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182-573: The Holocene correlates with the last maximum axial tilt of the Earth towards the Sun, and corresponds with the rapid proliferation, growth, and impacts of the human species worldwide, including all of its written history , technological revolutions , development of major civilizations , and overall significant transition towards urban living in the present. The human impact on modern-era Earth and its ecosystems may be considered of global significance for

364-471: A jigsaw puzzle . Rocks normally form relatively horizontal layers, with each layer younger than the one underneath it. If a fossil is found between two layers whose ages are known, the fossil's age must lie between the two known ages. Because rock sequences are not continuous, but may be broken up by faults or periods of erosion , it is very difficult to match up rock beds that are not directly next to one another. However, fossils of species that survived for

546-415: A multi-layered network of cooperating, distinct, or even competing social groups – from families and peer groups to corporations and political states . As such, social interactions between humans have established a wide variety of values, social norms , languages , and traditions (collectively termed institutions ), each of which bolsters human society . Humans are also highly curious , with

728-668: A theory of mind . The human mind is capable of introspection , private thought , imagination , volition , and forming views on existence . This has allowed great technological advancements and complex tool development through complex reasoning and the transmission of knowledge to subsequent generations through language . Humans have had a dramatic effect on the environment . They are apex predators , being rarely preyed upon by other species. Human population growth , industrialization, land development, overconsumption and combustion of fossil fuels have led to environmental destruction and pollution that significantly contributes to

910-552: A collision that formed the Moon about 40 million years later, may have cooled quickly enough to have oceans and an atmosphere about 4,440  million years ago . There is evidence on the Moon of a Late Heavy Bombardment by asteroids from 4,000 to 3,800 million years ago . If, as seems likely, such a bombardment struck Earth at the same time, the first atmosphere and oceans may have been stripped away. Paleontology traces

1092-602: A common ancestor. Ideally the "family tree" has only two branches leading from each node ("junction"), but sometimes there is too little information to achieve this, and paleontologists have to make do with junctions that have several branches. The cladistic technique is sometimes fallible, as some features, such as wings or camera eyes , evolved more than once, convergently  – this must be taken into account in analyses. Evolutionary developmental biology , commonly abbreviated to "Evo Devo", also helps paleontologists to produce "family trees", and understand fossils. For example,

1274-443: A consequence of better nutrition, healthcare, and living conditions. The average mass of an adult human is 59 kg (130 lb) for females and 77 kg (170 lb) for males. Like many other conditions, body weight and body type are influenced by both genetic susceptibility and environment and varies greatly among individuals. Humans have a far faster and more accurate throw than other animals. Humans are also among

1456-456: A constant rate. These " molecular clocks ", however, are fallible, and provide only a very approximate timing: for example, they are not sufficiently precise and reliable for estimating when the groups that feature in the Cambrian explosion first evolved, and estimates produced by different techniques may vary by a factor of two. Earth formed about 4,570  million years ago and, after

1638-403: A data source that is not limited to animals with easily fossilised hard parts, and they reflect organisms' behaviours. Also many traces date from significantly earlier than the body fossils of animals that are thought to have been capable of making them. Whilst exact assignment of trace fossils to their makers is generally impossible, traces may for example provide the earliest physical evidence of

1820-419: A few other places), saw the emergence of agriculture and permanent human settlement ; in turn, this led to the development of civilization and kickstarted a period of continuous (and ongoing) population growth and rapid technological change . Since then, a number of civilizations have risen and fallen, while a number of sociocultural and technological developments have resulted in significant changes to

2002-573: A fortunate accident during other research. For example, the 1980 discovery by Luis and Walter Alvarez of iridium , a mainly extraterrestrial metal, in the Cretaceous – Paleogene boundary layer made asteroid impact the most favored explanation for the Cretaceous–Paleogene extinction event – although debate continues about the contribution of volcanism. A complementary approach to developing scientific knowledge, experimental science ,

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2184-463: A great capacity for altering their habitats by means of technology, irrigation , urban planning , construction, deforestation and desertification . Human settlements continue to be vulnerable to natural disasters , especially those placed in hazardous locations and with low quality of construction. Grouping and deliberate habitat alteration is often done with the goals of providing protection, accumulating comforts or material wealth, expanding

2366-612: A higher body fat percentage. At puberty , humans develop secondary sex characteristics . Females are capable of pregnancy , usually between puberty, at around 12 years old, and menopause , around the age of 50. As omnivorous creatures, they are capable of consuming a wide variety of plant and animal material, and have used fire and other forms of heat to prepare and cook food since the time of Homo erectus . Humans can survive for up to eight weeks without food and several days without water . Humans are generally diurnal , sleeping on average seven to nine hours per day. Childbirth

2548-475: A human can lose personhood (such as by going into a persistent vegetative state ). Humans are apes ( superfamily Hominoidea ). The lineage of apes that eventually gave rise to humans first split from gibbons (family Hylobatidae) and orangutans (genus Pongo ), then gorillas (genus Gorilla ), and finally, chimpanzees and bonobos (genus Pan ). The last split, between the human and chimpanzee–bonobo lineages, took place around 8–4 million years ago, in

2730-673: A larger effect on the mid-to-low latitudes and mid-to-high latitudes after ~5600 B.P. Human activity through land use changes already by the Mesolithic had major ecological impacts; it was an important influence on Holocene climatic changes, and is believed to be why the Holocene is an atypical interglacial that has not experienced significant cooling over its course. From the start of the Industrial Revolution onwards, large-scale anthropogenic greenhouse gas emissions caused

2912-563: A marine transgression occurred in southeastern Africa; in the Lake Lungué basin, this sea level highstand occurred from 740 to 910 AD, or from 1,210 to 1,040 BP, as evidenced by the lake's connection to the Indian Ocean at this time. This transgression was followed by a period of transition that lasted until 590 BP, when the region experienced significant aridification and began to be extensively used by humans for livestock herding. In

3094-821: A minor group until the first jawed fish appeared in the Late Ordovician . The spread of animals and plants from water to land required organisms to solve several problems, including protection against drying out and supporting themselves against gravity . The earliest evidence of land plants and land invertebrates date back to about 476  million years ago and 490  million years ago respectively. Those invertebrates, as indicated by their trace and body fossils, were shown to be arthropods known as euthycarcinoids . The lineage that produced land vertebrates evolved later but very rapidly between 370  million years ago and 360  million years ago ; recent discoveries have overturned earlier ideas about

3276-581: A more barrel-shaped chests in contrast to the funnel shape of other apes, an adaptation for bipedal respiration. Apart from bipedalism and brain size, humans differ from chimpanzees mostly in smelling , hearing and digesting proteins . While humans have a density of hair follicles comparable to other apes, it is predominantly vellus hair , most of which is so short and wispy as to be practically invisible. Humans have about 2 million sweat glands spread over their entire bodies, many more than chimpanzees, whose sweat glands are scarce and are mainly located on

3458-649: A more sustainable sedentary lifestyle . This form of lifestyle change allowed humans to develop towns and villages in centralized locations, which gave rise to the world known today. It is believed that the domestication of plants and animals began in the early part of the Holocene in the tropical areas of the planet. Because these areas had warm, moist temperatures, the climate was perfect for effective farming. Culture development and human population change, specifically in South America, has also been linked to spikes in hydroclimate resulting in climate variability in

3640-504: A much clearer picture of life at the time. Between the 8th and 6th century BCE, Europe entered the classical antiquity age, a period when ancient Greece and ancient Rome flourished. Around this time other civilizations also came to prominence. The Maya civilization started to build cities and create complex calendars . In Africa, the Kingdom of Aksum overtook the declining Kingdom of Kush and facilitated trade between India and

3822-632: A much wetter climate from 11,400 to 11,100 BP due to intensification of the ISM. Over the Early Holocene, the region was very wet, but during the Middle Holocene from 6,200 to 3,900 BP, aridification occurred, with the subsequent Late Holocene being relatively arid as a whole. Coastal southwestern India experienced a stronger ISM from 9,690 to 7,560 BP, during the HCO. From 3,510 to 2,550 BP, during

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4004-555: A rapid increase in knowledge about the history of life on Earth and to progress in the definition of the geologic time scale , largely based on fossil evidence. Although she was rarely recognised by the scientific community, Mary Anning was a significant contributor to the field of palaeontology during this period; she uncovered multiple novel Mesozoic reptile fossils and deducted that what were then known as bezoar stones are in fact fossilised faeces . In 1822 Henri Marie Ducrotay de Blainville , editor of Journal de Physique , coined

