105-644: Coenonympha nipisiquit , the maritime ringlet , is a rare butterfly in the family Nymphalidae . It is a "species at risk" in Canada due to water pollution and its limited range. Its range is restricted in Canada to the Chaleur Bay region, between New Brunswick and the Gaspé Peninsula . The species was discovered near Bathurst, New Brunswick by J. McDunnough in 1939. Of the six salt marshes where
210-637: A diapause (resting) stage, and the hatching may take place only in spring. Some temperate region butterflies, such as the Camberwell beauty , lay their eggs in the spring and have them hatch in the summer. Butterfly larvae, or caterpillars, consume plant leaves and spend practically all of their time searching for and eating food. Although most caterpillars are herbivorous, a few species are predators : Spalgis epius eats scale insects , while lycaenids such as Liphyra brassolis are myrmecophilous , eating ant larvae. Some larvae, especially those of
315-584: A botanical garden, and a museum that was designed to have a national and permanent status. The former occupied space where the present Botanic Garden sits. The museum contained specimens of zoology, botany, archeology, fossils, etc., some of which were passed on to the Smithsonian Institution after its formation. The institute's charter expired in 1838, but its spirit lived on in the National Institution , founded in 1840. With
420-493: A butterfly cannot fly until the wings are unfolded. A newly emerged butterfly needs to spend some time inflating its wings with hemolymph and letting them dry, during which time it is extremely vulnerable to predators. The colourful patterns on many butterfly wings tell potential predators that they are toxic. Hence, the genetic basis of wing pattern formation can illuminate both the evolution of butterflies as well as their developmental biology . The colour of butterfly wings
525-570: A butterfly through metamorphosis has held great appeal to mankind. To transform from the miniature wings visible on the outside of the pupa into large structures usable for flight, the pupal wings undergo rapid mitosis and absorb a great deal of nutrients. If one wing is surgically removed early on, the other three will grow to a larger size. In the pupa, the wing forms a structure that becomes compressed from top to bottom and pleated from proximal to distal ends as it grows, so that it can rapidly be unfolded to its full adult size. Several boundaries seen in
630-417: A common family. In some species, such as the great spangled fritillary , the eggs are deposited close to but not on the food plant. This most likely happens when the egg overwinters before hatching and where the host plant loses its leaves in winter, as do violets in this example. The egg stage lasts a few weeks in most butterflies, but eggs laid close to winter, especially in temperate regions, go through
735-627: A conspicuous, fluttering flight. The group comprises the superfamilies Hedyloidea (moth-butterflies in the Americas) and Papilionoidea (all others). The oldest butterfly fossils have been dated to the Paleocene , about 56 million years ago, though molecular likely originated in the Cretaceous . Butterflies have a four-stage life cycle , and like other holometabolous insects they undergo complete metamorphosis . Winged adults lay eggs on
840-576: A few butterflies (e.g., harvesters ) eat harmful insects, and a few are predators of ants , while others live as mutualists in association with ants. Culturally, butterflies are a popular motif in the visual and literary arts. The Smithsonian Institution says "butterflies are certainly one of the most appealing creatures in nature". The Oxford English Dictionary derives the word straightforwardly from Old English butorflēoge , butter-fly; similar names in Old Dutch and Old High German show that
945-444: A few species. Some butterflies have organs of hearing and some species make stridulatory and clicking sounds. Many species of butterfly maintain territories and actively chase other species or individuals that may stray into them. Some species will bask or perch on chosen perches. The flight styles of butterflies are often characteristic and some species have courtship flight displays. Butterflies can only fly when their temperature
1050-606: A hard-ridged outer layer of shell, called the chorion . This is lined with a thin coating of wax which prevents the egg from drying out before the larva has had time to fully develop. Each egg contains a number of tiny funnel-shaped openings at one end, called micropyles ; the purpose of these holes is to allow sperm to enter and fertilize the egg. Butterfly eggs vary greatly in size and shape between species, but are usually upright and finely sculptured. Some species lay eggs singly, others in batches. Many females produce between one hundred and two hundred eggs. Butterfly eggs are fixed to
1155-508: A leaf before eggs are laid on it. Many butterflies use chemical signals, pheromones ; some have specialized scent scales ( androconia ) or other structures ( coremata or "hair pencils" in the Danaidae). Vision is well developed in butterflies and most species are sensitive to the ultraviolet spectrum. Many species show sexual dimorphism in the patterns of UV reflective patches. Colour vision may be widespread but has been demonstrated in only
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#17327808950541260-416: A leaf with a special glue which hardens rapidly. As it hardens it contracts, deforming the shape of the egg. This glue is easily seen surrounding the base of every egg forming a meniscus. The nature of the glue has been little researched but in the case of Pieris brassicae , it begins as a pale yellow granular secretion containing acidophilic proteins. This is viscous and darkens when exposed to air, becoming
1365-547: A longer lifespan of several months as adults. The thorax of the butterfly is devoted to locomotion. Each of the three thoracic segments has two legs (among nymphalids , the first pair is reduced and the insects walk on four legs). The second and third segments of the thorax bear the wings. The leading edges of the forewings have thick veins to strengthen them, and the hindwings are smaller and more rounded and have fewer stiffening veins. The forewings and hindwings are not hooked together ( as they are in moths ) but are coordinated by
