68-569: The Living Planet Index ( LPI ) is an indicator of the state of global biological diversity , based on trends in vertebrate populations of species from around the world. The Zoological Society of London (ZSL) manages the index in cooperation with the World Wide Fund for Nature (WWF). As of 2022, the index is statistically created from journal studies, online databases and government reports for 31,821 populations of 5,230 species of mammal, bird, reptile, amphibian and fish. According to
136-508: A weighted geometric mean method which places most weight on the largest (most species-rich) groups within a biogeographic realm. This is done to counteract the uneven spatial and taxonomic distribution of data in the LPD. The three system indices are then averaged to produce the global LPI. The fact that "all decreases in population size, regardless of whether they bring a population close to extinction, are equally accounted for" has been noted as
204-489: A 10% increase in biodiversity, which was canceled out by a loss in low-income countries. This is despite the fact that high-income countries use five times the ecological resources of low-income countries, which was explained as a result of a process whereby wealthy nations are outsourcing resource depletion to poorer nations, which are suffering the greatest ecosystem losses. A 2017 study published in PLOS One found that
272-501: A country, endangered species are initially supported on a national level then internationally. Ecotourism may be utilized to support the economy and encourages tourists to continue to visit and support species and ecosystems they visit, while they enjoy the available amenities provided. International biodiversity impacts global livelihood, food systems, and health. Problematic pollution, over consumption, and climate change can devastate international biodiversity. Nature-based solutions are
340-517: A critical tool for a global resolution. Many species are in danger of becoming extinct and need world leaders to be proactive with the Kunming-Montreal Global Biodiversity Framework . Terrestrial biodiversity is thought to be up to 25 times greater than ocean biodiversity. Forests harbour most of Earth's terrestrial biodiversity. The conservation of the world's biodiversity is thus utterly dependent on
408-530: A decline between the 1980s and 2000s, but a roughly positive trend after 2000. This extreme sensitivity to outliers indicates that the present approach of the Living Planet Index may be flawed. A 2024 study by Charles University focused on calculation method, found that calculation of the Living Planet Index is biased by several mathematical issues, leading to overestimation of vertebrate population declines. When those mathematical issues are fixed,
476-455: A first-order positive feedback (more ancestors, more descendants) and/or a negative feedback arising from resource limitation. Hyperbolic model implies a second-order positive feedback. Differences in the strength of the second-order feedback due to different intensities of interspecific competition might explain the faster rediversification of ammonoids in comparison to bivalves after the end-Permian extinction . The hyperbolic pattern of
544-460: A limit would also cap the number of species. While records of life in the sea show a logistic pattern of growth, life on land (insects, plants and tetrapods) shows an exponential rise in diversity. As one author states, "Tetrapods have not yet invaded 64 percent of potentially habitable modes and it could be that without human influence the ecological and taxonomic diversity of tetrapods would continue to increase exponentially until most or all of
612-485: A limitation. In 2005, WWF authors identified that the population data was potentially unrepresentative. As of 2009, the database was found to contain too much bird data and gaps in the population coverage of tropical species, although it showed "little evidence of bias toward threatened species". The 2016 report was criticized by a professor at Duke University for over-representing western Europe, where more data were available. Talking to National Geographic , he criticised
680-947: A mandate to facilitate the delivery of the global indicators under the CBD's 2010 Biodiversity Target on the rate of loss of biological diversity and works alongside the CITES Secretariat producing a range of reports and databases. It also manages the World Database of Protected Areas in collaboration with the IUCN World Commission on Protected Areas . A series of world atlases on biodiversity topics have been published by UNEP-WCMC through University of California Press . UNEP-WCMC has expertise across six thematic areas: These thematic areas are supported by cross-cutting expertise in science, economics, knowledge management and digital innovation. UNEP-WCMC has created
748-409: A re-analysis of the baseline data by McGill University showed that the overall estimated trend of a decline by 60% since 1970 was driven by less than 3% of the studied populations; when some outliers of extreme decline are removed, the decline still exists but is considerably less catastrophic, and when more outliers (roughly amounting to 2.4% of the populations) are removed, the trend shifts to that of
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#1732766272799816-405: A report saying that "biodiversity is being destroyed at a rate unprecedented in human history". The report claims that 68% of the population of the examined species were destroyed in the years 1970 – 2016. Of 70,000 monitored species, around 48% are experiencing population declines from human activity (in 2023), whereas only 3% have increasing populations. Rates of decline in biodiversity in
