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110-411: The estate included a Spanish colonial house, a sculpture park, and a complete zoo that included many kinds of animals from different continents such as antelope , elephants , exotic birds, giraffes , hippopotamuses , ostriches , and ponies . The ranch also boasted a large collection of old and luxury cars and bikes, a private airport , a brothel , and even a Formula 1 -racing track. Mounted atop

220-537: A bear and a bull , chained together, rolled in fierce combat across the sand ... Four hundred bears were killed in a single day under Caligula ... Under Nero , four hundred tigers fought with bulls and elephants. In a single day, at the dedication of the Colosseum by Titus , five thousand animals perished. Under Trajan ... lions , tigers, elephants, rhinoceroses , hippopotami, giraffes, bulls, stags, even crocodiles and serpents were employed to give novelty to

330-399: A gastrula with a digestive chamber and two separate germ layers , an external ectoderm and an internal endoderm . In most cases, a third germ layer, the mesoderm , also develops between them. These germ layers then differentiate to form tissues and organs. Repeated instances of mating with a close relative during sexual reproduction generally leads to inbreeding depression within

440-518: A "house of animals" with a large collection of birds, mammals and reptiles in a garden tended by more than 600 employees. The garden was described by several Spanish conquerors, including Hernán Cortés in 1520. After the Aztec revolt against the Spanish rule, and during the subsequent battle for the city, Cortés reluctantly ordered the zoo to be destroyed. The oldest zoo in the world still in existence

550-400: A Congolese pygmy , displayed in a cage with the chimpanzees, then with an orangutan named Dohong, and a parrot. The exhibit was intended as an example of the "missing link" between the orangutan and white man. It triggered protests from the city's clergymen, but the public reportedly flocked to see Benga. Humans were also displayed at various events, especially colonial expositions such as

660-561: A clade Xenambulacraria for the Xenacoelamorpha + Ambulacraria; this is either within Deuterostomia, as sister to Chordata, or the Deuterostomia are recovered as paraphyletic, and Xenambulacraria is sister to the proposed clade Centroneuralia , consisting of Chordata + Protostomia. Eumetazoa , a clade which contains Ctenophora and ParaHoxozoa , has been proposed as a sister group to Porifera . A competing hypothesis

770-482: A consequence, various management tools are used to preserve the space for the genetically most important individuals and to reduce the risk of inbreeding . Management of animal populations is typically through international organizations such as AZA and EAZA . Zoos have several different ways of managing the animal populations, such as moves between zoos, contraception , sale of excess animals and euthanization (culling). Contraception can be an effective way to limit

880-610: A difference in wildlife conservation and education. Humans were occasionally displayed in cages at zoos along with non-human animals, to illustrate the differences between people of European and non-European origin. In September 1906, William Hornaday , director of the Bronx Zoo in New York—;with the agreement of Madison Grant , head of the New York Zoological Society —had Ota Benga ,

990-676: A gas station. The animals may be trained to perform tricks, and visitors are able to get closer to them than in larger zoos. Since they are sometimes less regulated, roadside zoos are often subject to accusations of neglect and cruelty . In June 2014 the Animal Legal Defense Fund filed a lawsuit against the Iowa-based roadside Cricket Hollow Zoo for violating the Endangered Species Act by failing to provide proper care for its animals. Since filing

1100-458: A greenhouse with an aviary, aquarium, and museum of stuffed animals and birds), a pond, a bridge over a waterfall, an artificial lake with a fountain, a wood-ornamented greenhouse, a forest area, and enclosures and buildings. The first zoological garden in Australia was Melbourne Zoo in 1860. In German states leading roles came Berlin (1841), Frankfurt (1856), and Hamburg (1863). In 1907,

1210-509: A lack of natural wild habitat in which to reintroduce animals. This highlights the importance of in situ conservation, or preservation of natural spaces, in addition to the utility of zoo captive breeding and reintroduction programs. In situ conservation and reintroduction programs are key elements to obtaining certification by reputable organisations such as the Association of Zoos and Aquariums (AZA). Efforts to clone endangered species in

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1320-461: A matter of public interest in the 1970s, a few zoos began to consider making conservation their central role, with Gerald Durrell of the Jersey Zoo , George Rabb of Brookfield Zoo , and William Conway of the Bronx Zoo ( Wildlife Conservation Society ) leading the discussion. From then on, zoo professionals became increasingly aware of the need to engage themselves in conservation programs, and

1430-448: A model of cooperation for conservation. Loaned animals usually remain the property of the original park, and any offspring yielded by loaned animals are usually divided between the lending and holding institutions. For decades the capture of wild animals or purchasing of animals has been broadly considered unethical and has not been practiced by reputable zoos. Especially in large animals, a limited number of spaces are available in zoos. As

1540-418: A new strategy that sets out the aims and mission of zoological gardens of the 21st century. When studying behaviour of captive animals, several things should however be taken into account before drawing conclusions about wild populations. Including that captive populations are often smaller than wild ones and that the space available to each animal is often less than in the wild. Conservation programs all over

1650-434: A place of refuge . Today, many zoos are improving enclosures by including tactile and sensory features in the habitat that allow animals to encourage natural behaviors. These additions can prove to be effective in improving the lives of animals in captivity. The tactile and sensory features will vary depending on the species of animal. There are animals that are injured in the wild and are unable to survive on their own, but in

