimated at 39,000. Some settlements that are part historic county of Northumberland now fall under the county of Tyne and Wear : A precursor of modern football is still seen in the region at some annual Shrove Tuesday games at Alnwick . In 1280 at Ulgham near Morpeth Northumberland, records show that Henry of Ellington

3689-501: Is bordered by the Scottish Borders to the north, the North Sea to the east, Tyne and Wear and County Durham to the south, and Cumbria to the west. The town of Blyth is the largest settlement. Northumberland is the northernmost county in England. The county has an area of 5,013 km (1,936 sq mi) and a population of 320,274, making it the least-densely populated county in England. The south-east contains

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3808-674: Is divided into four parliamentary constituencies: Berwick-upon-Tweed, Blyth Valley, Wansbeck and Hexham. The 2019 general election produced the following total votes, with changes since previous election: On 23 June 2016 Northumberland took part in the UK-wide referendum on the UK's membership of the EU. In Northumberland a majority voted to leave the European Union. The only one of the four parliamentary constituencies in Northumberland to vote Remain

3927-833: Is dominated by conifers . Sitka spruce ( Picea sitchensis ) covers 75% of the planted area; this species thrives in the damp conditions afforded by northern Britain. Other species include Norway spruce ( Picea abies ) and lodgepole pine ( Pinus contorta ), which cover 9% of the area each. The remainder is made up of Scots pine ( Pinus sylvestris ), larch ( Larix spp.), Douglas-fir ( Pseudotsuga menziesii ), and broadleaves including birch ( Betula spp.), rowan ( Sorbus aucuparia ), cherry ( Prunus spp.), oak ( Quercus spp.), beech ( Fagus sylvatica ), and willow ( Salix spp.). 475,000 cubic metres of timber are harvested annually to supply local sawmilling , chipboard , pulp and wood fuel customers. Most of this volume comes from clearfelling areas; an increasing percentage however

4046-547: Is in the south of the county, surrounding Cramlington and other communities along the county border, to afford a protection from the Tyneside conurbation. The belt continues west along the border, past Darras Hall , and on to Hexham, stopping before Haydon Bridge . Its border there is shared with the North Pennines AONB. There are also some separated belt areas, for example to the east of Morpeth. The green belt

4165-610: Is next to the Anglo-Scottish border . The forest is owned and managed by Forestry England , which (as the Forestry Commission ) initiated the first plantings in the 1920s. During the 1930s, the Ministry of Labour supplied men from among the ranks of the unemployed. Many came from the mining communities and shipyards of North East England . They were housed in one of a number of instructional centres created by

4284-402: Is probably: first to apical growth and new needle formation, then to buds for the next year's growth, with the cambium in the older parts of the branches receiving sustenance last. In the white spruce studied by Fraser et al. (1964), the needles constituted 17.5% of the over-day weight. Undoubtedly, the proportions change with time. Wind and animal dispersals are two major mechanisms involved in

4403-508: Is sourced from stands harvested under continuous cover silviculture systems. Clear-felled areas are replanted with a mix of coniferous and broadleaf tree species; opportunities are also taken to increase the proportion of open space and to improve the riparian habitat. As with all Forestry England's woodlands, the timber is independently certified under the Forest Stewardship Council scheme. The forest contains

4522-432: Is the whitespotted sawyer ( Monochamus scutellatus ). Adults are found in summer on newly fallen or recently felled trees chewing tiny slits in the bark in which they lay eggs. The eggs hatch in about two weeks and the tiny larvae tunnel to the wood and score its surface with their feeding channels. With the onset of cooler weather, they bore into the wood, making oval entrance holes and tunnelling deeply. Feeding continues

4641-472: Is the fourth oldest in the country and is now Alnmouth Village Club; it is a 9 hole links course. Pinophyta Conifers are a group of cone-bearing seed plants , a subset of gymnosperms . Scientifically, they make up the division Pinophyta ( / p ɪ ˈ n ɒ f ɪ t ə , ˈ p aɪ n oʊ f aɪ t ə / ), also known as Coniferophyta ( / ˌ k ɒ n ɪ f ə ˈ r ɒ f ɪ t ə , - oʊ f aɪ t ə / ) or Coniferae . The division contains