884-574: A smaller scale. The latest edition of the Living Planet Report was released in October 2022. The index is often misinterpreted in the media, with incorrect suggestions that it shows we have lost 69% of all animals or species since 1970. This widespread misinterpretation has led to several articles being published which detail what the LPI does and doesn't show, and how to correctly interpret
952-590: A third of the Earth's land mass) and are home to approximately 80% of the world's biodiversity. About 1 billion hectares are covered by primary forests. Over 700 million hectares of the world's woods are officially protected. The biodiversity of forests varies considerably according to factors such as forest type, geography, climate and soils – in addition to human use. Most forest habitats in temperate regions support relatively few animal and plant species and species that tend to have large geographical distributions, while
1020-399: A variety of sources such as journals, online databases and government reports. A generalized additive modelling framework is used to determine the underlying trend in each population time-series. Average rates of change are calculated and aggregated to the species level. Each species trend is aggregated to produce an index for the terrestrial, marine and freshwater systems. This process uses
1088-1035: Is 'planned' diversity or 'associated' diversity. This is a functional classification that we impose and not an intrinsic feature of life or diversity. Planned diversity includes the crops which a farmer has encouraged, planted or raised (e.g. crops, covers, symbionts, and livestock, among others), which can be contrasted with the associated diversity that arrives among the crops, uninvited (e.g. herbivores, weed species and pathogens, among others). Associated biodiversity can be damaging or beneficial. The beneficial associated biodiversity include for instance wild pollinators such as wild bees and syrphid flies that pollinate crops and natural enemies and antagonists to pests and pathogens. Beneficial associated biodiversity occurs abundantly in crop fields and provide multiple ecosystem services such as pest control, nutrient cycling and pollination that support crop production. World Conservation Monitoring Centre The UN Environment Programme World Conservation Monitoring Centre (UNEP-WCMC)
1156-405: Is an increase in biodiversity from the poles to the tropics . Thus localities at lower latitudes have more species than localities at higher latitudes . This is often referred to as the latitudinal gradient in species diversity. Several ecological factors may contribute to the gradient, but the ultimate factor behind many of them is the greater mean temperature at the equator compared to that at
1224-491: Is estimated at 5.0 x 10 and weighs 50 billion tonnes . In comparison, the total mass of the biosphere has been estimated to be as much as four trillion tons of carbon . In July 2016, scientists reported identifying a set of 355 genes from the last universal common ancestor (LUCA) of all organisms living on Earth. The age of Earth is about 4.54 billion years. The earliest undisputed evidence of life dates at least from 3.7 billion years ago, during
1292-402: Is greater now than at any time in human history, with extinctions occurring at rates hundreds of times higher than background extinction rates. and expected to still grow in the upcoming years. As of 2012, some studies suggest that 25% of all mammal species could be extinct in 20 years. In absolute terms, the planet has lost 58% of its biodiversity since 1970 according to a 2016 study by
1360-427: Is local biodiversity, which directly impacts daily life, affecting the availability of fresh water, food choices, and fuel sources for humans. Regional biodiversity includes habitats and ecosystems that synergizes and either overlaps or differs on a regional scale. National biodiversity within a country determines the ability for a country to thrive according to its habitats and ecosystems on a national scale. Also, within
1428-458: Is no concrete definition for biodiversity, as its definition continues to be defined. Other definitions include (in chronological order): According to estimates by Mora et al. (2011), there are approximately 8.7 million terrestrial species and 2.2 million oceanic species. The authors note that these estimates are strongest for eukaryotic organisms and likely represent the lower bound of prokaryote diversity. Other estimates include: Since
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#17327662727991496-401: Is possible to build fractal hyper volumes, whose fractal dimension rises to three moving towards the equator . A biodiversity hotspot is a region with a high level of endemic species that have experienced great habitat loss . The term hotspot was introduced in 1988 by Norman Myers . While hotspots are spread all over the world, the majority are forest areas and most are located in
1564-561: Is so full, that that district produces the most variety which is the most examined." Biodiversity is the result of 3.5 billion years of evolution . The origin of life has not been established by science, however, some evidence suggests that life may already have been well-established only a few hundred million years after the formation of the Earth . Until approximately 2.5 billion years ago, all life consisted of microorganisms – archaea , bacteria , and single-celled protozoans and protists . Biodiversity grew fast during