4186-543: A relatively short time can be used to link up isolated rocks: this technique is called biostratigraphy . For instance, the conodont Eoplacognathus pseudoplanus has a short range in the Middle Ordovician period. If rocks of unknown age are found to have traces of E. pseudoplanus , they must have a mid-Ordovician age. Such index fossils must be distinctive, be globally distributed and have a short time range to be useful. However, misleading results are produced if

4368-451: A result, humans are a cosmopolitan species found in almost all regions of the world, including tropical rainforest , arid desert , extremely cold arctic regions , and heavily polluted cities; in comparison, most other species are confined to a few geographical areas by their limited adaptability. The human population is not, however, uniformly distributed on the Earth 's surface, because

4550-471: A role. Drangajökull, Iceland's northernmost glacier, melted shortly after 9,200 BP. In Northern Germany , the Middle Holocene saw a drastic increase in the amount of raised bogs, most likely related to sea level rise. Although human activity affected geomorphology and landscape evolution in Northern Germany throughout the Holocene, it only became a dominant influence in the last four centuries. In

4732-639: A similar species. Migrating out of Africa , they gradually replaced and interbred with local populations of archaic humans. Multiple hypotheses for the extinction of archaic human species such as Neanderthals include competition, violence , interbreeding with Homo sapiens , or inability to adapt to climate change . Humans began exhibiting behavioral modernity about 160,000–60,000 years ago. For most of their history, humans were nomadic hunter-gatherers. The Neolithic Revolution , which began in Southwest Asia around 13,000 years ago (and separately in

4914-515: A simple linear or branched progression but involved interbreeding between related species . Genomic research has shown that hybridization between substantially diverged lineages was common in human evolution. DNA evidence suggests that several genes of Neanderthal origin are present among all non sub-Saharan-African populations, and Neanderthals and other hominins, such as Denisovans , may have contributed up to 6% of their genome to present-day non sub-Saharan-African humans. Human evolution

5096-608: A steady increase in brain size after about 3  million years ago . There is a long-running debate about whether modern humans are descendants of a single small population in Africa , which then migrated all over the world less than 200,000 years ago and replaced previous hominine species, or arose worldwide at the same time as a result of interbreeding . Life on earth has suffered occasional mass extinctions at least since 542  million years ago . Despite their disastrous effects, mass extinctions have sometimes accelerated

5278-451: A successful labor increased significantly during the 20th century in wealthier countries with the advent of new medical technologies. In contrast, pregnancy and natural childbirth remain hazardous ordeals in developing regions of the world, with maternal death rates approximately 100 times greater than in developed countries. Both the mother and the father provide care for human offspring, in contrast to other primates, where parental care

5460-498: A variety of methods, with a view toward further verifying and refining the Blytt–Sernander sequence . This is a classification of climatic periods initially defined by plant remains in peat mosses . Though the method was once thought to be of little interest, based on C dating of peats that was inconsistent with the claimed chronozones, investigators have found a general correspondence across Eurasia and North America . The scheme

5642-406: A wide variety of plant and animal material. Human groups have adopted a range of diets from purely vegan to primarily carnivorous . In some cases, dietary restrictions in humans can lead to deficiency diseases ; however, stable human groups have adapted to many dietary patterns through both genetic specialization and cultural conventions to use nutritionally balanced food sources. The human diet

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5824-511: Is 38 weeks, but a normal pregnancy can vary by up to 37 days. Embryonic development in the human covers the first eight weeks of development; at the beginning of the ninth week the embryo is termed a fetus . Humans are able to induce early labor or perform a caesarean section if the child needs to be born earlier for medical reasons. In developed countries, infants are typically 3–4 kg (7–9 lb) in weight and 47–53 cm (19–21 in) in height at birth. However, low birth weight

6006-485: Is a mixture of the two parental sets. Among the 23 pairs of chromosomes, there are 22 pairs of autosomes and one pair of sex chromosomes . Like other mammals, humans have an XY sex-determination system , so that females have the sex chromosomes XX and males have XY. Genes and environment influence human biological variation in visible characteristics, physiology, disease susceptibility and mental abilities. The exact influence of genes and environment on certain traits

6188-424: Is a population decline in the winter months of this realm. Humans established nation-states in the other seven realms, such as South Africa , India , Russia , Australia , Fiji , United States and Brazil (each located in a different biogeographical realm). By using advanced tools and clothing , humans have been able to extend their tolerance to a wide variety of temperatures, humidities , and altitudes. As

6370-402: Is characterized by a number of morphological , developmental , physiological , and behavioral changes that have taken place since the split between the last common ancestor of humans and chimpanzees . The most significant of these adaptations are hairlessness , obligate bipedalism, increased brain size and decreased sexual dimorphism ( neoteny ). The relationship between all these changes

6552-456: Is common in developing countries, and contributes to the high levels of infant mortality in these regions. Compared with other species, human childbirth is dangerous, with a much higher risk of complications and death. The size of the fetus's head is more closely matched to the pelvis than in other primates. The reason for this is not completely understood, but it contributes to a painful labor that can last 24 hours or more. The chances of

6734-595: Is composed only of eukaryotic cells, and the earliest evidence for it is the Francevillian Group Fossils from 2,100  million years ago , although specialisation of cells for different functions first appears between 1,430  million years ago (a possible fungus) and 1,200  million years ago (a probable red alga ). Sexual reproduction may be a prerequisite for specialisation of cells, as an asexual multicellular organism might be at risk of being taken over by rogue cells that retain

6916-421: Is dangerous, with a high risk of complications and death . Often, both the mother and the father provide care for their children, who are helpless at birth . Humans have a large, highly developed, and complex prefrontal cortex , the region of the brain associated with higher cognition. Humans are highly intelligent and capable of episodic memory ; they have flexible facial expressions, self-awareness , and

7098-479: Is generally aging, with the median age around 40 years. In the developing world , the median age is between 15 and 20 years. While one in five Europeans is 60 years of age or older, only one in twenty Africans is 60 years of age or older. In 2012, the United Nations estimated that there were 316,600 living centenarians (humans of age 100 or older) worldwide. Humans are omnivorous , capable of consuming

7280-482: Is hard to decide at what level to place a new higher-level grouping, e.g. genus or family or order ; this is important since the Linnaean rules for naming groups are tied to their levels, and hence if a group is moved to a different level it must be renamed. Paleontologists generally use approaches based on cladistics , a technique for working out the evolutionary "family tree" of a set of organisms. It works by

7462-443: Is inherited only from the mother, geneticists have concluded that the last female common ancestor whose genetic marker is found in all modern humans, the so-called mitochondrial Eve , must have lived around 90,000 to 200,000 years ago. Most human reproduction takes place by internal fertilization via sexual intercourse , but can also occur through assisted reproductive technology procedures. The average gestation period

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7644-408: Is mostly done by the mother. Helpless at birth , humans continue to grow for some years, typically reaching sexual maturity at 15 to 17 years of age. The human life span has been split into various stages ranging from three to twelve. Common stages include infancy , childhood , adolescence , adulthood and old age . The lengths of these stages have varied across cultures and time periods but

7826-521: Is not well understood. While no humans – not even monozygotic twins  – are genetically identical, two humans on average will have a genetic similarity of 99.5%-99.9%. This makes them more homogeneous than other great apes, including chimpanzees. This small variation in human DNA compared to many other species suggests a population bottleneck during the Late Pleistocene (around 100,000 years ago), in which

8008-443: Is often said to work by conducting experiments to disprove hypotheses about the workings and causes of natural phenomena. This approach cannot prove a hypothesis, since some later experiment may disprove it, but the accumulation of failures to disprove is often compelling evidence in favor. However, when confronted with totally unexpected phenomena, such as the first evidence for invisible radiation , experimental scientists often use

8190-594: Is one that contained an extinct "crocodile-like" marine reptile, which eventually came to be known as the mosasaurid Mosasaurus of the Cretaceous period. The first half of the 19th century saw geological and paleontological activity become increasingly well organised with the growth of geologic societies and museums and an increasing number of professional geologists and fossil specialists. Interest increased for reasons that were not purely scientific, as geology and paleontology helped industrialists to find and exploit natural resources such as coal. This contributed to

8372-401: Is our only means of giving rocks greater than about 50 million years old an absolute age, and can be accurate to within 0.5% or better. Although radiometric dating requires very careful laboratory work, its basic principle is simple: the rates at which various radioactive elements decay are known, and so the ratio of the radioactive element to the element into which it decays shows how long ago