1760-1109: A population due to the increased prevalence of harmful recessive traits. Animals have evolved numerous mechanisms for avoiding close inbreeding . Some animals are capable of asexual reproduction , which often results in a genetic clone of the parent. This may take place through fragmentation ; budding , such as in Hydra and other cnidarians ; or parthenogenesis , where fertile eggs are produced without mating , such as in aphids . Animals are categorised into ecological groups depending on their trophic levels and how they consume organic material . Such groupings include carnivores (further divided into subcategories such as piscivores , insectivores , ovivores , etc.), herbivores (subcategorized into folivores , graminivores , frugivores , granivores , nectarivores , algivores , etc.), omnivores , fungivores , scavengers / detritivores , and parasites . Interactions between animals of each biome form complex food webs within that ecosystem . In carnivorous or omnivorous species, predation

1870-481: A population's breeding. However it may also have health repercussions and can be difficult or even impossible to reverse in some animals. Additionally, some species may lose their reproductive capability entirely if prevented from breeding for a period (whether through contraceptives or isolation), but further study is needed on the subject. Sale of surplus animals from zoos was once common and in some cases animals have ended up in substandard facilities. In recent decades

1980-502: A rare goat , one ostrich, and zebras . Escobar's hippopotamuses have escaped and become feral , living in at least four lakes in the area and spreading into neighbouring rivers. Contact between the hippos and local fishermen led to calls for the hippo population to be culled. By 2011, there were at least 30 animals roaming wild in the countryside; the large number of hippos makes it difficult to find zoos into which they can be resettled. There are also reportedly 40 hippopotamuses living on

2090-628: A small number of zoos. To solve this organizations like EAZA and AZA have begun to develop husbandry manuals. Many modern zoos attempt to improve animal welfare by providing more space and behavioural enrichments . This often involves housing the animals in naturalistic enclosures that allow the animals to express more of their natural behaviours, such as roaming and foraging. Whilst many zoos have been working hard on this change, in some zoos, some enclosures still remain barren concrete enclosures or other minimally enriched cages. Animal Animals are multicellular , eukaryotic organisms in

2200-547: A suitable alternative. Off-site conservation relies on zoos, national parks, or other care facilities to support the rehabilitation of the animals and their populations. Zoos benefit conservation by providing suitable habitats and care to endangered animals. When properly regulated, they present a safe, clean environment for the animals to increase populations sizes. A study on amphibian conservation and zoos addressed these problems by writing, Whilst addressing in situ threats, particularly habitat loss, degradation and fragmentation,

2310-553: A suitable environment for wild native animals such as herons to live in or visit. A colony of black-crowned night herons has regularly summered at the National Zoo in Washington, D.C. for more than a century. Some zoos may provide information to visitors on wild animals visiting or living in the zoo, or encourage them by directing them to specific feeding or breeding platforms. In modern, well-regulated zoos, breeding

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2420-589: A year and have special programs for schools. They are organized by the Association of Zoos and Aquariums . Japan's first modern zoo, Tokyo's Ueno Imperial Zoological Gardens , opened in 1882 based on European models. In World War II it was used to teach the Japanese people about the lands recently conquered by the Army. In 1943, fearing American bombing attacks, the government ordered the zoo to euthanize dangerous animals that might escape. When ecology emerged as

2530-992: Is "the advancement of Zoology and Animal Physiology and the introduction of new and curious subjects of the Animal Kingdom." It maintains two research institutes, the Nuffield Institute of Comparative Medicine and the Wellcome Institute of Comparative Physiology. In the United States, the Penrose Research Laboratory of the Philadelphia Zoo focuses on the study of comparative pathology . The World Association of Zoos and Aquariums produced its first conservation strategy in 1993, and in November 2004, it adopted

2640-464: Is a consumer–resource interaction where the predator feeds on another organism, its prey , who often evolves anti-predator adaptations to avoid being fed upon. Selective pressures imposed on one another lead to an evolutionary arms race between predator and prey, resulting in various antagonistic/ competitive coevolutions . Almost all multicellular predators are animals. Some consumers use multiple methods; for example, in parasitoid wasps ,

2750-433: Is a stage in embryonic development that is unique to animals, allowing cells to be differentiated into specialised tissues and organs. All animals are composed of cells, surrounded by a characteristic extracellular matrix composed of collagen and elastic glycoproteins . During development, the animal extracellular matrix forms a relatively flexible framework upon which cells can move about and be reorganised, making

2860-414: Is controlled to maintain a self-sustaining, global captive population. This is not the case in some less well-regulated zoos, often based in poorer regions. Overall "stock turnover" of animals during a year in a select group of poor zoos was reported as 20%-25% with 75% of wild caught apes dying in captivity within the first 20 months. The authors of the report stated that before successful breeding programs,

2970-463: Is disputed, as they might be water-escape or other structures. Animals are monophyletic , meaning they are derived from a common ancestor. Animals are the sister group to the choanoflagellates , with which they form the Choanozoa . The dates on the phylogenetic tree indicate approximately how many millions of years ago ( mya ) the lineages split. Ros-Rocher and colleagues (2021) trace

3080-450: Is known as zoology , and the study of animal behaviour is known as ethology . The animal kingdom is divided into five infrakingdoms / superphyla , namely Porifera , Ctenophora , Placozoa , Cnidaria and Bilateria . Most living animal species belong to the infrakingdom Bilateria, a highly proliferative clade whose members have a bilaterally symmetric body plan , and the vast majority of bilaterians belong to two large superphyla :