1632-586: Is the specialist biodiversity centre of UN Environment Programme , based in Cambridge in the United Kingdom . UNEP-WCMC has been part of UN Environment Programme since 2000 and has responsibility for biodiversity assessment and support to policy development and implementation. The "World Conservation Monitoring Centre" was previously an independent organisation jointly managed by IUCN , UN Environment Programme and WWF established in 1988. Before that,
1700-479: Is the variability of life on Earth . It can be measured on various levels. There is for example genetic variability , species diversity , ecosystem diversity and phylogenetic diversity. Diversity is not distributed evenly on Earth . It is greater in the tropics as a result of the warm climate and high primary productivity in the region near the equator . Tropical forest ecosystems cover less than one-fifth of Earth's terrestrial area and contain about 50% of
1768-405: Is uncertainty as to how strongly the fossil record is biased by the greater availability and preservation of recent geologic sections. Some scientists believe that corrected for sampling artifacts, modern biodiversity may not be much different from biodiversity 300 million years ago, whereas others consider the fossil record reasonably reflective of the diversification of life. Estimates of
1836-619: The Aichi Biodiversity Targets ). Informing the CBD 2020 strategic plan, the Indicators and Assessments Unit at ZSL is concerned with ensuring the most rigorous and robust methods are implemented for the measurement of population trends , expanding the coverage of the LPI to more broadly represent biodiversity, and disaggregating the index in meaningful ways (such as assessing the changes in exploited or invasive species ). Biological diversity Biodiversity
1904-513: The Carboniferous , rainforest collapse may have led to a great loss of plant and animal life. The Permian–Triassic extinction event , 251 million years ago, was the worst; vertebrate recovery took 30 million years. Human activities have led to an ongoing biodiversity loss and an accompanying loss of genetic diversity . This process is often referred to as Holocene extinction , or sixth mass extinction . For example, it
1972-792: The Eoarchean era after a geological crust started to solidify following the earlier molten Hadean eon. There are microbial mat fossils found in 3.48 billion-year-old sandstone discovered in Western Australia . Other early physical evidence of a biogenic substance is graphite in 3.7 billion-year-old meta-sedimentary rocks discovered in Western Greenland .. More recently, in 2015, "remains of biotic life " were found in 4.1 billion-year-old rocks in Western Australia . According to one of
2040-713: The Phanerozoic (the last 540 million years), especially during the so-called Cambrian explosion —a period during which nearly every phylum of multicellular organisms first appeared. However, recent studies suggest that this diversification had started earlier, at least in the Ediacaran , and that it continued in the Ordovician . Over the next 400 million years or so, invertebrate diversity showed little overall trend and vertebrate diversity shows an overall exponential trend. This dramatic rise in diversity
2108-461: The Stone Age , species loss has accelerated above the average basal rate, driven by human activity. Estimates of species losses are at a rate 100–10,000 times as fast as is typical in the fossil record. Loss of biodiversity results in the loss of natural capital that supplies ecosystem goods and services . Species today are being wiped out at a rate 100 to 1,000 times higher than baseline, and
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2176-846: The United Nations Environment Programme . WWF first published the index in 1998. Since 2006, the Zoological Society of London (ZSL) manages the index in cooperation with WWF. Results are presented biennially in the WWF Living Planet Report and in publications such as the Millennium Ecosystem Assessment and the UN Global Biodiversity Outlook. National and regional reports are now being produced to focus on relevant issues at
2244-504: The tropics . Brazil 's Atlantic Forest is considered one such hotspot, containing roughly 20,000 plant species, 1,350 vertebrates and millions of insects, about half of which occur nowhere else. The island of Madagascar and India are also particularly notable. Colombia is characterized by high biodiversity, with the highest rate of species by area unit worldwide and it has the largest number of endemics (species that are not found naturally anywhere else) of any country. About 10% of
2312-417: The world population growth arises from a second-order positive feedback between the population size and the rate of technological growth. The hyperbolic character of biodiversity growth can be similarly accounted for by a feedback between diversity and community structure complexity. The similarity between the curves of biodiversity and human population probably comes from the fact that both are derived from
2380-558: The 2022 report, monitored wildlife populations show an average decline of 69% between 1970 and 2018, suggesting that natural ecosystems are degrading at a rate unprecedented in human history The extent of declines varies with geographic region, with monitored vertebrate populations in Latin America and the Caribbean experiencing average declines of 94%. One of the key drivers of declines has been identified as land-use change and