1470-452: A necessity for their successful establishment. Many butterflies, such as the painted lady , monarch, and several danaine migrate for long distances. These migrations take place over a number of generations and no single individual completes the whole trip. The eastern North American population of monarchs can travel thousands of miles south-west to overwintering sites in Mexico . There is
1575-591: A paid, one-month internship. Pilar O'Leary launched the program when she served as executive director of the Smithsonian Latino Center. According to the Latino Center, O'Leary told the press in 2007: "Our goal is to help our Young Ambassadors become the next generation of leaders in the arts and culture fields. This program encourages students to be proud of their roots and learn more about their cultural heritage to inspire them to educate
1680-658: A reverse migration in the spring. It has recently been shown that the British painted lady undertakes a 9,000-mile round trip in a series of steps by up to six successive generations, from tropical Africa to the Arctic Circle — almost double the length of the famous migrations undertaken by monarch. Spectacular large-scale migrations associated with the monsoon are seen in peninsular India. Migrations have been studied in more recent times using wing tags and also using stable hydrogen isotopes . Butterflies navigate using
1785-412: A segmented palp. Adjoining these is the labium-hypopharynx which houses a tubular spinneret which is able to extrude silk. Caterpillars such as those in the genus Calpodes (family Hesperiidae) have a specialized tracheal system on the 8th segment that function as a primitive lung. Butterfly caterpillars have three pairs of true legs on the thoracic segments and up to six pairs of prolegs arising from
1890-435: A silken girdle may be spun to keep the pupa in a head-up position. Most of the tissues and cells of the larva are broken down inside the pupa, as the constituent material is rebuilt into the imago. The structure of the transforming insect is visible from the exterior, with the wings folded flat on the ventral surface and the two halves of the proboscis, with the antennae and the legs between them. The pupal transformation into
1995-665: A single generation, and a few in cold locations may take several years to pass through their entire life cycle. Butterflies are often polymorphic , and many species make use of camouflage , mimicry , and aposematism to evade their predators. Some, like the monarch and the painted lady , migrate over long distances. Many butterflies are attacked by parasites or parasitoids , including wasps , protozoans , flies , and other invertebrates, or are preyed upon by other organisms. Some species are pests because in their larval stages they can damage domestic crops or trees; other species are agents of pollination of some plants. Larvae of
2100-410: A time-compensated sun compass. They can see polarized light and therefore orient even in cloudy conditions. The polarized light near the ultraviolet spectrum appears to be particularly important. Many migratory butterflies live in semi-arid areas where breeding seasons are short. The life histories of their host plants also influence butterfly behaviour. Butterflies in their adult stage can live from
2205-541: A total of about 20,000 species. Traditionally, butterflies have been divided into the superfamilies Papilionoidea and the moth-like Hedyloidea . Recent work has discovered that Hedylidae, the only family within Hedyloidea, is nested within the Papilionoidea, meaning that Papilionoidea would be synonymous with Rhopalocera. The relationships between the rest of the 6 families are extremely well resolved, which
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#17327808950542310-470: A trend towards multivoltinism . Courtship is often aerial and often involves pheromones . Butterflies then land on the ground or on a perch to mate. Copulation takes place tail-to-tail and may last from minutes to hours. Simple photoreceptor cells located at the genitals are important for this and other adult behaviours. The male passes a spermatophore to the female; to reduce sperm competition, he may cover her with his scent, or in some species such as
2415-486: A tubular proboscis which is curled up at rest and expanded when needed to feed. The first and second maxillae bear palps which function as sensory organs. Some species have a reduced proboscis or maxillary palps and do not feed as adults. Many Heliconius butterflies also use their proboscis to feed on pollen; in these species only 20% of the amino acids used in reproduction come from larval feeding, which allow them to develop more quickly as caterpillars, and gives them
2520-437: A water-insoluble, rubbery material which soon sets solid. Butterflies in the genus Agathymus do not fix their eggs to a leaf; instead, the newly laid eggs fall to the base of the plant. Eggs are almost invariably laid on plants. Each species of butterfly has its own host plant range and while some species of butterfly are restricted to just one species of plant, others use a range of plant species, often including members of
2625-422: A week to nearly a year depending on the species. Many species have long larval life stages while others can remain dormant in their pupal or egg stages and thereby survive winters. The Melissa Arctic ( Oeneis melissa ) overwinters twice as a caterpillar. Butterflies may have one or more broods per year. The number of generations per year varies from temperate to tropical regions with tropical regions showing
2730-527: A year and includes as ex officio members the chief justice of the United States and the vice president of the United States . The nominal head of the institution is the chancellor , an office which has traditionally been held by the chief justice. In September 2007, the board created the position of chair of the Board of Regents, a position currently held by Risa Lavizzo-Mourey . Other members of