8554-520: Is prominently reflected in human culture and has led to the development of food science . Paleontologists Paleontology ( / ˌ p eɪ l i ɒ n ˈ t ɒ l ə dʒ i , ˌ p æ l i -, - ən -/ PAY -lee-on- TOL -ə-jee, PAL -ee-, -⁠ən- ), also spelled palaeontology or palæontology , is the scientific study of life that existed prior to the start of the Holocene epoch (roughly 11,700 years before present). It includes

8736-445: Is the subject of ongoing debate. Hylobatidae ( gibbons ) Pongo abelii Pongo tapanuliensis Pongo pygmaeus Gorilla gorilla Gorilla beringei Pan troglodytes Pan paniscus Homo sapiens (humans) Until about 12,000 years ago, all humans lived as hunter-gatherers . The Neolithic Revolution (the invention of agriculture ) first took place in Southwest Asia and spread through large parts of

8918-503: Is thought to have been propelled by coevolution with pollinating insects. Social insects appeared around the same time and, although they account for only small parts of the insect "family tree", now form over 50% of the total mass of all insects. Humans evolved from a lineage of upright-walking apes whose earliest fossils date from over 6  million years ago . Although early members of this lineage had chimp -sized brains, about 25% as big as modern humans', there are signs of

9100-654: Is typified by an unusually rapid growth spurt during adolescence. Human females undergo menopause and become infertile at around the age of 50. It has been proposed that menopause increases a woman's overall reproductive success by allowing her to invest more time and resources in her existing offspring, and in turn their children (the grandmother hypothesis ), rather than by continuing to bear children into old age. The life span of an individual depends on two major factors, genetics and lifestyle choices. For various reasons, including biological/genetic causes, women live on average about four years longer than men. As of 2018 ,

9282-756: The 9th millennium BC . The preceding period of the Late Pleistocene had already brought advancements such as the bow and arrow , creating more efficient forms of hunting and replacing spear throwers . In the Holocene, however, the domestication of plants and animals allowed humans to develop villages and towns in centralized locations. Archaeological data shows that between 10,000 and 7,000 BP rapid domestication of plants and animals took place in tropical and subtropical parts of Asia , Africa , and Central America . The development of farming allowed humans to transition away from hunter-gatherer nomadic cultures, which did not establish permanent settlements, to

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9464-548: The British Empire expanded to become the world's largest empire . A tenuous balance of power among European nations collapsed in 1914 with the outbreak of the First World War , one of the deadliest conflicts in history. In the 1930s, a worldwide economic crisis led to the rise of authoritarian regimes and a Second World War , involving almost all of the world's countries . The war's destruction led to

9646-580: The Caral–Supe civilization is the oldest complex civilization in the Americas. Astronomy and mathematics were also developed and the Great Pyramid of Giza was built. There is evidence of a severe drought lasting about a hundred years that may have caused the decline of these civilizations, with new ones appearing in the aftermath. Babylonians came to dominate Mesopotamia while others, such as

9828-511: The Church would provide centralized authority and education. In the Middle East, Islam became the prominent religion and expanded into North Africa. It led to an Islamic Golden Age , inspiring achievements in architecture , the revival of old advances in science and technology, and the formation of a distinct way of life. The Christian and Islamic worlds would eventually clash, with

10010-629: The French Alps , geochemistry and lithium isotope signatures in lake sediments have suggested gradual soil formation from the Last Glacial Period to the Holocene climatic optimum , and this soil development was altered by the settlement of human societies. Early anthropogenic activities such as deforestation and agriculture reinforced soil erosion, which peaked in the Middle Ages at an unprecedented level, marking human forcing as

10192-738: The Galápagos Islands shows that the El Niño–Southern Oscillation (ENSO) was significantly weaker during the Middle Holocene, but that the strength of ENSO became moderate to high over the Late Holocene. Animal and plant life have not evolved much during the relatively short Holocene, but there have been major shifts in the richness and abundance of plants and animals. A number of large animals including mammoths and mastodons , saber-toothed cats like Smilodon and Homotherium , and giant sloths went extinct in

10374-468: The Great Basin . Eastern North America underwent abrupt warming and humidification around 10,500 BP and then declined from 9,300 to 9,100 BP. The region has undergone a long term wettening since 5,500 BP occasionally interrupted by intervals of high aridity. A major cool event lasting from 5,500 to 4,700 BP was coeval with a major humidification before being terminated by a major drought and warming at

10556-562: The Gulf of Thailand was exposed above sea level and was very arid. A marine transgression occurred from 7,500 to 6,200 BP amidst global warming. During the Middle Holocene, western North America was drier than present, with wetter winters and drier summers. After the end of the thermal maximum of the HCO around 4,500 BP, the East Greenland Current underwent strengthening. A massive megadrought occurred from 2,800 to 1,850 BP in

10738-474: The Homo genus. The name " Homo sapiens " means 'wise man' or 'knowledgeable man'. There is disagreement if certain extinct members of the genus, namely Neanderthals , should be included as a separate species of humans or as a subspecies of H. sapiens . Human is a loanword of Middle English from Old French humain , ultimately from Latin hūmānus , the adjectival form of homō ('man' – in

10920-621: The Intertropical Convergence Zone (ITCZ) produced increased monsoon rainfall over North Africa. The lush vegetation of the Sahara brought an increase in pastoralism . The AHP ended around 5,500 BP, after which the Sahara began to dry and become the desert it is today. A stronger East African Monsoon during the Middle Holocene increased precipitation in East Africa and raised lake levels. Around 800 AD, or 1,150 BP,

11102-677: The Kalahari Desert , Holocene climate was overall very stable and environmental change was of low amplitude. Relatively cool conditions have prevailed since 4,000 BP. In the Middle East, the Holocene brought a warmer and wetter climate, in contrast to the preceding cold, dry Younger Dryas . The Early Holocene saw the advent and spread of agriculture in the Fertile Crescent — sheep , goat , cattle , and later pig were domesticated, as well as cereals, like wheat and barley , and legumes —which would later disperse into much of

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11284-579: The Kingdom of England , the Kingdom of France and the Holy Roman Empire declaring a series of holy wars to regain control of the Holy Land from Muslims . In the Americas, between 200 and 900 CE Mesoamerica was in its Classic Period , while further north, complex Mississippian societies would arise starting around 800 CE. The Mongol Empire would conquer much of Eurasia in the 13th and 14th centuries. Over this same time period,

11466-801: The Mali Empire in Africa grew to be the largest empire on the continent, stretching from Senegambia to Ivory Coast . Oceania would see the rise of the Tuʻi Tonga Empire which expanded across many islands in the South Pacific. By the late 15th century, the Aztecs and Inca had become the dominant power in Mesoamerica and the Andes , respectively. The early modern period in Europe and

11648-733: The Mesolithic , Neolithic , and Bronze Age , are usually used. However, the time periods referenced by these terms vary with the emergence of those technologies in different parts of the world. Some scholars have argued that a third epoch of the Quaternary, the Anthropocene , has now begun. This term has been used to denote the present time-interval in which many geologically significant conditions and processes have been profoundly altered by human activities. The 'Anthropocene' (a term coined by Paul J. Crutzen and Eugene Stoermer in 2000)

11830-470: The Middle Ages the Persian naturalist Ibn Sina , known as Avicenna in Europe, discussed fossils and proposed a theory of petrifying fluids on which Albert of Saxony elaborated in the 14th century. The Chinese naturalist Shen Kuo (1031–1095) proposed a theory of climate change based on the presence of petrified bamboo in regions that in his time were too dry for bamboo. In early modern Europe ,

12012-614: The Middle East and Anatolia , the term Epipaleolithic is preferred in place of Mesolithic, as they refer to approximately the same time period. Cultures in this period include Hamburgian , Federmesser , and the Natufian culture , during which the oldest inhabited places still existing on Earth were first settled, such as Tell es-Sultan (Jericho) in the Middle East . There is also evolving archeological evidence of proto-religion at locations such as Göbekli Tepe , as long ago as

12194-538: The Neogene - Quaternary . In deeper-level deposits in western Europe are early-aged mammals such as the palaeothere perissodactyl Palaeotherium and the anoplotheriid artiodactyl Anoplotherium , both of which were described earliest after the former two genera, which today are known to date to the Paleogene period. Cuvier figured out that even older than the two levels of deposits with extinct large mammals

12376-587: The Old World over the following millennia. It also occurred independently in Mesoamerica (about 6,000 years ago), China, Papua New Guinea , and the Sahel and West Savanna regions of Africa. Access to food surplus led to the formation of permanent human settlements , the domestication of animals and the use of metal tools for the first time in history. Agriculture and sedentary lifestyle led to