2448-606: The IUCN's critically endangered . Numerous scientists and the IPBES Global Assessment Report on Biodiversity and Ecosystem Services assert that human population growth and overconsumption are the primary factors in this decline. However, other scientists have criticized this finding and say that loss of habitat caused by "the growth of commodities for export" is the main driver. Some studies have however pointed out that habitat destruction for
2516-582: The World Wildlife Fund. The Living Planet Report 2014 claims that "the number of mammals, birds, reptiles, amphibians, and fish across the globe is, on average, about half the size it was 40 years ago". Of that number, 39% accounts for the terrestrial wildlife gone, 39% for the marine wildlife gone and 76% for the freshwater wildlife gone. Biodiversity took the biggest hit in Latin America , plummeting 83 percent. High-income countries showed
2584-462: The associated habitat loss and degradation, often linked to unsustainable agriculture, logging, or other development. The Living Planet Database ( LPD ) has been available online since 2013, and has been maintained by ZSL since 2016. The LPD contains more than 30,000 population trends for more than 5,200 species of fish, amphibians, reptiles, birds and mammals. The global LPI is calculated using these population time-series, which are gathered from
2652-479: The attempt to combine data from different regions and ecosystems into a single figure, arguing that such reports are likely motivated by a desire to grab attention and raise money. A 2017 investigation of the index by members of the ZSL team published in PLOS One found higher declines than had been estimated, and indications that in areas where less data is available, species might be declining more quickly. In 2020,
2720-461: The available eco-space is filled." It also appears that the diversity continues to increase over time, especially after mass extinctions. On the other hand, changes through the Phanerozoic correlate much better with the hyperbolic model (widely used in population biology , demography and macrosociology , as well as fossil biodiversity) than with exponential and logistic models. The latter models imply that changes in diversity are guided by
2788-553: The biomass of insect life in Germany had declined by three-quarters in the last 25 years. Dave Goulson of Sussex University stated that their study suggested that humans "appear to be making vast tracts of land inhospitable to most forms of life, and are currently on course for ecological Armageddon. If we lose the insects then everything is going to collapse." In 2020 the World Wildlife Foundation published
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2856-625: The centre was a part of the IUCN Secretariat. The activities of UNEP-WCMC include biodiversity assessment, support to international conventions such as the Convention on Biological Diversity (CBD) and the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES), capacity building and management of both aspatial and spatial data on species and habitats of conservation concern. UNEP-WCMC has
2924-540: The current sixth mass extinction match or exceed rates of loss in the five previous mass extinction events in the fossil record . Biodiversity loss is in fact "one of the most critical manifestations of the Anthropocene " (since around the 1950s); the continued decline of biodiversity constitutes "an unprecedented threat" to the continued existence of human civilization. The reduction is caused primarily by human impacts , particularly habitat destruction . Since
2992-512: The end of the Maastrichtian , just before that extinction event. However, many other taxa were affected by this crisis, which affected even marine taxa, such as ammonites , which also became extinct around that time. The biodiversity of the past is called Paleobiodiversity. The fossil record suggests that the last few million years featured the greatest biodiversity in history . However, not all scientists support this view, since there
3060-481: The end of the last ice age partly resulted from overhunting. Biologists most often define biodiversity as the "totality of genes , species and ecosystems of a region". An advantage of this definition is that it presents a unified view of the traditional types of biological variety previously identified: Biodiversity is most commonly used to replace the more clearly-defined and long-established terms, species diversity and species richness . However, there
3128-451: The estimated global value of ecosystem services (not captured in traditional markets) at an average of $ 33 trillion annually. With regards to provisioning services, greater species diversity has the following benefits: With regards to regulating services, greater species diversity has the following benefits: Greater species diversity Agricultural diversity can be divided into two categories: intraspecific diversity , which includes
3196-428: The expansion of agriculture and the overexploitation of wildlife are the more significant drivers of contemporary biodiversity loss, not climate change . Biodiversity is not evenly distributed, rather it varies greatly across the globe as well as within regions and seasons. Among other factors, the diversity of all living things ( biota ) depends on temperature , precipitation , altitude , soils , geography and
3264-491: The genetic variation within a single species, like the potato ( Solanum tuberosum ) that is composed of many different forms and types (e.g. in the U.S. they might compare russet potatoes with new potatoes or purple potatoes, all different, but all part of the same species, S. tuberosum ). The other category of agricultural diversity is called interspecific diversity and refers to the number and types of different species. Agricultural diversity can also be divided by whether it