2835-576: A zoo, and historical and architectural landmarks, mostly located in Washington, D.C. Additional facilities are located in Maryland , New York , and Virginia . More than 200 institutions and museums in 47 states , Puerto Rico , and Panama are Smithsonian Affiliates . Institution publications include Smithsonian and Air & Space magazines. Almost all of the institution's 30 million annual visitors are admitted without charge,
2940-450: Is above 27 °C (81 °F); when it is cool, they can position themselves to expose the underside of the wings to the sunlight to heat themselves up. If their body temperature reaches 40 °C (104 °F), they can orientate themselves with the folded wings edgewise to the sun. Basking is an activity which is more common in the cooler hours of the morning. Some species have evolved dark wingbases to help in gathering more heat and this
3045-458: Is derived from tiny structures called scales, each of which have their own pigments . In Heliconius butterflies, there are three types of scales: yellow/white, black, and red/orange/brown scales. Some mechanism of wing pattern formation are now being solved using genetic techniques. For instance, a gene called cortex determines the colour of scales: deleting cortex turned black and red scales yellow. Mutations, e.g. transposon insertions of
3150-403: Is especially evident in alpine forms. As in many other insects, the lift generated by butterflies is more than can be accounted for by steady-state, non-transitory aerodynamics . Studies using Vanessa atalanta in a wind tunnel show that they use a wide variety of aerodynamic mechanisms to generate force. These include wake capture , vortices at the wing edge, rotational mechanisms and
3255-410: Is extruded and inserted into the female's vagina. A spermatophore is deposited in the female, following which the sperm make their way to a seminal receptacle where they are stored for later use. In both sexes, the genitalia are adorned with various spines, teeth, scales and bristles, which act to prevent the butterfly from mating with an insect of another species. After it emerges from its pupal stage,
Coenonympha nipisiquit - Misplaced Pages Continue
3360-451: Is good, especially in some species in the blue/violet range. The antennae are composed of many segments and have clubbed tips (unlike moths that have tapering or feathery antennae). The sensory receptors are concentrated in the tips and can detect odours. Taste receptors are located on the palps and on the feet. The mouthparts are adapted to sucking and the mandibles are usually reduced in size or absent. The first maxillae are elongated into
3465-469: Is interested, might be interested or ought to be interested". Another Secretary of the Smithsonian, Walter Boyne, founded Air & Space . The organization publishes under the imprints Smithsonian Institution Press, Smithsonian Books, and Smithsonian Institution Scholarly Press. The Smithsonian makes a number of awards to acknowledge and support meritorious work. The Smithsonian Institution
3570-581: Is native to the Americas, but in the nineteenth century or before, spread across the world, and is now found in Australia, New Zealand, other parts of Oceania, and the Iberian Peninsula . It is not clear how it dispersed; adults may have been blown by the wind or larvae or pupae may have been accidentally transported by humans, but the presence of suitable host plants in their new environment was
3675-465: Is restricted to the males, and studies have suggested that the nutrients collected may be provided as a nuptial gift , along with the spermatophore, during mating. In hilltopping , males of some species seek hilltops and ridge tops, which they patrol in search for females. Since it usually occurs in species with low population density, it is assumed these landscape points are used as meeting places to find mates. Butterflies use their antennae to sense
3780-698: Is summarized in the below cladogram. Papilionidae [REDACTED] Hedylidae [REDACTED] Hesperiidae [REDACTED] Pieridae [REDACTED] Nymphalidae [REDACTED] Lycaenidae [REDACTED] Riodinidae [REDACTED] Butterfly adults are characterized by their four scale-covered wings, which give the Lepidoptera their name ( Ancient Greek λεπίς lepís, scale + πτερόν pterón, wing). These scales give butterfly wings their colour: they are pigmented with melanins that give them blacks and browns, as well as uric acid derivatives and flavones that give them yellows, but many of
3885-405: Is taken up by the gut, but there may also be large silk glands, and special glands which secrete distasteful or toxic substances. The developing wings are present in later stage instars and the gonads start development in the egg stage. When the larva is fully grown, hormones such as prothoracicotropic hormone (PTTH) are produced. At this point the larva stops feeding, and begins "wandering" in
3990-479: The Lycaenidae , form mutual associations with ants. They communicate with the ants using vibrations that are transmitted through the substrate as well as using chemical signals. The ants provide some degree of protection to these larvae and they in turn gather honeydew secretions . Large blue ( Phengaris arion ) caterpillars trick Myrmica ants into taking them back to the ant colony where they feed on
4095-544: The Marquis de Lafayette . Operating expenses were covered from the $ 5 yearly dues collected from each member. The institute proposed a number of undertakings. These included the study of plant life and the creation of a botanical garden on the Capitol Mall , an examination of the country's mineral production, improvement in the management and care of livestock, and the writing of a topographical and statistical history of
4200-777: The National Museum of African Art and the Arthur M. Sackler Gallery opened in a new, joint, underground museum between the Freer Gallery and the Smithsonian Castle. Reuse of another old building came in 1993 with the opening of the National Postal Museum in the 1904 former City Post Office building, a few city blocks from the Mall. In 2004, the Smithsonian opened the National Museum of