4760-949: The Brigantes , to the south. During the Roman occupation of Britain , most of the present county lay north of Hadrian's Wall . It was controlled by Rome only for the brief period of its extension of power north to the Antonine Wall . The Roman road Dere Street crosses the county from Corbridge over high moorland west of the Cheviot Hills to Melrose, Scottish Borders ( Latin : Trimontium ). As evidence of its border position through medieval times, Northumberland has more castles than any other county in England, including prominent ones at Alnwick , Bamburgh , Dunstanburgh , Newcastle , and Warkworth . Nearly 2000-year-old Roman boxing gloves were uncovered at Vindolanda in 2017 by

4879-623: The Cephalotaxaceae may be better included within the Taxaceae, and some authors additionally recognize Phyllocladaceae as distinct from Podocarpaceae (in which it is included here). The family Taxodiaceae is here included in the family Cupressaceae, but was widely recognized in the past and can still be found in many field guides. A new classification and linear sequence based on molecular data can be found in an article by Christenhusz et al. The conifers are an ancient group, with

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4998-549: The Cheviots to the north-east but is generally well-defined on other sides. It reaches a maximum height of 602 m (1,975 ft) at Peel Fell and also contains the Marilyns of Sighty Crag and Larriston Fells. These hills, despite not being very high, are particularly remote owing to the scarcity of settlement in the region. Indeed, Sighty Crag is the furthest hill in England from a road, at four miles' distance. Kielder

5117-583: The Cordaitales , Vojnovskyales , Voltziales and perhaps also the Czekanowskiales (possibly more closely related to the Ginkgophyta ). Pinaceae Araucariaceae Podocarpaceae Sciadopityaceae Cupressaceae Cephalotaxaceae Taxaceae Multiple studies also indicate that the Gnetophyta belong within the conifers despite their distinct appearances, either placing them as

5236-586: The Humber Estuary . The land has long been an English frontier zone, and it is now bordered to the north by Scotland . Northumberland has a rich prehistory with many instances of rock art , hillforts such as Yeavering Bell , and stone circles such as the Goatstones and Duddo Five Stones . Most of the area was occupied by the Brythonic - Celtic Votadini people, with another large tribe,

5355-684: The Industrial Revolution the area had significant coal mining, shipbuilding, and armaments industries. The name of Northumberland is recorded in the Anglo-Saxon Chronicle as norð hẏmbra land , meaning "the land north of the Humber ". The name of the kingdom of Northumbria derives from the Old English Norþan-hymbre meaning "the people or province north of the Humber", as opposed to Southumbria , south of

5474-743: The Mesozoic era. Modern groups of conifers emerged from the Voltziales during the Late Permian through Jurassic . Conifers underwent a major decline in the Late Cretaceous corresponding to the explosive adaptive radiation of flowering plants . All living conifers are woody plants, and most are trees, the majority having a monopodial growth form (a single, straight trunk with side branches) with strong apical dominance . Many conifers have distinctly scented resin , secreted to protect

5593-555: The North Pennines AONB is also in the county. There is a variety of nature reserves in Northumberland including Holy Island National Nature Reserve and Farne Islands National Nature Reserve. Moreover, 50% of England's red squirrel population lives in the Kielder Water and Forest Park. Natural England recognises the following natural regions, or national character areas , that lie wholly or partially within Northumberland: The Cheviot Hills , in

5712-599: The Restoration in 1660, the county was a centre for Roman Catholicism in England, as well as a focus of Jacobite support. Northumberland was long a wild county, where outlaws and Border Reivers hid from the law. However, the frequent cross-border skirmishes and accompanying local lawlessness largely subsided after the Union of the Crowns of Scotland and England under King James I and VI in 1603. Northumberland played

5831-593: The River Tyne . The west of Northumberland contains part of the Cheviot Hills and North Pennines , while to the east the land becomes flatter before reaching the coast. The Cheviot (815 m (2,674 ft)), after which the range of hills is named, is the county's highest point. The county contains the source of the River North Tyne and much of the South Tyne; near Hexham they combine to form

5950-646: The Treaty of York (1237). The earls of Northumberland once wielded significant power in English affairs because, as powerful and militaristic Marcher Lords , they had the task of protecting England from Scottish retaliation for English invasions. Northumberland has a history of revolt and rebellion against the government, as seen in the Rising of the North (1569–1570) against Elizabeth I . These revolts were usually led by

6069-460: The influence of environmental conditions, their anatomical characteristics record growth rate changes produced by these changing conditions. The microscopic structure of conifer wood consists of two types of cells : parenchyma , which have an oval or polyhedral shape with approximately identical dimensions in three directions, and strongly elongated tracheids. Tracheids make up more than 90% of timber volume. The tracheids of earlywood formed at