3190-462: Is of primary importance; for many amphibian species in situ conservation alone will not be enough, especially in light of current un-mitigatable threats that can impact populations very rapidly such as chytridiomycosis [an infectious fungal disease]. Ex situ programmes can complement in situ activities in a number of ways including maintaining genetically and demographically viable populations while threats are either better understood or mitigated in

3300-808: Is often used to refer only to nonhuman animals. The term metazoa is derived from Ancient Greek μετα ( meta ) 'after' (in biology, the prefix meta- stands for 'later') and ζῷᾰ ( zōia ) 'animals', plural of ζῷον zōion 'animal'. Animals have several characteristics that set them apart from other living things. Animals are eukaryotic and multicellular . Unlike plants and algae , which produce their own nutrients , animals are heterotrophic , feeding on organic material and digesting it internally. With very few exceptions, animals respire aerobically . All animals are motile (able to spontaneously move their bodies) during at least part of their life cycle , but some animals, such as sponges , corals , mussels , and barnacles , later become sessile . The blastula

3410-422: Is only produced by sponges and pelagophyte algae. Its likely origin is from sponges based on molecular clock estimates for the origin of 24-ipc production in both groups. Analyses of pelagophyte algae consistently recover a Phanerozoic origin, while analyses of sponges recover a Neoproterozoic origin, consistent with the appearance of 24-ipc in the fossil record. The first body fossils of animals appear in

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3520-547: Is the Benthozoa clade, which would consist of Porifera and ParaHoxozoa as a sister group of Ctenophora . Several animal phyla lack bilateral symmetry. These are the Porifera (sea sponges), Placozoa , Cnidaria (which includes jellyfish , sea anemones , and corals), and Ctenophora (comb jellies). Sponges are physically very distinct from other animals, and were long thought to have diverged first, representing

3630-584: Is the Tiergarten Schönbrunn in Vienna , Austria. It was constructed by Adrian van Stekhoven in 1752 at the order of Emperor Francis I , to serve as an imperial menagerie as part of Schönbrunn Palace . The menagerie was initially reserved for the viewing pleasure of the imperial family and the court, but was made accessible to the public in 1765. In 1775, a zoo was founded in Madrid, and in 1795,

3740-501: Is the menagerie , which has a long history from the ancient world to modern times. The oldest known zoological collection was revealed during excavations at Hierakonpolis , Egypt in 2009, of a c.  3500 BCE menagerie. The exotic animals included hippopotami , hartebeest , elephants , baboons and wildcats . King Ashur-bel-kala of the Middle Assyrian Empire created zoological and botanical gardens in

3850-690: The Africarium , the only themed oceanarium devoted solely to exhibiting the fauna of Africa , comprehensively presenting selected ecosystems from the continent of Africa . Housing over 10 thousand animals, the facility's breadth extends from housing insects such cockroaches to large mammals like elephants on an area of over 33 hectares . In the United States, the Philadelphia Zoo , opened on July 1, 1874, earning its motto "America's First Zoo." The Lincoln Park Zoological Gardens in Chicago and

3960-541: The American Zoo Association soon said that conservation was its highest priority. In order to stress conservation issues, many large zoos stopped the practice of having animals perform tricks for visitors. The Detroit Zoo , for example, stopped its elephant show in 1969, and its chimpanzee show in 1983, acknowledging that the trainers had probably abused the animals to get them to perform. Mass destruction of wildlife habitat has yet to cease all over

4070-622: The Cambrian explosion , which began around 539  million years ago (Mya), and most classes during the Ordovician radiation 485.4 Mya. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived about 650 Mya during the Cryogenian period. Historically, Aristotle divided animals into those with blood and those without . Carl Linnaeus created

4180-619: The Ediacaran , represented by forms such as Charnia and Spriggina . It had long been doubted whether these fossils truly represented animals, but the discovery of the animal lipid cholesterol in fossils of Dickinsonia establishes their nature. Animals are thought to have originated under low-oxygen conditions, suggesting that they were capable of living entirely by anaerobic respiration , but as they became specialized for aerobic metabolism they became fully dependent on oxygen in their environments. Many animal phyla first appear in

4290-485: The Wnt and TGF-beta signalling pathways which may have enabled animals to become multicellular by providing a pattern for the body's system of axes (in three dimensions), and another 7 are for transcription factors including homeodomain proteins involved in the control of development . Giribet and Edgecombe (2020) provide what they consider to be a consensus internal phylogeny of the animals, embodying uncertainty about

4400-1025: The Zoological Society of London in 1931 which today (2014) covers 600 acres (2.4 km ). Since the early 1970s, an 1,800 acre (7 km ) park in the San Pasqual Valley near San Diego has featured the San Diego Zoo Safari Park , run by the Zoological Society of San Diego. One of two state-supported zoo parks in North Carolina is the 2,000-acre (8.1 km ) North Carolina Zoo in Asheboro. The 500-acre (2.0 km ) Werribee Open Range Zoo in Melbourne, Australia, displays animals living in an artificial savannah . The first public aquarium

4510-893: The biological kingdom Animalia ( / ˌ æ n ɪ ˈ m eɪ l i ə / ). With few exceptions, animals consume organic material , breathe oxygen , have myocytes and are able to move , can reproduce sexually , and grow from a hollow sphere of cells, the blastula , during embryonic development . Animals form a clade , meaning that they arose from a single common ancestor . Over 1.5 million living animal species have been described , of which around 1.05 million are insects , over 85,000 are molluscs , and around 65,000 are vertebrates . It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from 8.5 μm (0.00033 in) to 33.6 m (110 ft). They have complex ecologies and interactions with each other and their environments, forming intricate food webs . The scientific study of animals