3332-596: The interactions between other species. The study of the spatial distribution of organisms , species and ecosystems , is the science of biogeography . Diversity consistently measures higher in the tropics and in other localized regions such as the Cape Floristic Region and lower in polar regions generally. Rain forests that have had wet climates for a long time, such as Yasuní National Park in Ecuador , have particularly high biodiversity. There
3400-737: The interference of the hyperbolic trend with cyclical and stochastic dynamics. Most biologists agree however that the period since human emergence is part of a new mass extinction, named the Holocene extinction event , caused primarily by the impact humans are having on the environment. It has been argued that the present rate of extinction is sufficient to eliminate most species on the planet Earth within 100 years. New species are regularly discovered (on average between 5–10,000 new species each year, most of them insects ) and many, though discovered, are not yet classified (estimates are that nearly 90% of all arthropods are not yet classified). Most of
3468-495: The majority of studied vertebrate populations shows balanced decline and growth (the only exception are the populations of amphibians , where the numbers show steady decline). The index was originally developed in 1997 by the World Wide Fund for Nature (WWF) in collaboration with the World Conservation Monitoring Centre (UNEP-WCMC), the biodiversity assessment and policy implementation arm of
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#17327662727993536-820: The montane forests of Africa, South America and Southeast Asia and lowland forests of Australia, coastal Brazil, the Caribbean islands, Central America and insular Southeast Asia have many species with small geographical distributions. Areas with dense human populations and intense agricultural land use, such as Europe , parts of Bangladesh, China, India and North America, are less intact in terms of their biodiversity. Northern Africa, southern Australia, coastal Brazil, Madagascar and South Africa, are also identified as areas with striking losses in biodiversity intactness. European forests in EU and non-EU nations comprise more than 30% of Europe's land mass (around 227 million hectares), representing an almost 10% growth since 1990. Generally, there
3604-468: The planet's species went extinct prior to the evolution of humans. Estimates on the number of Earth's current species range from 10 million to 14 million, of which about 1.2 million have been documented and over 86% have not yet been described. However, a May 2016 scientific report estimates that 1 trillion species are currently on Earth, with only one-thousandth of one percent described. The total amount of related DNA base pairs on Earth
3672-546: The poles. Even though terrestrial biodiversity declines from the equator to the poles, some studies claim that this characteristic is unverified in aquatic ecosystems , especially in marine ecosystems . The latitudinal distribution of parasites does not appear to follow this rule. Also, in terrestrial ecosystems the soil bacterial diversity has been shown to be highest in temperate climatic zones, and has been attributed to carbon inputs and habitat connectivity. In 2016, an alternative hypothesis ("the fractal biodiversity")
3740-412: The present global macroscopic species diversity vary from 2 million to 100 million, with a best estimate of somewhere near 9 million, the vast majority arthropods . Diversity appears to increase continually in the absence of natural selection. The existence of a global carrying capacity , limiting the amount of life that can live at once, is debated, as is the question of whether such
3808-729: The rate of extinction has increased, many extant species may become extinct before they are described. Not surprisingly, in the animalia the most studied groups are birds and mammals , whereas fishes and arthropods are the least studied animals groups. During the last century, decreases in biodiversity have been increasingly observed. It was estimated in 2007 that up to 30% of all species will be extinct by 2050. Of these, about one eighth of known plant species are threatened with extinction . Estimates reach as high as 140,000 species per year (based on Species-area theory ). This figure indicates unsustainable ecological practices, because few species emerge each year. The rate of species loss
3876-584: The rate of extinctions is increasing. This process destroys the resilience and adaptability of life on Earth. In 2006, many species were formally classified as rare or endangered or threatened ; moreover, scientists have estimated that millions more species are at risk which have not been formally recognized. About 40 percent of the 40,177 species assessed using the IUCN Red List criteria are now listed as threatened with extinction —a total of 16,119. As of late 2022 9251 species were considered part of
3944-913: The researchers, "If life arose relatively quickly on Earth...then it could be common in the universe ." There have been many claims about biodiversity's effect on the ecosystem services , especially provisioning and regulating services . Some of those claims have been validated, some are incorrect and some lack enough evidence to draw definitive conclusions. Ecosystem services have been grouped in three types: Experiments with controlled environments have shown that humans cannot easily build ecosystems to support human needs; for example insect pollination cannot be mimicked, though there have been attempts to create artificial pollinators using unmanned aerial vehicles . The economic activity of pollination alone represented between $ 2.1–14.6 billion in 2003. Other sources have reported somewhat conflicting results and in 1997 Robert Costanza and his colleagues reported