4305-661: The National Museum of African Art , as well as the National Gallery of Art , were formally returned to Nigerian cultural officials in a ceremony held in Washington D.C. The Nigerian Minister of Information and Culture, Lai Mohammed , and Prince Aghatise Erediauwa, representing the Oba of Benin Kingdom, spoke at the ceremony. Mohammed said the "decision to return the timeless artworks is worth emulating." In February 2020,
Coenonympha nipisiquit - Misplaced Pages Continue
4410-498: The Renwick Gallery . A smaller amount of funds would go to educational initiatives and digitization of collections. As of September 2017, the Smithsonian claimed to have raised $ 1.79 billion, with three months left in the formal campaign calendar. Separately from the major capital campaign, the Smithsonian has begun fundraising through Kickstarter . An example is a campaign to fund the preservation and maintenance of
4515-541: The Smithsonian American Women's History Museum . In 2011, the Smithsonian undertook its first-ever capital fundraising campaign. The $ 1.5 billion effort raised $ 1 billion at the three-year mark. Smithsonian officials made the campaign public in October 2014 in an effort to raise the remaining $ 500 million. More than 60,000 individuals and organizations donated money to the campaign by
4620-632: The South Pacific Ocean . These specimens and artifacts became part of the Smithsonian collections, as did those collected by several military and civilian surveys of the American West , including the Mexican Boundary Survey and Pacific Railroad Surveys , which assembled many Native American artifacts and natural history specimens. In 1846, the regents developed a plan for weather observation; in 1847, money
4725-474: The Weis-Fogh ' clap-and-fling ' mechanism. Butterflies are able to change from one mode to another rapidly. Butterflies are threatened in their early stages by parasitoids and in all stages by predators, diseases and environmental factors. Braconid and other parasitic wasps lay their eggs in lepidopteran eggs or larvae and the wasps' parasitoid larvae devour their hosts, usually pupating inside or outside
4830-455: The ZW sex-determination system where females are the heterogametic sex (ZW) and males homogametic (ZZ). Butterflies are distributed worldwide except Antarctica, totalling some 18,500 species. Of these, 775 are Nearctic ; 7,700 Neotropical ; 1,575 Palearctic ; 3,650 Afrotropical ; and 4,800 are distributed across the combined Oriental and Australian / Oceania regions. The monarch butterfly
4935-458: The deaccession and restitution of items collected under circumstances considered unethical by contemporary standards and thus places moral over legal arguments. A month before, the Smithsonian's National Museum of African Art had announced the planned return of most of its 39 Benin Bronzes to Nigeria , as well as of other cultural items to Turkey . On October 11, 2022, Benin Bronzes from
5040-600: The non-coding DNA around the cortex gene can turn a black-winged butterfly into a butterfly with a yellow wing band. When the butterfly Bicyclus anynana is subjected to repeated inbreeding in the laboratory, there is a dramatic decrease in egg hatching. This severe inbreeding depression is considered to be likely due to a relatively high mutation rate to recessive alleles with substantial damaging effects and infrequent episodes of inbreeding in nature that might otherwise purge such mutations. Although B. anynana experiences inbreeding depression when forcibly inbred in
5145-542: The privilege of the floor at the United States Senate . On September 18, 2013, Secretary G. Wayne Clough announced he would retire in October 2014. The Smithsonian Board of Regents said it asked regent John McCarter, Jr., to lead a search committee. On March 10, 2014, the Smithsonian Board selected David Skorton , a physician and president of Cornell University , as the thirteenth secretary of
5250-1074: The ruby slippers worn by Judy Garland for her role as Dorothy Gale in the 1939 film The Wizard of Oz . Nineteen museums and galleries, as well as the National Zoological Park , comprise the Smithsonian museums. Eleven are on the National Mall , the park that runs between the Lincoln Memorial and the United States Capitol . Other museums are located elsewhere in Washington, D.C., with two more in New York City and one in Chantilly, Virginia . The Smithsonian has close ties with 168 other museums in 39 states, Panama , and Puerto Rico . These museums are known as Smithsonian Affiliated museums. Collections of artifacts are given to these museums in
5355-611: The American Indian in a new building near the United States Capitol . Twelve years later almost to the day, in 2016, the latest museum opened: the National Museum of African American History and Culture , in a new building near the Washington Monument . Two more museums have been established and are being planned for eventual construction on the mall: the National Museum of the American Latino and
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#17327808950545460-540: The Apollos ( Parnassius ) plugs her genital opening to prevent her from mating again. The vast majority of butterflies have a four-stage life cycle: egg , larva (caterpillar), pupa (chrysalis) and imago (adult). In the genera Colias , Erebia , Euchloe , and Parnassius , a small number of species are known that reproduce semi-parthenogenetically ; when the female dies, a partially developed larva emerges from her abdomen. Butterfly eggs are protected by
5565-567: The Board of Regents are three members of the U.S. House of Representatives appointed by the speaker of the House ; three members of the Senate, appointed by the president pro tempore of the Senate ; and nine citizen members, nominated by the board and approved by the Congress in a joint resolution signed by the president of the United States. Regents who are senators or representatives serve for
5670-602: The GDP at the time it may be more comparable to $ 220 million in the year 2007. Once the money was in hand, eight years of congressional haggling ensued over how to interpret Smithson's rather vague mandate "for the increase and diffusion of knowledge." The money was invested by the US Treasury in bonds issued by the state of Arkansas, which soon defaulted. After heated debate, Massachusetts representative (and former president) John Quincy Adams persuaded Congress to restore