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4620-665: The evolutionary relationships between taxa . Humans make use of many other animal species for food (including meat , eggs , and dairy products ), for materials (such as leather , fur , and wool ), as pets and as working animals for transportation , and services . Dogs , the first domesticated animal, have been used in hunting , in security and in warfare , as have horses , pigeons and birds of prey ; while other terrestrial and aquatic animals are hunted for sports, trophies or profits. Non-human animals are also an important cultural element of human evolution , having appeared in cave arts and totems since

4730-473: The fossil record during the Cambrian explosion , starting about 539 million years ago, in beds such as the Burgess shale . Extant phyla in these rocks include molluscs , brachiopods , onychophorans , tardigrades , arthropods , echinoderms and hemichordates , along with numerous now-extinct forms such as the predatory Anomalocaris . The apparent suddenness of the event may however be an artifact of

4840-497: The lobe-finned fish Tiktaalik started to move on to land in the late Devonian , about 375 million years ago. Animals occupy virtually all of earth's habitats and microhabitats, with faunas adapted to salt water, hydrothermal vents, fresh water, hot springs, swamps, forests, pastures, deserts, air, and the interiors of other organisms. Animals are however not particularly heat tolerant ; very few of them can survive at constant temperatures above 50 °C (122 °F) or in

4950-500: The protostomes , which includes organisms such as arthropods , molluscs , flatworms , annelids and nematodes ; and the deuterostomes , which include echinoderms , hemichordates and chordates , the latter of which contains the vertebrates . The much smaller basal phylum Xenacoelomorpha have an uncertain position within Bilateria. Animals first appear in the fossil record in the late Cryogenian period and diversified in

5060-593: The 11th century BCE. In the 2nd century BCE, the Chinese Empress Tanki had a "house of deer" built, and King Wen of Zhou kept a 1,500-acre (6.1 km ) zoo called Ling-Yu, or the Garden of Intelligence. Other well-known collectors of animals included King Solomon of the Kingdom of Israel and Judah , Queen Semiramis and King Ashurbanipal of Assyria , and King Nebuchadnezzar of Babylonia . By

5170-517: The 1931 Paris Colonial Exposition , with the practice continuing in Belgium at least to as late as 1958 in a "Congolese village" display at Expo '58 in Brussels. These displays, while sometimes called "human zoos", usually did not take place in zoos or use cages. Zoo animals live in enclosures that often attempt to replicate their natural habitats or behavioral patterns, for the benefit of both

5280-571: The 4th century BCE, zoos existed in most of the Greek city states; Alexander the Great is known to have sent animals that he found on his military expeditions back to Greece. The Roman emperors kept private collections of animals for study or for use in the arena, the latter faring notoriously poorly. The 19th-century historian W. E. H. Lecky wrote of the Roman games , first held in 366 BCE: At one time,

5390-715: The Cambrian explosion, possibly as early as 1 billion years ago. Early fossils that might represent animals appear for example in the 665-million-year-old rocks of the Trezona Formation of South Australia . These fossils are interpreted as most probably being early sponges . Trace fossils such as tracks and burrows found in the Tonian period (from 1 gya) may indicate the presence of triploblastic worm-like animals, roughly as large (about 5 mm wide) and complex as earthworms. However, similar tracks are produced by

5500-552: The Cincinnati Zoo opened in 1875. In the 1930s, federal relief programs provided financial aid to most local zoos. The Works Progress Administration and similar New Deal government agencies helped greatly in the construction, renovation, and expansion of zoos when the Great Depression severely reduced local budgets. It was "a new deal for animals." The Atlanta Zoo , founded in 1886, suffered neglect. By 1984 it

5610-717: The Cnidaria) never grow larger than 20 μm , and one of the smallest species ( Myxobolus shekel ) is no more than 8.5 μm when fully grown. The following table lists estimated numbers of described extant species for the major animal phyla, along with their principal habitats (terrestrial, fresh water, and marine), and free-living or parasitic ways of life. Species estimates shown here are based on numbers described scientifically; much larger estimates have been calculated based on various means of prediction, and these can vary wildly. For instance, around 25,000–27,000 species of nematodes have been described, while published estimates of

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5720-1025: The U.S. and Canada by Species Survival Plans; in Australasia, by the Australasian Species Management Program; in Europe, by the European Endangered Species Program; and in Japan, South Asia, and South East Asia, by the Japanese Association of Zoos and Aquariums, the South Asian Zoo Association for Regional Cooperation , and the South East Asian Zoo Association. Besides conservation of captive species, large zoos may form

5830-550: The United States, Europe, and Asia are frequently embedded in zoos and zoological parks. The position of most modern zoos in Australasia , Asia , Europe , and North America , particularly those with scientific societies, is that they display wild animals primarily for the conservation of endangered species , as well as for research purposes and education, and secondarily for the entertainment of visitors. The Zoological Society of London states in its charter that its aim

5940-649: The Zoo " by music-hall artist Alfred Vance . The term "zoological park" was used for more expansive facilities in Halifax, Nova Scotia , Washington, D.C. , and the Bronx in New York, which opened in 1846, 1891 and 1899 respectively. Relatively new terms for zoos in the late 20th century are " conservation park" or "bio park". Adopting a new name is a strategy used by some zoo professionals to distance their institutions from