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6188-402: The wind . In some (e.g. firs and cedars ), the cones disintegrate to release the seeds, and in others (e.g. the pines that produce pine nuts ) the nut-like seeds are dispersed by birds (mainly nutcrackers , and jays ), which break up the specially adapted softer cones. Ripe cones may remain on the plant for a varied amount of time before falling to the ground; in some fire-adapted pines,

6307-409: The 16 nutrient elements known to be essential to plants, 13 of which are obtained from the soil, including nitrogen , phosphorus , potassium , calcium , magnesium , and sulfur , all used in relatively large amounts. Nutrient concentrations in conifers also vary with season, age, and kind of tissue sampled, and analytical technique. The ranges of concentrations occurring in well-grown plants provide

6426-701: The English. The monastery at Lindisfarne was the centre of production of the Lindisfarne Gospels (around 700). It became the home of St Cuthbert (c. 634–687, abbot from c. 665), who is buried in Durham Cathedral . The Kingdom of Northumbria fragmented into a series of successor states following the Viking invasion. In the south, Viking settlers established the Kingdom of York between the Humber and Tees. However, Viking influence petered out at

6545-632: The Ministry, most of them on Forestry England property; by 1938, the Ministry had 38 Instructional Centres across Britain. The hutted camp in Kielder is now under Kielder Water. Numerous purpose-built villages were also constructed for workers' families, including Stonehaugh . Prior to the 1920s, the land was predominantly open moorland, managed for grouse shooting and sheep grazing with remnants of native upland woodland existing along stream sides and in isolated craggy areas. Forestry England, funded from

6664-743: The North American Forest Tree Nursery Soils Workshop at Syracuse in 1980 provided strong contrary evidence: Bob Eastman, President of the Western Maine Forest Nursery Co. stated that for 15 years he has been successful in avoiding winter “burn” to Norway spruce and white spruce in his nursery operation by fertilizing with 50–80 lb/ac (56–90 kg/ha) nitrogen in September, whereas previously winter burn had been experienced annually, often severely. Eastman also stated that

6783-464: The Pinales without Taxales as paraphyletic , and the latter order is no longer considered distinct. A more accurate subdivision would be to split the class into three orders, Pinales containing only Pinaceae, Araucariales containing Araucariaceae and Podocarpaceae, and Cupressales containing the remaining families (including Taxaceae), but there has not been any significant support for such a split, with

6902-668: The Tees, with the river serving as the northern boundary of the Danelaw. Between the Tyne and Tees rivers, the Community of St. Cuthbert emerged as the successors to the Bishops of Lindisfarne; north of the Tyne, the earls of Bamburgh, who traced their lineage back to the Kings of Northumbria, continued to exercise authority and governance over this northern expanse. York was eventually integrated as

7021-468: The Town Moor. The 'Pitmen's Derby' or Northumberland Plate was held from 1833 and moved to Gosforth in 1882. Modern day horse racing still takes place at Newcastle Racecourse. Golf is a Scottish import to many countries, but it is said to have been played in this region by St Cuthbert on the dunes of the Northumberland coast. The oldest club in Northumberland was at Alnmouth , founded in 1869, it

7140-683: The Tyne, which exits into Tyne and Wear shortly downstream. The other major rivers in Northumberland are, from south to north, the Blyth , Coquet , Aln , Wansbeck and Tweed , the last of which forms part of the Scottish border. The county contains Northumberland National Park and two national landscapes : the Northumberland Coast and part of the North Pennines . Much of the county's history has been defined by its position on

7259-531: The Vindolanda Trust experts, led by Andrew Birley . Being similar in style and function to the full-hand modern boxing gloves, those found at Vindolanda look like leather bands dating back to 120 AD. It is suggested that, based on their difference from gladiator gloves, the gloves were not used in mortal combat, but rather in a sport for promoting fighting skills. The gloves are currently displayed at Vindolanda's museum. Present-day Northumberland formed

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7378-558: The beginning of a growing season have large radial sizes and smaller, thinner cell walls . Then, the first tracheids of the transition zone are formed, where the radial size of cells and the thickness of their cell walls changes considerably. Finally, latewood tracheids are formed, with small radial sizes and greater cell wall thickness. This is the basic pattern of the internal cell structure of conifer tree rings. Most conifers are monoecious , but some are subdioecious or dioecious ; all are wind-pollinated . Conifer seeds develop inside