5775-510: The National Museum of African-American Heritage and Culture. The opening of the center was prompted, in part, by the publishing of a report called "Willful Neglect: The Smithsonian and U.S. Latinos". According to documents obtained by The Washington Post , when former Latino Center executive director Pilar O'Leary first took the job, the center faced employees who had "serious performance issues". No performance plans existed for
5880-489: The Smithsonian Libraries and Smithsonian Archives merged into one research center in 2020. The Smithsonian Institution includes three cultural centers among its units: In 1997, the Smithsonian Latino Center was created as a way to recognize Latinos across the Smithsonian Institution. The primary purpose of the center is to place Latino contributions to the arts, history, science, and national culture across
5985-646: The Smithsonian and funds to build the museum to hold it (which was named the Freer Gallery ), it was among the Smithsonian's first major donations from a private individual. The gallery opened in 1923. More than 40 years would pass before the next museum, the Museum of History and Technology (renamed the National Museum of American History in 1980), opened in 1964. It was designed by the world-renowned firm of McKim, Mead & White . The Anacostia Community Museum , an "experimental store-front" museum created at
6090-501: The Smithsonian made 2.8 million digital items available to the public under a Creative Commons Zero Public Domain Dedication, with a commitment to release further items in the future. The Smithsonian has eight research centers, located in Washington, D.C.; Front Royal, Virginia ; Edgewater, Maryland ; Suitland, Maryland ; Fort Pierce, Florida ; Cambridge, Massachusetts ; and Panama . Formerly two separate entities,
6195-683: The Smithsonian's Department of Living Animals. The park was designed by landscape architect Frederick Law Olmsted . The National Museum of Natural History opened in June 1911 to similarly accommodate the Smithsonian's United States National Museum, which had previously been housed in the Castle and then the Arts and Industries Building. This structure was designed by the D.C. architectural firm of Hornblower & Marshall . When Detroit philanthropist Charles Lang Freer donated his private collection to
6300-411: The Smithsonian's museums and research centers. The center is a division of the Smithsonian Institution. As of May 2016, the center is run by an executive director, Eduardo Díaz. At the time of its creation, the Smithsonian Institution had other entities dedicated to other minority groups: National Museum of the American Indian, Freer-Sackler Gallery for Asian Arts and Culture, African Art Museum, and
6405-484: The Smithsonian, slightly increased from previous years. Institution exhibits are free of charge, though in 2010 the Deficit Commission recommended admission fees. As approved by Congress on August 10, 1846, the legislation that created the Smithsonian Institution called for the creation of a Board of Regents to govern and administer the organization. This seventeen-member board meets at least four times
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#17327808950546510-532: The Smithsonian. Though the Smithsonian's first secretary, Joseph Henry , wanted the institution to be a center for scientific research, it also became the depository for various Washington and U.S. government collections. The United States Exploring Expedition by the U.S. Navy circumnavigated the globe between 1838 and 1842. The voyage amassed thousands of animal specimens, an herbarium of 50,000 plant specimens, and diverse shells and minerals, tropical birds, jars of seawater, and ethnographic artifacts from
6615-595: The United States. Reports were to be published periodically to share this knowledge with the greater public, but due to a lack of funds, this initially did not occur. The institute first met in Blodget's Hotel, later in the Treasury Department and City Hall, before being assigned a permanent home in 1824 in the Capitol building. Beginning in 1825, weekly sittings were arranged during sessions of Congress for
6720-430: The abdomen, generally with short prolegs on segments 3–6 and 10; the three pairs of true legs on the thorax have five segments each. Many are well camouflaged; others are aposematic with bright colours and bristly projections containing toxic chemicals obtained from their food plants. The pupa or chrysalis, unlike that of moths, is not wrapped in a cocoon. Many butterflies are sexually dimorphic . Most butterflies have
6825-405: The abdominal segments. These prolegs have rings of tiny hooks called crochets that are engaged hydrostatically and help the caterpillar grip the substrate. The epidermis bears tufts of setae , the position and number of which help in identifying the species. There is also decoration in the form of hairs, wart-like protuberances, horn-like protuberances and spines. Internally, most of the body cavity
6930-502: The adult colour pattern are marked by changes in the expression of particular transcription factors in the early pupa. The reproductive stage of the insect is the winged adult or imago . The surface of both butterflies and moths is covered by scales, each of which is an outgrowth from a single epidermal cell. The head is small and dominated by the two large compound eyes . These are capable of distinguishing flower shapes or motion but cannot view distant objects clearly. Colour perception
7035-467: The air for wind and scents. The antennae come in various shapes and colours; the hesperiids have a pointed angle or hook to the antennae, while most other families show knobbed antennae. The antennae are richly covered with sensory organs known as sensillae . A butterfly's sense of taste is coordinated by chemoreceptors on the tarsi , or feet, which work only on contact, and are used to determine whether an egg-laying insect's offspring will be able to feed on