12558-637: The Permian–Triassic extinction event . Amphibians Extinct Synapsids Mammals Extinct reptiles Lizards and snakes Extinct Archosaurs Crocodilians Extinct Dinosaurs Birds Naming groups of organisms in a way that is clear and widely agreed is important, as some disputes in paleontology have been based just on misunderstandings over names. Linnaean taxonomy is commonly used for classifying living organisms, but runs into difficulties when dealing with newly discovered organisms that are significantly different from known ones. For example: it

12740-516: The Permian–Triassic extinction event . A relatively recent discipline, molecular phylogenetics , compares the DNA and RNA of modern organisms to re-construct the "family trees" of their evolutionary ancestors. It has also been used to estimate the dates of important evolutionary developments, although this approach is controversial because of doubts about the reliability of the " molecular clock ". Techniques from engineering have been used to analyse how

12922-652: The Poverty Point culture , Minoans and the Shang dynasty , rose to prominence in new areas. The Late Bronze Age collapse around 1200 BCE resulted in the disappearance of a number of civilizations and the beginning of the Greek Dark Ages . During this period iron started replacing bronze, leading to the Iron Age . In the 5th century BCE, history started being recorded as a discipline , which provided

13104-581: The Tien Shan , sedimentological evidence from Swan Lake suggests the period between 8,500 and 6,900 BP was relatively warm, with steppe meadow vegetation being predominant. An increase in Cyperaceae from 6,900 to 2,600 BP indicates cooling and humidification of the Tian Shan climate that was interrupted by a warm period between 5,500 and 4,500 BP. After 2,600 BP, an alpine steppe climate prevailed across

13286-795: The colonization of the Americas and the Columbian Exchange . This expansion led to the Atlantic slave trade and the genocide of Native American peoples . This period also marked the Scientific Revolution , with great advances in mathematics , mechanics , astronomy and physiology . The late modern period (1800–present) saw the Technological and Industrial Revolution bring such discoveries as imaging technology , major innovations in transport and energy development . Influenced by Enlightenment ideals,

13468-454: The embryological development of some modern brachiopods suggests that brachiopods may be descendants of the halkieriids , which became extinct in the Cambrian period. Paleontology seeks to map out how living things have changed through time. A substantial hurdle to this aim is the difficulty of working out how old fossils are. Beds that preserve fossils typically lack the radioactive elements needed for radiometric dating . This technique

13650-958: The last glacial period . Local names for the last glacial period include the Wisconsinan in North America , the Weichselian in Europe, the Devensian in Britain, the Llanquihue in Chile and the Otiran in New Zealand. The Holocene can be subdivided into five time intervals, or chronozones , based on climatic fluctuations: Geologists working in different regions are studying sea levels, peat bogs, and ice-core samples, using

13832-526: The sixth mass extinction or Anthropocene extinction , is an ongoing extinction event of species during the present Holocene epoch (with the more recent time sometimes called Anthropocene) as a result of human activity . The included extinctions span numerous families of fungi , plants , and animals , including mammals , birds , reptiles , amphibians , fish and invertebrates . With widespread degradation of highly biodiverse habitats such as coral reefs and rainforests , as well as other areas,

14014-526: The " jigsaw puzzles " of biostratigraphy (arrangement of rock layers from youngest to oldest). Classifying ancient organisms is also difficult, as many do not fit well into the Linnaean taxonomy classifying living organisms, and paleontologists more often use cladistics to draw up evolutionary "family trees". The final quarter of the 20th century saw the development of molecular phylogenetics , which investigates how closely organisms are related by measuring

14196-518: The Americas and Europe experienced a period of political revolutions known as the Age of Revolution . The Napoleonic Wars raged through Europe in the early 1800s, Spain lost most of its colonies in the New World , while Europeans continued expansion into Africa  – where European control went from 10% to almost 90% in less than 50 years  – and Oceania. In the 19th century,

14378-555: The Early Cambrian , along with several "weird wonders" that bear little obvious resemblance to any modern animals. There is a long-running debate about whether this Cambrian explosion was truly a very rapid period of evolutionary experimentation; alternative views are that modern-looking animals began evolving earlier but fossils of their precursors have not yet been found, or that the "weird wonders" are evolutionary "aunts" and "cousins" of modern groups. Vertebrates remained

14560-613: The Earth to warm. Likewise, climatic changes have induced substantial changes in human civilisation over the course of the Holocene. During the transition from the last glacial to the Holocene, the Huelmo–Mascardi Cold Reversal in the Southern Hemisphere began before the Younger Dryas, and the maximum warmth flowed south to north from 11,000 to 7,000 years ago. It appears that this was influenced by

14742-411: The Earth's organic and inorganic past". William Whewell (1794–1866) classified paleontology as one of the historical sciences, along with archaeology , geology, astronomy , cosmology , philology and history itself: paleontology aims to describe phenomena of the past and to reconstruct their causes. Hence it has three main elements: description of past phenomena; developing a general theory about

14924-588: The Holocene Epoch. A 1,500-year cycle corresponding to the North Atlantic oceanic circulation may have had widespread global distribution in the Late Holocene. From 8,500 BP to 6,700 BP, North Atlantic climate oscillations were highly irregular and erratic because of perturbations from substantial ice discharge into the ocean from the collapsing Laurentide Ice Sheet. The Greenland ice core records indicate that climate changes became more regional and had

15106-474: The Holocene has shown significant variability despite ice core records from Greenland suggesting a more stable climate following the preceding ice age. Marine chemical fluxes during the Holocene were lower than during the Younger Dryas, but were still considerable enough to imply notable changes in the climate. The temporal and spatial extent of climate change during the Holocene is an area of considerable uncertainty, with radiative forcing recently proposed to be

15288-679: The International Union of Geological Sciences later formally confirmed, by a near unanimous vote, the rejection of the working group's Anthropocene Epoch proposal for inclusion in the Geologic Time Scale. The Holocene is a geologic epoch that follows directly after the Pleistocene . Continental motions due to plate tectonics are less than a kilometre over a span of only 10,000 years. However, ice melt caused world sea levels to rise about 35 m (115 ft) in

15470-434: The Late Holocene, the ISM became weaker, although this weakening was interrupted by an interval of unusually high ISM strength from 3,400 to 3,200 BP. Southwestern China experienced long-term warming during the Early Holocene up until ~7,000 BP. Northern China experienced an abrupt aridification event approximately 4,000 BP. From around 3,500 to 3,000 BP, northeastern China underwent a prolonged cooling, manifesting itself with

15652-409: The Late Holocene. The Northwest Australian Summer Monsoon was in a strong phase from 8,500 to 6,400 BP, from 5,000 to 4,000 BP (possibly until 3,000 BP), and from 1,300 to 900 BP, with weak phases in between and the current weak phase beginning around 900 BP after the end of the last strong phase. Ice core measurements imply that the sea surface temperature (SST) gradient east of New Zealand, across

15834-523: The MWP. A warming of +1 degree Celsius occurs 5–40 times more frequently in modern years than during the MWP. The major forcing during the MWP was due to greater solar activity, which led to heterogeneity compared to the greenhouse gas forcing of modern years that leads to more homogeneous warming. This was followed by the Little Ice Age (LIA) from the 13th or 14th century to the mid-19th century. The LIA

16016-817: The Mediterranean. In West Asia, the Achaemenid Empire 's system of centralized governance became the precursor to many later empires, while the Gupta Empire in India and the Han dynasty in China have been described as golden ages in their respective regions. Following the fall of the Western Roman Empire in 476, Europe entered the Middle Ages . During this period, Christianity and

16198-639: The Middle to Late Holocene, the coastline of the Levant and Persian Gulf receded, prompting a shift in human settlement patterns following this marine regression. Central Asia experienced glacial-like temperatures until about 8,000 BP, when the Laurentide Ice Sheet collapsed. In Xinjiang , long-term Holocene warming increased meltwater supply during summers, creating large lakes and oases at low altitudes and inducing enhanced moisture recycling. In

16380-832: The Near East ( c.  1450 –1800) began with the final defeat of the Byzantine Empire , and the rise of the Ottoman Empire . Meanwhile, Japan entered the Edo period , the Qing dynasty rose in China and the Mughal Empire ruled much of India. Europe underwent the Renaissance , starting in the 15th century, and the Age of Discovery began with the exploring and colonizing of new regions. This included