7497-506: The city became a part of the Tyne and Wear metropolitan county in 1974. The county council has been governed from Morpeth since 1981. In 2009 the county council became a unitary authority when the county's districts were abolished and the county council took on their functions. The unitary authority of Northumberland Council was established in April 2009 . Previously, the county had

7616-726: The coast of Northumberland are the Farne Islands , another dolerite outcrop, famous for their bird life. The Northumberland Coalfield extends across the southeast corner of the county, from the River Tyne as far north as Shilbottle . There were smaller-scale workings for coal within the Tyne Limestone Formation as far north as Scremerston . The term 'sea coal' likely originated from chunks of coal, found washed up on beaches, that wave action had broken from coastal outcroppings. Northumberland's green belt

7735-453: The collieries to the Tyne led to the development of the first railways. Shipbuilding and armaments manufacture were other important industries before the deindustrialisation of the 1980s. Northumberland remains largely rural, and is the least densely populated county in England. In recent years, the county has had considerable growth in tourism. Visitors are attracted both to its scenic beauty and to its historical sites. Northumberland has

7854-587: The cone consists of several fused scales, while in others (e.g. Taxaceae), the cone is reduced to just one seed scale or (e.g. Cephalotaxaceae) the several scales of a cone develop into individual arils, giving the appearance of a cluster of berries. The male cones have structures called microsporangia that produce yellowish pollen through meiosis. Pollen is released and carried by the wind to female cones. Pollen grains from living pinophyte species produce pollen tubes, much like those of angiosperms. The gymnosperm male gametophytes (pollen grains) are carried by wind to

7973-585: The core of the Anglian kingdom of Bernicia from about 547. It was united with Deira (south of the River Tees ) to form the kingdom of Northumbria in the 7th century. The boundaries of Northumbria under King Edwin (reigned 616–633) stretched from the Humber in the south to the Forth in the north. Northumberland is often called the "cradle of Christianity" in England because Christianity flourished on Lindisfarne —a tidal island north of Bamburgh , also called Holy Island—after King Oswald of Northumbria (r. 634–642) invited monks from Iona to come to convert

8092-402: The county, pre Second World War and immediate post war greats were George Camsell and Hughie Gallacher , these were described in the "Clown Prince of Football" by Len Shackleton . The author played for Newcastle United and Northumberland County Cricket Club . Shackleton's book was controversial when it was first published because chapter 9, named "The Average Director's Knowledge of Football",

8211-402: The dispersal of conifer seeds. Wind-born seed dispersal involves two processes, namely; local neighborhood dispersal and long-distance dispersal. Long-distance dispersal distances range from 11.9–33.7 kilometres (7.4–20.9 mi) from the source. Birds of the crow family, Corvidae , are the primary distributor of the conifer seeds. These birds are known to cache 32,000 pine seeds and transport

8330-419: The earls of Northumberland, the Percy family. Shakespeare makes one of the Percys, the dashing Harry Hotspur (1364–1403), the hero of his Henry IV, Part 1 . The Percys were often aided in conflict by other powerful Northern families, such as the Nevilles and the Patchetts. The latter were stripped of all power and titles by the victorious Parliamentarians after the English Civil War of 1642–1651. After

8449-567: The female cone is termed fruit , which undergoes ripening (maturation). It was found recently that the pollen of conifers transfers the mitochondrial organelles to the embryo , a sort of meiotic drive that perhaps explains why Pinus and other conifers are so productive, and perhaps also has bearing on observed sex-ratio bias. Conifers are heterosporous , generating two different types of spores: male microspores and female megaspores . These spores develop on separate male and female sporophylls on separate male and female cones. In

8568-449: The first year spring and become conelets. The conelet goes through another winter rest and, in the spring of the second year archegonia form in the conelet. Fertilization of the archegonia occurs by early summer of the second year, so the pollination-fertilization interval exceeds a year. After fertilization, the conelet is considered an immature cone. Maturation occurs by autumn of the second year, at which time seeds are shed. In summary,

8687-466: The following spring. Female strobili emerge then pollination occurs in spring of the second year then the pollinated strobili become conelets in the same year (i.e. the second year). The female gametophytes in the conelet develop so slowly that the megaspore does not go through free-nuclear divisions until autumn of the third year. The conelet then overwinters again in the free-nuclear female gametophyte stage. Fertilization takes place by early summer of