6050-737: The animals and visitors. Nocturnal animals are often housed in buildings with a reversed light-dark cycle, i.e. only dim white or red lights are on during the day so the animals are active during visitor hours, and brighter lights on at night when the animals sleep. Special climate conditions may be created for animals living in extreme environments, such as penguins. Special enclosures for birds , mammals , insects , reptiles , fish , and other aquatic life forms have also been developed. Some zoos have walk-through exhibits where visitors enter enclosures of non-aggressive species, such as lemurs , marmosets , birds, lizards , and turtles . Visitors are asked to keep to paths and avoid showing or eating foods that

6160-493: The animals might snatch. Some zoos keep animals in larger, outdoor enclosures, confining them with moats and fences, rather than in cages. Safari parks , also known as zoo parks and lion farms, allow visitors to drive through them and come in close proximity to the animals. Sometimes, visitors are able to feed animals through the car windows. The first safari park was Whipsnade Park in Bedfordshire, England, opened by

6270-541: The animals was too expensive for the government, so it was decided that most of the animals would be donated to various South American zoos. Other original features include dinosaur statues as well as decommissioned military vehicles and a giant hand sculpture. By November 2006, ownership of the property had passed to the Colombian government and was valued at 5 billion Colombian pesos (approximately $ 2.23 million). The hacienda's zoo as of February 2019 hosts bison ,

6380-671: The animals were moved to the Bulwark, renamed the Lion Tower, near the main western entrance of the Tower. It was opened to the public during the reign of Elizabeth I in the 16th century. During the 18th century, the price of admission was three half-pence, or the supply of a cat or dog for feeding to the lions. The animals were moved to the London Zoo when it opened. Aztec emperor Moctezuma had in his capital city of Tenochtitlan

6490-434: The animals' needs, but constraints such as size and expense can complicate this. The type of enclosure and the husbandry are of great importance in determining the welfare of animals. Substandard enclosures can lead to decreased lifespans, caused by factors as human diseases, unsafe materials in the cages and possible escape attempts (Bendow 382). However, when zoos take time to think about the animal's welfare, zoos can become

6600-589: The caves in Colombia's Cueva de los Guácharos National Park . In December 2018, a day ticket to the park cost 42,000 pesos (around $ 15). The Escobar museum, his burned private car collection, and the abandoned "ruins" of his house are still publicly accessible, but are reported to have collapsed in February 2015. Escobar kept four hippos in a private menagerie at Hacienda Nápoles. They were deemed too difficult to seize and move after Escobar's death, and hence left on

6710-654: The classical zoo, such as stage shows, roller coasters, and mythical creatures. Some examples are Busch Gardens Tampa Bay in Tampa, Florida , both Disney's Animal Kingdom and Gatorland in Orlando, Florida , Flamingo Land in North Yorkshire, England , and Six Flags Discovery Kingdom in Vallejo, California . By 2000 most animals being displayed in zoos were the offspring of other zoo animals. This trend, however

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6820-525: The culling proceeded. Although zoos in some countries have been open about culling, the controversy of the subject and pressure from the public has resulted in others being closed. This stands in contrast to most zoos publicly announcing animal births. Furthermore, while many zoos are willing to cull smaller and/or low-profile animals, fewer are willing to do it with larger high-profile species. Many animals breed readily in captivity. Zoos frequently are forced to intentionally limit captive breeding because of

6930-463: The dark sea floor consume organic matter produced through chemosynthesis (via oxidizing inorganic compounds such as hydrogen sulfide ) by archaea and bacteria . Animals evolved in the sea. Lineages of arthropods colonised land around the same time as land plants , probably between 510 and 471 million years ago during the Late Cambrian or Early Ordovician . Vertebrates such as

7040-512: The different organizing principle of geography, as opposed to taxonomy. The Wrocław Zoo ( Polish : Ogród Zoologiczny we Wrocławiu ) is the oldest zoo in Poland, opened in 1865 when the city was part of Prussia , and was home to about 10,500 animals representing about 1,132 species (in terms of the number of animal species, it is the third largest in the world ). In 2014 the Wrocław Zoo opened

7150-454: The earliest times, and are frequently featured in mythology , religion , arts , literature , heraldry , politics , and sports . The word animal comes from the Latin noun animal of the same meaning, which is itself derived from Latin animalis 'having breath or soul'. The biological definition includes all members of the kingdom Animalia. In colloquial usage, the term animal

7260-568: The enclosures and come into close proximity with the animals. The Zoological Society of London was founded in 1826 by Stamford Raffles and established the London Zoo in Regent's Park two years later in 1828. At its founding, it was the world's first scientific zoo. Originally intended to be used as a collection for scientific study , it was opened to the public in 1847. The Zoo was located in Regent's Park —then undergoing development at

7370-537: The entrepreneur Carl Hagenbeck founded the Tierpark Hagenbeck in Stellingen, now a quarter of Hamburg . His zoo was a radical departure from the layout of the zoo that had been established in 1828. It was the first zoo to use open enclosures surrounded by moats, rather than barred cages, to better approximate animals' natural environments. He also set up mixed-species exhibits and based the layout on

7480-570: The first hierarchical biological classification for animals in 1758 with his Systema Naturae , which Jean-Baptiste Lamarck expanded into 14 phyla by 1809. In 1874, Ernst Haeckel divided the animal kingdom into the multicellular Metazoa (now synonymous with Animalia) and the Protozoa , single-celled organisms no longer considered animals. In modern times, the biological classification of animals relies on advanced techniques, such as molecular phylogenetics , which are effective at demonstrating