16562-521: The Oort, Wolf , Spörer , and Maunder Minima . A notable cooling event in southeastern China occurred 3,200 BP. Strengthening of the winter monsoon occurred around 5,500, 4,000, and 2,500 BP. Monsoonal regions of China became more arid in the Late Holocene. In the Sea of Japan, the Middle Holocene was notable for its warmth, with rhythmic temperature fluctuations every 400-500 and 1,000 years. Before 7,500 BP,

16744-412: The ability to reproduce. The earliest known animals are cnidarians from about 580  million years ago , but these are so modern-looking that they must be descendants of earlier animals. Early fossils of animals are rare because they had not developed mineralised , easily fossilized hard parts until about 548  million years ago . The earliest modern-looking bilaterian animals appear in

16926-515: The appearance of moderately complex animals (comparable to earthworms ). Geochemical observations may help to deduce the global level of biological activity at a certain period, or the affinity of certain fossils. For example, geochemical features of rocks may reveal when life first arose on Earth, and may provide evidence of the presence of eukaryotic cells, the type from which all multicellular organisms are built. Analyses of carbon isotope ratios may help to explain major transitions such as

17108-625: The atmosphere increased their effectiveness as nurseries of evolution. While eukaryotes , cells with complex internal structures, may have been present earlier, their evolution speeded up when they acquired the ability to transform oxygen from a poison to a powerful source of metabolic energy. This innovation may have come from primitive eukaryotes capturing oxygen-powered bacteria as endosymbionts and transforming them into organelles called mitochondria . The earliest evidence of complex eukaryotes with organelles (such as mitochondria) dates from 1,850  million years ago . Multicellular life

17290-478: The available food, improving aesthetics , increasing knowledge or enhancing the exchange of resources. Humans are one of the most adaptable species, despite having a low or narrow tolerance for many of the earth's extreme environments. Currently the species is present in all eight biogeographical realms , although their presence in the Antarctic realm is very limited to research stations and annually there

17472-561: The best long-distance runners in the animal kingdom, but slower over short distances. Humans' thinner body hair and more productive sweat glands help avoid heat exhaustion while running for long distances. Compared to other apes, the human heart produces greater stroke volume and cardiac output and the aorta is proportionately larger. Like most animals, humans are a diploid and eukaryotic species. Each somatic cell has two sets of 23 chromosomes , each set received from one parent; gametes have only one set of chromosomes, which

17654-415: The bodies of ancient organisms might have worked, for example the running speed and bite strength of Tyrannosaurus , or the flight mechanics of Microraptor . It is relatively commonplace to study the internal details of fossils using X-ray microtomography . Paleontology, biology, archaeology, and paleoneurobiology combine to study endocranial casts (endocasts) of species related to humans to clarify

17836-729: The border between biology and geology , but it differs from archaeology in that it excludes the study of anatomically modern humans . It now uses techniques drawn from a wide range of sciences, including biochemistry , mathematics , and engineering. Use of all these techniques has enabled paleontologists to discover much of the evolutionary history of life , almost back to when Earth became capable of supporting life, nearly 4 billion years ago. As knowledge has increased, paleontology has developed specialised sub-divisions, some of which focus on different types of fossil organisms while others study ecology and environmental history, such as ancient climates . Body fossils and trace fossils are

18018-401: The causes of various types of change; and applying those theories to specific facts. When trying to explain the past, paleontologists and other historical scientists often construct a set of one or more hypotheses about the causes and then look for a " smoking gun ", a piece of evidence that strongly accords with one hypothesis over any others. Sometimes researchers discover a "smoking gun" by

18200-763: The characteristics and evolution of humans as a species. When dealing with evidence about humans, archaeologists and paleontologists may work together – for example paleontologists might identify animal or plant fossils around an archaeological site , to discover the people who lived there, and what they ate; or they might analyze the climate at the time of habitation. In addition, paleontology often borrows techniques from other sciences, including biology, osteology , ecology, chemistry , physics and mathematics. For example, geochemical signatures from rocks may help to discover when life first arose on Earth, and analyses of carbon isotope ratios may help to identify climate changes and even to explain major transitions such as

18382-520: The chronological order in which rocks were formed, is useful to both paleontologists and geologists. Biogeography studies the spatial distribution of organisms, and is also linked to geology, which explains how Earth's geography has changed over time. Although paleontology became established around 1800, earlier thinkers had noticed aspects of the fossil record. The ancient Greek philosopher Xenophanes (570–480 BCE) concluded from fossil sea shells that some areas of land were once under water. During

18564-665: The collapse of most global empires, leading to widespread decolonization. Following the conclusion of the Second World War in 1945, the United States and the USSR emerged as the remaining global superpowers . This led to a Cold War that saw a struggle for global influence, including a nuclear arms race and a space race , ending in the collapse of the Soviet Union. The current Information Age , spurred by

18746-787: The combined eastern and western Roman Empire in the 4th century AD. Bubonic plagues , first recorded in the 6th century AD, reduced the population by 50%, with the Black Death killing 75–200 million people in Eurasia and North Africa alone. Human population is believed to have reached one billion in 1800. It has since then increased exponentially, reaching two billion in 1930 and three billion in 1960, four in 1975, five in 1987 and six billion in 1999. It passed seven billion in 2011 and passed eight billion in November 2022. It took over two million years of human prehistory and history for

18928-478: The continent, gradually replacing or interbreeding with local populations of archaic humans. Humans began exhibiting behavioral modernity about 160,000–70,000 years ago, and possibly earlier. This development was likely selected amidst natural climate change in Middle to Late Pleistocene Africa. The "out of Africa" migration took place in at least two waves, the first around 130,000 to 100,000 years ago,

19110-445: The date when lineages first appeared. For instance, if fossils of B or C date to X million years ago and the calculated "family tree" says A was an ancestor of B and C, then A must have evolved more than X million years ago. It is also possible to estimate how long ago two living clades diverged – i.e. approximately how long ago their last common ancestor must have lived – by assuming that DNA mutations accumulate at

19292-423: The desire to understand and influence phenomena having motivated humanity's development of science , technology , philosophy , mythology , religion , and other frameworks of knowledge ; humans also study themselves through such domains as anthropology , social science , history , psychology , and medicine . There are estimated to be more than eight billion living humans . Although some scientists equate

19474-594: The development of mammalian traits such as endothermy and hair. After the Cretaceous–Paleogene extinction event 66  million years ago killed off all the dinosaurs except the birds, mammals increased rapidly in size and diversity, and some took to the air and the sea. Fossil evidence indicates that flowering plants appeared and rapidly diversified in the Early Cretaceous between 130  million years ago and 90  million years ago . Their rapid rise to dominance of terrestrial ecosystems

19656-516: The development of the Internet and Artificial Intelligence systems, sees the world becoming increasingly globalized and interconnected. Early human settlements were dependent on proximity to water and – depending on the lifestyle – other natural resources used for subsistence , such as populations of animal prey for hunting and arable land for growing crops and grazing livestock. Modern humans, however, have

19838-561: The development of the body plans of most animal phyla . The discovery of fossils of the Ediacaran biota and developments in paleobiology extended knowledge about the history of life back far before the Cambrian. Increasing awareness of Gregor Mendel 's pioneering work in genetics led first to the development of population genetics and then in the mid-20th century to the modern evolutionary synthesis , which explains evolution as

20020-426: The different levels of deposits represented different time periods in the early 19th century. The surface-level deposits in the Americas contained later mammals like the megatheriid ground sloth Megatherium and the mammutid proboscidean Mammut (later known informally as a "mastodon"), which were some of the earliest-named fossil mammal genera with official taxonomic authorities. They today are known to date to

20202-567: The disruption of Bronze Age civilisations in the region. Eastern and southern China, the monsoonal regions of China, were wetter than present in the Early and Middle Holocene. Lake Huguangyan's TOC, δC wax , δC org , δN values suggest the period of peak moisture lasted from 9,200 to 1,800 BP and was attributable to a strong East Asian Summer Monsoon (EASM). Late Holocene cooling events in the region were dominantly influenced by solar forcing, with many individual cold snaps linked to solar minima such as

20384-640: The early 20th century, there has been continuous human presence in Antarctica through research stations and, since 2000, in space through habitation on the International Space Station . All modern humans are classified into the species Homo sapiens , coined by Carl Linnaeus in his 1735 work Systema Naturae . The generic name " Homo " is a learned 18th-century derivation from Latin homō , which refers to humans of either sex. The word human can refer to all members of

20566-472: The early part of the Holocene and another 30 m in the later part of the Holocene. In addition, many areas above about 40 degrees north latitude had been depressed by the weight of the Pleistocene glaciers and rose as much as 180 m (590 ft) due to post-glacial rebound over the late Pleistocene and Holocene, and are still rising today. The sea-level rise and temporary land depression allowed temporary marine incursions into areas that are now far from