8806-492: The following summer when larvae occasionally return to the surface of the wood and extend the feeding channels generally in a U-shaped configuration. During this time, small piles of frass extruded by the larvae accumulate under logs. Early in the spring of the second year following egg-laying, the larvae, about 30 mm long, pupate in the tunnel enlargement just below the wood surface. The resulting adults chew their way out in early summer, leaving round exit holes, so completing

8925-512: The forest. In 2020, an episode of series 11 of Michael Portillo 's Great British Railway Journeys featured Kielder Forest, referring to the 1936 Bradshaw's Guide . [REDACTED] Media related to Kielder Forest at Wikimedia Commons Northumberland Northumberland ( / n ɔːr ˈ θ ʌ m b ər l ə n d / nor- THUM -bər-lənd ) is a ceremonial county in North East England , bordering Scotland . It

9044-483: The fourth attempt in 1910. Today top quality professional football remains in Northumberland. Since the 2017 – 18 season, Newcastle United is a Premier League team. St James' Park in Newcastle is a first class football venue, often used for international games at all levels. Blyth Spartans A.F.C. have had success and public attention through Football Association Cup runs. There are many notable footballers from

9163-399: The fourth year and seeds mature in the cones by autumn of the fourth year. The growth and form of a forest tree are the result of activity in the primary and secondary meristems , influenced by the distribution of photosynthate from its needles and the hormonal gradients controlled by the apical meristems. External factors also influence growth and form. Fraser recorded the development of

9282-419: The growing embryo) and its surrounding integument, becomes a seed . Eventually, the seed may fall to the ground and, if conditions permit, grow into a new plant. In forestry , the terminology of flowering plants has commonly though inaccurately been applied to cone-bearing trees as well. The male cone and unfertilized female cone are called male flower and female flower , respectively. After fertilization,

9401-530: The immense conifer forests of the world represent the largest terrestrial carbon sink . Conifers are of great economic value for softwood lumber and paper production. Conifer is a Latin word, a compound of conus (cone) and ferre (to bear), meaning "the one that bears (a) cone(s)". The division name Pinophyta conforms to the rules of the International Code of Nomenclature for algae, fungi, and plants (ICN), which state (Article 16.1) that

9520-623: The land previously invaded by the Anglian kingdom. In 1018, its northern part, the region between the Tweed and the Forth (including Lothian , which includes present-day Edinburgh ), was ceded to the Kingdom of Scotland. The Earldom of Northumberland was briefly held by the Scottish royal family by marriage, between 1139–1157 and 1215–1217. Scotland relinquished all claims to the region as part of

9639-535: The largest and economically most important component group of gymnosperms, but nevertheless they comprise only one of the four groups. The division Pinophyta consists of just one class, Pinopsida, which includes both living and fossil taxa. Subdivision of the living conifers into two or more orders has been proposed from time to time. The most commonly seen in the past was a split into two orders, Taxales (Taxaceae only) and Pinales (the rest), but recent research into DNA sequences suggests that this interpretation leaves

9758-429: The largest towns: Blyth (37,339), Cramlington (27,683), Ashington (27,670), and Morpeth (14,304), which is the administrative centre. The remainder of the county is rural, and the largest towns are Berwick-upon-Tweed (12,043) in the far north and Hexham (13,097) in the west. For local government purposes the county is a unitary authority area. The county historically included the parts of Tyne and Wear north of

9877-594: The leading academic institutions nearby. The local industry includes commercial or academic activity in pre-clinical research and development, clinical research and development, pilot-scale manufacturing, full-scale active pharmaceutical ingredient/intermediate manufacturing, formulation, packaging, and distribution. Having no large population centres, the county's mainstream media outlets are served from nearby Tyne and Wear , including radio stations and television channels (such as BBC Look North , BBC Radio Newcastle , ITV Tyne Tees and Hits Radio North East ), along with

9996-417: The leaf bases are twisted to present the leaves in a very flat plane for maximum light capture. Leaf size varies from 2 mm in many scale-leaved species, up to 400 mm long in the needles of some pines (e.g. Apache pine, Pinus engelmannii ). The stomata are in lines or patches on the leaves and can be closed when it is very dry or cold. The leaves are often dark green in colour, which may help absorb

10115-447: The majority of daily newspapers covering the area ( The Journal , Evening Chronicle ). It is worth remembering however that although Northumberland, like many administrative areas in England, has been shorn of its geographical regional centre, that centre—Newcastle upon Tyne—remains an essential element within the entity we know as Northumberland. Newcastle's newspapers are as widely read in its Northumbrian hinterland as any of those of