7590-458: The food is supplied by the zoo, either from vending machines or a kiosk nearby. An animal theme park is a combination of an amusement park and a zoo, mainly for entertaining and commercial purposes. Marine mammal parks such as Sea World and Marineland are more elaborate dolphinariums keeping whales , and containing additional entertainment attractions. Another kind of animal theme park contains more entertainment and amusement elements than

7700-448: The formation of complex structures possible. This may be calcified, forming structures such as shells , bones , and spicules . In contrast, the cells of other multicellular organisms (primarily algae, plants, and fungi ) are held in place by cell walls, and so develop by progressive growth. Animal cells uniquely possess the cell junctions called tight junctions , gap junctions , and desmosomes . With few exceptions—in particular,

7810-520: The fossil record, rather than showing that all these animals appeared simultaneously. That view is supported by the discovery of Auroralumina attenboroughii , the earliest known Ediacaran crown-group cnidarian (557–562 mya, some 20 million years before the Cambrian explosion) from Charnwood Forest , England. It is thought to be one of the earliest predators , catching small prey with its nematocysts as modern cnidarians do. Some palaeontologists have suggested that animals appeared much earlier than

7920-680: The giant single-celled protist Gromia sphaerica , so the Tonian trace fossils may not indicate early animal evolution. Around the same time, the layered mats of microorganisms called stromatolites decreased in diversity, perhaps due to grazing by newly evolved animals. Objects such as sediment-filled tubes that resemble trace fossils of the burrows of wormlike animals have been found in 1.2 gya rocks in North America, in 1.5 gya rocks in Australia and North America, and in 1.7 gya rocks in Australia. Their interpretation as having an animal origin

8030-408: The grounds of the hacienda itself; as of June 2014, the park's mascot, a live female hippo named Vanessa (who responds to her name), remains at the site. In 2014, a "Jurassic Park"-style African theme park was operating on the grounds, which have been rented by a private company. "Parque Temático Hacienda Nápoles" comes complete with a water park, a guided safari attraction, aquariums, and a replica of

8140-599: The hacienda's entrance gate is a replica of the Piper PA-18 Super Cub airplane ( tail number HK-617-P). After Escobar was shot and killed by Colombian police in 1993, his family entered a legal struggle with the Colombian government over the property. The government prevailed, and the neglected property is now managed by the Municipality of Puerto Triunfo . The cost of maintenance for the zoo and

8250-463: The hands of the architect John Nash . What set the London zoo apart from its predecessors was its focus on society at large. The zoo was established in the middle of a city for the public, and its layout was designed to cater for the large London population. The London zoo was widely copied as the archetype of the public city zoo. In 1853, the Zoo opened the world's first public aquarium . Dublin Zoo

8360-410: The high mortality rate was the reason for the "massive scale of importations." One 2-year study indicated that of 19,361 mammals that left accredited zoos in the U.S. between 1992 and 1998, 7,420 (38%) went to dealers, auctions, hunting ranches, unaccredited zoos and individuals, and game farms. The welfare of zoo animals varies widely. Many zoos work to improve their animal enclosures and make it fit

8470-492: The hippos which found disagreement among environmentalists on whether they were having a positive or negative impact, but that conservationists and locals — particularly those in the tourism industry — were mostly in support of their continued presence. In January 2021, scientists proposed to euthanize the now approximately 100 hippos, as they have dispersed over the Magdalena river basin. Some scientists have suggested castrating

8580-560: The hippos' uncontrolled reproduction poses a threat to humans and native wildlife and estimates suggest that the population could reach 1,000 by 2035. 5°55′33.60″N 74°43′18.57″W  /  5.9260000°N 74.7218250°W  / 5.9260000; -74.7218250 Zoo A zoo (short for zoological garden ; also called an animal park or menagerie ) is a facility in which animals are kept within enclosures for public exhibition and often bred for conservation purposes. The term zoological garden refers to zoology ,

8690-424: The importance of animal conservation, often through letting visitors witness the animals firsthand. Some critics, and the majority of animal rights activists, say that zoos, no matter their intentions, or how noble these intentions, are immoral and serve as nothing but to fulfill human leisure at the expense of the animals (an opinion that has spread over the years). However, zoo advocates argue that their efforts make

8800-441: The larvae feed on the hosts' living tissues, killing them in the process, but the adults primarily consume nectar from flowers. Other animals may have very specific feeding behaviours , such as hawksbill sea turtles which mainly eat sponges . Most animals rely on biomass and bioenergy produced by plants and phytoplanktons (collectively called producers ) through photosynthesis . Herbivores, as primary consumers , eat

8910-520: The lawsuit, ALDF has obtained records from investigations conducted by the USDA's Animal and Plant Health Inspection Services; these records show that the zoo is also violating the Animal Welfare Act. A petting zoo, also called petting farms or children's zoos, features a combination of domestic animals and wild species that are docile enough to touch and feed. To ensure the animals' health,

9020-502: The male hippos to prevent further breeding. In February 2022, the Colombian government announced that hippos will be declared an invasive species . The hippo was officially added to the list of invasive alien species on March 25 2022. On November 3 2023, Environment Minister of Colombia, Susana Muhamad , said that, of the 166 hippos descended from Escobar's herd, 20 would be sterilised, others would be transferred abroad - and "some" would be euthanised. Colombian experts have long warned that