20748-685: The emergence of early civilizations . An urban revolution took place in the 4th millennium BCE with the development of city-states , particularly Sumerian cities located in Mesopotamia . It was in these cities that the earliest known form of writing, cuneiform script , appeared around 3000 BCE. Other major civilizations to develop around this time were Ancient Egypt and the Indus Valley Civilisation . They eventually traded with each other and invented technology such as wheels, plows and sails. Emerging by 3000 BCE,

20930-573: The end of that interval. During the Early Holocene, relative sea level rose in the Bahia region, causing a landward expansion of mangroves. During the Late Holocene, the mangroves declined as sea level dropped and freshwater supply increased. In the Santa Catarina region, the maximum sea level highstand was around 2.1 metres above present and occurred about 5,800 to 5,000 BP. Sea levels at Rocas Atoll were likewise higher than present for much of

21112-409: The end of the 20th century have been particularly important as they have provided new information about the earliest evolution of animals, early fish, dinosaurs and the evolution of birds. The last few decades of the 20th century saw a renewed interest in mass extinctions and their role in the evolution of life on Earth. There was also a renewed interest in the Cambrian explosion that apparently saw

21294-551: The end of the HCO to before the Industrial Revolution . From the 10th-14th century, the climate was similar to that of modern times during a period known as the Mediaeval Warm Period (MWP), also known as the Mediaeval Climatic Optimum (MCO). It was found that the warming that is taking place in current years is both more frequent and more spatially homogeneous than what was experienced during

21476-410: The evolution of the human brain. Paleontology even contributes to astrobiology , the investigation of possible life on other planets , by developing models of how life may have arisen and by providing techniques for detecting evidence of life. As knowledge has increased, paleontology has developed specialised subdivisions. Vertebrate paleontology concentrates on fossils from the earliest fish to

21658-466: The evolutionary history of life back to over 3,000  million years ago , possibly as far as 3,800  million years ago . The oldest clear evidence of life on Earth dates to 3,000  million years ago , although there have been reports, often disputed, of fossil bacteria from 3,400  million years ago and of geochemical evidence for the presence of life 3,800  million years ago . Some scientists have proposed that life on Earth

21840-555: The exceptional events that cause quick burial make it difficult to study the normal environments of the animals. The sparseness of the fossil record means that organisms are expected to exist long before and after they are found in the fossil record – this is known as the Signor–Lipps effect . Trace fossils consist mainly of tracks and burrows, but also include coprolites (fossil feces ) and marks left by feeding. Trace fossils are particularly significant because they represent

22022-463: The focus of paleontology shifted to understanding evolutionary paths, including human evolution , and evolutionary theory. The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection. Fossils found in China near

22204-449: The following: At the end of the 18th century Georges Cuvier 's work established comparative anatomy as a scientific discipline and, by proving that some fossil animals resembled no living ones, demonstrated that animals could become extinct , leading to the emergence of paleontology. The expanding knowledge of the fossil record also played an increasing role in the development of geology, particularly stratigraphy . Cuvier proved that

22386-580: The fossil record: different environments are more favorable to the preservation of different types of organism or parts of organisms. Further, only the parts of organisms that were already mineralised are usually preserved, such as the shells of molluscs. Since most animal species are soft-bodied, they decay before they can become fossilised. As a result, although there are 30-plus phyla of living animals, two-thirds have never been found as fossils. Occasionally, unusual environments may preserve soft tissues. These lagerstätten allow paleontologists to examine

22568-531: The future evolution of living species, including approximately synchronous lithospheric evidence, or more recently hydrospheric and atmospheric evidence of the human impact. In July 2018, the International Union of Geological Sciences split the Holocene Epoch into three distinct ages based on the climate, Greenlandian (11,700 years ago to 8,200 years ago), Northgrippian (8,200 years ago to 4,200 years ago) and Meghalayan (4,200 years ago to

22750-566: The genomes of 1,184 individuals from 11 populations and identified 1.6 million single nucleotide polymorphisms . African populations harbor the highest number of private genetic variants. While many of the common variants found in populations outside of Africa are also found on the African continent, there are still large numbers that are private to these regions, especially Oceania and the Americas . By 2010 estimates, humans have approximately 22,000 genes. By comparing mitochondrial DNA , which

22932-595: The global average life expectancy at birth of a girl is estimated to be 74.9 years compared to 70.4 for a boy. There are significant geographical variations in human life expectancy, mostly correlated with economic development – for example, life expectancy at birth in Hong Kong is 87.6 years for girls and 81.8 for boys, while in the Central African Republic , it is 55.0 years for girls and 50.6 for boys. The developed world

23114-687: The history and driving forces behind their evolution. Land plants were so successful that their detritus caused an ecological crisis in the Late Devonian , until the evolution of fungi that could digest dead wood. During the Permian period, synapsids , including the ancestors of mammals , may have dominated land environments, but this ended with the Permian–Triassic extinction event 251  million years ago , which came very close to wiping out all complex life. The extinctions were apparently fairly sudden, at least among vertebrates. During

23296-533: The history of Earth's climate and the mechanisms that have changed it  – which have sometimes included evolutionary developments, for example the rapid expansion of land plants in the Devonian period removed more carbon dioxide from the atmosphere, reducing the greenhouse effect and thus helping to cause an ice age in the Carboniferous period. Biostratigraphy , the use of fossils to work out

23478-443: The human lifestyle. Genes and the environment influence human biological variation in visible characteristics, physiology , disease susceptibility, mental abilities, body size, and life span. Though humans vary in many traits, humans are among the least genetically diverse species. Any two humans are at least 99.5% genetically similar. Humans are sexually dimorphic : generally, males have greater body strength and females have

23660-555: The human population to reach one billion and only 207 years more to grow to 7 billion. The combined biomass of the carbon of all the humans on Earth in 2018 was estimated at 60 million tons, about 10 times larger than that of all non-domesticated mammals. In 2018, 4.2 billion humans (55%) lived in urban areas, up from 751 million in 1950. The most urbanized regions are Northern America (82%), Latin America (81%), Europe (74%) and Oceania (68%), with Africa and Asia having nearly 90% of

23842-517: The human population was reduced to a small number of breeding pairs. The forces of natural selection have continued to operate on human populations, with evidence that certain regions of the genome display directional selection in the past 15,000 years. The human genome was first sequenced in 2001 and by 2020 hundreds of thousands of genomes had been sequenced. In 2012 the International HapMap Project had compared

24024-542: The immediate ancestors of modern mammals . Invertebrate paleontology deals with fossils such as molluscs , arthropods , annelid worms and echinoderms . Paleobotany studies fossil plants , algae , and fungi. Palynology , the study of pollen and spores produced by land plants and protists , straddles paleontology and botany , as it deals with both living and fossil organisms. Micropaleontology deals with microscopic fossil organisms of all kinds. Instead of focusing on individual organisms, paleoecology examines

24206-434: The index fossils turn out to have longer fossil ranges than first thought. Stratigraphy and biostratigraphy can in general provide only relative dating ( A was before B ), which is often sufficient for studying evolution. However, this is difficult for some time periods, because of the problems involved in matching up rocks of the same age across different continents . Family-tree relationships may also help to narrow down

24388-538: The interactions between different ancient organisms, such as their food chains , and the two-way interactions with their environments.   For example, the development of oxygenic photosynthesis by bacteria caused the oxygenation of the atmosphere and hugely increased the productivity and diversity of ecosystems . Together, these led to the evolution of complex eukaryotic cells, from which all multicellular organisms are built. Paleoclimatology , although sometimes treated as part of paleoecology, focuses more on

24570-463: The internal anatomy of animals that in other sediments are represented only by shells, spines, claws, etc. – if they are preserved at all. However, even lagerstätten present an incomplete picture of life at the time. The majority of organisms living at the time are probably not represented because lagerstätten are restricted to a narrow range of environments, e.g. where soft-bodied organisms can be preserved very quickly by events such as mudslides; and

24752-456: The investigation of evolutionary "family trees" by techniques derived from biochemistry , began to make an impact, particularly when it was proposed that the human lineage had diverged from apes much more recently than was generally thought at the time. Although this early study compared proteins from apes and humans, most molecular phylogenetics research is now based on comparisons of RNA and DNA . Fossils of organisms' bodies are usually