10234-402: The majority of opinion preferring retention of all the families within a single order Pinales, despite their antiquity and diverse morphology . There were seven families of conifers c.  2011 , with 65–70 genera and over 600 living species ( c.  2002 ). The seven most distinct families are linked in the box above right and phylogenetic diagram left. In other interpretations,

10353-403: The male cones, microspores are produced from microsporocytes by meiosis . The microspores develop into pollen grains, which contain the male gametophytes. Large amounts of pollen are released and carried by the wind. Some pollen grains will land on a female cone for pollination. The generative cell in the pollen grain divides into two haploid sperm cells by mitosis leading to the development of

10472-466: The names of higher taxa in plants (above the rank of family) are either formed from the name of an included family (usually the most common and/or representative), in this case Pinaceae (the pine family), or are descriptive. A descriptive name in widespread use for the conifers (at whatever rank is chosen) is Coniferae (Art 16 Ex 2). According to the ICN, it is possible to use a name formed by replacing

10591-531: The northwest of the county, consist mainly of resistant Devonian granite and andesite lava . A second area of igneous rock underlies the Whin Sill (on which Hadrian's Wall runs), an intrusion of Carboniferous dolerite . Both ridges support a rather bare moorland landscape. Either side of the Whin Sill the county lies on Carboniferous Limestone , giving some areas of karst landscape . Lying off

10710-486: The one 11 m tall white spruce, Fraser et al. (1964) speculated that if the photosynthate used in making apical growth in 1961 was manufactured the previous year, then the 4 million needles that were produced up to 1960 manufactured food for about 600,000 mm of apical growth or 730 g dry weight, over 12 million mm of wood for the 1961 annual ring, plus 1 million new needles, in addition to new tissue in branches, bark, and roots in 1960. Added to this would be

10829-424: The one-year and the two-year cycles differ mainly in the duration of the pollination-fertilization interval. Three-year reproductive cycle : Three of the conifer species are pine species ( Pinus pinea , Pinus leiophylla , Pinus torreyana ) which have pollination and fertilization events separated by a two-year interval. Female strobili initiated during late summer or autumn of a year, then overwinter until

10948-425: The overwintering storage capacity of stock thus treated was much improved (Eastman 1980). The concentrations of nutrients in plant tissues depend on many factors, including growing conditions. Interpretation of concentrations determined by analysis is easy only when a nutrient occurs in excessively low or occasionally excessively high concentration. Values are influenced by environmental factors and interactions among

11067-569: The paraphyletic assemblage of " walchian conifers ", which were small trees, and probably originated in dry upland habitats. The range of conifers expanded during the Early Permian ( Cisuralian ) to lowlands due to increasing aridity. Walchian conifers were gradually replaced by more advanced voltzialean or "transition" conifers. Conifers were largely unaffected by the Permian–Triassic extinction event , and were dominant land plants of

11186-429: The photosynthate to produce energy to sustain respiration over this period, an amount estimated to be about 10% of the total annual photosynthate production of a young healthy tree. On this basis, one needle produced food for about 0.19 mg dry weight of apical growth, 3 mm wood, one-quarter of a new needle, plus an unknown amount of branch wood, bark and roots. The order of priority of photosynthate distribution

11305-561: The plant for several (2–40) years before falling, but five genera ( Larix , Pseudolarix , Glyptostrobus , Metasequoia and Taxodium ) are deciduous , shedding their leaves in autumn. The seedlings of many conifers, including most of the Cupressaceae, and Pinus in Pinaceae, have a distinct juvenile foliage period where the leaves are different, often markedly so, from the typical adult leaves. Tree rings are records of

11424-416: The pollen tube. At fertilization, one of the sperm cells unites its haploid nucleus with the haploid nucleus of an egg cell. The female cone develops two ovules, each of which contains haploid megaspores. A megasporocyte is divided by meiosis in each ovule. Each winged pollen grain is a four celled male gametophyte . Three of the four cells break down leaving only a single surviving cell which will develop into

11543-446: The public purse, purchased land across the country with the brief of establishing a strategic reserve of timber for the nation. This single objective held sway until the 1960s. Since that time, management principles have changed in order to reflect rising awareness of environmental needs and to provide recreational facilities whilst seeking to maintain a sustainable supply of timber. Kielder today remains state-owned and its development from

11662-551: The region. Tourism is a major source of employment and income in Northumberland. In the early 2000s the county annually received 1.1 million British visitors and 50,000 foreign tourists, who spent a total of £162 million. Coal mining in the county goes back to Tudor times . Coal mines continue to operate today; many of them are open-cast mines. Planning approval was given in January 2014 for an open-cast mine at Halton Lea Gate near Lambley . A major employer in Northumberland