7140-434: The ant eggs and larvae in a parasitic relationship. Caterpillars mature through a series of developmental stages known as instars . Near the end of each stage, the larva undergoes a process called apolysis , mediated by the release of a series of neurohormones . During this phase, the cuticle , a tough outer layer made of a mixture of chitin and specialized proteins , is released from the softer epidermis beneath, and
7245-658: The blues, greens, reds and iridescent colours are created by structural coloration produced by the micro-structures of the scales and hairs. As in all insects, the body is divided into three sections: the head, thorax , and abdomen . The thorax is composed of three segments, each with a pair of legs. In most families of butterfly the antennae are clubbed, unlike those of moths which may be threadlike or feathery. The long proboscis can be coiled when not in use for sipping nectar from flowers. Nearly all butterflies are diurnal , have relatively bright colours, and hold their wings vertically above their bodies when at rest, unlike
7350-615: The construction of the Arts and Industries Building in 1881. Congress had promised to build a new structure for the museum if the 1876 Philadelphia Centennial Exposition generated enough income. It did, and the building was designed by architects Adolf Cluss and Paul Schulze, based on original plans developed by Major General Montgomery C. Meigs of the United States Army Corps of Engineers . It opened in 1881. The National Zoological Park opened in 1889 to accommodate
7455-441: The desiccated husk. Most wasps are very specific about their host species and some have been used as biological controls of pest butterflies like the large white butterfly . When the small cabbage white was accidentally introduced to New Zealand, it had no natural enemies. In order to control it, some pupae that had been parasitised by a chalcid wasp were imported, and natural control was thus regained. Some flies lay their eggs on
7560-401: The duration of their elected terms, while citizen Regents serve a maximum of two six-year terms. Regents are compensated on a part-time basis. The chief executive officer (CEO) of the Smithsonian is the secretary, who is appointed by the Board of Regents. The secretary also serves as secretary to the Board of Regents but is not a voting member of that body. The secretary of the Smithsonian has
7665-419: The epidermis begins to form a new cuticle. At the end of each instar, the larva moults , the old cuticle splits and the new cuticle expands, rapidly hardening and developing pigment. Development of butterfly wing patterns begins by the last larval instar. Caterpillars have short antennae and several simple eyes . The mouthparts are adapted for chewing with powerful mandibles and a pair of maxillae, each with
7770-413: The exception being Cooper Hewitt, Smithsonian Design Museum in New York City, which charges an admissions fee. The Smithsonian's annual budget is around $ 1.25 billion, with two-thirds coming from annual federal appropriations. Other funding comes from the institution's endowment , private and corporate contributions, membership dues, and earned retail, concession, and licensing revenue. As of 2021,
7875-741: The family Hesperiidae (skippers). Molecular clock estimates suggest that butterflies originated sometime in the Late Cretaceous , but only significantly diversified during the Cenozoic, with one study suggesting a North American origin for the group. The oldest American butterfly is the Late Eocene Prodryas persephone from the Florissant Fossil Beds , approximately 34 million years old. Butterflies are divided into seven families that contain
7980-403: The food plant on which their larvae , known as caterpillars , will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in a chrysalis . When metamorphosis is complete, the pupal skin splits, the adult insect climbs out, expands its wings to dry, and flies off. Some butterflies, especially in the tropics, have several generations in a year, while others have
8085-944: The form of long-term loans. The Smithsonian also has a large number of traveling exhibitions, operated through the Smithsonian Institution Traveling Exhibition Service (SITES). In 2008, 58 of these traveling exhibitions went to 510 venues across the country. Smithsonian collections include 156 million artworks, artifacts, and specimens. The National Museum of Natural History houses 145 million of these specimens and artifacts, which are mostly animals preserved in formaldehyde. The Collections Search Center has 9.9 million digital records available online. The Smithsonian Institution Libraries hold 2 million library volumes. Smithsonian Archives hold 156,830 cubic feet (4,441 m ) of archival material. The Smithsonian Institution has many categories of displays that can be visited at
8190-409: The friction of their overlapping parts. The front two segments have a pair of spiracles which are used in respiration. The abdomen consists of ten segments and contains the gut and genital organs. The front eight segments have spiracles and the terminal segment is modified for reproduction. The male has a pair of clasping organs attached to a ring structure, and during copulation, a tubular structure
8295-555: The grass was growing. The earliest Lepidoptera fossils date to the Triassic - Jurassic boundary, around 200 million years ago. Butterflies evolved from moths, so while the butterflies are monophyletic (forming a single clade ), the moths are not. The oldest known butterfly is Protocoeliades kristenseni from the Palaeocene aged Fur Formation of Denmark, approximately 55 million years old, which belongs to
8400-857: The initiative of Smithsonian Secretary S. Dillon Ripley , opened in the Anacostia neighborhood of Washington, D.C., in 1967. That same year, the Smithsonian signed an agreement to take over the Cooper Union Museum for the Arts of Decoration (now the Cooper Hewitt, Smithsonian Design Museum ). The National Portrait Gallery and the Smithsonian American Art Museum opened in the Old Patent Office Building (built in 1867) on October 7, 1968. The reuse of an older building continued with