24934-567: The late Miocene epoch. During this split, chromosome 2 was formed from the joining of two other chromosomes, leaving humans with only 23 pairs of chromosomes, compared to 24 for the other apes. Following their split with chimpanzees and bonobos, the hominins diversified into many species and at least two distinct genera. All but one of these lineages – representing the genus Homo and its sole extant species Homo sapiens  – are now extinct. The genus Homo evolved from Australopithecus . Though fossils from

25116-459: The late Pleistocene and early Holocene. These extinctions can be mostly attributed to people. In America, it coincided with the arrival of the Clovis people; this culture was known for " Clovis points " which were fashioned on spears for hunting animals. Shrubs, herbs, and mosses had also changed in relative abundance from the Pleistocene to Holocene, identified by permafrost core samples. Throughout

25298-409: The logic that, if groups B and C have more similarities to each other than either has to group A, then B and C are more closely related to each other than either is to A. Characters that are compared may be anatomical , such as the presence of a notochord , or molecular , by comparing sequences of DNA or proteins . The result of a successful analysis is a hierarchy of clades – groups that share

25480-714: The mid-Holocene (8.2 - 4.2 k cal BP). Climate change on seasonality and available moisture also allowed for favorable agricultural conditions which promoted human development for Maya and Tiwanaku regions. In the Korean Peninsula , climatic changes fostered a population boom during the Middle Chulmun period from 5,500 to 5,000 BP, but contributed to a subsequent bust during the Late and Final Chulmun periods, from 5,000 to 4,000 BP and from 4,000 to 3,500 BP respectively. The Holocene extinction , otherwise referred to as

25662-501: The most common and widespread species of primate , and the last surviving species of the genus Homo . They are great apes characterized by their hairlessness , bipedalism , and high intelligence . Humans have large brains , enabling more advanced cognitive skills that enable them to thrive and adapt in varied environments, develop highly complex tools , and form complex social structures and civilizations . Humans are highly social , with individual humans tending to belong to

25844-409: The most informative type of evidence. The most common types are wood, bones, and shells. Fossilisation is a rare event, and most fossils are destroyed by erosion or metamorphism before they can be observed. Hence the fossil record is very incomplete, increasingly so further back in time. Despite this, it is often adequate to illustrate the broader patterns of life's history. There are also biases in

26026-582: The most powerful factor affecting surface processes. The sedimentary record from Aitoliko Lagoon indicates that wet winters locally predominated from 210 to 160 BP, followed by dry winter dominance from 160 to 20 BP. North Africa, dominated by the Sahara Desert in the present, was instead a savanna dotted with large lakes during the Early and Middle Holocene, regionally known as the African Humid Period (AHP). The northward migration of

26208-509: The ongoing mass extinction of other forms of life. Within the last century, humans have explored challenging environments such as Antarctica , the deep sea , and outer space . Human habitation within these hostile environments is restrictive and expensive, typically limited in duration, and restricted to scientific , military , or industrial expeditions. Humans have visited the Moon and made their presence known on other celestial bodies through human-made robotic spacecraft . Since

26390-442: The only primates to have short, relatively flush canine teeth . Humans have characteristically crowded teeth, with gaps from lost teeth usually closing up quickly in young individuals. Humans are gradually losing their third molars , with some individuals having them congenitally absent. Humans share with chimpanzees a vestigial tail, appendix , flexible shoulder joints, grasping fingers and opposable thumbs . Humans also have

26572-534: The origin of cycles identified in the North Atlantic region. Climate cyclicity through the Holocene ( Bond events ) has been observed in or near marine settings and is strongly controlled by glacial input to the North Atlantic. Periodicities of ≈2500, ≈1500, and ≈1000 years are generally observed in the North Atlantic. At the same time spectral analyses of the continental record, which is remote from oceanic influence, reveal persistent periodicities of 1,000 and 500 years that may correspond to solar activity variations during

26754-414: The outcome of events such as mutations and horizontal gene transfer , which provide genetic variation , with genetic drift and natural selection driving changes in this variation over time. Within the next few years the role and operation of DNA in genetic inheritance were discovered, leading to what is now known as the "Central Dogma" of molecular biology . In the 1960s molecular phylogenetics ,

26936-474: The palm of the hand and on the soles of the feet. It is estimated that the worldwide average height for an adult human male is about 171 cm (5 ft 7 in), while the worldwide average height for adult human females is about 159 cm (5 ft 3 in). Shrinkage of stature may begin in middle age in some individuals but tends to be typical in the extremely aged . Throughout history, human populations have universally become taller, probably as

27118-501: The period exceeds any likely tectonic uplift of non-glacial origin. Post-glacial rebound in the Scandinavia region resulted in a shrinking Baltic Sea . The region continues to rise, still causing weak earthquakes across Northern Europe. An equivalent event in North America was the rebound of Hudson Bay , as it shrank from its larger, immediate post-glacial Tyrrell Sea phase, to its present boundaries. The climate throughout

27300-515: The population density varies from one region to another, and large stretches of surface are almost completely uninhabited, like Antarctica and vast swathes of the ocean. Most humans (61%) live in Asia; the remainder live in the Americas (14%), Africa (14%), Europe (11%), and Oceania (0.5%). Estimates of the population at the time agriculture emerged in around 10,000 BC have ranged between 1 million and 15 million. Around 50–60 million people lived in

27482-512: The present), as proposed by International Commission on Stratigraphy . The oldest age, the Greenlandian, was characterized by a warming following the preceding ice age. The Northgrippian Age is known for vast cooling due to a disruption in ocean circulations that was caused by the melting of glaciers. The most recent age of the Holocene is the present Meghalayan, which began with extreme drought that lasted around 200 years. The word Holocene

27664-452: The principal types of evidence about ancient life, and geochemical evidence has helped to decipher the evolution of life before there were organisms large enough to leave body fossils. Estimating the dates of these remains is essential but difficult: sometimes adjacent rock layers allow radiometric dating , which provides absolute dates that are accurate to within 0.5%, but more often paleontologists have to rely on relative dating by solving

27846-432: The radioactive element was incorporated into the rock. Radioactive elements are common only in rocks with a volcanic origin, and so the only fossil-bearing rocks that can be dated radiometrically are a few volcanic ash layers. Consequently, paleontologists must usually rely on stratigraphy to date fossils. Stratigraphy is the science of deciphering the "layer-cake" that is the sedimentary record, and has been compared to

28028-608: The region. Sand dune evolution in the Bayanbulak Basin shows that the region was very dry from the Holocene's beginning until around 6,500 BP, when a wet interval began. In the Tibetan Plateau, the moisture optimum spanned from around 7,500 to 5,500 BP. The Tarim Basin records the onset of significant aridification around 3,000-2,000 BP. After 11,800 BP, and especially between 10,800 and 9,200 BP, Ladakh experienced tremendous moisture increase most likely related to

28210-734: The residual glacial ice remaining in the Northern Hemisphere until the later date. The first major phase of Holocene climate was the Preboreal . At the start of the Preboreal occurred the Preboreal Oscillation (PBO). The Holocene Climatic Optimum (HCO) was a period of warming throughout the globe but was not globally synchronous and uniform. Following the HCO, the global climate entered a broad trend of very gradual cooling known as Neoglaciation , which lasted from

28392-701: The same approach as historical scientists: construct a set of hypotheses about the causes and then look for a "smoking gun". Paleontology lies between biology and geology since it focuses on the record of past life, but its main source of evidence is fossils in rocks. For historical reasons, paleontology is part of the geology department at many universities: in the 19th and early 20th centuries, geology departments found fossil evidence important for dating rocks, while biology departments showed little interest. Paleontology also has some overlap with archaeology , which primarily works with objects made by humans and with human remains, while paleontologists are interested in

28574-400: The sea. For example, marine fossils from the Holocene epoch have been found in locations such as Vermont and Michigan . Other than higher-latitude temporary marine incursions associated with glacial depression, Holocene fossils are found primarily in lakebed, floodplain , and cave deposits. Holocene marine deposits along low-latitude coastlines are rare because the rise in sea levels during

28756-404: The second ( Southern Dispersal ) around 70,000 to 50,000 years ago. H. sapiens proceeded to colonize all the continents and larger islands, arriving in Eurasia 125,000 years ago, Australia around 65,000 years ago, the Americas around 15,000 years ago, and remote islands such as Hawaii , Easter Island , Madagascar , and New Zealand in the years 300 to 1280 CE. Human evolution was not

28938-547: The sense of humanity). The native English term man can refer to the species generally (a synonym for humanity ) as well as to human males. It may also refer to individuals of either sex. Despite the fact that the word animal is colloquially used as an antonym for human , and contrary to a common biological misconception , humans are animals . The word person is often used interchangeably with human , but philosophical debate exists as to whether personhood applies to all humans or all sentient beings , and further if