11781-514: The removal of individual plants beyond plantations is encouraged. At least 20 species of roundheaded borers of the family Cerambycidae feed on the wood of spruce , fir , and hemlock (Rose and Lindquist 1985). Borers rarely bore tunnels in living trees, although when populations are high, adult beetles feed on tender twig bark, and may damage young living trees. One of the most common and widely distributed borer species in North America

11900-578: The road and the often rough nature of the road surfaces. Notable Motorsport UK marshalls, such as Anthony Jamieson, hold the area in very high regard. Historically, the WRC RAC Rally would include several stages set in Kielder, often nicknamed "Killer Kielder" by commentators due to the high number of rally cars which could be expected to retire from the event at Kielder through accidents and mechanical failures. National rallies, Historic rallies, and British Rally Championship events continue to use

12019-556: The same decade the local authority began a reduction of late night street lights to enhance the dark skies of the park and sponsored observation facilities and advertising to attract visitors to the darkest skies in the mainland of Britain. The Kielder forest has been used extensively for rallying . The forest's tracks are regarded among crews and enthusiasts as particularly challenging, featuring lengthy straight sections on which very high speeds can be reached, but also deceptive corners, exacerbated by ditches that are very deep and close to

12138-453: The same effect in 105-day-old white spruce. The general short-term effect of nitrogen fertilization on coniferous seedlings is to stimulate shoot growth more so than root growth (Armson and Carman 1961). Over a longer period, root growth is also stimulated. Many nursery managers were long reluctant to apply nitrogenous fertilizers late in the growing season, for fear of increased danger of frost damage to succulent tissues. A presentation at

12257-927: The seeds as far as 12–22 km (7.5–13.7 mi) from the source. The birds store the seeds in the soil at depths of 2–3 cm ( 3 ⁄ 4 – 1 + 1 ⁄ 4  in) under conditions which favor germination . Conifers are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. A number of conifers originally introduced for forestry have become invasive species in parts of New Zealand , including radiata pine ( Pinus radiata ), lodgepole pine ( P. contorta ), Douglas fir ( Pseudotsuga mensiezii ) and European larch ( Larix decidua ). In parts of South Africa , maritime pine ( Pinus pinaster ), patula pine ( P. patula ) and radiata pine have been declared invasive species. These wilding conifers are

12376-545: The seeds may be stored in closed cones for up to 60–80 years, being released only when a fire kills the parent tree. In the families Podocarpaceae , Cephalotaxaceae , Taxaceae , and one Cupressaceae genus ( Juniperus ), the scales are soft, fleshy, sweet, and brightly colored, and are eaten by fruit-eating birds, which then pass the seeds in their droppings. These fleshy scales are (except in Juniperus ) known as arils . In some of these conifers (e.g. most Podocarpaceae),

12495-429: The sole source of nitrogen guanidine compounds were less prominent. Durzan and Steward noted that their results, drawn from determinations made in late summer, did not rule out the occurrence of different interim responses at other times of the year. Ammonium nitrogen produced significantly heavier (dry weight) seedlings with a higher nitrogen content after 5 weeks than did the same amount of nitrate nitrogen. Swan found

12614-682: The success and history of their respective clubs. They both became permanent fixtures in Alf Ramsey's 1966 England World Cup winning team. Malcolm Macdonald was a successful Newcastle player of the 1970s. Great national players who played at Northumberland clubs in the 1980s and 1990s include Peter Beardsley , Paul Gascoigne , Chris Waddle and Alan Shearer . Shearer remains the highest scoring player in Premier League history with 260 goals in 441 appearances. Early races were held at Newcastle's Killingworth Moor from 1632 before moving to

12733-517: The termination -aceae in the name of an included family, in this case preferably Pinaceae , by the appropriate termination, in the case of this division -ophyta . Alternatively, " descriptive botanical names " may also be used at any rank above family. Both are allowed. This means that if conifers are considered a division, they may be called Pinophyta or Coniferae. As a class, they may be called Pinopsida or Coniferae. As an order they may be called Pinales or Coniferae or Coniferales . Conifers are

12852-570: The total number of species is relatively small, conifers are ecologically important. They are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere , but also in similar cool climates in mountains further south. Boreal conifers have many wintertime adaptations. The narrow conical shape of northern conifers, and their downward-drooping limbs, help them shed snow. Many of them seasonally alter their biochemistry to make them more resistant to freezing. While tropical rainforests have more biodiversity and turnover,