8505-552: The institution's endowment had a total value of about $ 5.4 billion. In many ways, the origin of the Smithsonian Institution can be traced to a group of Washington citizens who, being "impressed with the importance of forming an association for promoting useful knowledge," met on June 28, 1816, to establish the Columbian Institute for the Promotion of Arts and Sciences . Officers were elected in October 1816, and
8610-1030: The laboratory it recovers within a few generation when allowed to breed freely. During mate selection, adult females do not innately avoid or learn to avoid siblings, implying that such detection may not be critical to reproductive fitness. Inbreeding may persist in B anynana because the probability of encountering close relatives is rare in nature; that is, movement ecology may mask the deleterious effect of inbreeding resulting in relaxation of selection for active inbreeding avoidance behaviors. Butterflies feed primarily on nectar from flowers. Some also derive nourishment from pollen , tree sap, rotting fruit, dung, decaying flesh, and dissolved minerals in wet sand or dirt. Butterflies are important as pollinators for some species of plants. In general, they do not carry as much pollen load as bees , but they are capable of moving pollen over greater distances. Flower constancy has been observed for at least one species of butterfly. Adult butterflies consume only liquids, ingested through
8715-719: The largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge." Founded on August 10, 1846, it operates as a trust instrumentality and is not formally a part of any of the three branches of the federal government . The institution is named after its founding donor, British scientist James Smithson . It was originally organized as the United States National Museum , but that name ceased to exist administratively in 1967. The Smithsonian Institution has historical holdings of over 157 million items, 21 museums, 21 libraries, 14 education and research centers,
8820-531: The legacy bequeathed to the nation and pledged the faith of the United States to the charitable trust on July 1, 1836. The American diplomat Richard Rush was dispatched to England by President Andrew Jackson to collect the bequest. Rush returned in August 1838 with 105 sacks containing 104,960 gold sovereigns . This is approximately $ 500,000 at the time, which is equivalent to $ 14,000,000 in 2023 or equivalent to £12,000,000 in 2023. However, when considering
8925-400: The lost funds with interest and, despite designs on the money for other purposes, convinced his colleagues to preserve it for an institution of science and learning. Finally, on August 10, 1846, President James K. Polk signed the legislation that established the Smithsonian Institution as a trust instrumentality of the United States, to be administered by a Board of Regents and a secretary of
9030-506: The majority of moths which fly by night, are often cryptically coloured (well camouflaged), and either hold their wings flat (touching the surface on which the moth is standing) or fold them closely over their bodies. Some day-flying moths, such as the hummingbird hawk-moth , are exceptions to these rules. Butterfly larvae , caterpillars , have a hard ( sclerotised ) head with strong mandibles used for cutting their food, most often leaves. They have cylindrical bodies, with ten segments to
9135-481: The maritime ringlet is found in New Brunswick, four of these sites are located within only a 10 km radius in or near Bathurst Harbour . The maritime ringlet is preyed on by birds and larger insects. The larvae feed only on salt-meadow cordgrass ( Spartina patens ). Sea lavender ( Limonium nashii ) is its preferred nectar plant. Dr. Reginald Webster, a self-employed Fredericton -based entomologist,
9240-452: The mission to "promote science and the useful arts, and to establish a national museum of natural history," this organization continued to press Congress to establish a museum that would be structured in terms that were very similar to those finally incorporated into the founding of the Smithsonian Institution. Its work helped to develop an underlying philosophy that pushed for the pursuit and development of scientific knowledge that would benefit
9345-629: The museums. In 1912, First Lady Helen Herron Taft donated her inauguration gown to the museum to begin the First Ladies' Gown display at the National Museum of American History , one of the Smithsonian's most popular exhibits. The museum displays treasures such as the Star-Spangled Banner , the stove pipe hat that was worn by President Abraham Lincoln , the ruby slippers worn by Judy Garland in The Wizard Of Oz , and
9450-401: The name is ancient, but modern Dutch and German use different words ( vlinder and Schmetterling ) and the common name often varies substantially between otherwise closely related languages. A possible source of the name is the bright yellow male of the brimstone ( Gonepteryx rhamni ); another is that butterflies were on the wing in meadows during the spring and summer butter season while
9555-473: The nation, and edify its citizens at the same time. The British scientist James Smithson (1765–1829) left most of his wealth to his nephew Henry James Hungerford. When Hungerford died childless in 1835, the estate passed "to the United States of America, to found at Washington, under the name of the Smithsonian Institution, an Establishment for the increase & diffusion of knowledge among men", in accordance with Smithson's will. Congress officially accepted
9660-621: The opening of the Renwick Gallery in 1972 in the 1874 Renwick-designed art gallery originally built by local philanthropist William Wilson Corcoran to house the Corcoran Gallery of Art . The first new museum building to open since the National Museum of History and Technology was the Hirshhorn Museum and Sculpture Garden , which opened in 1974. The National Air and Space Museum , the Smithsonian's largest in terms of floor space, opened in June 1976. Eleven years later,