29120-478: The similarity of the DNA in their genomes . Molecular phylogenetics has also been used to estimate the dates when species diverged, but there is controversy about the reliability of the molecular clock on which such estimates depend. The simplest definition of "paleontology" is "the study of ancient life". The field seeks information about several aspects of past organisms: "their identity and origin, their environment and evolution, and what they can tell us about

29302-470: The slow recovery from this catastrophe a previously obscure group, archosaurs , became the most abundant and diverse terrestrial vertebrates. One archosaur group, the dinosaurs, were the dominant land vertebrates for the rest of the Mesozoic , and birds evolved from one group of dinosaurs. During this time mammals' ancestors survived only as small, mainly nocturnal insectivores , which may have accelerated

29484-697: The strengthening of the Indian Summer Monsoon (ISM). From 9,200 to 6,900 BP, relative aridity persisted in Ladakh. A second major humid phase occurred in Ladakh from 6,900 to 4,800 BP, after which the region was again arid. From 900 to 1,200 AD, during the MWP, the ISM was again strong as evidenced by low δO values from the Ganga Plain. The sediments of Lonar Lake in Maharashtra record dry conditions around 11,400 BP that transitioned into

29666-625: The study of fossils to classify organisms and study their interactions with each other and their environments (their paleoecology ). Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier 's work on comparative anatomy , and developed rapidly in the 19th century. The term has been used since 1822 formed from Greek παλαιός ( 'palaios' , "old, ancient"), ὄν ( 'on' , ( gen. 'ontos' ), "being, creature"), and λόγος ( 'logos' , "speech, thought, study"). Paleontology lies on

29848-511: The subtropical front (STF), was around 2 degrees Celsius during the HCO. This temperature gradient is significantly less than modern times, which is around 6 degrees Celsius. A study utilizing five SST proxies from 37°S to 60°S latitude confirmed that the strong temperature gradient was confined to the area immediately south of the STF, and is correlated with reduced westerly winds near New Zealand. Since 7,100 BP, New Zealand experienced 53 cyclones similar in magnitude to Cyclone Bola . Evidence from

30030-510: The systematic study of fossils emerged as an integral part of the changes in natural philosophy that occurred during the Age of Reason . In the Italian Renaissance, Leonardo da Vinci made various significant contributions to the field as well as depicted numerous fossils. Leonardo's contributions are central to the history of paleontology because he established a line of continuity between the two main branches of paleontology – ichnology and body fossil paleontology. He identified

30212-433: The term "humans" with all members of the genus Homo , in common usage it generally refers to Homo sapiens , the only extant member. All other members of the genus Homo , which are now extinct, are known as archaic humans , and the term "modern human" is used to distinguish Homo sapiens from archaic humans. Anatomically modern humans emerged around 300,000 years ago in Africa, evolving from Homo heidelbergensis or

30394-399: The term 'recent' as an incorrect way of referring to the Holocene, preferring the term 'modern' instead to describe current processes. It also observes that the term 'Flandrian' may be used as a synonym for Holocene, although it is becoming outdated. The International Commission on Stratigraphy, however, considers the Holocene to be an epoch following the Pleistocene and specifically following

30576-427: The transition are scarce, the earliest members of Homo share several key traits with Australopithecus . The earliest record of Homo is the 2.8 million-year-old specimen LD 350-1 from Ethiopia , and the earliest named species are Homo habilis and Homo rudolfensis which evolved by 2.3 million years ago. H. erectus (the African variant is sometimes called H. ergaster ) evolved 2 million years ago and

30758-428: The vast majority of these extinctions are thought to be undocumented, as the species are undiscovered at the time of their extinction, or no one has yet discovered their extinction. The current rate of extinction of species is estimated at 100 to 1,000 times higher than natural background extinction rates . Human species Humans ( Homo sapiens , meaning "thinking man" or "wise man") or modern humans are

30940-406: The word "palaeontology" to refer to the study of ancient living organisms through fossils. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. This encouraged early evolutionary theories on the transmutation of species . After Charles Darwin published Origin of Species in 1859, much of

31122-440: The world's 3.4 billion rural population. Problems for humans living in cities include various forms of pollution and crime , especially in inner city and suburban slums . Most aspects of human physiology are closely homologous to corresponding aspects of animal physiology. The dental formula of humans is: 2.1.2.3 2.1.2.3 . Humans have proportionately shorter palates and much smaller teeth than other primates. They are

31304-408: The world, ecosystems in cooler climates that were previously regional have been isolated in higher altitude ecological "islands". The 8.2-ka event , an abrupt cold spell recorded as a negative excursion in the δ O record lasting 400 years, is the most prominent climatic event occurring in the Holocene Epoch, and may have marked a resurgence of ice cover. It has been suggested that this event

31486-448: The world. This ' Neolithic Revolution ', likely influenced by Holocene climatic changes, included an increase in sedentism and population, eventually resulting in the world's first large-scale state societies in Mesopotamia and Egypt . During the Middle Holocene, the Intertropical Convergence Zone , which governs the incursion of monsoon precipitation through the Arabian Peninsula, shifted southwards, resulting in increased aridity. In

31668-454: Was "seeded" from elsewhere , but most research concentrates on various explanations of how life could have arisen independently on Earth. For about 2,000 million years microbial mats , multi-layered colonies of different bacteria, were the dominant life on Earth. The evolution of oxygenic photosynthesis enabled them to play the major role in the oxygenation of the atmosphere from about 2,400  million years ago . This change in

31850-450: Was caused by the final drainage of Lake Agassiz , which had been confined by the glaciers, disrupting the thermohaline circulation of the Atlantic . This disruption was the result of an ice dam over Hudson Bay collapsing sending cold lake Agassiz water into the North Atlantic ocean . Furthermore, studies show that the melting of Lake Agassiz led to sea-level rise which flooded the North American coastal landscape. The basal peat plant

32032-413: Was defined for Northern Europe , but the climate changes were claimed to occur more widely. The periods of the scheme include a few of the final pre-Holocene oscillations of the last glacial period and then classify climates of more recent prehistory . Paleontologists have not defined any faunal stages for the Holocene. If subdivision is necessary, periods of human technological development, such as

32214-520: Was formed from two Ancient Greek words. Hólos ( ὅλος ) is the Greek word for "whole". "Cene" comes from the Greek word kainós ( καινός ), meaning "new". The concept is that this epoch is "entirely new". The suffix '-cene' is used for all the seven epochs of the Cenozoic Era. The International Commission on Stratigraphy has defined the Holocene as starting approximately 11,700 years before 2000 CE (11,650 cal years BP , or 9,700 BCE). The Subcommission on Quaternary Stratigraphy (SQS) regards

32396-429: Was never a formally defined geological unit. The Subcommission on Quaternary Stratigraphy (SQS) of the International Commission on Stratigraphy (ICS) had a working group to determine whether it should be. In May 2019, members of the working group voted in favour of recognizing the Anthropocene as formal chrono-stratigraphic unit, with stratigraphic signals around the mid-twentieth century CE as its base. The exact criteria

32578-428: Was still to be determined, after which the recommendation also had to be approved by the working group's parent bodies (ultimately the International Union of Geological Sciences ). In March 2024, after 15 years of deliberation, the Anthropocene Epoch proposal of the working group was voted down by a wide margin by the SQS, owing largely to its shallow sedimentary record and extremely recent proposed start date. The ICS and

32760-424: Was the coldest interval of time of the past two millennia. Following the Industrial Revolution, warm decadal intervals became more common relative to before as a consequence of anthropogenic greenhouse gases, resulting in progressive global warming. In the late 20th century, anthropogenic forcing superseded variations in solar activity as the dominant driver of climate change, though solar activity has continued to play

32942-402: Was the first archaic human species to leave Africa and disperse across Eurasia. H. erectus also was the first to evolve a characteristically human body plan . Homo sapiens emerged in Africa around 300,000 years ago from a species commonly designated as either H. heidelbergensis or H. rhodesiensis , the descendants of H. erectus that remained in Africa. H. sapiens migrated out of

33124-442: Was then used to determine the resulting local sea-level rise of 0.20-0.56m in the Mississippi Delta . Subsequent research, however, suggested that the discharge was probably superimposed upon a longer episode of cooler climate lasting up to 600 years and observed that the extent of the area affected was unclear. The beginning of the Holocene corresponds with the beginning of the Mesolithic age in most of Europe . In regions such as

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