9130-753: The most extreme cold deserts of continental Antarctica . The blue whale ( Balaenoptera musculus ) is the largest animal that has ever lived, weighing up to 190 tonnes and measuring up to 33.6 metres (110 ft) long. The largest extant terrestrial animal is the African bush elephant ( Loxodonta africana ), weighing up to 12.25 tonnes and measuring up to 10.67 metres (35.0 ft) long. The largest terrestrial animals that ever lived were titanosaur sauropod dinosaurs such as Argentinosaurus , which may have weighed as much as 73 tonnes, and Supersaurus which may have reached 39 meters. Several animals are microscopic; some Myxozoa ( obligate parasites within

9240-620: The oldest animal phylum and forming a sister clade to all other animals. Despite their morphological dissimilarity with all other animals, genetic evidence suggests sponges may be more closely related to other animals than the comb jellies are. Sponges lack the complex organization found in most other animal phyla; their cells are differentiated, but in most cases not organised into distinct tissues, unlike all other animals. They typically feed by drawing in water through pores, filtering out small particles of food. The comb jellies and Cnidaria are radially symmetric and have digestive chambers with

9350-525: The origins of animals to unicellular ancestors, providing the external phylogeny shown in the cladogram. Uncertainty of relationships is indicated with dashed lines. Holomycota (inc. fungi) [REDACTED] Ichthyosporea [REDACTED] Pluriformea [REDACTED] Filasterea [REDACTED] [REDACTED] [REDACTED] The most basal animals, the Porifera , Ctenophora , Cnidaria , and Placozoa , have body plans that lack bilateral symmetry . Their relationships are still disputed;

9460-519: The plant material directly to digest and absorb the nutrients, while carnivores and other animals on higher trophic levels indirectly acquire the nutrients by eating the herbivores or other animals that have eaten the herbivores. Animals oxidize carbohydrates , lipids , proteins and other biomolecules, which allows the animal to grow and to sustain basal metabolism and fuel other biological processes such as locomotion . Some benthic animals living close to hydrothermal vents and cold seeps on

9570-496: The practice of selling animals from certified zoos has declined. A large number of animals are culled each year in zoos, but this is controversial. A highly publicized culling as part of population management was that of a healthy giraffe at Copenhagen Zoo in 2014. The zoo argued that his genes already were well-represented in captivity, making the giraffe unsuitable for future breeding. There were offers to adopt him and an online petition to save him had many thousand signatories, but

9680-753: The sister group to all other animals could be the Porifera or the Ctenophora, both of which lack hox genes , which are important for body plan development . Hox genes are found in the Placozoa, Cnidaria, and Bilateria. 6,331 groups of genes common to all living animals have been identified; these may have arisen from a single common ancestor that lived 650 million years ago in the Precambrian . 25 of these are novel core gene groups, found only in animals; of those, 8 are for essential components of

9790-407: The smaller, motile gametes are spermatozoa and the larger, non-motile gametes are ova . These fuse to form zygotes , which develop via mitosis into a hollow sphere, called a blastula. In sponges, blastula larvae swim to a new location, attach to the seabed, and develop into a new sponge. In most other groups, the blastula undergoes more complicated rearrangement. It first invaginates to form

9900-654: The spectacle. Charlemagne had an elephant named Abul-Abbas that was given to him by the Abbasid caliph. King Henry I of England kept a collection of animals at his palace in Woodstock which reportedly included lions, leopards, and camels. The most prominent collection in medieval England was in the Tower of London , created as early as 1204 by King John I . Henry III received a wedding gift in 1235 of three leopards from Frederick II, Holy Roman Emperor , and in 1264,

10010-474: The sponges and placozoans —animal bodies are differentiated into tissues . These include muscles , which enable locomotion, and nerve tissues , which transmit signals and coordinate the body. Typically, there is also an internal digestive chamber with either one opening (in Ctenophora, Cnidaria, and flatworms) or two openings (in most bilaterians). Nearly all animals make use of some form of sexual reproduction. They produce haploid gametes by meiosis ;

10120-553: The stereotypical and nowadays criticized zoo concept of the 19th century. The term "bio park" was first coined and developed by the National Zoo in Washington D.C. in the late 1980s. In 1993, the New York Zoological Society changed its name to the Wildlife Conservation Society and re branded the zoos under its jurisdiction as "wildlife conservation parks". The predecessor of the zoological garden

10230-411: The structure at the base of the tree (dashed lines). Porifera [REDACTED] Ctenophora [REDACTED] Placozoa [REDACTED] Cnidaria [REDACTED] Xenacoelomorpha [REDACTED] Ambulacraria [REDACTED] Chordata [REDACTED] Ecdysozoa [REDACTED] Spiralia [REDACTED] An alternative phylogeny, from Kapli and colleagues (2021), proposes

10340-748: The study of animals. The term is derived from the Ancient Greek ζῷον , zōion , 'animal', and the suffix -λογία , -logia , 'study of'. The abbreviation zoo was first used of the London Zoological Gardens , which was opened for scientific study in 1828, and to the public in 1847. The first modern zoo was the Tierpark Hagenbeck by Carl Hagenbeck in Germany . In the United States alone, zoos are visited by over 181 million people annually. The London Zoo , which

10450-522: The subsequent Ediacaran period in what is known as the Avalon explosion . Earlier evidence of animals is still controversial; the sponge -like organism Otavia has been dated back to the Tonian period at the start of the Neoproterozoic , but its identity as an animal is heavily contested. Nearly all modern animal phyla became clearly established in the fossil record as marine species during