12971-475: The tree against insect infestation and fungal infection of wounds. Fossilized resin hardens into amber , which has been commercially exploited historically (for example, in New Zealand's 19th-century kauri gum industry). The size of mature conifers varies from less than one metre to over 100 metres in height. The world's tallest, thickest, largest, and oldest living trees are all conifers. The tallest

13090-416: The usual 2-year life cycle. Conifers – notably Abies (fir), Cedrus , Chamaecyparis lawsoniana (Lawson's cypress), Cupressus (cypress), juniper , Picea (spruce), Pinus (pine), Taxus (yew), Thuja (cedar) – have been the subject of selection for ornamental purposes. Plants with unusual growth habits, sizes, and colours are propagated and planted in parks and gardens throughout

13209-437: The west. About a quarter of the county forms the Northumberland National Park , an area of outstanding landscape that has largely been protected from development and agriculture. The park stretches south from the Scottish border and includes Hadrian's Wall . Most of the park is over 240 metres (790 feet) above sea level. The Northumberland Coast is also a designated Area of Outstanding Natural Beauty (AONB). A small part of

13328-971: The wider county: the Northumberland Gazette , Morpeth Herald , Berwick Advertiser , Hexham Courant and the News Post Leader . Lionheart Radio , a community radio station based in Alnwick , has recently been awarded a five-year community broadcasting licence by Ofcom . Ashington has the former Alcan Lynemouth Aluminium Smelter , next to the Lynemouth Power Station . Hammerite and Cuprinol are made in Prudhoe by ICI Paints . A Procter & Gamble factory in Seaton Delaval makes Hugo Boss aftershave and Clairol and Nice 'n Easy hair dye at

13447-468: The world. Conifers can absorb nitrogen in either the ammonium (NH 4 ) or nitrate (NO 3 ) form, but the forms are not physiologically equivalent. Form of nitrogen affected both the total amount and relative composition of the soluble nitrogen in white spruce tissues (Durzan and Steward). Ammonium nitrogen was shown to foster arginine and amides and lead to a large increase of free guanidine compounds, whereas in leaves nourished by nitrate as

13566-403: The years 1955 through 1961, respectively. The total number of needles of all ages present on the 36-year-old tree in 1961 was 5.25 million weighing 14.25 kg. In 1961, needles as old as 13 years remained on the tree. The ash weight of needles increased progressively with age from about 4% in first-year needles in 1961 to about 8% in needles 10 years old. In discussing the data obtained from

13685-421: Was Hexham . At the 2001 UK Census Northumberland registered a population of 307,190, estimated to be 309,237 in 2003, The 2011 UK Census gave a population of 316,028. In 2001 there were 130,780 households, 10% of the population were retired, and one-third rented their homes. Northumberland has an ethnic minority population at 0.985% of the population, far lower compared to the average of 9.1% for England as

13804-513: Was first drawn up in the 1950s. The historic county town was Alnwick ; assizes were mainly held in Newcastle , with the county gaol in Morpeth . Newcastle became a city in 1400, with county corporate status, with both areas having joint assizes. From the county council 's formation in 1889 until 1981 Newcastle was the county town , being briefly the county town of two counties when

13923-488: Was killed playing football when David Le Keu's knife went into Henry's belly and killed him. Organised football teams as we know today did not appear until the 1870s. Newcastle United Football Club was formed in 1892 by uniting Newcastle West End FC with Newcastle East End. Newcastle United were first division champions three times in the early 20th century, reaching the FA Cup Final three times before winning it at

14042-598: Was produced as a blank page. Notable players after the Second World War included Joe Harvey , Jackie Milburn , Brian Clough and Newcastle's Bobby Moncur who led his team to win the Inter City Fairs Cup in 1969. Two of Jackie Milburn's nephews from Ashington , Bobby Charlton and Jackie Charlton are perhaps the two most significant players for England . Bobby joined Manchester United and Jackie Leeds United both contributing much to

14161-745: Was proposed in July 2016 as one of the preferred reintroduction sites for the Eurasian lynx , which has been extinct in Britain for 1,300 years. Interest in reintroducing the species was further bolstered in 2016 in relation to a successful breeding program for the Iberian lynx in Spain. Kielder Castle Visitor Centre is an 18th-century hunting lodge built by the Duke of Northumberland , which has been converted into

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