4012-693: The species of the Earth can be found in Colombia, including over 1,900 species of bird, more than in Europe and North America combined, Colombia has 10% of the world's mammals species, 14% of the amphibian species and 18% of the bird species of the world. Madagascar dry deciduous forests and lowland rainforests possess a high ratio of endemism . Since the island separated from mainland Africa 66 million years ago, many species and ecosystems have evolved independently. Indonesia 's 17,000 islands cover 735,355 square miles (1,904,560 km ) and contain 10% of
4080-419: The terrestrial diversity is found in tropical forests and in general, the land has more species than the ocean; some 8.7 million species may exist on Earth, of which some 2.1 million live in the ocean. It is estimated that 5 to 50 billion species have existed on the planet. Assuming that there may be a maximum of about 50 million species currently alive, it stands to reason that greater than 99% of
4148-498: The trend. In April 2002, and again in 2006, at the Convention on Biological Diversity (CBD), 188 nations committed themselves to actions to: "… achieve, by 2010, a significant reduction of the current rate of biodiversity loss at the global, regional and national levels…" The LPI played a pivotal role in measuring progress towards the CBD's 2010 target. It has also been adopted by the CBD as an indicator of progress towards its Nagoya Protocol 2011-2020 targets 5, 6, and 12 (part of
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#17327662727994216-773: The way in which we interact with and use the world's forests. A new method used in 2011, put the total number of species on Earth at 8.7 million, of which 2.1 million were estimated to live in the ocean. However, this estimate seems to under-represent the diversity of microorganisms. Forests provide habitats for 80 percent of amphibian species , 75 percent of bird species and 68 percent of mammal species. About 60 percent of all vascular plants are found in tropical forests. Mangroves provide breeding grounds and nurseries for numerous species of fish and shellfish and help trap sediments that might otherwise adversely affect seagrass beds and coral reefs, which are habitats for many more marine species. Forests span around 4 billion acres (nearly
4284-630: The world's flowering plants , 12% of mammals and 17% of reptiles , amphibians and birds —along with nearly 240 million people. Many regions of high biodiversity and/or endemism arise from specialized habitats which require unusual adaptations, for example, alpine environments in high mountains , or Northern European peat bogs . Accurately measuring differences in biodiversity can be difficult. Selection bias amongst researchers may contribute to biased empirical research for modern estimates of biodiversity. In 1768, Rev. Gilbert White succinctly observed of his Selborne, Hampshire "all nature
4352-652: The world's species. There are latitudinal gradients in species diversity for both marine and terrestrial taxa. Since life began on Earth , six major mass extinctions and several minor events have led to large and sudden drops in biodiversity. The Phanerozoic aeon (the last 540 million years) marked a rapid growth in biodiversity via the Cambrian explosion . In this period, the majority of multicellular phyla first appeared. The next 400 million years included repeated, massive biodiversity losses. Those events have been classified as mass extinction events. In
4420-507: Was estimated in 2007 that up to 30% of all species will be extinct by 2050. Destroying habitats for farming is a key reason why biodiversity is decreasing today. Climate change also plays a role. This can be seen for example in the effects of climate change on biomes . This anthropogenic extinction may have started toward the end of the Pleistocene , as some studies suggest that the megafaunal extinction event that took place around
4488-483: Was marked by periodic, massive losses of diversity classified as mass extinction events. A significant loss occurred in anamniotic limbed vertebrates when rainforests collapsed in the Carboniferous , but amniotes seem to have been little affected by this event; their diversification slowed down later, around the Asselian / Sakmarian boundary, in the early Cisuralian (Early Permian ), about 293 Ma ago. The worst
4556-401: Was proposed to explain the biodiversity latitudinal gradient. In this study, the species pool size and the fractal nature of ecosystems were combined to clarify some general patterns of this gradient. This hypothesis considers temperature , moisture , and net primary production (NPP) as the main variables of an ecosystem niche and as the axis of the ecological hypervolume . In this way, it
4624-567: Was the Permian-Triassic extinction event , 251 million years ago. Vertebrates took 30 million years to recover from this event. The most recent major mass extinction event, the Cretaceous–Paleogene extinction event , occurred 66 million years ago. This period has attracted more attention than others because it resulted in the extinction of the non-avian dinosaurs , which were represented by many lineages at
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