10560-467: The total number of nematode species include 10,000–20,000; 500,000; 10 million; and 100 million. Using patterns within the taxonomic hierarchy, the total number of animal species—including those not yet described—was calculated to be about 7.77 million in 2011. 3,000–6,500 4,000–25,000 Evidence of animals is found as long ago as the Cryogenian period. 24-Isopropylcholestane (24-ipc) has been found in rocks from roughly 650 million years ago; it

10670-463: The tremendous expansion in the urbanization of London, led to a heightened demand for a greater variety of public forms of entertainment to be made available. The need for public entertainment, as well as the requirements of scholarly research, came together in the founding of the first modern zoos. Whipsnade Park Zoo in Bedfordshire , England, opened in 1931. It allowed visitors to drive through

10780-474: The untended estate. By 2007, the animals had multiplied to 16 and had taken to roaming the area for food in the nearby Magdalena River . The National Geographic Channel produced a documentary about them titled Cocaine Hippos . A report published in a Yale student magazine noted that local environmentalists are campaigning to protect the animals, although there is no clear plan for what will happen to them. In 2018, National Geographic published another article on

10890-513: The wild The breeding of endangered species is coordinated by cooperative breeding programmes containing international studbooks and coordinators, who evaluate the roles of individual animals and institutions from a global or regional perspective, and there are regional programmes all over the world for the conservation of endangered species . In Africa, conservation is handled by the African Preservation Program (APP); in

11000-403: The world and many species such as elephants , big cats, penguins , tropical birds, primates, rhinos , exotic reptiles, and many others are in danger of dying out. Many of today's zoos hope to stop or slow the decline of many endangered species and see their primary purpose as breeding endangered species in captivity and reintroducing them into the wild. Modern zoos also aim to help teach visitors

11110-516: The world fight to protect species from going extinct , but many conservation programs are underfunded and under-represented. Conservation programs can struggle to fight bigger issues like habitat loss and illness. It often takes significant funding and long time periods to rebuild degraded habitats, both of which are scarce in conservation efforts. The current state of conservation programs cannot rely solely in situ (on-site conservation) plans alone, ex situ (off-site conservation) may therefore provide

11220-556: The zoo inside the Jardin des Plantes in Paris was founded by Jacques-Henri Bernardin , with animals from the royal menagerie at Versailles, primarily for scientific research and education. The planning about a space for the conservation and observation of animals was expressed in connection with the political construction of republican citizenship. The Kazan Zoo , the first zoo in Russia

11330-432: The zoos they can live out the rest of their lives healthy and happy (McGaffin). In recent years, some zoos have chosen to move out some larger animals because they do not have the space available to provide an adequate enclosure for them (Lemonic, McDowell, and Bjerklie 50). An issue with animal welfare in zoos is that best animal husbandry practices are often not completely known, especially for species that are only kept in

11440-571: Was and still is somewhat species-specific. When animals are transferred between zoos, they usually spend time in quarantine, and are given time to acclimatize to their new enclosures which are often designed to mimic their natural environment. For example, some species of penguins may require refrigerated enclosures. Guidelines on necessary care for such animals is published in the International Zoo Yearbook . Animal exchanges between facilities are usually made voluntarily, based on

11550-530: Was founded in 1806 by the Professor of Kazan State University Karl Fuchs . Until the early 19th century, the function of the zoo was often to symbolize royal power, like King Louis XIV 's menagerie at Versailles . Major cities in Europe set up zoos in the 19th century, usually using London and Paris as models. The transition was made from princely menageries designed to entertain high society with strange novelties into public zoological gardens. The new goal

11660-609: Was opened at the London Zoo in 1853. This was followed by the opening of public aquaria in continental Europe (e.g. Paris in 1859, Hamburg in 1864, Berlin in 1869, and Brighton in 1872) and the United States (e.g. Boston in 1859, Washington in 1873, San Francisco Woodward's Garden in 1873, and the New York Aquarium at Battery Park in 1896). Roadside zoos are found throughout North America , particularly in remote locations. They are often small, for-profit zoos, often intended to attract visitors to some other facility, such as

11770-531: Was opened in 1828, was initially known as the "Gardens and Menagerie of the Zoological Society of London", and it described itself as a menagerie or "zoological forest". The abbreviation "zoo" first appeared in print in the United Kingdom around 1847, when it was used for the Clifton Zoo , but it was not until some 20 years later that the shortened form became popular in the rhyming song " Walking in

11880-583: Was opened in 1831 by members of the medical profession interested in studying animals while they were alive and more particularly getting hold of them when they were dead. Downs' Zoological Gardens created by Andrew Downs and opened to the Nova Scotia public in 1847. It was originally intended to be used as a collection for scientific study. By the early 1860s, the zoo grounds covered 40 hectares with many fine flowers and ornamental trees, picnic areas, statues, walking paths, The Glass House (which contained

11990-439: Was ranked among the ten worst zoos in the United States. Systematic reform by 2000 put it on the list of the ten best. By 2020, the United States featured 230 accredited zoos and aquariums across 45 states, accommodating 800,000 animals, and 6,000 species out of which about 1,000 are endangered. The zoos provide 208,000 jobs, and with an annual budget of $ 230 million for wildlife conservation . They attract over 200 million visits

12100-485: Was to educate the entire population with information along modern scientific lines. Zoos were supported by local commercial or scientific societies. The modern zoo that emerged in the 19th century in the United Kingdom , was focused on providing scientific study and later educational exhibits to the public for entertainment and inspiration. A growing fascination for natural history and zoology , coupled with

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