9765-809: The organization was granted a charter by Congress on April 20, 1818 (this charter expired in 1838). Benjamin Latrobe , who was architect for the US Capitol after the War of 1812, and William Thornton , the architect who designed the Octagon House and Tudor Place , would serve as officers. Other prominent members, who numbered from 30 to 70 during the institute's existence, included John Quincy Adams , Andrew Jackson , Henry Clay , Judge William Cranch , and James Hoban . Honorary members included James Madison , James Monroe , John Adams , Thomas Jefferson , and
9870-443: The original Teddy Bear that was named after President Theodore Roosevelt . In 2016, the Smithsonian's Air & Space museum curators restored the large model Enterprise from the original Star Trek TV series. Following international debates about the decolonisation of museums and the legal and moral justifications of their acquisitions, the Smithsonian adopted a new "ethical returns policy" on April 29, 2022. This will permit
9975-502: The outside of caterpillars and the newly hatched fly larvae bore their way through the skin and feed in a similar way to the parasitoid wasp larvae. Predators of butterflies include ants, spiders, wasps, and birds. Smithsonian Institution The Smithsonian Institution ( / s m ɪ θ ˈ s oʊ n i ə n / smith- SOH -nee-ən ), or simply the Smithsonian , is a group of museums , education and research centers ,
10080-529: The proboscis. They sip water from damp patches for hydration and feed on nectar from flowers, from which they obtain sugars for energy, and sodium and other minerals vital for reproduction. Several species of butterflies need more sodium than that provided by nectar and are attracted by sodium in salt; they sometimes land on people, attracted by the salt in human sweat. Some butterflies also visit dung and scavenge rotting fruit or carcasses to obtain minerals and nutrients. In many species, this mud-puddling behaviour
10185-422: The public in their own communities about how Latinos are enriching America's cultural fabric." The institution publishes Smithsonian magazine monthly and Air & Space magazine bimonthly. Smithsonian was the result of Secretary of the Smithsonian S. Dillon Ripley asking the retired editor of Life magazine Edward K. Thompson to produce a magazine "about things in which the Smithsonian Institution
10290-431: The quest for a suitable pupation site, often the underside of a leaf or other concealed location. There it spins a button of silk which it uses to fasten its body to the surface and moults for a final time. While some caterpillars spin a cocoon to protect the pupa, most species do not. The naked pupa, often known as a chrysalis, usually hangs head down from the cremaster, a spiny pad at the posterior end, but in some species
10395-419: The reading of scientific and literary productions, but this was continued for only a short time, as the number attending declined rapidly. Eighty-five communications by 26 people were made to Congress during the entire life of the society, with more than a half relating to astronomy or mathematics. Among all the activities planned by the institute, only a few were actually implemented. Two were the establishment of
10500-640: The staff and unfulfilled financial obligations to sponsors existed. The website's quality was poor, and the center did not have a public affairs manager, a programs director, adequate human resources support, or cohesive mission statement. After difficult times in the first few years, the center improved. According to the Smithsonian, the center "support[s] scholarly research, exhibitions, public and educational programs, web-based content and virtual platforms, and collections and archives. [It] also manage[s] leadership and professional development programs for Latino youth, emerging scholars and museum professionals." Today,
10605-473: The time it went public. This included 192 gifts of at least $ 1 million. Members of the boards of directors of various Smithsonian museums donated $ 372 million. The Smithsonian said that funds raised would go toward completion of the National Museum of African American History and Culture building, and renovations of the National Air and Space Museum , National Museum of American History , and
10710-546: The website features a high-tech virtual museum including self-guided virtual tours of past and present exhibits. The Smithsonian Latino Center's Young Ambassadors Program (YAP) is a program within the Latino Center that reaches out to Latino high school students with the goal of encouraging them to become leaders in arts, sciences, and the humanities. Students selected for the program travel to Washington, D.C. for an "enrichment seminar" that lasts approximately five days. Afterwards, students return to their communities to serve in
10815-850: Was appropriated for meteorological research. The institution became a magnet for young scientists from 1857 to 1866, who formed a group called the Megatherium Club . The Smithsonian played a critical role as the US partner institution in early bilateral scientific exchanges with the Academy of Sciences of Cuba . Construction began on the Smithsonian Institution Building ("the Castle") in 1849. Designed by architect James Renwick Jr. , its interiors were completed by general contractor Gilbert Cameron. The building opened in 1855. The Smithsonian's first expansion came with
10920-559: Was established as a trust instrumentality by act of Congress. More than two-thirds of the Smithsonian's workforce of some 6,300 persons are employees of the federal government. The Smithsonian Institution Office of Protection Services oversees security at the Smithsonian facilities and enforces laws and regulations for National Capital Parks together with the United States Park Police . The president's 2011 budget proposed just under $ 800 million in support for
11025-599: Was responsible from 1993 on for a WWF -funded project that led to the implementation of a recovery plan, monitoring protocol, and successful introduction of a new colony in the Acadian Peninsula . Coenonympha tullia – common ringlet This Satyrini article is a stub . You can help Misplaced Pages by expanding it . Butterfly Butterflies are winged insects from the lepidopteran suborder Rhopalocera , characterized by large, often brightly coloured wings that often fold together when at rest, and
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